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<mods:title id="DC84776C69E8256DCBCBBAA8DF4F31CB">A revision of the South American species of the Morelloid clade (Solanum L., Solanaceae)</mods:title>
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11.
<taxonomicName id="AFA00FA86C4493F12BFAE73F981B42EE" ID-CoL="4XZX5" ID-ENA="795972" authority="Lam., Tabl. Encycl. 2: 18. 1794." authorityName="Lam., Tabl. Encycl. 2: 18. 1794." class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum chenopodioides" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="chenopodioides">Solanum chenopodioides Lam., Tabl. Encycl. 2: 18. 1794.</taxonomicName>
</paragraph>
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<paragraph id="58135B0E844CB4A6486AA73E9BE199F3" pageId="0" pageNumber="1">
<figureCitation id="A8C73D4333D482055651445CFC34004C" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Solanum chenopodioides A habit B detail of adaxial leaf surface C detail of abaxial leaf surface D opening bud E dissected flower F fruiting branch G detail of infructescence H maturing fruit I fully mature fruit (A-E Fox s. n. F-I Hieronymus s. n.). Illustration by R. Wise. Previously published in Barboza et al. (2013: 242), Saerkinen et al. (2018: 67) and Knapp et al. (2019: 46)." figureDoi="10.3897/phytokeys.231.100894.figure35" httpUri="https://binary.pensoft.net/fig/897496" pageId="0" pageNumber="1">Figs 35</figureCitation>
<figureCitation id="4CFB3E5EBE27C8334D562FAC0DF93E01" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Solanum chenopodioides A habit B buds C flowers at full anthesis D fully mature dull black fruits with appressed calyx lobes (A-D Nijmegen acc. # A 14750051). Photos by S. Knapp and G. van der Weerden. Previously published in Saerkinen et al. (2018: 68) and Knapp et al. (2019: 47)." figureDoi="10.3897/phytokeys.231.100894.figure36" httpUri="https://binary.pensoft.net/fig/897497" pageId="0" pageNumber="1">, 36</figureCitation>
</paragraph>
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<paragraph id="63AEB940D07D6185C015B61F37E3708D" pageId="0" pageNumber="1">
<taxonomicName id="1C8A2CECEE5A0D6B7C53B3EAD0A04D54" authorityName="Willd. ex Roemer &amp; Schultes" authorityYear="1819" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum sublobatum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="sublobatum">Solanum sublobatum</taxonomicName>
Willd. ex Roem. &amp; Schult., Syst. Veg., ed. 15 bis [Roemer &amp; Schultes] 4: 664. 1819. Type. Argentina. Buenos Aires,
<emphasis id="55CD21657B5F09DD0B35E2602B051F6F" italics="true" pageId="0" pageNumber="1">Anon. s.n.</emphasis>
[probably
<emphasis id="DBC9E062DD32BFCC6B3FE6AB3F38F546" italics="true" pageId="0" pageNumber="1">P. Commerson</emphasis>
] (
<emphasis id="28F3DE49E41E3E644BC4FBC99130B0E7" italics="true" pageId="0" pageNumber="1">Herb. Willdenow 4336</emphasis>
) (lectotype, designated by
<bibRefCitation id="27BD91A1C1FB4B6B4BEF821A5484373D" DOI="https://doi.org/10.2307/4117874" author="Edmonds, JM" journalOrPublisher="Kew Bulletin" pageId="0" pageNumber="1" pagination="95 - 114" refId="B73" refString="Edmonds, JM, 1972. A synopsis of the taxonomy of Solanum sect. Solanum (Maurella) in South America. Kew Bulletin 27 (1): 95 - 114, DOI: https://doi.org/10.2307/4117874" title="A synopsis of the taxonomy of Solanum sect. Solanum (Maurella) in South America." url="https://doi.org/10.2307/4117874" volume="27" year="1972">Edmonds 1972</bibRefCitation>
, pg. 105 [as type ex photo]: B [B-W04336-01-0]).
</paragraph>
<paragraph id="A2C0131114068BBFFE89A6BDD7AAD9AC" pageId="0" pageNumber="1">
<taxonomicName id="F921670D4F3B487C13F7D95600B4B17C" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum besseri" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="besseri">Solanum besseri</taxonomicName>
Weinm., Syst. Veg., ed. 15 bis [Roemer &amp; Schultes] 4: 593. 1819. Type. &quot;In America&quot; [cultivated in Europe?],
<emphasis id="03453C69F2E53B4A0222B3F02AFC89E6" italics="true" pageId="0" pageNumber="1">Anon. s.n.</emphasis>
(no specimens cited; no original material located; neotype, designated by
<bibRefCitation id="D6C95FC1D31FE77B0C2903D1C1A229F7" DOI="https://doi.org/10.3897/phytokeys.106.21991" author="Saerkinen, T" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="1 - 223" refId="B209" refString="Saerkinen, T, Poczai, P, Barboza, GE, van der Weerden, GM, Baden, M, Knapp, S, 2018. A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae). PhytoKeys 106: 1 - 223, DOI: https://doi.org/10.3897/phytokeys.106.21991" title="A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae)." url="https://doi.org/10.3897/phytokeys.106.21991" volume="106" year="2018">
<normalizedToken id="548CFF2F6054DD901FD29E8A016D9F13" originalValue="Särkinen">Saerkinen</normalizedToken>
et al. 2018
</bibRefCitation>
, pg. 65: G-DC [G00144198]).
</paragraph>
<paragraph id="4523BE9E3EC89FE79BDC2C54EF5BFBAD" pageId="0" pageNumber="1">
<taxonomicName id="AE4675A704BE5097BADD928F6EE0893B" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum subspatulatum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="subspatulatum">Solanum subspatulatum</taxonomicName>
Sendtn., Fl. Bras. (Martius) 10: 45, tab. 4, figs 16-18. 1846. Type. Brazil. Sin. loc.,
<emphasis id="4362998EE2610D2AE8A2B587C4780E4E" italics="true" pageId="0" pageNumber="1">F. Sellow s.n.</emphasis>
(holotype: B, destroyed [F neg. 3183]; lectotype, designated by
<bibRefCitation id="2FDD6364DFB437B2918E9A9425448210" DOI="https://doi.org/10.2307/2395139" author="D'Arcy, WG" journalOrPublisher="Annals of the Missouri Botanical Garden" pageId="0" pageNumber="1" pagination="573 - 780" refId="B53" refString="D'Arcy, WG, 1974a. [1973]) Solanaceae, Flora of Panama. Part 9. Annals of the Missouri Botanical Garden 60: 573 - 780, DOI: https://doi.org/10.2307/2395139" title="[1973]) Solanaceae, Flora of Panama. Part 9." url="https://doi.org/10.2307/2395139" volume="60" year="1974 a">
<normalizedToken id="449329905420049E4B59E38AB97D2629" originalValue="DArcy">D'Arcy</normalizedToken>
1974a
</bibRefCitation>
, pg. 735 [as type]: P [P00384051]; isolectotype: F [v0361921F, acc. # 621700, fragment]).
</paragraph>
<paragraph id="0603C052CE9105AE5506B291EF7BDCB3" pageId="0" pageNumber="1">
<taxonomicName id="D492FC23DC168BFDF1F72EF0AD25221E" authorityName="J.Remy" authorityYear="1849" baseAuthorityName="Lam." class="Magnoliopsida" family="Solanaceae" genus="Witheringia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Witheringia chenopodioides" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="chenopodioides">Witheringia chenopodioides</taxonomicName>
(Lam.) J.
<normalizedToken id="B0246817DFD157A5EA7B318F9876C1E7" originalValue="Rémy">Remy</normalizedToken>
, Fl. Chil. [Gay] 5: 69. 1849. Type. Based on
<taxonomicName id="88D4D1C2BA72237B1D08AE96D2CA6BAA" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum chenopodioides" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="chenopodioides">Solanum chenopodioides</taxonomicName>
Lam.
</paragraph>
<paragraph id="C178EF406248C0FFE7956F8BE084A6C2" pageId="0" pageNumber="1">
<taxonomicName id="8E17363D118815203BB06C81A11DA9EB" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum chenopodiifolium" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="chenopodiifolium">Solanum chenopodiifolium</taxonomicName>
Dunal, Prodr. [A. P. de Candolle] 13(1): 44. 1852. Type. Argentina/Uruguay. &quot;Buenos Aires et Montevideo&quot;,
<emphasis id="FABB4DFA976CFE1FA0E7A9A489D71710" italics="true" pageId="0" pageNumber="1">P. Commerson s.n.</emphasis>
(lectotype, designated by
<bibRefCitation id="46C3FF6A3EF06A3BC08071DAFFA63AC5" DOI="https://doi.org/10.2307/4117874" author="Edmonds, JM" journalOrPublisher="Kew Bulletin" pageId="0" pageNumber="1" pagination="95 - 114" refId="B73" refString="Edmonds, JM, 1972. A synopsis of the taxonomy of Solanum sect. Solanum (Maurella) in South America. Kew Bulletin 27 (1): 95 - 114, DOI: https://doi.org/10.2307/4117874" title="A synopsis of the taxonomy of Solanum sect. Solanum (Maurella) in South America." url="https://doi.org/10.2307/4117874" volume="27" year="1972">Edmonds 1972</bibRefCitation>
, pg. 108 [as holotype], second step designated by
<bibRefCitation id="4BE46C710219DCA70D7EA6D9FF581BC2" DOI="https://doi.org/10.3897/phytokeys.106.21991" author="Saerkinen, T" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="1 - 223" refId="B209" refString="Saerkinen, T, Poczai, P, Barboza, GE, van der Weerden, GM, Baden, M, Knapp, S, 2018. A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae). PhytoKeys 106: 1 - 223, DOI: https://doi.org/10.3897/phytokeys.106.21991" title="A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae)." url="https://doi.org/10.3897/phytokeys.106.21991" volume="106" year="2018">
<normalizedToken id="8CDF12CF8573E0B5C52A10CCFAF6E560" originalValue="Särkinen">Saerkinen</normalizedToken>
et al. 2018
</bibRefCitation>
, pg. 65: P [P00384081]).
</paragraph>
<paragraph id="6B52A461AA1B1CFFD08EC47FCAE27B92" pageId="0" pageNumber="1">
<taxonomicName id="A537BB269757B843A7468BF171E0AEBF" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum gracile" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="gracile">Solanum gracile</taxonomicName>
Dunal, Prodr. [A.P. de Candolle] 13(1): 54. 1852, nom. illeg., not
<taxonomicName id="14A7C5861E2E2D2C9CA025152568FCDB" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum gracile" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="gracile">Solanum gracile</taxonomicName>
Sendtn. (1846). Type. Brazil. Rio de Janeiro: &quot;Rio de Janeiro&quot;, 1831-1833,
<emphasis id="5D7558050D499C16A96376C338FFD557" italics="true" pageId="0" pageNumber="1">C. Gaudichaud 520</emphasis>
(lectotype, designated by
<bibRefCitation id="B740209132368BC3DA4A3664D3707D3A" author="Henderson, RJF" journalOrPublisher="Contributions from the Queensland Herbarium" pageId="0" pageNumber="1" pagination="1 - 78" refId="B105" refString="Henderson, RJF, 1974. Solanum nigrum L. (Solanaceae) and related species in Australia. Contributions from the Queensland Herbarium 16: 1 - 78" title="Solanum nigrum L. (Solanaceae) and related species in Australia." volume="16" year="1974">Henderson 1974</bibRefCitation>
, pg. 46: G-DC [G00144391]; isolectotypes: G [G00343457], P [P00384052, P00384053]).
</paragraph>
<paragraph id="EBFCB199EA88D753F5878A0329F51CFC" pageId="0" pageNumber="1">
<taxonomicName id="B1789049C2B5BA4DF02EB1D63338E3AD" authorityName="Dunal" authorityYear="1852" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum gracile subsp. var. var. microphyllum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="gracile" subSpecies="var." variety="microphyllum">Solanum gracile Dunal var. microphyllum</taxonomicName>
Dunal, Prodr. [A. P. de Candolle] 13(1): 54. 1852. Type. Argentina/Uruguay. &quot;Circa Buenos Ayres et Montevideo&quot;,
<emphasis id="FA9648AAAA79F40BCDE7C32CBE2C8BA6" italics="true" pageId="0" pageNumber="1">P. Commerson s.n.</emphasis>
(lectotype, designated by
<bibRefCitation id="F6988657FDF79296CD01407BBBC81DE9" author="Morton, CV" journalOrPublisher="Biodiversity and Conservation" pageId="0" pageNumber="1" refId="B166" refString="Morton, CV, 1976. A revision of the Argentine species of Solanum. Academia Nacional de Ciencias, Cordoba." title="A revision of the Argentine species of Solanum. Academia Nacional de Ciencias, Cordoba." year="1976">Morton 1976</bibRefCitation>
, pg. 151: P [P00384061, Morton neg. 8207]; possible isolectotype: F [v0073283F, acc. # 976485, fragment only]).
</paragraph>
<paragraph id="01A1892367BDBBB868027B0B2DB274DB" pageId="0" pageNumber="1">
<taxonomicName id="52E172802CBAF2EDB2BAD68166256658" authorityName="Dunal" authorityYear="1852" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum isabellei" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="isabellei">Solanum isabellei</taxonomicName>
Dunal, Prodr. [A. P. de Candolle] 13(1): 153. 1852. Type. Uruguay. Montevideo, Lat. aust. 34°45'08&quot;, 1838,
<emphasis id="D92CB373DB89CA34D0A5A382EE60ABEC" italics="true" pageId="0" pageNumber="1">A. Isabelle s.n.</emphasis>
(lectotype, designated by
<bibRefCitation id="85ED70931F2A45C91C72CDCEF37CA03F" DOI="https://doi.org/10.3897/phytokeys.106.21991" author="Saerkinen, T" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="1 - 223" refId="B209" refString="Saerkinen, T, Poczai, P, Barboza, GE, van der Weerden, GM, Baden, M, Knapp, S, 2018. A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae). PhytoKeys 106: 1 - 223, DOI: https://doi.org/10.3897/phytokeys.106.21991" title="A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae)." url="https://doi.org/10.3897/phytokeys.106.21991" volume="106" year="2018">
<normalizedToken id="9ABE756F833A969E6BA053B4D9A40732" originalValue="Särkinen">Saerkinen</normalizedToken>
et al. 2018
</bibRefCitation>
, pg. 65: G-DC (G00145645); isolectotypes: F [v0073298F, acc. # 680251; v0073299F, acc. # 680253], K [K000585686], P [P00384071], W [acc. # 1889-115034]).
</paragraph>
<paragraph id="423568C1CE83C83A3E5D4ACECD7A0191" pageId="0" pageNumber="1">
<taxonomicName id="E90F48966F6F9544656638D0160387A8" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum nodiflorum subsp. var. var. microphyllum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="nodiflorum" subSpecies="var." variety="microphyllum">Solanum nodiflorum Jacq. var. microphyllum</taxonomicName>
Hassl., Repert. Spec. Nov. Regni Veg. 9: 118. 1911. Type. Paraguay. Estrella: Mar,
<emphasis id="46474EF99543D2CCAA94B6BD90B08DDA" italics="true" pageId="0" pageNumber="1">
<normalizedToken id="AA66B5F824B18DB015BE899B2DB89565" originalValue="É">E</normalizedToken>
. Hassler 10271
</emphasis>
(holotype: G [n.v.], Morton photo 8612).
</paragraph>
<paragraph id="C18584DF84BC4EBD3C840BF688001C20" pageId="0" pageNumber="1">
<taxonomicName id="5DD53543DDB37DA9DAAB65737C391A55" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum vile" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="vile">Solanum vile</taxonomicName>
Bitter, Repert. Spec. Nov. Regni Veg. 11: 221. 1912. Type. Brazil. Rio de Janeiro: Restinga do Harpoador,
<emphasis id="081F4A342EC2BD676E680068540245B5" italics="true" pageId="0" pageNumber="1">E. Ule 4310</emphasis>
(lectotype, designated by
<bibRefCitation id="5F09D33B677AC232F0A1DB25350F01C5" DOI="https://doi.org/10.3897/phytokeys.106.21991" author="Saerkinen, T" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="1 - 223" refId="B209" refString="Saerkinen, T, Poczai, P, Barboza, GE, van der Weerden, GM, Baden, M, Knapp, S, 2018. A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae). PhytoKeys 106: 1 - 223, DOI: https://doi.org/10.3897/phytokeys.106.21991" title="A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae)." url="https://doi.org/10.3897/phytokeys.106.21991" volume="106" year="2018">
<normalizedToken id="C456A93A9DCD8C3F7EDE21B663C8B903" originalValue="Särkinen">Saerkinen</normalizedToken>
et al. 2018
</bibRefCitation>
, pg. 66: CORD [CORD00004277]; isolectotype: HBG [HBG511507]).
</paragraph>
<paragraph id="8E9AEBBE5326070514A024B2F0CDC2C1" pageId="0" pageNumber="1">
<taxonomicName id="B64EFC929BB715589C6F9E70F5AEEA7E" authorityName="Herter" authorityYear="1943" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum gracilius" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="gracilius">Solanum gracilius</taxonomicName>
Herter, Rev. Sudamer. Bot. 7: 266. 1943. Type: Based on (replacement name for)
<taxonomicName id="EB3B269C218EB152EAB912086A4E548C" family="Solanaceae" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="S. gracile" order="Solanales" pageId="0" pageNumber="1" rank="species" species="gracile">S. gracile</taxonomicName>
Dunal.
</paragraph>
<paragraph id="D3570678FDD7643136CFF43E6E0ED12F" pageId="0" pageNumber="1">
<taxonomicName id="4E27197F35394983CFD2F2DEFED7C7DF" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum ottonis" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="ottonis">Solanum ottonis</taxonomicName>
Hyl., Uppsala Univ.
<normalizedToken id="8F43A0A44D5273AB555240233D9C3E29" originalValue="Årsskr">Arsskr</normalizedToken>
. 7: 279. 1945. Type. Based on (replacement name for)
<taxonomicName id="7BD613B70FDE619BEC8037DC7AE68E2E" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum gracile" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="gracile">Solanum gracile</taxonomicName>
Dunal.
</paragraph>
</subSubSection>
<subSubSection id="EB37B3E3F85CCA522B6DE04B262D49E9" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="2A1BC1BE4FFC47F64784D897489EBC81" pageId="0" pageNumber="1">
<typeStatus id="AB3739BCD53703D50D1A4FB600AF0B17">Type</typeStatus>
.
</paragraph>
<paragraph id="FE493290E6472EC111590CE53C8C6EEB" pageId="0" pageNumber="1">
<collectingCountry id="8B43F2F86E145F528507815E6E5024EA" name="Mauritius">Mauritius</collectingCountry>
. &quot;Ex ins. Mauritiana&quot;,
<emphasis id="88F7DCB15947D2602F07B72821010814" italics="true" pageId="0" pageNumber="1">Herb. Lamarck s.n.</emphasis>
(
<typeStatus id="CDBA369C9EEF3A0647B13CA31E0CBD96">lectotype</typeStatus>
, designated by
<bibRefCitation id="3E0147FA7CC2C65E81BB730D0735CCF8" author="Barboza, GE" editor="Anton, AM" journalOrPublisher=") Flora Argentina (Vol. 13), Solanaceae. IOBDA- IMBIV, CONICET: Buenos Aires &amp; Cordoba, Argentina" pageId="0" pageNumber="1" pagination="231 - 264" refId="B10" refString="Barboza, GE, Knapp, S, Saerkinen, T, 2013. Grupo VII. Moreloide. In: Anton, AM, Zuloaga, FO, Eds., Barboza GE (Coord.) Flora Argentina (Vol. 13), Solanaceae. IOBDA- IMBIV, CONICET: Buenos Aires &amp; Cordoba, Argentina: 231 - 264" title="Grupo VII. Moreloide." volumeTitle="Barboza GE (Coord." year="2013">Barboza et al. 2013</bibRefCitation>
, pg. 242: P [P00357629]).
</paragraph>
<caption id="023925B9ABEEC2993A347D2CCB22C572" doi="10.3897/phytokeys.231.100894.figure35" httpUri="https://binary.pensoft.net/fig/897496" pageId="0" pageNumber="1" start="Figure 35" startId="F35">
<paragraph id="77704833C546775B9796B7B4A646439E" pageId="0" pageNumber="1">
<emphasis id="7E7A28931DC9D56DFFED95349C5002AF" bold="true" pageId="0" pageNumber="1">Figure 35.</emphasis>
<taxonomicName id="6BEE55BA955785048EC0614D4454CCAB" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum chenopodioides" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="chenopodioides">
<emphasis id="9EDBD2A5BD057081CAEEFAE670432EDD" italics="true" pageId="0" pageNumber="1">Solanum chenopodioides</emphasis>
</taxonomicName>
<emphasis id="AAA781FD53714C031A1545D0C1B0FF0B" bold="true" pageId="0" pageNumber="1">A</emphasis>
habit
<emphasis id="C098910724FDDFB34B144C7EF5F30B03" bold="true" pageId="0" pageNumber="1">B</emphasis>
detail of adaxial leaf surface
<emphasis id="ECF117C59F290CECB434FF1A8FCDBF3A" bold="true" pageId="0" pageNumber="1">C</emphasis>
detail of abaxial leaf surface
<emphasis id="4FA1AE15BB56E51B10BC5BA2776BF8F8" bold="true" pageId="0" pageNumber="1">D</emphasis>
opening bud
<emphasis id="238E3C1F660C08629CDEE694DC053373" bold="true" pageId="0" pageNumber="1">E</emphasis>
dissected flower
<emphasis id="36790AEED5E64D0FC5671C9A5F4A6DB4" bold="true" pageId="0" pageNumber="1">F</emphasis>
fruiting branch
<emphasis id="A8BE379654A6ED6D7651D1D295FBF16A" bold="true" pageId="0" pageNumber="1">G</emphasis>
detail of infructescence
<emphasis id="EE5E9024446470A5FD3D69351C569765" bold="true" pageId="0" pageNumber="1">H</emphasis>
maturing fruit
<emphasis id="1BD7B9FEBE03B5775A0AEAC1040A3CC2" bold="true" pageId="0" pageNumber="1">I</emphasis>
fully mature fruit (
<emphasis id="82B82179E143C00A250A635A90C5BA06" bold="true" pageId="0" pageNumber="1">A-E</emphasis>
<emphasis id="83009AB7D2402E12E77FA5A40FA2D3AC" italics="true" pageId="0" pageNumber="1">Fox s.n.</emphasis>
<emphasis id="3FA0CC296B187F15E06D7727150B49EA" bold="true" pageId="0" pageNumber="1">F-I</emphasis>
<emphasis id="95C47338DC0FB094B4ED719893CB63A9" italics="true" pageId="0" pageNumber="1">Hieronymus s.n.</emphasis>
). Illustration by R. Wise. Previously published in
<bibRefCitation id="4FB853F5CF7B083F788564130F620EF8" author="Barboza, GE" editor="Anton, AM" journalOrPublisher=") Flora Argentina (Vol. 13), Solanaceae. IOBDA- IMBIV, CONICET: Buenos Aires &amp; Cordoba, Argentina" pageId="0" pageNumber="1" pagination="231 - 264" refId="B10" refString="Barboza, GE, Knapp, S, Saerkinen, T, 2013. Grupo VII. Moreloide. In: Anton, AM, Zuloaga, FO, Eds., Barboza GE (Coord.) Flora Argentina (Vol. 13), Solanaceae. IOBDA- IMBIV, CONICET: Buenos Aires &amp; Cordoba, Argentina: 231 - 264" title="Grupo VII. Moreloide." volumeTitle="Barboza GE (Coord." year="2013">Barboza et al. (2013</bibRefCitation>
: 242),
<bibRefCitation id="0D782CD92A3248A5CF37CC97A2E7F777" DOI="https://doi.org/10.3897/phytokeys.106.21991" author="Saerkinen, T" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="1 - 223" refId="B209" refString="Saerkinen, T, Poczai, P, Barboza, GE, van der Weerden, GM, Baden, M, Knapp, S, 2018. A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae). PhytoKeys 106: 1 - 223, DOI: https://doi.org/10.3897/phytokeys.106.21991" title="A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae)." url="https://doi.org/10.3897/phytokeys.106.21991" volume="106" year="2018">
<normalizedToken id="C8DF5F805A9CB466605ED7C609B5266E" originalValue="Särkinen">Saerkinen</normalizedToken>
et al. (2018
</bibRefCitation>
: 67) and
<bibRefCitation id="C65E57ED2C5DF6F4A844161AB4FDC183" DOI="https://doi.org/10.3897/phytokeys.123.31738" author="Knapp, S" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="1 - 174" refId="B136" refString="Knapp, S, Barboza, GE, Bohs, L, Saerkinen, T, 2019. A revision of the Morelloid Clade of Solanum L. (Solanaceae) in the Caribbean and North and Central America. PhytoKeys 123: 1 - 174, DOI: https://doi.org/10.3897/phytokeys.123.31738" title="A revision of the Morelloid Clade of Solanum L. (Solanaceae) in the Caribbean and North and Central America." url="https://doi.org/10.3897/phytokeys.123.31738" volume="123" year="2019">Knapp et al. (2019</bibRefCitation>
: 46).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="186FDAE589E5CAA735E1A404F8A4EF1A" pageId="0" pageNumber="1" type="description">
<paragraph id="D32C88080CCDDA58534E57E4C9F9D1C6" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="4D0A79F077429B0280162AB40D22530A" pageId="0" pageNumber="1">
Annual herbs to short-lived perennial shrubs up to 1 m high, subwoody and branching at base. Stems terete, green-grey to straw colour, sprawling, somewhat weak and decumbent, not markedly hollow; new growth usually densely pubescent with simple, uniseriate appressed 1-6-celled eglandular trichomes, these 0.1-0.6 mm long; older stems more sparsely pubescent, glabrescent. Sympodial units difoliate, the leaves not geminate. Leaves simple, the blades 1.5-5.5(-7) cm long, 0.5-3(-3.5) cm wide, lanceolate to narrowly ovate, rarely ovate, widest at the middle or slightly below, membranous, discolorous; adaxial surface green, sparsely pubescent with appressed 1-4-celled translucent, simple, uniseriate trichomes like those on stem, these denser along the veins; abaxial surface pale grey, more densely pubescent with trichomes like those of the upper surface evenly distributed across lamina and veins; major veins 3-6 pairs, not clearly evident abaxially; base attenuate, decurrent on the petiole; margins entire or sinuate; apex acute to obtuse; petioles (0.5-)1-1.5(-3.5) cm long, sparsely pubescent with simple uniseriate trichomes like those of the stems and leaves. Inflorescences generally internodal but appearing to arise opposite the leaves on young shoots, unbranched or rarely forked, 1-2.5(-4) cm long, with 3-7(-10) flowers clustered near the tips (sub-umbelliform), sparsely pubescent with appressed 1-2-celled simple uniseriate trichomes; peduncle 1-2.3(-4) cm long, strongly deflexed downwards in fruit; pedicels 5-10 mm long, ca. 0.5 mm in diameter at the base and 1 mm in diameter at the apex, straight and spreading, articulated at the base; pedicel scars spaced ca. 0-1 mm apart. Buds elongate-oblong, the corolla only slightly exserted from the calyx tube before anthesis. Flowers 5-merous, cosexual (hermaphroditic). Calyx tube 2-3 mm long, conical, the lobes 0.6-1.2 mm long, less than 1 mm wide, broadly deltate to triangular with acute to obtuse apices, sparsely pubescent with 1-4-celled appressed hairs like those on stem but shorter. Corolla 0.6-1.2 cm in diameter, white with a black and yellow-green central portion near the base, the black colour usually distal to the yellow green, deeply stellate, lobed 4/5 of the way to the base, the lobes 3.5-4 mm long, 1.5-1.9 mm wide, strongly reflexed at anthesis, later spreading, densely puberulent-papillate abaxially with 1-4-celled simple uniseriate trichomes, these denser on the tips and margins. Stamens equal; filament tube minute; free portion of the filaments 0.6-1 mm long, adaxially pubescent with simple tangled uniseriate 4-6-celled simple trichomes; anthers (2-)2.3-2.8 mm long, 0.5-0.8 mm wide, narrowly ellipsoid, yellow, poricidal at the tips, the pores lengthening to slits with age and drying, the connective becoming darker brown with age in dry plants. Ovary globose, glabrous; style 3.7-4.5 mm long, straight, exserted beyond the anther cone, densely pubescent with 2-3-celled simple uniseriate trichomes in the lower half where it is included in the anther cone, exserted up to 1.5 mm beyond the anther cone; stigma capitate, minutely papillate, green in live plants. Fruit a globose berry, 0.4-0.9 cm in diameter, dull purplish black at maturity, the pericarp thin, matte and somewhat glaucous, opaque, glabrous; fruiting pedicels 0.6-1.3 cm long, (0.4)0.8-1.4 mm in diameter at the base, 1-2 mm in diameter at the apex, deflexed and slightly curving, not persistent, but the downwards pointing peduncle often persistent on older stems; fruiting calyx not accrescent, the tube less than 1 mm long, the lobes 1-1.5 mm long, appressed against the berry. Seeds (13-)20-35(-50) per berry, 1.2-1.4 mm long, 1-1.2 mm wide, flattened and teardrop shaped with a subapical hilum, pale yellow, the surfaces minutely pitted, the testal cells pentagonal in outline. Stone cells absent. Chromosome number:
<emphasis id="1BD1A477A6325164F468A6F874FC4B0B" italics="true" pageId="0" pageNumber="1">2n</emphasis>
= 24 (see
<bibRefCitation id="B99B6D6B6899275D17C78D877E4C938C" DOI="https://doi.org/10.3897/phytokeys.106.21991" author="Saerkinen, T" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="1 - 223" refId="B209" refString="Saerkinen, T, Poczai, P, Barboza, GE, van der Weerden, GM, Baden, M, Knapp, S, 2018. A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae). PhytoKeys 106: 1 - 223, DOI: https://doi.org/10.3897/phytokeys.106.21991" title="A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae)." url="https://doi.org/10.3897/phytokeys.106.21991" volume="106" year="2018">
<normalizedToken id="A427CCDD463BC22B43023DF9106792EA" originalValue="Särkinen">Saerkinen</normalizedToken>
et al. 2018
</bibRefCitation>
).
</paragraph>
<caption id="EE15D485D9B152B1B7DC2F77C46A0526" doi="10.3897/phytokeys.231.100894.figure36" httpUri="https://binary.pensoft.net/fig/897497" pageId="0" pageNumber="1" start="Figure 36" startId="F36">
<paragraph id="79383FC21AFDB16DD087B2EB7B87036A" pageId="0" pageNumber="1">
<emphasis id="950A7CA307CD980B3B66509E7C02893A" bold="true" pageId="0" pageNumber="1">Figure 36.</emphasis>
<taxonomicName id="248240ADE00109687AB7EA74FA757F1B" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum chenopodioides" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="chenopodioides">
<emphasis id="1C990F4CFC0E55451A7C563266DDAB3E" italics="true" pageId="0" pageNumber="1">Solanum chenopodioides</emphasis>
</taxonomicName>
<emphasis id="E6183CFFB35E1EF01A575D8FCFDFB776" bold="true" pageId="0" pageNumber="1">A</emphasis>
habit
<emphasis id="83562BE191A5726598833C2FF4E9A1D3" bold="true" pageId="0" pageNumber="1">B</emphasis>
buds
<emphasis id="78993A3213750B672D9E63BBB271082F" bold="true" pageId="0" pageNumber="1">C</emphasis>
flowers at full anthesis
<emphasis id="26F74AD14153855D2902F0C0519856D4" bold="true" pageId="0" pageNumber="1">D</emphasis>
fully mature dull black fruits with appressed calyx lobes (
<emphasis id="25088BBB6E429513EC9EB699E453AAC8" bold="true" pageId="0" pageNumber="1">A-D</emphasis>
Nijmegen acc. # A14750051). Photos by S. Knapp and G. van der Weerden. Previously published in
<bibRefCitation id="F9B88716BE3AFD175E67F65F9A06DAD0" DOI="https://doi.org/10.3897/phytokeys.106.21991" author="Saerkinen, T" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="1 - 223" refId="B209" refString="Saerkinen, T, Poczai, P, Barboza, GE, van der Weerden, GM, Baden, M, Knapp, S, 2018. A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae). PhytoKeys 106: 1 - 223, DOI: https://doi.org/10.3897/phytokeys.106.21991" title="A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae)." url="https://doi.org/10.3897/phytokeys.106.21991" volume="106" year="2018">
<normalizedToken id="592108209A4A1C46E2132505732D5ED5" originalValue="Särkinen">Saerkinen</normalizedToken>
et al. (2018
</bibRefCitation>
: 68) and
<bibRefCitation id="BD63122D80DA5CA22486BE70D4F5C110" DOI="https://doi.org/10.3897/phytokeys.123.31738" author="Knapp, S" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="1 - 174" refId="B136" refString="Knapp, S, Barboza, GE, Bohs, L, Saerkinen, T, 2019. A revision of the Morelloid Clade of Solanum L. (Solanaceae) in the Caribbean and North and Central America. PhytoKeys 123: 1 - 174, DOI: https://doi.org/10.3897/phytokeys.123.31738" title="A revision of the Morelloid Clade of Solanum L. (Solanaceae) in the Caribbean and North and Central America." url="https://doi.org/10.3897/phytokeys.123.31738" volume="123" year="2019">Knapp et al. (2019</bibRefCitation>
: 47).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="0D7B18C755495CC34448440D1A07A119" pageId="0" pageNumber="1" type="distribution">
<paragraph id="306BFB4EFF7FD1A9010584ADDBB97C51" pageId="0" pageNumber="1">Distribution</paragraph>
<paragraph id="A083B6B5F23F80268573C10BB040A513" pageId="0" pageNumber="1">
<emphasis id="75FE5515D48C59D9F724F013BA1FE6E8" bold="true" pageId="0" pageNumber="1">
(Fig.
<figureCitation id="5199366AB4722C9D112D1C2BE9CD28BF" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Distribution map of Solanum chenopodioides in South America. For distribution elsewhere see Saerkinen et al. (2018: 70) and Knapp et al. (2019: 50)." figureDoi="10.3897/phytokeys.231.100894.figure37" httpUri="https://binary.pensoft.net/fig/897498" pageId="0" pageNumber="1">37</figureCitation>
).
</emphasis>
<taxonomicName id="D97E68EB7B073694C7016834989F69E7" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum chenopodioides" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="chenopodioides">
<emphasis id="7A3898EC2C4EF4184825B97A46228401" italics="true" pageId="0" pageNumber="1">Solanum chenopodioides</emphasis>
</taxonomicName>
is native to southern South America, and has been introduced globally, largely with the wool trade. In South America it is known from the littoral of Argentina (Provs. Buenos Aires, Chaco,
<normalizedToken id="B49B0DA58533F1BE97B62DC1C91E5017" originalValue="Córdoba">Cordoba</normalizedToken>
, Corrientes, Entre Rios, Jujuy, La Pampa, La Rioja, Mendoza,
<normalizedToken id="6058BE3AE89192AE84DD839D76589AA4" originalValue="Río">Rio</normalizedToken>
Negro, Salta, San Luis, Santa
<normalizedToken id="A635F3A529F7E8EA059C0D30569820E9" originalValue="Fé">Fe</normalizedToken>
,
<normalizedToken id="18C8A16709424383A38717A82DDCB127" originalValue="Tucumán">Tucuman</normalizedToken>
), southern Brazil (Provs. Mato Grosso do Sul, Minas Gerais, Rio de Janeiro,
<normalizedToken id="35D888135954C0C3460AD67F60A76369" originalValue="Paraná">Parana</normalizedToken>
, Rio Grande do Sul,
<normalizedToken id="8846B9BA6B71B483A5F90D3C5CAF3227" originalValue="São">Sao</normalizedToken>
Paulo and Santa Catarina), Paraguay (Depts. Amambay,
<normalizedToken id="283CAE4C189A8306B6CD7D1BA9F165D0" originalValue="Canindeyú">Canindeyu</normalizedToken>
,
<normalizedToken id="B9CEEF01979A8AB6CD513CE1D38D71BD" originalValue="Itapúa">Itapua</normalizedToken>
, Presidente Hayes) and Uruguay (Colonia, Florida, Lavalleja, Maldonado, Montevideo, Rivera, Rocha, Salto, San
<normalizedToken id="3349700EB9A7E15CC8A9F54136D96F55" originalValue="José">Jose</normalizedToken>
,
<normalizedToken id="C5ED8847BAC7DBCC6804188A4E937A57" originalValue="Tacuarembó">Tacuarembo</normalizedToken>
) with sporadic occurrences elsewhere, where it may be introduced.
</paragraph>
<caption id="F87DF1BBC66A371FBD75A704BF0CCC5F" doi="10.3897/phytokeys.231.100894.figure37" httpUri="https://binary.pensoft.net/fig/897498" pageId="0" pageNumber="1" start="Figure 37" startId="F37">
<paragraph id="D72DF302A8C8D64FDEF6B3F625301FD5" pageId="0" pageNumber="1">
<emphasis id="057A4248B3FE7091D3A516D29F00E888" bold="true" pageId="0" pageNumber="1">Figure 37.</emphasis>
Distribution map of
<taxonomicName id="BBAFDE08914CFAEAFF4B404C5228221F" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum chenopodioides" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="chenopodioides">
<emphasis id="D6881D0B62D1E40498F16C7895C3C3D0" italics="true" pageId="0" pageNumber="1">Solanum chenopodioides</emphasis>
</taxonomicName>
in South America. For distribution elsewhere see
<bibRefCitation id="7A097162C475799F86C4D2FE62AC7F5A" DOI="https://doi.org/10.3897/phytokeys.106.21991" author="Saerkinen, T" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="1 - 223" refId="B209" refString="Saerkinen, T, Poczai, P, Barboza, GE, van der Weerden, GM, Baden, M, Knapp, S, 2018. A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae). PhytoKeys 106: 1 - 223, DOI: https://doi.org/10.3897/phytokeys.106.21991" title="A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae)." url="https://doi.org/10.3897/phytokeys.106.21991" volume="106" year="2018">
<normalizedToken id="3CCB16798E5D9108446090943F1BCF9E" originalValue="Särkinen">Saerkinen</normalizedToken>
et al. (2018
</bibRefCitation>
: 70) and
<bibRefCitation id="EDA00A54E0EF3452B54B78D1B0F888C6" DOI="https://doi.org/10.3897/phytokeys.123.31738" author="Knapp, S" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="1 - 174" refId="B136" refString="Knapp, S, Barboza, GE, Bohs, L, Saerkinen, T, 2019. A revision of the Morelloid Clade of Solanum L. (Solanaceae) in the Caribbean and North and Central America. PhytoKeys 123: 1 - 174, DOI: https://doi.org/10.3897/phytokeys.123.31738" title="A revision of the Morelloid Clade of Solanum L. (Solanaceae) in the Caribbean and North and Central America." url="https://doi.org/10.3897/phytokeys.123.31738" volume="123" year="2019">Knapp et al. (2019</bibRefCitation>
: 50).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="7B97D3C787DA77749F0E4E3D48D3BB83" pageId="0" pageNumber="1" type="ecology">
<paragraph id="97BBBF02700E71D5EA24B42DBC535D5F" pageId="0" pageNumber="1">Ecology and habitat.</paragraph>
<paragraph id="86BBC839697AD68B818F556A4CF42384" pageId="0" pageNumber="1">
<taxonomicName id="596E7CD56F133FD190A00B8DA2FB14D6" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum chenopodioides" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="chenopodioides">
<emphasis id="2B3BEAEA64E084E5D9EB07F7B041FD86" italics="true" pageId="0" pageNumber="1">Solanum chenopodioides</emphasis>
</taxonomicName>
is a weedy species, growing in disturbed areas in many different vegetation types, close to urban areas and human-altered habitats; from 0 and 2,400 m elevation.
</paragraph>
</subSubSection>
<subSubSection id="46EE6D4FACD7B80ED2654B3B96639F38" pageId="0" pageNumber="1" type="common names and uses">
<paragraph id="F5F4EB4505944138C867A7B604631B21" pageId="0" pageNumber="1">Common names and uses.</paragraph>
<paragraph id="2A28E2DABB7B53E478B35C8166468721" pageId="0" pageNumber="1">
Argentina. Buenos Aires:
<normalizedToken id="06FBCC927397CC5C9B1D0A13401CB5D0" originalValue="kushú-kushú">kushu-kushu</normalizedToken>
(as
<taxonomicName id="3834025B787B347294B9D71B1B87940B" family="Solanaceae" kingdom="Plantae" lsidName="S. sublobatum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="sublobatum">
<emphasis id="A5CCAFA0361A302CFFBBCB31F6732A3B" italics="true" pageId="0" pageNumber="1">S. sublobatum</emphasis>
</taxonomicName>
,
<bibRefCitation id="12FC412B7BC3847FCB9AE4ABE67FD269" DOI="https://doi.org/10.30972/etn.0122361" author="Martinez Crovetto, R" journalOrPublisher="Etnobiologica" pageId="0" pageNumber="1" pagination="1 - 24" refId="B155" refString="Martinez Crovetto, R, 1968. Estudios etnobotanicos III. Nombres de plantas y su utilidad, segun los indios araucano-pampas del Oeste de Buenos Aires (Republica Argentina). Etnobiologica 12: 1 - 24, DOI: https://doi.org/10.30972/etn.0122361" title="Estudios etnobotanicos III. Nombres de plantas y su utilidad, segun los indios araucano-pampas del Oeste de Buenos Aires (Republica Argentina)." url="https://doi.org/10.30972/etn.0122361" volume="12" year="1968">
<normalizedToken id="EB76C8DBA88B2BFA3361F8C6FF1CE5CA" originalValue="Martínez">Martinez</normalizedToken>
Crovetto 1968
</bibRefCitation>
), yerba mora (
<emphasis id="A16D780C21CD87087120DC04D0A6E4CE" italics="true" pageId="0" pageNumber="1">Robles et al. 1869</emphasis>
);
<normalizedToken id="162E9DF2464D7C082D84AC413DC55EB8" originalValue="Córdoba">Cordoba</normalizedToken>
: yerba mora (
<emphasis id="62C7507B093290B45E900B8BF112F8F1" italics="true" pageId="0" pageNumber="1">
<normalizedToken id="C747AA36AB4EB84178F65130BDEE502E" originalValue="Müller">Mueller</normalizedToken>
et al. 308
</emphasis>
); Salta:
<normalizedToken id="6E7E27CC5CC180810265874D04AF0ABA" originalValue="iój">ioj</normalizedToken>
s(l) I s(l) I (Vilela, as
<taxonomicName id="8D34FB54FC408A2552DF28AAF228917D" family="Solanaceae" kingdom="Plantae" lsidName="S. gracile" order="Solanales" pageId="0" pageNumber="1" rank="species" species="gracile">
<emphasis id="C2231A0A4AC10539C3AFBA8D8B35C11D" italics="true" pageId="0" pageNumber="1">S. gracile</emphasis>
</taxonomicName>
,
<bibRefCitation id="9497CADE7CA632DFE0FDC0D42ECF1093" DOI="https://doi.org/10.30972/bon.21-41531" author="Martinez Crovetto, R" journalOrPublisher="Bonplandia" pageId="0" pageNumber="1" pagination="1 - 28" refId="B154" refString="Martinez Crovetto, R, 1965. Estudios etnobotanicos II. Nombres de plantas y su utilidad, segun los indios Vilelas del Chaco. Bonplandia 2 (1): 1 - 28, DOI: https://doi.org/10.30972/bon.21-41531" title="Estudios etnobotanicos II. Nombres de plantas y su utilidad, segun los indios Vilelas del Chaco." url="https://doi.org/10.30972/bon.21-41531" volume="2" year="1965">
<normalizedToken id="662BAE8FF2E336E9691548839D0BB5AE" originalValue="Martínez">Martinez</normalizedToken>
Crovetto 1965
</bibRefCitation>
). Uruguay. Montevideo: yerba mora (
<emphasis id="E4586D8CF0A94F9A7476A8DB3351E432" italics="true" pageId="0" pageNumber="1">Barattini s.n.</emphasis>
).
<taxonomicName id="7F97D7C53C207AA094BC4F334FFB6803" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum chenopodioides" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="chenopodioides">
<emphasis id="11DAE0F7E2CAF42F9816A31150114CF7" italics="true" pageId="0" pageNumber="1">Solanum chenopodioides</emphasis>
</taxonomicName>
is considered to be toxic for cattle due to the high solanidine content of unripe fruits (
<bibRefCitation id="CEFA584645F8EA95218DAC4A4DC5965D" author="Marzocca, A" journalOrPublisher="Miscelanea - Fundacion Miguel Lillo" pageId="0" pageNumber="1" refId="B157" refString="Marzocca, A, 1994. Guia Descriptiva de Malezas del Cono Sur. INTA, Buenos Aires." title="Guia Descriptiva de Malezas del Cono Sur. INTA, Buenos Aires." year="1994">Marzocca 1994</bibRefCitation>
), but as fruits ripen the alkaloid content decreases and fruits are apparently eaten without effect (
<bibRefCitation id="7CC6B0CB6596FA8EDE2A5BDC39A29DCE" author="Gallo, GG" journalOrPublisher="American Journal of Botany" pageId="0" pageNumber="1" refId="B93" refString="Gallo, GG, 1987. Plantas toxicas para el ganado en el Cono Sur de America (2nd edn.). Editiones Hemisferio Sur S.A., Buenos Aires." title="Plantas toxicas para el ganado en el Cono Sur de America (2 nd edn.). Editiones Hemisferio Sur S. A., Buenos Aires." year="1987">Gallo 1987</bibRefCitation>
). In Argentina, fruits of
<taxonomicName id="4DE68F8A37744C1193DAC2B80087F746" family="Solanaceae" kingdom="Plantae" lsidName="S. chenopodioides" order="Solanales" pageId="0" pageNumber="1" rank="species" species="chenopodioides">
<emphasis id="4BFCEC3340B913B5399D31C847807272" italics="true" pageId="0" pageNumber="1">S. chenopodioides</emphasis>
</taxonomicName>
are eaten by children of the Araucarian peoples (
<bibRefCitation id="E3A3B7F21EE3023C1B68A83F89E50498" DOI="https://doi.org/10.30972/etn.0122361" author="Martinez Crovetto, R" journalOrPublisher="Etnobiologica" pageId="0" pageNumber="1" pagination="1 - 24" refId="B155" refString="Martinez Crovetto, R, 1968. Estudios etnobotanicos III. Nombres de plantas y su utilidad, segun los indios araucano-pampas del Oeste de Buenos Aires (Republica Argentina). Etnobiologica 12: 1 - 24, DOI: https://doi.org/10.30972/etn.0122361" title="Estudios etnobotanicos III. Nombres de plantas y su utilidad, segun los indios araucano-pampas del Oeste de Buenos Aires (Republica Argentina)." url="https://doi.org/10.30972/etn.0122361" volume="12" year="1968">
<normalizedToken id="D25EF969775570752FA539E549F3DF2D" originalValue="Martínez">Martinez</normalizedToken>
Crovetto 1968
</bibRefCitation>
) and the Vilela people use them as a purple dye (
<bibRefCitation id="1BDD9F83A0510FE2F6C42D02B38DFD04" DOI="https://doi.org/10.30972/bon.21-41531" author="Martinez Crovetto, R" journalOrPublisher="Bonplandia" pageId="0" pageNumber="1" pagination="1 - 28" refId="B154" refString="Martinez Crovetto, R, 1965. Estudios etnobotanicos II. Nombres de plantas y su utilidad, segun los indios Vilelas del Chaco. Bonplandia 2 (1): 1 - 28, DOI: https://doi.org/10.30972/bon.21-41531" title="Estudios etnobotanicos II. Nombres de plantas y su utilidad, segun los indios Vilelas del Chaco." url="https://doi.org/10.30972/bon.21-41531" volume="2" year="1965">
<normalizedToken id="620AA10497F2DB31E9BE236E06BDD9B7" originalValue="Martínez">Martinez</normalizedToken>
Crovetto 1965
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="80653745DAF4AF70AE919057266FD338" pageId="0" pageNumber="1" type="preliminary conservation status">
<paragraph id="309B3FC05532613EE06691BC578115C7" pageId="0" pageNumber="1">Preliminary conservation status</paragraph>
<paragraph id="A197056CC5DD69630EBCC14C57587E8B" pageId="0" pageNumber="1">
<emphasis id="FE0CB616876461E1CDAC4123C89BDE24" bold="true" pageId="0" pageNumber="1">
(
<bibRefCitation id="7B02280F489391578F4CAC6ED0C75D4F" author="Ibarrola, DA" journalOrPublisher="Nueva Serie" pageId="0" pageNumber="1" publicationUrl="https://www.iucnredlist.org/resources/redlistguidelines" refId="B116" refString="2022. . https://www.iucnredlist.org/resources/redlistguidelines" url="https://www.iucnredlist.org/resources/redlistguidelines" year="2022">IUCN 2022</bibRefCitation>
).
</emphasis>
Least Concern [LC]. Worldwide distribution: EOO = 95,008,211 km2 [LC]; AOO = 1,560 km2 [LC].
<taxonomicName id="7507EA7DCC9ECA43B9A4CD996E8B2F0E" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum chenopodioides" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="chenopodioides">
<emphasis id="BE5638035441546BE02884274955E253" italics="true" pageId="0" pageNumber="1">Solanum chenopodioides</emphasis>
</taxonomicName>
is a widespread weed of disturbed areas (see
<bibRefCitation id="56313DE16ADEC005DF9ED511515DCF22" author="Barboza, GE" editor="Anton, AM" journalOrPublisher=") Flora Argentina (Vol. 13), Solanaceae. IOBDA- IMBIV, CONICET: Buenos Aires &amp; Cordoba, Argentina" pageId="0" pageNumber="1" pagination="231 - 264" refId="B10" refString="Barboza, GE, Knapp, S, Saerkinen, T, 2013. Grupo VII. Moreloide. In: Anton, AM, Zuloaga, FO, Eds., Barboza GE (Coord.) Flora Argentina (Vol. 13), Solanaceae. IOBDA- IMBIV, CONICET: Buenos Aires &amp; Cordoba, Argentina: 231 - 264" title="Grupo VII. Moreloide." volumeTitle="Barboza GE (Coord." year="2013">Barboza et al. 2013</bibRefCitation>
;
<bibRefCitation id="2D49B9ACDAF3C0A982DDAAB8EA2FC2CE" DOI="https://doi.org/10.3897/phytokeys.106.21991" author="Saerkinen, T" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="1 - 223" refId="B209" refString="Saerkinen, T, Poczai, P, Barboza, GE, van der Weerden, GM, Baden, M, Knapp, S, 2018. A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae). PhytoKeys 106: 1 - 223, DOI: https://doi.org/10.3897/phytokeys.106.21991" title="A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae)." url="https://doi.org/10.3897/phytokeys.106.21991" volume="106" year="2018">
<normalizedToken id="D14825CA33CF3F5DDD6937C0CC4340BB" originalValue="Särkinen">Saerkinen</normalizedToken>
et al. 2018
</bibRefCitation>
;
<bibRefCitation id="CC7020698FFD1927B9EA378CF73FC353" DOI="https://doi.org/10.3897/phytokeys.123.31738" author="Knapp, S" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="1 - 174" refId="B136" refString="Knapp, S, Barboza, GE, Bohs, L, Saerkinen, T, 2019. A revision of the Morelloid Clade of Solanum L. (Solanaceae) in the Caribbean and North and Central America. PhytoKeys 123: 1 - 174, DOI: https://doi.org/10.3897/phytokeys.123.31738" title="A revision of the Morelloid Clade of Solanum L. (Solanaceae) in the Caribbean and North and Central America." url="https://doi.org/10.3897/phytokeys.123.31738" volume="123" year="2019">Knapp et al. 2019</bibRefCitation>
) and is widely introduced outside of and very common within its native range.
</paragraph>
</subSubSection>
<subSubSection id="928832E3DEE9408AC0B2422D97913FA5" pageId="0" pageNumber="1" type="discussion">
<paragraph id="D427AA6B7AAC6638E3CBDB70F222509E" pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph id="D2397C826A6962BCD5CA5094E678BC2A" pageId="0" pageNumber="1">
<taxonomicName id="687C30CFD0342E4B406465498F13DACD" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum chenopodioides" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="chenopodioides">
<emphasis id="AFA2D3128F6F8C5D72362C4B20BC6AD4" italics="true" pageId="0" pageNumber="1">Solanum chenopodioides</emphasis>
</taxonomicName>
is a weedy, ruderal species occurring in open disturbed areas throughout its range. It is somewhat similar morphologically to
<taxonomicName id="940F09B05ED8F4D0BB1F918D1105295E" family="Solanaceae" kingdom="Plantae" lsidName="S. pilcomayense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="pilcomayense">
<emphasis id="FD3B60478E4526631BE4C430129F399F" italics="true" pageId="0" pageNumber="1">S. pilcomayense</emphasis>
</taxonomicName>
, with which it is sympatric in Argentina, but differs in its elliptic leaves with acute to attenuate bases (versus triangular leaves with truncate to hastate bases), smaller anthers (2-2.8 mm long versus 3-4 mm long), and deltate or triangular versus spathulate calyx lobes. The fruiting peduncle of
<taxonomicName id="C7BDB2401C13D38EE9218F2915FA85E9" family="Solanaceae" kingdom="Plantae" lsidName="S. chenopodioides" order="Solanales" pageId="0" pageNumber="1" rank="species" species="chenopodioides">
<emphasis id="E03BA945D839EC3BF7B4E237E3142F82" italics="true" pageId="0" pageNumber="1">S. chenopodioides</emphasis>
</taxonomicName>
bends downwards at the base so it is held at an angle of ca. 45-degree with respect to the stem (see Figs
<figureCitation id="C455BC38A20636B65983565524998C5A" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Solanum chenopodioides A habit B detail of adaxial leaf surface C detail of abaxial leaf surface D opening bud E dissected flower F fruiting branch G detail of infructescence H maturing fruit I fully mature fruit (A-E Fox s. n. F-I Hieronymus s. n.). Illustration by R. Wise. Previously published in Barboza et al. (2013: 242), Saerkinen et al. (2018: 67) and Knapp et al. (2019: 46)." figureDoi="10.3897/phytokeys.231.100894.figure35" httpUri="https://binary.pensoft.net/fig/897496" pageId="0" pageNumber="1">35G</figureCitation>
,
<figureCitation id="4D236033467313EA549F0B97B9D1B2B3" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Solanum chenopodioides A habit B buds C flowers at full anthesis D fully mature dull black fruits with appressed calyx lobes (A-D Nijmegen acc. # A 14750051). Photos by S. Knapp and G. van der Weerden. Previously published in Saerkinen et al. (2018: 68) and Knapp et al. (2019: 47)." figureDoi="10.3897/phytokeys.231.100894.figure36" httpUri="https://binary.pensoft.net/fig/897497" pageId="0" pageNumber="1">36D</figureCitation>
), but this character is not always obvious in herbarium specimens. Anthers in
<taxonomicName id="B378258516F8B4380C1F7D22C1F92611" family="Solanaceae" kingdom="Plantae" lsidName="S. chenopodioides" order="Solanales" pageId="0" pageNumber="1" rank="species" species="chenopodioides">
<emphasis id="645E2E42B39804EBF398A8ECECDA8640" italics="true" pageId="0" pageNumber="1">S. chenopodioides</emphasis>
</taxonomicName>
are always much longer (2-2.8 mm) than in
<taxonomicName id="66300EA3BE896B3409E11B08C7DC22AE" family="Solanaceae" kingdom="Plantae" lsidName="S. americanum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="americanum">
<emphasis id="75B0C4C2A8510693F65C2FFCF6EFC924" italics="true" pageId="0" pageNumber="1">S. americanum</emphasis>
</taxonomicName>
(0.8-1.5 mm), and the berries are matte (versus shiny) in texture and always lack stone cells (versus often with 2-4 stone cells per berry in
<taxonomicName id="412F47562E73D563CC38A98BB3ECBAC4" family="Solanaceae" kingdom="Plantae" lsidName="S. americanum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="americanum">
<emphasis id="284CCE76B406812CC6C30E05618FF769" italics="true" pageId="0" pageNumber="1">S. americanum</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="88388924608D7E3CF38BE1BDDED3F6F6" pageId="0" pageNumber="1">
Typification details for the synonyms of
<taxonomicName id="676377E9B66E79D6B7B143011DF73084" family="Solanaceae" kingdom="Plantae" lsidName="S. chenopodioides" order="Solanales" pageId="0" pageNumber="1" rank="species" species="chenopodioides">
<emphasis id="E5C2CF0EBA188B0AF28025870C2855A9" italics="true" pageId="0" pageNumber="1">S. chenopodioides</emphasis>
</taxonomicName>
and a more comprehensive discussion of its worldwide distribution as a weed of wool waste used in agriculture can be found in
<bibRefCitation id="648792CA21EC9FD3F48BB5C15865326F" DOI="https://doi.org/10.3897/phytokeys.106.21991" author="Saerkinen, T" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="1 - 223" refId="B209" refString="Saerkinen, T, Poczai, P, Barboza, GE, van der Weerden, GM, Baden, M, Knapp, S, 2018. A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae). PhytoKeys 106: 1 - 223, DOI: https://doi.org/10.3897/phytokeys.106.21991" title="A revision of the Old World black nightshades (Morelloid clade of Solanum L., Solanaceae)." url="https://doi.org/10.3897/phytokeys.106.21991" volume="106" year="2018">
<normalizedToken id="1516139E1344CB34EBF20950D043EBE7" originalValue="Särkinen">Saerkinen</normalizedToken>
et al. (2018)
</bibRefCitation>
and
<bibRefCitation id="28979A1244DEF8A75452F0E7B81EF437" DOI="https://doi.org/10.3897/phytokeys.123.31738" author="Knapp, S" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="1 - 174" refId="B136" refString="Knapp, S, Barboza, GE, Bohs, L, Saerkinen, T, 2019. A revision of the Morelloid Clade of Solanum L. (Solanaceae) in the Caribbean and North and Central America. PhytoKeys 123: 1 - 174, DOI: https://doi.org/10.3897/phytokeys.123.31738" title="A revision of the Morelloid Clade of Solanum L. (Solanaceae) in the Caribbean and North and Central America." url="https://doi.org/10.3897/phytokeys.123.31738" volume="123" year="2019">Knapp et al. (2019)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>