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<mods:title id="687D07B1725C3599791C3FAFC1790C13">A review of Paratranes Zimmerman, 1994, Xanthorrhoea-associated weevils of the Tranes group (Coleoptera, Curculionidae, Molytinae), with description of a new species</mods:title>
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<figureCitation id="218D8088FFCDFF9D0B90520CFCFF9C1D" box="[735,852,815,842]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStart-7="Fig" captionStart-8="Fig" captionStart-9="Fig" captionStartId-0="3.[189,232,1920,1946]" captionStartId-1="4.[191,234,1958,1984]" captionStartId-2="7.[189,232,1952,1978]" captionStartId-3="8.[189,232,1952,1978]" captionStartId-4="10.[190,233,1948,1974]" captionStartId-5="11.[189,232,1852,1878]" captionStartId-6="12.[189,232,1852,1878]" captionStartId-7="16.[189,232,1884,1910]" captionStartId-8="17.[191,234,1879,1905]" captionStartId-9="18.[191,234,1879,1905]" captionTargetBox-0="[189,1398,493,1882]" captionTargetBox-1="[189,1375,733,1900]" captionTargetBox-2="[292,1272,288,1838]" captionTargetBox-3="[206,1347,314,1844]" captionTargetBox-4="[199,1398,879,1841]" captionTargetBox-5="[189,1398,605,1814]" captionTargetBox-6="[283,1309,267,1783]" captionTargetBox-7="[340,1238,280,1818]" captionTargetBox-8="[189,1398,947,1841]" captionTargetBox-9="[197,1395,506,1801]" captionTargetId-0="figure-88@3.[189,1398,493,1882]" captionTargetId-1="figure-171@4.[189,1398,732,1920]" captionTargetId-2="figure-12@7.[274,1314,265,1915]" captionTargetId-3="figure-14@8.[189,1398,293,1886]" captionTargetId-4="figure-171@10.[189,1398,879,1911]" captionTargetId-5="figure-88@11.[189,1398,602,1815]" captionTargetId-6="figure-14@12.[256,1332,265,1815]" captionTargetId-7="figure-15@16.[317,1271,265,1843]" captionTargetId-8="figure-205@17.[189,1398,947,1841]" captionTargetId-9="figure-85@18.[189,1398,490,1841]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionTargetPageId-7="16" captionTargetPageId-8="17" captionTargetPageId-9="18" captionText-0="Fig. 1. Diagnostic external characters of Paratranes Zimmerman, 1994. A. Elytral surface and setae. B. Eyes and prothorax, ventral view. C. Head and base of rostrum, dorsal view. D. Pronotal surface. Scale bars = 1.0 mm." captionText-1="Fig. 2. Diagnostic external characters and terminalia of Paratranes Zimmerman, 1994. A. Elytral apex. B. Right metatibia. C. Tergite VII of female. D. Tergite VIII of male. Scale bars = 0.5 mm." captionText-2="Fig. 3. Habitus, dorsal view. A–B. Paratranes monopticus (Pascoe, 1870). C–D. P. zimmermani sp. nov. (C. Holotype (ANIC); D. Paratype (ANIC)). A, C. Male. B, D. Female. Scale bars = 5.0 mm." captionText-3="Fig. 4. Habitus, lateral view. A–B. Paratranes monopticus (Pascoe, 1870). C–D. P. zimmermani sp. nov. (C. Holotype (ANIC); D. Paratype (ANIC)). A, C. Male. B, D. Female. Scale bars = 5.0 mm." captionText-4="Fig. 5. Left antenna, dorsal view. A. Paratranes monopticus (Pascoe, 1870). B. P. zimmermani sp. nov. Scale bars = 0.5 mm." captionText-5="Fig. 6. Diagnostic differences of external characters between Paratranes monopticus (Pascoe, 1870) and P. zimmermani sp. nov.A. P. monopticus, prosternum.B. P. zimmermani sp. nov., ditto. C. P. monopticus, profemur. D. P. zimmermani sp. nov., ditto. E. P. monopticus, protibial uncus, indicated by black arrow. F. P. zimmermani sp. nov., ditto. G. P. monopticus, terminal tarsal segment with claws, lateral view. H. P. zimmermani sp. nov., ditto. Scale bars: A–D = 0.5 mm; E–H = 0.2 mm." captionText-6="Fig. 7. Terminalia of Paratranes Zimmerman, 1994. A. P. monopticus (Pascoe, 1870), tergite VIII of female. B. P. monopticus, sternite VIII of female. C. P. zimmermani sp. nov., tergite VIII of female. D. P. zimmermani sp. nov., sternite VIII of female. E. P. monopticus, sternite VIII of male. F. P. zimmermani sp. nov., sternite VIII of male. G. P. monopticus, spiculum gastrale. H. P. zimmermani sp. nov., ditto. I. P. monopticus, tegmen. J. P. zimmermani sp. nov., ditto. Scale bars = 0.5 mm." captionText-7="Fig. 8. Penis of Paratranes Zimmerman, 1994. A. P. monopticus (Pascoe, 1870), dorsal view (Griffith Uni., Qld). B. P. monopticus, ventral view (Griffith Uni., Qld). C. P. monopticus, lateral view (Griffith Uni., Qld). D. P. zimmermani sp. nov., dorsal view (Kalbarri, WA). E. P. zimmermani sp. nov., ventral view (Kalbarri, WA). F. P. zimmermani sp. nov., lateral view (Kalbarri, WA). Scale bars = 1.0 mm." captionText-8="Fig. 9. Endophallus and apex of penis of Paratranes Zimmerman, 1994. A. P. monopticus (Pascoe, 1870), apical sclerite (Griffith Uni., Qld). B. P. monopticus, ditto (Lucindale, SA). C. P. monopticus, ditto (Deepdene, WA). D. P. zimmermani sp. nov., ditto (Kalbarri, WA). E. P. zimmermani sp. nov., apex of penis (Kalbarri, WA). F. P. zimmermani sp. nov., ditto (NSW). Scale bars = 0.2 mm." captionText-9="Fig. 10. Female genital structures of Paratranes Zimmerman, 1994. A. P. monopticus (Pascoe, 1870), ovipositor (Homebush, NSW). B. P. zimmermani sp. nov., ditto (Wembley, WA). C. P. monopticus, spermatheca with gland (Homebush, NSW). D. P. zimmermani sp. nov., ditto (Wembley, WA). Scale bars: A–B = 0.5 mm; C–D = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5514623" figureDoi-1="http://doi.org/10.5281/zenodo.5514627" figureDoi-2="http://doi.org/10.5281/zenodo.5514629" figureDoi-3="http://doi.org/10.5281/zenodo.5514631" figureDoi-4="http://doi.org/10.5281/zenodo.5514635" figureDoi-5="http://doi.org/10.5281/zenodo.5514637" figureDoi-6="http://doi.org/10.5281/zenodo.5514639" figureDoi-7="http://doi.org/10.5281/zenodo.5514643" figureDoi-8="http://doi.org/10.5281/zenodo.5514645" figureDoi-9="http://doi.org/10.5281/zenodo.5514649" httpUri-0="https://zenodo.org/record/5514623/files/figure.png" httpUri-1="https://zenodo.org/record/5514627/files/figure.png" httpUri-2="https://zenodo.org/record/5514629/files/figure.png" httpUri-3="https://zenodo.org/record/5514631/files/figure.png" httpUri-4="https://zenodo.org/record/5514635/files/figure.png" httpUri-5="https://zenodo.org/record/5514637/files/figure.png" httpUri-6="https://zenodo.org/record/5514639/files/figure.png" httpUri-7="https://zenodo.org/record/5514643/files/figure.png" httpUri-8="https://zenodo.org/record/5514645/files/figure.png" httpUri-9="https://zenodo.org/record/5514649/files/figure.png" pageId="2" pageNumber="119">Figs 1–10</figureCitation>
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– 
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(catalogue); 2014: 560 (classification, host associations). — 
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(classification, host associations). — 
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Pullen 
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. 2014: 289
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</treatmentCitation>
(catalogue). — 
<treatmentCitation id="3817BA1CFFCDFF9D0D8A52DAFEE89B61" author="Anderson R. S. &amp; Oberprieler R. G. &amp; Setliff G. P." page="2" pageId="2" pageNumber="119" year="2018">
<bibRefCitation id="DD27E1FCFFCDFF9D0D8A52DAFEE89B61" author="Anderson R. S. &amp; Oberprieler R. G. &amp; Setliff G. P." pageId="2" pageNumber="119" pagination="54" refId="ref11423" refString="Anderson R. S., Oberprieler R. G. &amp; Setliff G. P. 2018. A review of the Araucaria - associated weevils of the tribe Orthorhinini (Coleoptera: Curculionidae: Molytinae), with description of new species of Ilacuris Pascoe, 1865 and Notopissodes Zimmerman &amp; Oberprieler, 2014 and a new genus, Kuschelorhinus Anderson &amp; Setliff. Diversity 10 (3): 54. https: // doi. org / 10.3390 / d 10030054" type="journal article" year="2018">
Anderson 
<emphasis id="8BC2401FFFCDFF9D0C7252DAFAC49B44" box="[1341,1391,1017,1043]" italics="true" pageId="2" pageNumber="119">et al</emphasis>
. 2018: 2
</bibRefCitation>
</treatmentCitation>
(classification, host associations). — 
<treatmentCitation id="3817BA1CFFCDFF9D0BB0553FFC769B60" author="Legalov A. A." box="[767,989,1052,1079]" page="345" pageId="2" pageNumber="119" year="2018">
<bibRefCitation id="DD27E1FCFFCDFF9D0BB0553FFC769B60" author="Legalov A. A." box="[767,989,1052,1079]" pageId="2" pageNumber="119" pagination="340 - 350" refId="ref11872" refString="Legalov A. A. 2018. Annotated key to weevils of the world. Part 2. Subfamily Molytinae (Coleoptera, Curculionidae). Ukrainian Journal of Ecology 8: 340 - 350." type="journal article" year="2018">Legalov 2018: 345</bibRefCitation>
</treatmentCitation>
(key, catalogue, classification).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F1ACCF86FFCDFF9D09F2557EFD459BF1" pageId="2" pageNumber="119" type="type_taxon">
<paragraph id="B9099C0DFFCDFF9D09F2557EFEF39B20" blockId="2.[189,344,1117,1143]" box="[189,344,1117,1143]" pageId="2" pageNumber="119">
<heading id="E2412B61FFCDFF9D09F2557EFEF39B20" bold="true" box="[189,344,1117,1143]" fontSize="11" level="3" pageId="2" pageNumber="119" reason="3">
<emphasis id="8BC2401FFFCDFF9D09F2557EFEF39B20" bold="true" box="[189,344,1117,1143]" pageId="2" pageNumber="119">
<typeStatus id="660D22AFFFCDFF9D09F2557EFF519B20" box="[189,250,1117,1143]" pageId="2" pageNumber="119">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="B9099C0DFFCDFF9D09F255AEFD459BF1" blockId="2.[189,750,1164,1190]" box="[189,750,1164,1190]" pageId="2" pageNumber="119">
<taxonomicName id="7EB6E78EFFCDFF9D09F255AEFDEB9BF1" ID-CoL="57XGY" authority="Pascoe, 1870" authorityName="Pascoe" authorityYear="1870" box="[189,576,1164,1190]" class="Insecta" family="Curculionidae" genus="Tranes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="monopticus">
<emphasis id="8BC2401FFFCDFF9D09F255AEFE309BF1" box="[189,411,1165,1190]" italics="true" pageId="2" pageNumber="119">Tranes monopticus</emphasis>
<bibRefCitation id="DD27E1FCFFCDFF9D08EC55AFFDEB9BF1" author="Pascoe F. P." box="[419,576,1164,1190]" pageId="2" pageNumber="119" pagination="445 - 484" refId="ref12264" refString="Pascoe F. P. 1870. Descriptions of some genera and species of Australian Curculionidae. Transactions of the Entomological Society of London 1870 (2): 181 - 209, 445 - 484." type="journal article" year="1870">Pascoe, 1870</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="F1ACCF86FFCDFF9B09F255EEFB5D9EDA" lastPageId="4" lastPageNumber="121" pageId="2" pageNumber="119" type="description">
<paragraph id="B9099C0DFFCDFF9D09F255EEFEC09BB0" blockId="2.[189,363,1229,1255]" box="[189,363,1229,1255]" pageId="2" pageNumber="119">
<heading id="E2412B61FFCDFF9D09F255EEFEC09BB0" bold="true" box="[189,363,1229,1255]" fontSize="11" level="3" pageId="2" pageNumber="119" reason="3">
<emphasis id="8BC2401FFFCDFF9D09F255EEFEC09BB0" bold="true" box="[189,363,1229,1255]" pageId="2" pageNumber="119">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="B9099C0DFFCDFF9B09F255DFFB5D9EDA" blockId="2.[187,1401,1276,2007]" lastBlockId="4.[189,1399,265,397]" lastPageId="4" lastPageNumber="121" pageId="2" pageNumber="119">
Body slender and depressed, completely shiny black; pronotal and elytral derm nearly nude, covered with very short setae (
<figureCitation id="218D8088FFCDFF9D08CB543CFE4A9A6E" box="[388,481,1311,1337]" captionStart="Fig" captionStartId="3.[189,232,1920,1946]" captionTargetBox="[189,1398,493,1882]" captionTargetId="figure-88@3.[189,1398,493,1882]" captionTargetPageId="3" captionText="Fig. 1. Diagnostic external characters of Paratranes Zimmerman, 1994. A. Elytral surface and setae. B. Eyes and prothorax, ventral view. C. Head and base of rostrum, dorsal view. D. Pronotal surface. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514623" httpUri="https://zenodo.org/record/5514623/files/figure.png" pageId="2" pageNumber="119">Fig. 1A</figureCitation>
). Rostrum moderately long, equally long in both sexes (
<figureCitation id="218D8088FFCDFF9D0D38543CFB6B9A6E" box="[1143,1216,1311,1337]" captionStart="Fig" captionStartId="8.[189,232,1952,1978]" captionTargetBox="[206,1347,314,1844]" captionTargetId="figure-14@8.[189,1398,293,1886]" captionTargetPageId="8" captionText="Fig. 4. Habitus, lateral view. A–B. Paratranes monopticus (Pascoe, 1870). C–D. P. zimmermani sp. nov. (C. Holotype (ANIC); D. Paratype (ANIC)). A, C. Male. B, D. Female. Scale bars = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514631" httpUri="https://zenodo.org/record/5514631/files/figure.png" pageId="2" pageNumber="119">Fig. 4</figureCitation>
). Eyes dorsally well separated (
<figureCitation id="218D8088FFCDFF9D083B5461FE659A0B" box="[372,462,1346,1372]" captionStart="Fig" captionStartId="3.[189,232,1920,1946]" captionTargetBox="[189,1398,493,1882]" captionTargetId="figure-88@3.[189,1398,493,1882]" captionTargetPageId="3" captionText="Fig. 1. Diagnostic external characters of Paratranes Zimmerman, 1994. A. Elytral surface and setae. B. Eyes and prothorax, ventral view. C. Head and base of rostrum, dorsal view. D. Pronotal surface. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514623" httpUri="https://zenodo.org/record/5514623/files/figure.png" pageId="2" pageNumber="119">Fig. 1C</figureCitation>
); ventrally very narrowly separated, as narrowly as procoxae (
<figureCitation id="218D8088FFCDFF9D0DEF5461FB519A0B" box="[1184,1274,1346,1372]" captionStart="Fig" captionStartId="3.[189,232,1920,1946]" captionTargetBox="[189,1398,493,1882]" captionTargetId="figure-88@3.[189,1398,493,1882]" captionTargetPageId="3" captionText="Fig. 1. Diagnostic external characters of Paratranes Zimmerman, 1994. A. Elytral surface and setae. B. Eyes and prothorax, ventral view. C. Head and base of rostrum, dorsal view. D. Pronotal surface. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514623" httpUri="https://zenodo.org/record/5514623/files/figure.png" pageId="2" pageNumber="119">Fig. 1B</figureCitation>
); forehead slightly narrower than basal width of rostrum (
<figureCitation id="218D8088FFCDFF9D0B9C5446FC869AD7" box="[723,813,1381,1408]" captionStart="Fig" captionStartId="3.[189,232,1920,1946]" captionTargetBox="[189,1398,493,1882]" captionTargetId="figure-88@3.[189,1398,493,1882]" captionTargetPageId="3" captionText="Fig. 1. Diagnostic external characters of Paratranes Zimmerman, 1994. A. Elytral surface and setae. B. Eyes and prothorax, ventral view. C. Head and base of rostrum, dorsal view. D. Pronotal surface. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514623" httpUri="https://zenodo.org/record/5514623/files/figure.png" pageId="2" pageNumber="119">Fig. 1C</figureCitation>
). Antennae inserted at about distal third of rostrum in both sexes, not sexually dimorphic (
<figureCitation id="218D8088FFCDFF9D0B3654AAFD6B9AF4" box="[633,704,1417,1443]" captionStart="Fig" captionStartId="8.[189,232,1952,1978]" captionTargetBox="[206,1347,314,1844]" captionTargetId="figure-14@8.[189,1398,293,1886]" captionTargetPageId="8" captionText="Fig. 4. Habitus, lateral view. A–B. Paratranes monopticus (Pascoe, 1870). C–D. P. zimmermani sp. nov. (C. Holotype (ANIC); D. Paratype (ANIC)). A, C. Male. B, D. Female. Scale bars = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514631" httpUri="https://zenodo.org/record/5514631/files/figure.png" pageId="2" pageNumber="119">Fig. 4</figureCitation>
); funicles 7-segmented but segment 7 closely approximated to club and nearly continuous in outline with it, segments 1 and 2 short, 2 slightly longer or as long as apical width of scape and slightly shorter than segments 3+4 (
<figureCitation id="218D8088FFCDFF9D0AEB54ECFC479ABD" box="[932,1004,1487,1514]" captionStart="Fig" captionStartId="10.[190,233,1948,1974]" captionTargetBox="[199,1398,879,1841]" captionTargetId="figure-171@10.[189,1398,879,1911]" captionTargetPageId="10" captionText="Fig. 5. Left antenna, dorsal view. A. Paratranes monopticus (Pascoe, 1870). B. P. zimmermani sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514635" httpUri="https://zenodo.org/record/5514635/files/figure.png" pageId="2" pageNumber="119">Fig. 5</figureCitation>
); clubs stout and short, distinctly shorter than funicles, 4-segmented, with small conical apical segment, appearing 5-segmented due to funicle segment 7 being closely approximated to basal club segment. Pronotum distinctly narrower than elytra at humeri, with sides weakly to moderately arcuate (
<figureCitation id="218D8088FFCDFF9D0A21571AFC1E9904" box="[878,949,1593,1619]" captionStart="Fig" captionStartId="7.[189,232,1952,1978]" captionTargetBox="[292,1272,288,1838]" captionTargetId="figure-12@7.[274,1314,265,1915]" captionTargetPageId="7" captionText="Fig. 3. Habitus, dorsal view. A–B. Paratranes monopticus (Pascoe, 1870). C–D. P. zimmermani sp. nov. (C. Holotype (ANIC); D. Paratype (ANIC)). A, C. Male. B, D. Female. Scale bars = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514629" httpUri="https://zenodo.org/record/5514629/files/figure.png" pageId="2" pageNumber="119">Fig. 3</figureCitation>
); surface punctate, punctures separate on disc but confluent and vague laterally (
<figureCitation id="218D8088FFCDFF9D0BEE577FFD579921" box="[673,764,1628,1654]" captionStart="Fig" captionStartId="3.[189,232,1920,1946]" captionTargetBox="[189,1398,493,1882]" captionTargetId="figure-88@3.[189,1398,493,1882]" captionTargetPageId="3" captionText="Fig. 1. Diagnostic external characters of Paratranes Zimmerman, 1994. A. Elytral surface and setae. B. Eyes and prothorax, ventral view. C. Head and base of rostrum, dorsal view. D. Pronotal surface. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514623" httpUri="https://zenodo.org/record/5514623/files/figure.png" pageId="2" pageNumber="119">Fig. 1D</figureCitation>
). Elytra elongate, jointly ca 0.5–0.6× as broad as long, sides subparallel or slightly diverging apicad, subapically slightly indented (
<figureCitation id="218D8088FFCDFF9D0D7C575CFB3A99CE" box="[1075,1169,1663,1689]" captionStart="Fig" captionStartId="4.[191,234,1958,1984]" captionTargetBox="[189,1375,733,1900]" captionTargetId="figure-171@4.[189,1398,732,1920]" captionTargetPageId="4" captionText="Fig. 2. Diagnostic external characters and terminalia of Paratranes Zimmerman, 1994. A. Elytral apex. B. Right metatibia. C. Tergite VII of female. D. Tergite VIII of male. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514627" httpUri="https://zenodo.org/record/5514627/files/figure.png" pageId="2" pageNumber="119">Fig. 2A</figureCitation>
); surface nearly flat (
<figureCitation id="218D8088FFCDFF9D098A5781FE9B99EB" box="[197,304,1698,1724]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="7.[189,232,1952,1978]" captionStartId-1="8.[189,232,1952,1978]" captionTargetBox-0="[292,1272,288,1838]" captionTargetBox-1="[206,1347,314,1844]" captionTargetId-0="figure-12@7.[274,1314,265,1915]" captionTargetId-1="figure-14@8.[189,1398,293,1886]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Fig. 3. Habitus, dorsal view. A–B. Paratranes monopticus (Pascoe, 1870). C–D. P. zimmermani sp. nov. (C. Holotype (ANIC); D. Paratype (ANIC)). A, C. Male. B, D. Female. Scale bars = 5.0 mm." captionText-1="Fig. 4. Habitus, lateral view. A–B. Paratranes monopticus (Pascoe, 1870). C–D. P. zimmermani sp. nov. (C. Holotype (ANIC); D. Paratype (ANIC)). A, C. Male. B, D. Female. Scale bars = 5.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5514629" figureDoi-1="http://doi.org/10.5281/zenodo.5514631" httpUri-0="https://zenodo.org/record/5514629/files/figure.png" httpUri-1="https://zenodo.org/record/5514631/files/figure.png" pageId="2" pageNumber="119">Figs 3–4</figureCitation>
). Prothorax with anterior margin laterally extended into weak ocular lobes ventrally separated by shallow emargination (
<figureCitation id="218D8088FFCDFF9D08A457E6FDEF99B7" box="[491,580,1733,1760]" captionStart="Fig" captionStartId="3.[189,232,1920,1946]" captionTargetBox="[189,1398,493,1882]" captionTargetId="figure-88@3.[189,1398,493,1882]" captionTargetPageId="3" captionText="Fig. 1. Diagnostic external characters of Paratranes Zimmerman, 1994. A. Elytral surface and setae. B. Eyes and prothorax, ventral view. C. Head and base of rostrum, dorsal view. D. Pronotal surface. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514623" httpUri="https://zenodo.org/record/5514623/files/figure.png" pageId="2" pageNumber="119">Fig. 1B</figureCitation>
); prosternum before procoxal cavities impunctate or sparsely punctate medially; procoxal cavities confluent, with procoxae contiguous; metanepisterna without sclerolepidia. Femora distinctly sulcate beneath; meso- and metatibiae with distal setal combs continued around apex and extending to almost middle of tibia (
<figureCitation id="218D8088FFCDFF9D0BA5560CFCE3981E" box="[746,840,1839,1865]" captionStart="Fig" captionStartId="4.[191,234,1958,1984]" captionTargetBox="[189,1375,733,1900]" captionTargetId="figure-171@4.[189,1398,732,1920]" captionTargetPageId="4" captionText="Fig. 2. Diagnostic external characters and terminalia of Paratranes Zimmerman, 1994. A. Elytral apex. B. Right metatibia. C. Tergite VII of female. D. Tergite VIII of male. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514627" httpUri="https://zenodo.org/record/5514627/files/figure.png" pageId="2" pageNumber="119">Fig. 2B</figureCitation>
), metatibiae with dorso-apical corner rounded. Terminalia: tergite VII of female subtrapezoidal (
<figureCitation id="218D8088FFCDFF9D0A465671FCCC983B" box="[777,871,1874,1900]" captionStart="Fig" captionStartId="4.[191,234,1958,1984]" captionTargetBox="[189,1375,733,1900]" captionTargetId="figure-171@4.[189,1398,732,1920]" captionTargetPageId="4" captionText="Fig. 2. Diagnostic external characters and terminalia of Paratranes Zimmerman, 1994. A. Elytral apex. B. Right metatibia. C. Tergite VII of female. D. Tergite VIII of male. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514627" httpUri="https://zenodo.org/record/5514627/files/figure.png" pageId="2" pageNumber="119">Fig. 2C</figureCitation>
), with posterior margin subtruncate, anterior margin ca 1.5–1.9× as wide as posterior margin; tergite VIII of male subquadratic (
<figureCitation id="218D8088FFCDFF9D0D885656FA8398C7" box="[1223,1320,1909,1936]" captionStart="Fig" captionStartId="4.[191,234,1958,1984]" captionTargetBox="[189,1375,733,1900]" captionTargetId="figure-171@4.[189,1398,732,1920]" captionTargetPageId="4" captionText="Fig. 2. Diagnostic external characters and terminalia of Paratranes Zimmerman, 1994. A. Elytral apex. B. Right metatibia. C. Tergite VII of female. D. Tergite VIII of male. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514627" httpUri="https://zenodo.org/record/5514627/files/figure.png" pageId="2" pageNumber="119">Fig. 2D</figureCitation>
), with posterior margin subtruncate, of female subtrapezoidal (
<figureCitation id="218D8088FFCDFF9D0A2C56BAFC4198E4" box="[867,1002,1945,1971]" captionStart="Fig" captionStartId="12.[189,232,1852,1878]" captionTargetBox="[283,1309,267,1783]" captionTargetId="figure-14@12.[256,1332,265,1815]" captionTargetPageId="12" captionText="Fig. 7. Terminalia of Paratranes Zimmerman, 1994. A. P. monopticus (Pascoe, 1870), tergite VIII of female. B. P. monopticus, sternite VIII of female. C. P. zimmermani sp. nov., tergite VIII of female. D. P. zimmermani sp. nov., sternite VIII of female. E. P. monopticus, sternite VIII of male. F. P. zimmermani sp. nov., sternite VIII of male. G. P. monopticus, spiculum gastrale. H. P. zimmermani sp. nov., ditto. I. P. monopticus, tegmen. J. P. zimmermani sp. nov., ditto. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514639" httpUri="https://zenodo.org/record/5514639/files/figure.png" pageId="2" pageNumber="119">Fig. 7A, C</figureCitation>
), very long in 
<taxonomicName id="7EB6E78EFFCDFF9D0DEC56BAFAE198E5" authorityName="Pascoe" authorityYear="1870" box="[1187,1354,1945,1970]" class="Insecta" family="Curculionidae" genus="Tranes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="monopticus">
<emphasis id="8BC2401FFFCDFF9D0DEC56BAFAE198E5" box="[1187,1354,1945,1970]" italics="true" pageId="2" pageNumber="119">P. monopticus</emphasis>
</taxonomicName>
, ca 0.9–1.8 × as long as width at anterior margin, posterior margin subtruncate, anterior margin ca 1.4–1.5 × as wide as posterior margin; sternite VIII of female with sclerotised parts of apical lobes slender, linear (
<figureCitation id="218D8088FFCCFF9C0989500FFEE89E10" box="[198,323,300,327]" captionStart="Fig" captionStartId="12.[189,232,1852,1878]" captionTargetBox="[283,1309,267,1783]" captionTargetId="figure-14@12.[256,1332,265,1815]" captionTargetPageId="12" captionText="Fig. 7. Terminalia of Paratranes Zimmerman, 1994. A. P. monopticus (Pascoe, 1870), tergite VIII of female. B. P. monopticus, sternite VIII of female. C. P. zimmermani sp. nov., tergite VIII of female. D. P. zimmermani sp. nov., sternite VIII of female. E. P. monopticus, sternite VIII of male. F. P. zimmermani sp. nov., sternite VIII of male. G. P. monopticus, spiculum gastrale. H. P. zimmermani sp. nov., ditto. I. P. monopticus, tegmen. J. P. zimmermani sp. nov., ditto. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514639" httpUri="https://zenodo.org/record/5514639/files/figure.png" pageId="3" pageNumber="120">Fig. 7B, D</figureCitation>
); tegmen with oval to rounded ring (
<figureCitation id="218D8088FFCCFF9C0BAB500FFCE49E10" box="[740,847,300,327]" captionStart="Fig" captionStartId="12.[189,232,1852,1878]" captionTargetBox="[283,1309,267,1783]" captionTargetId="figure-14@12.[256,1332,265,1815]" captionTargetPageId="12" captionText="Fig. 7. Terminalia of Paratranes Zimmerman, 1994. A. P. monopticus (Pascoe, 1870), tergite VIII of female. B. P. monopticus, sternite VIII of female. C. P. zimmermani sp. nov., tergite VIII of female. D. P. zimmermani sp. nov., sternite VIII of female. E. P. monopticus, sternite VIII of male. F. P. zimmermani sp. nov., sternite VIII of male. G. P. monopticus, spiculum gastrale. H. P. zimmermani sp. nov., ditto. I. P. monopticus, tegmen. J. P. zimmermani sp. nov., ditto. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514639" httpUri="https://zenodo.org/record/5514639/files/figure.png" pageId="3" pageNumber="120">Fig. 7I–J</figureCitation>
), manubrium distinctly shorter than parameroid lobes; penis subparallel (
<figureCitation id="218D8088FFCCFF9C08A6506CFD989E3E" box="[489,563,335,361]" captionStart="Fig" captionStartId="16.[189,232,1884,1910]" captionTargetBox="[340,1238,280,1818]" captionTargetId="figure-15@16.[317,1271,265,1843]" captionTargetPageId="16" captionText="Fig. 8. Penis of Paratranes Zimmerman, 1994. A. P. monopticus (Pascoe, 1870), dorsal view (Griffith Uni., Qld). B. P. monopticus, ventral view (Griffith Uni., Qld). C. P. monopticus, lateral view (Griffith Uni., Qld). D. P. zimmermani sp. nov., dorsal view (Kalbarri, WA). E. P. zimmermani sp. nov., ventral view (Kalbarri, WA). F. P. zimmermani sp. nov., lateral view (Kalbarri, WA). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514643" httpUri="https://zenodo.org/record/5514643/files/figure.png" pageId="3" pageNumber="120">Fig. 8</figureCitation>
), dorsum strongly sclerotised at both sides, forming a broad median groove, apical margin roundly subtruncate or slightly medially emarginate, body distinctly shorter than temones (ca 2.0–3.0 ×); endophallus (
<figureCitation id="218D8088FFCCFF9C0BCF50B5FCE39EE7" box="[640,840,406,432]" captionStart="Fig" captionStartId="16.[189,232,1884,1910]" captionTargetBox="[340,1238,280,1818]" captionTargetId="figure-15@16.[317,1271,265,1843]" captionTargetPageId="16" captionText="Fig. 8. Penis of Paratranes Zimmerman, 1994. A. P. monopticus (Pascoe, 1870), dorsal view (Griffith Uni., Qld). B. P. monopticus, ventral view (Griffith Uni., Qld). C. P. monopticus, lateral view (Griffith Uni., Qld). D. P. zimmermani sp. nov., dorsal view (Kalbarri, WA). E. P. zimmermani sp. nov., ventral view (Kalbarri, WA). F. P. zimmermani sp. nov., lateral view (Kalbarri, WA). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514643" httpUri="https://zenodo.org/record/5514643/files/figure.png" pageId="3" pageNumber="120">Fig. 8A–B, D–E</figureCitation>
) membranous, long, extending below body of penis, apically with anchor-shaped symmetrical armature composed of two strongly hooked sclerites joined medially to form a common stalk (
<figureCitation id="218D8088FFCBFF9B0BF0500FFCEE9E11" box="[703,837,300,326]" captionStart="Fig" captionStartId="17.[191,234,1879,1905]" captionTargetBox="[189,1398,947,1841]" captionTargetId="figure-205@17.[189,1398,947,1841]" captionTargetPageId="17" captionText="Fig. 9. Endophallus and apex of penis of Paratranes Zimmerman, 1994. A. P. monopticus (Pascoe, 1870), apical sclerite (Griffith Uni., Qld). B. P. monopticus, ditto (Lucindale, SA). C. P. monopticus, ditto (Deepdene, WA). D. P. zimmermani sp. nov., ditto (Kalbarri, WA). E. P. zimmermani sp. nov., apex of penis (Kalbarri, WA). F. P. zimmermani sp. nov., ditto (NSW). Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5514645" httpUri="https://zenodo.org/record/5514645/files/figure.png" pageId="4" pageNumber="121">Fig. 9A–D</figureCitation>
); ovipositor short (
<taxonomicName id="7EB6E78EFFCBFF9B0D7F500EFB4A9E11" authority="Hsiao &amp; Oberprieler, 2021" authorityName="Hsiao &amp; Oberprieler" authorityYear="2021" box="[1072,1249,301,326]" class="Insecta" family="Curculionidae" genus="Paratranes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="zimmermani" status="sp. nov.">
<emphasis id="8BC2401FFFCBFF9B0D7F500EFB4A9E11" box="[1072,1249,301,326]" italics="true" pageId="4" pageNumber="121">P. zimmermani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90F1FD64FFCBFF9B0DA2500EFAE79E10" box="[1261,1356,301,327]" pageId="4" pageNumber="121" rank="species">sp. nov.</taxonomicNameLabel>
) to long (
<taxonomicName id="7EB6E78EFFCBFF9B084B5073FE029E3E" authorityName="Pascoe" authorityYear="1870" box="[260,425,336,361]" class="Insecta" family="Curculionidae" genus="Tranes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="monopticus">
<emphasis id="8BC2401FFFCBFF9B084B5073FE029E3E" box="[260,425,336,361]" italics="true" pageId="4" pageNumber="121">P. monopticus</emphasis>
</taxonomicName>
), ca 1.5–5.7 × as long as wide, proximal gonocoxite ca 0.8–1.4 × as long as distal gonocoxite (
<figureCitation id="218D8088FFCBFF9B081E5051FE4B9EDA" box="[337,480,370,397]" captionStart="Fig" captionStartId="18.[191,234,1879,1905]" captionTargetBox="[197,1395,506,1801]" captionTargetId="figure-85@18.[189,1398,490,1841]" captionTargetPageId="18" captionText="Fig. 10. Female genital structures of Paratranes Zimmerman, 1994. A. P. monopticus (Pascoe, 1870), ovipositor (Homebush, NSW). B. P. zimmermani sp. nov., ditto (Wembley, WA). C. P. monopticus, spermatheca with gland (Homebush, NSW). D. P. zimmermani sp. nov., ditto (Wembley, WA). Scale bars: A–B = 0.5 mm; C–D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5514649" httpUri="https://zenodo.org/record/5514649/files/figure.png" pageId="4" pageNumber="121">Fig. 10A–B</figureCitation>
), gonostyli subapical, broad, apically truncate, with few long setae.
</paragraph>
</subSubSection>
<caption id="EDC9CC85FFCCFF9C09F256A3FE1398B6" ID-DOI="http://doi.org/10.5281/zenodo.5514623" ID-Zenodo-Dep="5514623" httpUri="https://zenodo.org/record/5514623/files/figure.png" pageId="3" pageNumber="120" startId="3.[189,232,1920,1946]" targetBox="[189,1398,493,1882]" targetPageId="3">
<paragraph id="B9099C0DFFCCFF9C09F256A3FE1398B6" blockId="3.[189,1399,1920,2017]" pageId="3" pageNumber="120">
<emphasis id="8BC2401FFFCCFF9C09F256A3FEA698CD" bold="true" box="[189,269,1920,1946]" pageId="3" pageNumber="120">Fig. 1.</emphasis>
Diagnostic external characters of 
<taxonomicName id="7EB6E78EFFCCFF9C0BE456A2FBBD98CC" authority="Zimmerman, 1994" authorityName="Zimmerman" authorityYear="1994" box="[683,1046,1920,1947]" class="Insecta" family="Curculionidae" genus="Paratranes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCCFF9C0BE456A2FC8498CD" box="[683,815,1921,1946]" italics="true" pageId="3" pageNumber="120">Paratranes</emphasis>
<bibRefCitation id="DD27E1FCFFCCFF9C0A7956A3FBBD98CC" author="Zimmerman E. C." box="[822,1046,1920,1947]" pageId="3" pageNumber="120" refId="ref12493" refString="Zimmerman E. C. 1994. Australian Weevils (Coleoptera: Curculionoidea). Volume I. Orthoceri. Anthribidae to Attelabidae. The Primitive Weevils. CSIRO, Melbourne." type="book" year="1994">Zimmerman, 1994</bibRefCitation>
</taxonomicName>
. 
<emphasis id="8BC2401FFFCCFF9C0D6A56A3FB9398CD" bold="true" box="[1061,1080,1920,1946]" pageId="3" pageNumber="120">A</emphasis>
. Elytral surface and setae. 
<emphasis id="8BC2401FFFCCFF9C09F05687FF7B98E9" bold="true" box="[191,208,1956,1982]" pageId="3" pageNumber="120">B</emphasis>
. Eyes and prothorax, ventral view. 
<emphasis id="8BC2401FFFCCFF9C0BCE5680FD3F98EA" bold="true" box="[641,660,1955,1981]" pageId="3" pageNumber="120">C</emphasis>
. Head and base of rostrum, dorsal view. 
<emphasis id="8BC2401FFFCCFF9C0DC65687FB3798E9" bold="true" box="[1161,1180,1956,1982]" pageId="3" pageNumber="120">D</emphasis>
. Pronotal surface. Scale bars = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="F1ACCF86FFCBFF9A09F25097FB539DF6" lastPageId="5" lastPageNumber="122" pageId="4" pageNumber="121" type="discussion">
<paragraph id="B9099C0DFFCBFF9B09F25097FE849E99" blockId="4.[189,303,436,462]" box="[189,303,436,462]" pageId="4" pageNumber="121">
<heading id="E2412B61FFCBFF9B09F25097FE849E99" bold="true" box="[189,303,436,462]" fontSize="11" level="3" pageId="4" pageNumber="121" reason="3">
<emphasis id="8BC2401FFFCBFF9B09F25097FE849E99" bold="true" box="[189,303,436,462]" pageId="4" pageNumber="121">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="B9099C0DFFCBFF9A09F250C0FE969E3E" blockId="4.[187,1399,482,685]" lastBlockId="5.[189,1399,265,361]" lastPageId="5" lastPageNumber="122" pageId="4" pageNumber="121">
<taxonomicName id="7EB6E78EFFCBFF9B09F250C0FEEA9EAB" authorityName="Zimmerman" authorityYear="1994" box="[189,321,483,508]" class="Insecta" family="Curculionidae" genus="Paratranes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCBFF9B09F250C0FEEA9EAB" box="[189,321,483,508]" italics="true" pageId="4" pageNumber="121">Paratranes</emphasis>
</taxonomicName>
is here classified in the tribe 
<taxonomicName id="7EB6E78EFFCBFF9B0BC550C1FC8B9EAB" authorityName="Jekel" authorityYear="1865" box="[650,800,482,508]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="tribe" tribe="Orthorhinini">Orthorhinini</taxonomicName>
, following 
<bibRefCitation id="DD27E1FCFFCBFF9B0AEF50C1FB0C9EAA" author="Anderson R. S. &amp; Oberprieler R. G. &amp; Setliff G. P." box="[928,1191,482,509]" pageId="4" pageNumber="121" pagination="54" refId="ref11423" refString="Anderson R. S., Oberprieler R. G. &amp; Setliff G. P. 2018. A review of the Araucaria - associated weevils of the tribe Orthorhinini (Coleoptera: Curculionidae: Molytinae), with description of new species of Ilacuris Pascoe, 1865 and Notopissodes Zimmerman &amp; Oberprieler, 2014 and a new genus, Kuschelorhinus Anderson &amp; Setliff. Diversity 10 (3): 54. https: // doi. org / 10.3390 / d 10030054" type="journal article" year="2018">
Anderson 
<emphasis id="8BC2401FFFCBFF9B0D5850C0FBE29EAB" box="[1047,1097,482,508]" italics="true" pageId="4" pageNumber="121">et al</emphasis>
. (2018)
</bibRefCitation>
, who included the 
<taxonomicName id="7EB6E78EFFCBFF9B09F45324FEA09D77" authorityName="Schoenherr" authorityYear="1843" box="[187,267,519,544]" class="Insecta" family="Curculionidae" genus="Tranes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCBFF9B09F45324FEA09D77" box="[187,267,519,544]" italics="true" pageId="4" pageNumber="121">Tranes</emphasis>
</taxonomicName>
group in this tribe based on the strongly supported relationship between 
<taxonomicName id="7EB6E78EFFCBFF9B0D1C5324FADC9D77" authority="Schoenherr, 1843" authorityName="Schoenherr" authorityYear="1843" box="[1107,1399,518,544]" class="Insecta" family="Curculionidae" genus="Tranes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCBFF9B0D1C5324FB089D77" box="[1107,1187,519,544]" italics="true" pageId="4" pageNumber="121">Tranes</emphasis>
Schoenherr, 1843
</taxonomicName>
and the orthorhinine genus 
<taxonomicName id="7EB6E78EFFCBFF9B08B85309FCB69D13" authority="Pouillaude, 1915" authorityName="Pouillaude" authorityYear="1915" box="[503,797,553,580]" class="Insecta" family="Curculionidae" genus="Vanapa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCBFF9B08B85309FDE49D14" box="[503,591,554,579]" italics="true" pageId="4" pageNumber="121">Vanapa</emphasis>
Pouillaude, 1915
</taxonomicName>
found by 
<bibRefCitation id="DD27E1FCFFCBFF9B0ADF530AFBF29D14" author="Shin S. &amp; Clarke D. J. &amp; Lemmon A. R. &amp; Moriarty-Lemmon E. &amp; Aitken A. L. &amp; Haddad S. &amp; Farrell B. D. &amp; Marvaldi A. E. &amp; Oberprieler R. G. &amp; McKenna D. D." box="[912,1113,553,579]" pageId="4" pageNumber="121" pagination="823 - 836" refId="ref12400" refString="Shin S., Clarke D. J., Lemmon A. R., Moriarty-Lemmon E., Aitken A. L., Haddad S., Farrell B. D., Marvaldi A. E., Oberprieler R. G. &amp; McKenna D. D. 2017. Phylogenomic data yield new and robust insights into the phylogeny and evolution of weevils. Molecular Biology and Evolution 35: 823 - 836. https: // doi. org / 10.1093 / molbev / msx 324" type="journal article" year="2017">
Shin 
<emphasis id="8BC2401FFFCBFF9B0A845309FC559D14" box="[971,1022,553,579]" italics="true" pageId="4" pageNumber="121">et al</emphasis>
. (2017)
</bibRefCitation>
in a robust phylogenetic analysis of 522 protein-coding genes. Without considering these results, 
<bibRefCitation id="DD27E1FCFFCBFF9B0D03536FFAA39D31" author="Legalov A. A." box="[1100,1288,588,614]" pageId="4" pageNumber="121" pagination="340 - 350" refId="ref11872" refString="Legalov A. A. 2018. Annotated key to weevils of the world. Part 2. Subfamily Molytinae (Coleoptera, Curculionidae). Ukrainian Journal of Ecology 8: 340 - 350." type="journal article" year="2018">Legalov (2018)</bibRefCitation>
recently proposed a separate tribe for the 
<taxonomicName id="7EB6E78EFFCBFF9B0B0F5353FD3B9DDE" authorityName="Schoenherr" authorityYear="1843" box="[576,656,624,649]" class="Insecta" family="Curculionidae" genus="Tranes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCBFF9B0B0F5353FD3B9DDE" box="[576,656,624,649]" italics="true" pageId="4" pageNumber="121">Tranes</emphasis>
</taxonomicName>
group, Tranini (incorrectly formed as ‘Tranesini’; see 
<bibRefCitation id="DD27E1FCFFCBFF9B0C5C534CFE039DFA" author="Hsiao Y. &amp; Oberprieler R. G." pageId="4" pageNumber="121" pagination="677 - 700" refId="ref11651" refString="Hsiao Y. &amp; Oberprieler R. G. 2020. A review of the trunk-boring cycad weevils in Australia, with description of a second species of Demyrsus Pascoe, 1872 (Coleoptera: Curculionidae: Molytinae). Austral Entomology 59 (4): 677 - 700. https: // doi. org / 10.1111 / aen. 12498" type="journal article" year="2020">Hsiao &amp; Oberprieler (2020))</bibRefCitation>
, but his diagnosis of Tranini did not distinguish the group from 
<taxonomicName id="7EB6E78EFFCBFF9B0DE553B1FAEB9DFB" authorityName="Jekel" authorityYear="1865" box="[1194,1344,658,684]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="140" phylum="Arthropoda" rank="tribe" tribe="Orthorhinini">Orthorhinini</taxonomicName>
, and it further contained characters not present in all the included genera. Pending further and more detailed analysis of the phylogenetic relationships of the 
<taxonomicName id="7EB6E78EFFCAFF9A0BBC500EFCE89E11" authorityName="Schoenherr" authorityYear="1843" box="[755,835,301,326]" class="Insecta" family="Curculionidae" genus="Tranes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCAFF9A0BBC500EFCE89E11" box="[755,835,301,326]" italics="true" pageId="5" pageNumber="122">Tranes</emphasis>
</taxonomicName>
group, it is premature to afford it tribal status in 
<taxonomicName id="7EB6E78EFFCAFF9A09F2506CFE929E3E" authorityName="Schoenherr" authorityYear="1823" box="[189,313,335,361]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="subFamily" subFamily="Molytinae">Molytinae</taxonomicName>
.
</paragraph>
<caption id="EDC9CC85FFCBFF9B09F05685FB4F98B3" ID-DOI="http://doi.org/10.5281/zenodo.5514627" ID-Zenodo-Dep="5514627" httpUri="https://zenodo.org/record/5514627/files/figure.png" pageId="4" pageNumber="121" startId="4.[191,234,1958,1984]" targetBox="[189,1375,733,1900]" targetPageId="4">
<paragraph id="B9099C0DFFCBFF9B09F05685FB4F98B3" blockId="4.[189,1399,1958,2020]" pageId="4" pageNumber="121">
<emphasis id="8BC2401FFFCBFF9B09F05685FEA79897" bold="true" box="[191,268,1958,1984]" pageId="4" pageNumber="121">Fig. 2.</emphasis>
Diagnostic external characters and terminalia of 
<taxonomicName id="7EB6E78EFFCBFF9B0A065684FB199897" authority="Zimmerman, 1994" authorityName="Zimmerman" authorityYear="1994" box="[841,1202,1958,1984]" class="Insecta" family="Curculionidae" genus="Paratranes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCBFF9B0A065684FC669897" box="[841,973,1959,1984]" italics="true" pageId="4" pageNumber="121">Paratranes</emphasis>
<bibRefCitation id="DD27E1FCFFCBFF9B0A9A5685FB199897" author="Zimmerman E. C." box="[981,1202,1958,1984]" pageId="4" pageNumber="121" refId="ref12493" refString="Zimmerman E. C. 1994. Australian Weevils (Coleoptera: Curculionoidea). Volume I. Orthoceri. Anthribidae to Attelabidae. The Primitive Weevils. CSIRO, Melbourne." type="book" year="1994">Zimmerman, 1994</bibRefCitation>
</taxonomicName>
. 
<emphasis id="8BC2401FFFCBFF9B0DF25685FB799897" bold="true" box="[1213,1234,1958,1984]" pageId="4" pageNumber="121">A</emphasis>
. Elytral apex. 
<emphasis id="8BC2401FFFCBFF9B09F256E9FF7A98B3" bold="true" box="[189,209,1994,2020]" pageId="4" pageNumber="121">B</emphasis>
. Right metatibia. 
<emphasis id="8BC2401FFFCBFF9B08EB56EAFE1298B4" bold="true" box="[420,441,1993,2019]" pageId="4" pageNumber="121">C</emphasis>
. Tergite VII of female. 
<emphasis id="8BC2401FFFCBFF9B0B8156E9FD4898B3" bold="true" box="[718,739,1994,2020]" pageId="4" pageNumber="121">D</emphasis>
. Tergite VIII of male. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="B9099C0DFFCAFF9A09F250B2FB539DF6" blockId="5.[187,1399,401,674]" pageId="5" pageNumber="122">
The relationships of 
<taxonomicName id="7EB6E78EFFCAFF9A08F950B1FD919EFC" authorityName="Zimmerman" authorityYear="1994" box="[438,570,402,427]" class="Insecta" family="Curculionidae" genus="Paratranes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCAFF9A08F950B1FD919EFC" box="[438,570,402,427]" italics="true" pageId="5" pageNumber="122">Paratranes</emphasis>
</taxonomicName>
to the other genera of the 
<taxonomicName id="7EB6E78EFFCAFF9A0A3550B1FC619EFC" authorityName="Schoenherr" authorityYear="1843" box="[890,970,402,427]" class="Insecta" family="Curculionidae" genus="Tranes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCAFF9A0A3550B1FC619EFC" box="[890,970,402,427]" italics="true" pageId="5" pageNumber="122">Tranes</emphasis>
</taxonomicName>
group also remain uncertain for the moment. It agrees with 
<taxonomicName id="7EB6E78EFFCAFF9A089A5096FD8F9E99" authorityName="Schoenherr" authorityYear="1843" box="[469,548,437,462]" class="Insecta" family="Curculionidae" genus="Tranes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCAFF9A089A5096FD8F9E99" box="[469,548,437,462]" italics="true" pageId="5" pageNumber="122">Tranes</emphasis>
</taxonomicName>
, 
<taxonomicName id="7EB6E78EFFCAFF9A0B7B5097FC349E99" authority="Zimmerman, 1994" authorityName="Zimmerman" authorityYear="1994" box="[564,927,436,462]" class="Insecta" family="Curculionidae" genus="Miltotranes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCAFF9A0B7B5097FD169E99" box="[564,701,436,462]" italics="true" pageId="5" pageNumber="122">Miltotranes</emphasis>
<bibRefCitation id="DD27E1FCFFCAFF9A0B8B5097FC349E99" author="Zimmerman E. C." box="[708,927,436,462]" pageId="5" pageNumber="122" refId="ref12493" refString="Zimmerman E. C. 1994. Australian Weevils (Coleoptera: Curculionoidea). Volume I. Orthoceri. Anthribidae to Attelabidae. The Primitive Weevils. CSIRO, Melbourne." type="book" year="1994">Zimmerman, 1994</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="7EB6E78EFFCAFF9A0A965096FAF39E99" authority="Zimmerman, 1994" authorityName="Zimmerman" authorityYear="1994" box="[985,1368,436,462]" class="Insecta" family="Curculionidae" genus="Howeotranes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCAFF9A0A965096FBDE9E99" box="[985,1141,437,462]" italics="true" pageId="5" pageNumber="122">Howeotranes</emphasis>
<bibRefCitation id="DD27E1FCFFCAFF9A0D325097FAF39E99" author="Zimmerman E. C." box="[1149,1368,436,462]" pageId="5" pageNumber="122" refId="ref12493" refString="Zimmerman E. C. 1994. Australian Weevils (Coleoptera: Curculionoidea). Volume I. Orthoceri. Anthribidae to Attelabidae. The Primitive Weevils. CSIRO, Melbourne." type="book" year="1994">
Zimmerman, 
<quantity id="7E4E31E8FFCAFF9A0C525097FAF39E99" box="[1309,1368,436,462]" metricMagnitude="1" metricUnit="m" metricValue="5.06476" pageId="5" pageNumber="122" unit="in" value="1994.0">1994</quantity>
</bibRefCitation>
</taxonomicName>
in lacking sclerolepidia along its metanepisternal sutures, which are present in 
<taxonomicName id="7EB6E78EFFCAFF9A0D6050FBFAED9EA6" authority="Pascoe, 1872" authorityName="Pascoe" authorityYear="1872" box="[1071,1350,471,498]" class="Insecta" family="Curculionidae" genus="Demyrsus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCAFF9A0D6050FBFB0F9EA6" box="[1071,1188,472,497]" italics="true" pageId="5" pageNumber="122">Demyrsus</emphasis>
Pascoe, 1872
</taxonomicName>
and 
<taxonomicName id="7EB6E78EFFCAFF9A09F350D8FE789D42" authority="Hustache, 1934" authorityName="Hustache" authorityYear="1934" box="[188,467,506,533]" class="Insecta" family="Brachyceridae" genus="Siraton" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCAFF9A09F350D8FEB99D42" box="[188,274,507,533]" italics="true" pageId="5" pageNumber="122">Siraton</emphasis>
Hustache, 1934
</taxonomicName>
(
<bibRefCitation id="DD27E1FCFFCAFF9A08AD50D9FC9F9D42" author="Oberprieler R. G. &amp; Caldara R." box="[482,820,506,533]" pageId="5" pageNumber="122" pagination="55 - 58" refId="ref12131" refString="Oberprieler R. G. &amp; Caldara R. 2012. Siraton devillei Hustache (Coleoptera: Curculionidae), the mysterious weevil from the Isle of Elba: exiled no longer. Zootaxa 3573 (1): 55 - 58. https: // doi. org / 10.11646 / zootaxa. 3573.1.6" type="journal article" year="2012">Oberprieler &amp; Caldara 2012</bibRefCitation>
; 
<bibRefCitation id="DD27E1FCFFCAFF9A0A0E50D9FC109D42" author="Lyal C. H. C." box="[833,955,506,533]" pageId="5" pageNumber="122" pagination="529 - 570" refId="ref11908" refString="Lyal C. H. C. 2014. 3.7. 7 Molytinae Schoenherr, 1823. In: Leschen R. A. B. &amp; Beutel R. G. (eds) Handbook of Zoology. Arthropoda: Insecta: Coleoptera, Beetles. Volume 3: Morphology and Systematics (Phytophaga): 529 - 570. Walter de Gruyter, Berlin." type="book chapter" year="2014">Lyal 2014</bibRefCitation>
; 
<bibRefCitation id="DD27E1FCFFCAFF9A0A8750D9FAA99D42" author="Hsiao Y. &amp; Oberprieler R. G." box="[968,1282,506,533]" pageId="5" pageNumber="122" pagination="677 - 700" refId="ref11651" refString="Hsiao Y. &amp; Oberprieler R. G. 2020. A review of the trunk-boring cycad weevils in Australia, with description of a second species of Demyrsus Pascoe, 1872 (Coleoptera: Curculionidae: Molytinae). Austral Entomology 59 (4): 677 - 700. https: // doi. org / 10.1111 / aen. 12498" type="journal article" year="2020">Hsiao &amp; Oberprieler 2020</bibRefCitation>
), but how closely related it may be to the cycad-associated genera, 
<taxonomicName id="7EB6E78EFFCAFF9A0A02533DFC369D60" authorityName="Schoenherr" authorityYear="1843" box="[845,925,542,567]" class="Insecta" family="Curculionidae" genus="Tranes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCAFF9A0A02533DFC369D60" box="[845,925,542,567]" italics="true" pageId="5" pageNumber="122">Tranes</emphasis>
</taxonomicName>
and 
<taxonomicName id="7EB6E78EFFCAFF9A0A9B533DFBF69D6F" authorityName="Zimmerman" authorityYear="1994" box="[980,1117,542,568]" class="Insecta" family="Curculionidae" genus="Miltotranes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCAFF9A0A9B533DFBF69D6F" box="[980,1117,542,568]" italics="true" pageId="5" pageNumber="122">Miltotranes</emphasis>
</taxonomicName>
, is as yet unknown. The similarity of the anchor-shaped sclerite of the endophallus in 
<taxonomicName id="7EB6E78EFFCAFF9A0ACF5361FBAF9D0C" authorityName="Zimmerman" authorityYear="1994" box="[896,1028,578,603]" class="Insecta" family="Curculionidae" genus="Paratranes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCAFF9A0ACF5361FBAF9D0C" box="[896,1028,578,603]" italics="true" pageId="5" pageNumber="122">Paratranes</emphasis>
</taxonomicName>
and 
<taxonomicName id="7EB6E78EFFCAFF9A0D735361FEAA9D28" authority="(Pascoe, 1875)" baseAuthorityName="Pascoe" baseAuthorityYear="1875" class="Insecta" family="Curculionidae" genus="Tranes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="lyterioides">
<emphasis id="8BC2401FFFCAFF9A0D735361FAA59D0C" box="[1084,1294,577,603]" italics="true" pageId="5" pageNumber="122">Tranes lyterioides</emphasis>
(Pascoe, 1875)
</taxonomicName>
suggests a close relationship between these taxa and that 
<taxonomicName id="7EB6E78EFFCAFF9A0AD85346FC4C9D29" authorityName="Schoenherr" authorityYear="1843" box="[919,999,613,638]" class="Insecta" family="Curculionidae" genus="Tranes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCAFF9A0AD85346FC4C9D29" box="[919,999,613,638]" italics="true" pageId="5" pageNumber="122">Tranes</emphasis>
</taxonomicName>
in its current concept may be nonmonophyletic. The taxonomy of 
<taxonomicName id="7EB6E78EFFCAFF9A0B7153ABFD259DF6" authorityName="Schoenherr" authorityYear="1843" box="[574,654,648,673]" class="Insecta" family="Curculionidae" genus="Tranes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="8BC2401FFFCAFF9A0B7153ABFD259DF6" box="[574,654,648,673]" italics="true" pageId="5" pageNumber="122">Tranes</emphasis>
</taxonomicName>
is currently under investigation (unpubl. data 2020).
</paragraph>
</subSubSection>
</treatment>
</document>