154 lines
18 KiB
XML
154 lines
18 KiB
XML
<document id="D8B35608058C15F153074D2C0A118D33" ID-DOI="http://dx.doi.org/10.3897/nl.39.6760" ID-GBIF-Dataset="a61fd129-b970-45d9-8e52-c87b2d158fde" ID-Pensoft-Pub="2367-5365-1-13" ID-ZooBank="2C6F4BBA55404EA7ACA644A99BA71023" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="2367-5365-1-13" ModsDocOrigin="Nota Lepidopterologica 39 (1)" ModsDocTitle="Taxonomy of two montane Dichrorampha species from the Balkans and Caucasus (Lepidoptera, Tortricidae)" checkinTime="1454707307449" checkinUser="pensoft" docAuthor="Zlatkov, Boyan" docDate="2016" docId="3EF38246B52B5BC231186EF0E3EC8D3D" docLanguage="en" docName="NotaLepidopt 39(1): 13-20" docOrigin="Nota Lepidopterologica 39 (1)" docSource="http://dx.doi.org/10.3897/nl.39.6760" docTitle="Dichrorampha sakartvelana Zlatkov, 2016, sp. n." docType="treatment" docUuid="1919B149-1E92-49DA-90BB-4E1689F9C42D" docUuidSource="ZooBank" docVersion="6" lastPageNumber="19" masterDocId="FFBFFFB7BC69DB7D0735FFC5FFE8FFEC" masterDocTitle="Taxonomy of two montane Dichrorampha species from the Balkans and Caucasus (Lepidoptera, Tortricidae)" masterLastPageNumber="20" masterPageNumber="13" pageNumber="16" updateTime="1701316646520" updateUser="plazi">
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<mods:title id="0F6982B20CB310C3190D5099DE30F1F0">Taxonomy of two montane Dichrorampha species from the Balkans and Caucasus (Lepidoptera, Tortricidae)</mods:title>
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<mods:namePart id="83A5C427AC97C82B2B9D31B533E017D6">Zlatkov, Boyan</mods:namePart>
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<mods:title id="309E2D7A5CA5468C32974595A327F0D3">Nota Lepidopterologica</mods:title>
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<mods:date id="D3A4A5CB474088013D31F8C5EC4C0A2C">2016</mods:date>
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<mods:number id="1AB26C986A69B60C1E3171813D99E679">39</mods:number>
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<mods:url id="FE496CBFA9B766801A5A0C36B8E70A55">http://dx.doi.org/10.3897/nl.39.6760</mods:url>
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<mods:identifier id="9F8309B9CC3D5A007DEB296435E6C450" type="DOI">http://dx.doi.org/10.3897/nl.39.6760</mods:identifier>
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<treatment id="3EF38246B52B5BC231186EF0E3EC8D3D" ID-GBIF-Taxon="127896411" LSID="urn:lsid:zoobank.org:act:1919B149-1E92-49DA-90BB-4E1689F9C42D" httpUri="http://treatment.plazi.org/id/3EF38246B52B5BC231186EF0E3EC8D3D" lastPageId="6" lastPageNumber="19" pageId="3" pageNumber="16">
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<subSubSection id="7782B61F6BD03690F331F6A61CB9455B" pageId="3" pageNumber="16" type="multiple">
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<paragraph id="1061A4B1EE101BA78490C70120ABD968" pageId="3" pageNumber="16">Taxon classification Animalia Lepidoptera Tortricidae</paragraph>
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</subSubSection>
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<subSubSection id="8D15FDB079D5569BB3C1D57B3A90B7B1" pageId="3" pageNumber="16" type="nomenclature">
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<paragraph id="944001DBFF7A511DFA3A578326D2824A" pageId="3" pageNumber="16">
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<taxonomicName id="F56247F9B39A4B482E042E6FE5CEA3F2" ID-CoL="B4VDW" LSID="http://zoobank.org/1919B149-1E92-49DA-90BB-4E1689F9C42D" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha sakartvelana" order="Lepidoptera" pageId="3" pageNumber="16" phylum="Arthropoda" rank="species" species="sakartvelana">Dichrorampha sakartvelana</taxonomicName>
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<taxonomicNameLabel id="8B3A3E66FB3A5449A93273345A020AE4" pageId="3" pageNumber="16">sp. n.</taxonomicNameLabel>
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Figs 5-6, 11-13, 15
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</paragraph>
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</subSubSection>
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<subSubSection id="C1BF1631391F56C3786526E8625A9F41" pageId="4" pageNumber="17" type="materials_examined">
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<paragraph id="2D0C32A6B9C2C04B3019525D01F5E14C" pageId="4" pageNumber="17">
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<pageBreakToken id="44BE089DC12D5D608AED36BF3E2E1A61" pageId="4" pageNumber="17" start="start">Material</pageBreakToken>
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.
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</paragraph>
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<paragraph id="DFAF777C9564F7CC1FD7EC81D7D6719D" pageId="4" pageNumber="17">
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Holotype ♀, with three labels: "Georgia, Great Caucasus | Mountains, Caucasus Range, | near Abano pass | 2280 m
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<geoCoordinate id="713029FF4FC51129A48992F4200EB348" direction="north" orientation="latitude" precision="15" value="42.265">
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N 42°15
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<normalizedToken id="0EB6521DFA7ABC3C0A66BC489F5CBC8B" originalValue="’54’’">'54''</normalizedToken>
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</geoCoordinate>
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<geoCoordinate id="FF57553EDF7CA8B61928E2549793C6E7" direction="east" orientation="longitude" precision="15" value="45.504723">
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E 45°30
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<normalizedToken id="FC028BDD867BC977B1C3CB7F52FF993C" originalValue="’17’’">'17''</normalizedToken>
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</geoCoordinate>
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| 27.vii.2014, netting | leg. B. Zlatkov", "Holotypus ♀ |
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<taxonomicName id="2824CD27F8EC683A80CF917AF48513E2" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha" order="Lepidoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="genus">Dichrorampha</taxonomicName>
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<taxonomicName id="3CD765032BEE343437D4E8E392916C2D" lsidName="sakartvelana" pageId="4" pageNumber="17" rank="species" species="sakartvelana">sakartvelana</taxonomicName>
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| det. B. Zlatkov 2015 [red label]", " BFUS | ♀ | Genitalia slide | No. 27.vii.2014/1", BFUS. - Paratype ♂, with three labels: the first as in Holotype; the other two: "Paratypus ♂ |
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<taxonomicName id="5799450E8A5805B8DD657CE6DF199801" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha" order="Lepidoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="genus">Dichrorampha</taxonomicName>
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<taxonomicName id="8C9B387291133D4EB9E672043FA058BB" lsidName="sakartvelana" pageId="4" pageNumber="17" rank="species" species="sakartvelana">sakartvelana</taxonomicName>
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| det. B. Zlatkov 2015 [red label]", "NMNHS | ♂ | Genitalia slide | No. 27.vii.2014/2", NMNHS.
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</paragraph>
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</subSubSection>
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<subSubSection id="A663CD0F3F98FE2D66DCEBE18CEBBAFC" lastPageId="6" lastPageNumber="19" pageId="4" pageNumber="17" type="description">
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<paragraph id="C94C7B0D4292D2C0BBA15FA2312B336D" pageId="4" pageNumber="17">Description.</paragraph>
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<paragraph id="86CEDE0A4E8E976B95249F0BC2D8BB16" lastPageId="5" lastPageNumber="18" pageId="4" pageNumber="17">
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Adult (Figs 5-6). Sexual dimorphism subtle. Head: Frons and vertex covered with brown-grey scales. Palpus labialis with whitish basal and brown-grey distal segment; the second segment with whitish base and brown tuft at the distal end. Antennae covered with dark grey scales. Thorax: Nota, patagia and tegulae uniformly grey, thorax underside (including coxae) whitish, legs brown. Forewings comparatively wide, in male with costal fold with 1/5 of the length of the costal edge. Forewing length male 8.65, female 8.90 mm, wingspan in set specimens
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<pageBreakToken id="50A4CE7CA595DE8F95DF25C1F5173C90" pageId="5" pageNumber="18" start="start">18.5</pageBreakToken>
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-19 mm. The specimens are worn, but preserved areas on the forewing upperside have pale greenish-grey overlaying scales which apparently do not form an obvious pattern. No markings are visible with the exception of five pairs of faint distal costal strigulae (pairs 5-9 sensu Baixeras, 2002). Three black terminal dots are present. Underside uniformly brown. Cilia creamy with pale brown margin. Hindwings with pale brown upperside and whitish underside. Cilia whitish. Abdomen covered with pale grey scales.
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</paragraph>
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<paragraph id="8D042B4FBB60CA24782E0FC5A53C6672" pageId="5" pageNumber="18">Male genitalia (Figs 11-12): Tegumen bearing a small lobe as uncus. Valva broad basally with wide basal cavity. Costal edge slightly convex. Sacculus nearly parallel to costal edge, indistinctly concave, ending with nearly straight angle. Ventral incision elongated, trapezoidal. Neck of valva slender, more than two times narrower than the basal part of valva and relatively long, 3/4 of its length. Cucullus with large dorsal lobe, densely covered with long setae, and a small rectangular ventral prominence. Phallus slender, ca. 3/5 of the length of valva, bent ventrally in the basal part, with membranous area extending at first on right, then on dorsal side. A large triangular prominence pointed dorsally at the right side of the tip is present. Circa 20 sockets of deciduous cornuti are counted.</paragraph>
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<caption id="2B14586189549D036DAFF5E37E604032" pageId="5" pageNumber="18">
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<paragraph id="80747E2128DB6CA25637FDD3D9B9F9FB" pageId="5" pageNumber="18">
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Figures 11-13. Genitalia of
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<taxonomicName id="DCDC3F6DF113F840DB5A7308B8D39C77" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha sakartvelana" order="Lepidoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="sakartvelana">Dichrorampha sakartvelana</taxonomicName>
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sp. n. 11. Male genitalia, paratype; 12. Phallus, dorsal view, paratype; 13. Female genitalia, holotype. Scale bar: 0.5 mm.
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</paragraph>
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</caption>
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<paragraph id="7AFA3E58E278220ADE9E3C2E06AA5C51" pageId="6" pageNumber="19">
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<pageBreakToken id="6182775A53099AD640BCFC648A880BF6" pageId="6" pageNumber="19" start="start">Female</pageBreakToken>
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genitalia (Fig. 13): Papillae anales wide. Apophyses posteriores equal in length to the apophyses anteriores, the latter look naturally deformed. Sterigma sclerotised, slightly asymmetrical, trapezoidal, with two incisions on the posterior margin of the postostial part. Ostium wide. Subgenital plate trapezoidal with distinctly sclerotised lateroposterior margins and rounded posterior angles. Antrum with the length of the membranous part of ductus bursae, well sclerotised, remarkably wide, nearly symmetrical, wineglass shaped and enveloped in a thin cuticular membrane visible after staining. The proximal sclerotisation of ductus bursae is barely discernible only under higher magnification. Ductus seminalis emerging at the middle of the membranous ductus bursae. Corpus bursae ovoid. A single moderately sized signum is present.
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</paragraph>
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</subSubSection>
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<subSubSection id="A31B6BC5BA7D1DF68B85759A02167A3D" pageId="6" pageNumber="19" type="diagnosis">
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<paragraph id="025F79571177B3ABCB851C89539561A9" pageId="6" pageNumber="19">Diagnosis.</paragraph>
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<paragraph id="737E42E23F242CFF5BEF1128BC3D466B" pageId="6" pageNumber="19">
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The new species is characterized externally by uniformly coloured forewings (though the specimens are not fresh) and large size. The male genitalia resemble some species of the "section petiverellae" (sensu
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<bibRefCitation id="B4828BC9E589DA90FAC1F11FAD1459CA" author="Danilevsky, A" editor="Byhovsky, BE" journalOrPublisher="Nauka, Leningrad" pageId="7" pageNumber="20" title="Listoviortki-Tortricidae triba plodozhorki Laspeyresiini [Tortrix Moths Tortricidae: Tribe Laspeyresiini]." volumeTitle="Fauna SSSR. Nasekomyie cheshuyekrylyie" year="1968">Danilevsky and Kuznetzov 1968</bibRefCitation>
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), which apparently are closely related to
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<taxonomicName id="B09A26B01620D5D776836FF0C6914A88" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha sakartvelana" order="Lepidoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="sakartvelana">Dichrorampha sakartvelana</taxonomicName>
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sp. n.
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<taxonomicName id="CC81646329F65880414A3F4B6B8DF0B0" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha flavidorsana" order="Lepidoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="flavidorsana">Dichrorampha flavidorsana</taxonomicName>
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Knaggs, 1867 and
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<taxonomicName id="D8D16A02CA6929CCF387C0B5FFDE05C9" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha filipjevi" order="Lepidoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="filipjevi">Dichrorampha filipjevi</taxonomicName>
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(Danilevsky, 1948) have similarly shaped valva, but the apical prominence of the phallus is pointed ventrally, and the ventral process of the cucullus is much larger in
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<taxonomicName id="01C77090D1895077B919BC2E4772C96A" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha filipjevi" order="Lepidoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="filipjevi">Dichrorampha filipjevi</taxonomicName>
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.
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<taxonomicName id="74AD1782A3E4A06FD4C07BE369B0EFE7" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha petiverella" order="Lepidoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="petiverella">Dichrorampha petiverella</taxonomicName>
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(Linnaeus, 1758) and
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<taxonomicName id="905A4DC652A1A0DDD6362F0B15081105" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha proxima" order="Lepidoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="proxima">Dichrorampha proxima</taxonomicName>
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(Danilevsky, 1948) are also similar in general, but they have two distinct processes of the cucullus. Female genitalia do not demonstrate clear affinities to any Palaearctic
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<taxonomicName id="1DA805E9D0C95A6B62C4E65768463574" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha" order="Lepidoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="genus">Dichrorampha</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="3FEA0A436BD7D2EC656A80D83AC36C88" pageId="6" pageNumber="19" type="preimaginal stages">
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<paragraph id="FED93C4E9ACB04F2DF4890F3E0885A20" pageId="6" pageNumber="19">Preimaginal stages.</paragraph>
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<paragraph id="DCD5E951EE3A8B64B0DF18768A1B1DDA" pageId="6" pageNumber="19">Unknown.</paragraph>
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</subSubSection>
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<subSubSection id="DD8E3BB8D71A4FF46F28E818741FEED7" pageId="6" pageNumber="19" type="biology">
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<paragraph id="04F816746F2BCB6D8AAFA123CAF1005E" pageId="6" pageNumber="19">Biology.</paragraph>
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<paragraph id="70902CBFCEC9754CFDC6052CBA046F0E" pageId="6" pageNumber="19">
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The moths were collected at the end of July, but their condition presumes that they are on the wing earlier. The larval host plant is most probably
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<taxonomicName id="8DB9CF14BD1487C52646ED874C0E1CFF" class="Magnoliopsida" family="Asteraceae" genus="Achillea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Achillea" order="Asterales" pageId="6" pageNumber="19" phylum="Tracheophyta" rank="genus">Achillea</taxonomicName>
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sp. above which the moths were flying (plant material was not preserved and an exact identification is thus not possible). The habitat (Fig. 15) is a subalpine meadow at an elevation of ca. 2300 m with denuded rocks where the host plant grows.
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</paragraph>
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<caption id="DB56B6308D30EC76F42971C6FF34B409" pageId="6" pageNumber="19">
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<paragraph id="3A3DAFA60D03277E14E079D2E36AAF68" pageId="6" pageNumber="19">
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Figures 14-15. Habitats. 14. Habitat of
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<taxonomicName id="2FDAFCDF802044041636334DB6FC1F90" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha pentheriana" order="Lepidoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="pentheriana">Dichrorampha pentheriana</taxonomicName>
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(Rebel, 1917), Vitosha Mts, 13.vii.2012, foreground: the suspected larval host plant
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<taxonomicName id="A02AE66B1F553423B724E8A74218821F" class="Magnoliopsida" family="Asteraceae" genus="Achillea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Achillea lingulata" order="Asterales" pageId="6" pageNumber="19" phylum="Tracheophyta" rank="species" species="lingulata">Achillea lingulata</taxonomicName>
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(photo: D. Gradinarov); 15. Habitat of
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<taxonomicName id="F0E40C66E7F7D07BC0B3388E3F9DA38D" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha sakartvelana" order="Lepidoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="sakartvelana">Dichrorampha sakartvelana</taxonomicName>
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sp. n., Great Caucasus Mts, Tusheti Range, 27.vii.2014 (photo: M. Ilieva).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="A7BA8EF7099916CE3B26DCD3F1D09870" pageId="6" pageNumber="19" type="distribution">
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<paragraph id="E52628459B9FCA3F5BE5F8E94471053D" pageId="6" pageNumber="19">Distribution.</paragraph>
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<paragraph id="9CFF7631A9A7EBAC2A8B281E67371EA2" pageId="6" pageNumber="19">Known from the type locality only.</paragraph>
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</subSubSection>
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<subSubSection id="1E5289C3FB5DECBBA934B50E2AE5F00E" pageId="6" pageNumber="19" type="etymology">
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<paragraph id="99F816AF9AF76CE2F3F5681B767445B7" pageId="6" pageNumber="19">Etymology.</paragraph>
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<paragraph id="735C24F9DE45849A1A0F5088B530CCEC" pageId="6" pageNumber="19">
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The name of the species is an adjective, derived from the autonym for Georgia, Sakartvelo, and the specific ending for
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<taxonomicName id="854FA6C211148404D9A944443CA02256" family="Tortricidae" lsidName="" pageId="6" pageNumber="19" rank="family">Tortricidae</taxonomicName>
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-ana.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |