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<document id="8B3B14E45BB6DB62459BC6368FD252BD" ID-DOI="10.11646/zootaxa.3846.1.1" ID-GBIF-Dataset="01a9d979-7b10-4c36-b1af-4acc2c0cb9a7" ID-ISSN="1175-5326" ID-Zenodo-Dep="286792" ID-ZooBank="97D4A04A-D75E-45CC-8A70-3EB3A4E94D9B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460464243854" checkinUser="plazi" docAuthor="Zwick, Peter" docDate="2014" docId="113287AD9B53FFD7FF13F8B2FAD2FC3B" docLanguage="en" docName="zt03846p041.pdf" docOrigin="Zootaxa 3846 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calvarium (Calvariellum) cochlearifer Zwick, 2014, n. sp." docType="treatment" docVersion="6" lastPageNumber="12" masterDocId="ED0BFFD59B59FFDCFF84FFF7FFB5FFF3" masterDocTitle="Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer" masterLastPageNumber="41" masterPageNumber="1" pageNumber="11" updateTime="1698304595582" updateUser="plazi">
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<mods:title id="2249D4999A8F9A8BEC9C20393A213B1C">Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer</mods:title>
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<mods:namePart id="123CB39D323D20F7092326F84B7AAF95">Zwick, Peter</mods:namePart>
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<mods:title id="CDAF83956B398ACC10567FAEFB247556">Zootaxa</mods:title>
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<mods:part id="B82BE2943491029EF8955A97E1765064">
<mods:date id="3277D647B4386B983B7921CB7FF83487">2014</mods:date>
<mods:detail id="CAB9B7DD102E01EB64A25F46972AB09F" type="volume">
<mods:number id="D7930B7F363DEA9C204379BF6CF44951">3846</mods:number>
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<mods:classification id="07432652ECDEC9CC661B452E60792935">journal article</mods:classification>
<mods:identifier id="82F517CF34A7999DE18AF8EF7956FB59" type="DOI">10.11646/zootaxa.3846.1.1</mods:identifier>
<mods:identifier id="1A764143F5587AD3BAF837D0ABD46DE8" type="GBIF-Dataset">01a9d979-7b10-4c36-b1af-4acc2c0cb9a7</mods:identifier>
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<treatment id="113287AD9B53FFD7FF13F8B2FAD2FC3B" ID-DOI="http://doi.org/10.5281/zenodo.6126811" ID-GBIF-Taxon="127669617" ID-Zenodo-Dep="6126811" LSID="urn:lsid:plazi:treatment:113287AD9B53FFD7FF13F8B2FAD2FC3B" httpUri="http://treatment.plazi.org/id/113287AD9B53FFD7FF13F8B2FAD2FC3B" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<subSubSection id="D18165309B53FFD6FF13F8B2FE9BF873" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="992436BB9B53FFD6FF13F8B2FD0CF8AC" blockId="10.[151,697,1861,1920]" box="[151,697,1861,1887]" pageId="10" pageNumber="11">
<heading id="C26C81D79B53FFD6FF13F8B2FD0CF8AC" bold="true" box="[151,697,1861,1887]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="ABEFEAA99B53FFD6FF13F8B2FD0CF8AC" bold="true" box="[151,697,1861,1887]" pageId="10" pageNumber="11">
<emphasis id="ABEFEAA99B53FFD6FF13F8B2FDC5F8AC" bold="true" box="[151,624,1861,1887]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="5E9B4D389B53FFD6FF13F8B2FDDFF8AC" box="[151,618,1861,1887]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cochlearifer" status="sp. nov." subGenus="Calvariellum">Calvarium (Calvariellum) cochlearifer</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="B0DC57D29B53FFD6FDF3F8B2FD0CF8AC" box="[631,697,1861,1887]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="992436BB9B53FFD6FF13F890FE9BF873" blockId="10.[151,697,1861,1920]" box="[151,302,1895,1920]" pageId="10" pageNumber="11">
(
<figureCitation id="01A02A3E9B53FFD6FF1BF890FE93F873" box="[159,294,1895,1920]" captionStart-0="FIGURES 26 − 28" captionStart-1="FIGURES 29 − 37" captionStartId-0="9.[151,264,1931,1954]" captionStartId-1="10.[151,264,1707,1730]" captionTargetBox-0="[253,1329,906,1914]" captionTargetBox-1="[246,1345,203,1671]" captionTargetId-0="figure@9.[245,1339,900,1920]" captionTargetId-1="figure@10.[228,1358,197,1686]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 26 − 28. Calvarium (Calvariellum) spp., females: 26, C. hamifer, n. sp. (paratype from Station Ck), abdomen with protracted ovipositor and prehensor in situ; 27, prehensor enlarged, diagrammatic. 28, C. cochlearifer, n. sp. (from Bamboo Ck), prehensor. Abbreviation: cf, circular fold. 27 and 28 to the same scale." captionText-1="FIGURES 29 − 37. Calvarium (Calvariellum) spp., males. C. cochlearifer, n. sp.: 29, T 8 and S 8, superimposed; 30, penis; 31, T 9, S 9 and tegmen (tn) with parameres (p) superimposed. C. bellendenker, n. sp.: 32, T 8; 33, S 8; 34, T 9; 35, S 9; 36, penis; 37, tegmen and parameres. Figures for a given species to the same scale." httpUri-0="https://zenodo.org/record/286798/files/figure.png" httpUri-1="https://zenodo.org/record/286799/files/figure.png" pageId="10" pageNumber="11">Figs. 28–31</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D18165309B53FFD7FF13F858FBEEFEB3" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="992436BB9B53FFD7FF13F858FD29FEEE" blockId="10.[151,1436,1967,2029]" lastBlockId="11.[151,1437,151,968]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="ABEFEAA99B53FFD6FF13F858FE08F83B" bold="true" box="[151,445,1967,1992]" pageId="10" pageNumber="11">
<typeStatus id="462088199B53FFD6FF13F858FF67F83B" box="[151,210,1967,1992]" pageId="10" pageNumber="11">Type</typeStatus>
material.
<typeStatus id="462088199B53FFD6FED4F858FE08F83B" box="[336,445,1967,1992]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="8F9DFD329B53FFD6FE4AF847FE41F83B" box="[462,500,1968,1992]" pageId="10" pageNumber="11" type="generic">1♂</specimenCount>
: “
<geoCoordinate id="FCAF507C9B53FFD6FD89F858FDE8F83B" box="[525,605,1967,1992]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="555" value="-15.03">15.03S</geoCoordinate>
<geoCoordinate id="FCAF507C9B53FFD6FDE2F858FD71F83B" box="[614,708,1967,1992]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="555" value="145.09">145.09E</geoCoordinate>
<quantity id="5E639B5E9B53FFD6FD4AF858FD4AF83B" box="[718,767,1967,1992]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="11" unit="km" value="3.0">3km</quantity>
NE of Mt.
<collectionCode id="FF8AAE7E9B53FFD6FC0DF847FC77F83B" box="[905,962,1968,1992]" pageId="10" pageNumber="11">QLD</collectionCode>
Webb
<date id="ED25107B9B53FFD6FB92F847FB2BF83B" box="[1046,1182,1968,1992]" pageId="10" pageNumber="11" value="1980-10-02">2 Oct. 1980</date>
D.H.Colless (Malaise Trap)” (
<collectionCode id="FF8AAE7E9B53FFD6FF7CF823FEF7F81F" LSID="urn:lsid:biocol.org:col:32981" box="[248,322,2004,2028]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="10" pageNumber="11">ANIC</collectionCode>
).
<emphasis id="ABEFEAA99B53FFD6FED9F823FE61F81E" bold="true" box="[349,468,2004,2029]" pageId="10" pageNumber="11">
<typeStatus id="462088199B53FFD6FED9F823FE61F81E" box="[349,468,2004,2029]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
</emphasis>
(all
<collectionCode id="FF8AAE7E9B53FFD6FD9CF823FDD7F81F" LSID="urn:lsid:biocol.org:col:32981" box="[536,610,2004,2028]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="10" pageNumber="11">ANIC</collectionCode>
):
<specimenCount id="8F9DFD329B53FFD6FDF9F823FD17F81F" box="[637,674,2004,2028]" pageId="10" pageNumber="11" type="generic">1♂</specimenCount>
: “
<geoCoordinate id="FCAF507C9B53FFD6FD45F823FCA4F81E" box="[705,785,2004,2029]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="555" value="-15.03">15.03S</geoCoordinate>
<geoCoordinate id="FCAF507C9B53FFD6FCA5F823FCCAF81E" box="[801,895,2004,2029]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="555" value="145.09">145.09E</geoCoordinate>
<quantity id="5E639B5E9B53FFD6FC0AF823FC0AF81E" box="[910,959,2004,2029]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="11" unit="km" value="3.0">3km</quantity>
NE of Mt.
<collectionCode id="FF8AAE7E9B53FFD6FBE4F823FB2FF81F" box="[1120,1178,2004,2028]" pageId="10" pageNumber="11">QLD</collectionCode>
Webb
<date id="ED25107B9B53FFD6FB7DF823FA29F81F" box="[1273,1436,2004,2028]" pageId="10" pageNumber="11" value="1981-04-30">30 Apr. 1981</date>
D.H.Colless (at light)”;
<specimenCount id="8F9DFD329B52FFD7FE27FF6FFE7CFF43" box="[419,457,152,176]" pageId="11" pageNumber="12" type="generic">1♂</specimenCount>
: “Moses Ck
<quantity id="5E639B5E9B52FFD7FDDDFF6FFD27FF43" box="[601,658,151,176]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="12" unit="km" value="4.0">4 km</quantity>
NbyE of Mt Finnigan
<collectionCode id="FF8AAE7E9B52FFD7FC16FF6FFC79FF43" box="[914,972,152,176]" pageId="11" pageNumber="12">QLD</collectionCode>
<date id="ED25107B9B52FFD7FC57FF60FBD1FF43" box="[979,1124,151,176]" pageId="11" pageNumber="12" value="1980-10-15">15 Oct. 1980</date>
D.H.Colless”;
<specimenCount id="8F9DFD329B52FFD7FA8BFF6FFA80FF43" box="[1295,1333,152,176]" pageId="11" pageNumber="12" type="generic">1♂</specimenCount>
: “
<geoCoordinate id="FCAF507C9B52FFD7FAC8FF60FA29FF43" box="[1356,1436,151,176]" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="555" value="-15.3">15.30S</geoCoordinate>
<geoCoordinate id="FCAF507C9B52FFD7FF13FF4BFF40FF26" box="[151,245,188,213]" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="555" value="145.16">145.16E</geoCoordinate>
1 k SE of Mt. Cook
<collectionCode id="FF8AAE7E9B52FFD7FE58FF4BFDA3FF27" box="[476,534,188,212]" pageId="11" pageNumber="12">QLD</collectionCode>
<date id="ED25107B9B52FFD7FD98FF4BFD13FF27" box="[540,678,188,212]" pageId="11" pageNumber="12" value="1989-10-13">13 Oct.1989</date>
D.H.Colless”;
<specimenCount id="8F9DFD329B52FFD7FCD4FF4BFCC2FF27" box="[848,887,188,212]" pageId="11" pageNumber="12" type="generic">2♂</specimenCount>
: “
<geoCoordinate id="FCAF507C9B52FFD7FC09FF4BFC69FF26" box="[909,988,188,213]" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="555" value="-15.03">15.03S</geoCoordinate>
<geoCoordinate id="FCAF507C9B52FFD7FC67FF4BFBF4FF26" box="[995,1089,188,213]" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="555" value="145.09">145.09E</geoCoordinate>
<quantity id="5E639B5E9B52FFD7FBCCFF4BFBCFFF26" box="[1096,1146,188,213]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="12" unit="km" value="3.0">3km</quantity>
NE of Mt. Webb
<collectionCode id="FF8AAE7E9B52FFD7FAC4FF4BFACFFF27" box="[1344,1402,188,212]" pageId="11" pageNumber="12">QLD</collectionCode>
<date id="ED25107B9B52FFD7FA04FF4BFEB9FF0B" pageId="11" pageNumber="12" value="1981-04-30">30 Apr. 1981</date>
D.H.Colless (at light)”;
<specimenCount id="8F9DFD329B52FFD7FDB0FF17FDE2FF0B" box="[564,599,224,248]" pageId="11" pageNumber="12" type="generic">1♂</specimenCount>
1♀: “Bamboo Ck., near Miallo [
<geoCoordinate id="FCAF507C9B52FFD7FC6DFF28FB3EFF0B" box="[1001,1163,223,248]" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="1" value="-16.389866">16° 23.392' S</geoCoordinate>
<geoCoordinate id="FCAF507C9B52FFD7FB13FF28FAFCFF04" box="[1175,1353,223,248]" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="1" value="145.37505">145° 22.503' E</geoCoordinate>
] N. of Mossman N.Qld.
<date id="ED25107B9B52FFD7FEDAFEF3FE44FEEE" box="[350,497,260,285]" pageId="11" pageNumber="12" value="1967-04-25">25 Apr. 1967</date>
D.H. Colless”.
</paragraph>
<paragraph id="992436BB9B52FFD7FF43FED0FBEEFEB3" blockId="11.[151,1437,151,968]" box="[199,1115,295,320]" pageId="11" pageNumber="12">
<emphasis id="ABEFEAA99B52FFD7FF43FED0FE93FEB3" bold="true" box="[199,294,295,320]" pageId="11" pageNumber="12">Habitus</emphasis>
. Similar to
<taxonomicName id="5E9B4D389B52FFD7FE2EFEDFFD95FEB3" box="[426,544,296,320]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="hamifer">
<emphasis id="ABEFEAA99B52FFD7FE2EFEDFFD95FEB3" box="[426,544,296,320]" italics="true" pageId="11" pageNumber="12">C. hamifer</emphasis>
</taxonomicName>
.
<collectionCode id="FF8AAE7E9B52FFD7FDAAFEDFFDE7FECC" box="[558,594,296,319]" pageId="11" pageNumber="12">BL</collectionCode>
<quantity id="5E639B5E9B52FFD7FDDDFEDFFD56FECC" box="[601,739,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="11" pageNumber="12" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.9–2.2 mm</quantity>
. Dark brown, appendages lighter.
</paragraph>
</subSubSection>
<subSubSection id="D18165309B52FFD7FF43FEBBFD9DFCE7" pageId="11" pageNumber="12" type="description">
<paragraph id="992436BB9B52FFD7FF43FEBBFDBAFE23" blockId="11.[151,1437,151,968]" pageId="11" pageNumber="12">
<emphasis id="ABEFEAA99B52FFD7FF43FEBBFEB1FE96" bold="true" box="[199,260,332,357]" pageId="11" pageNumber="12">Male</emphasis>
. T8 only with microtrichia, caudal edge slightly angular. Disc with grouped microtrichia, very short setae only near the caudolateral angles. Apodemes less than twice as long as plate, front edge of plate barely defined. Ventrally near the end of each apodeme is an irregularly shaped sclerotized pocket. S8 with wide U-shaped base, disc unsclerotized, bare (
<figureCitation id="01A02A3E9B52FFD7FE2AFE40FDB7FE23" box="[430,514,439,464]" captionStart="FIGURES 29 − 37" captionStartId="10.[151,264,1707,1730]" captionTargetBox="[246,1345,203,1671]" captionTargetId="figure@10.[228,1358,197,1686]" captionTargetPageId="10" captionText="FIGURES 29 − 37. Calvarium (Calvariellum) spp., males. C. cochlearifer, n. sp.: 29, T 8 and S 8, superimposed; 30, penis; 31, T 9, S 9 and tegmen (tn) with parameres (p) superimposed. C. bellendenker, n. sp.: 32, T 8; 33, S 8; 34, T 9; 35, S 9; 36, penis; 37, tegmen and parameres. Figures for a given species to the same scale." httpUri="https://zenodo.org/record/286799/files/figure.png" pageId="11" pageNumber="12">Fig. 29</figureCitation>
).
</paragraph>
<paragraph id="992436BB9B52FFD7FF43FE2BFD7DFD3F" blockId="11.[151,1437,151,968]" pageId="11" pageNumber="12">
T9 represented only by long rod-shaped apodemes ending in sharp tips. S9 with paired sclerotized strips in the narrow base, oval plate caudally vaguely bilobed, with lateral groups of fine setae. Tegmen with unpaired slender base, parameres in the form of slender gently inwardly curved barely sclerotized horns (
<figureCitation id="01A02A3E9B52FFD7FBF4FDD3FB76FDCF" box="[1136,1219,548,573]" captionStart="FIGURES 29 − 37" captionStartId="10.[151,264,1707,1730]" captionTargetBox="[246,1345,203,1671]" captionTargetId="figure@10.[228,1358,197,1686]" captionTargetPageId="10" captionText="FIGURES 29 − 37. Calvarium (Calvariellum) spp., males. C. cochlearifer, n. sp.: 29, T 8 and S 8, superimposed; 30, penis; 31, T 9, S 9 and tegmen (tn) with parameres (p) superimposed. C. bellendenker, n. sp.: 32, T 8; 33, S 8; 34, T 9; 35, S 9; 36, penis; 37, tegmen and parameres. Figures for a given species to the same scale." httpUri="https://zenodo.org/record/286799/files/figure.png" pageId="11" pageNumber="12">Fig. 31</figureCitation>
). Penis base squarish, front slightly concave, lateral corners blunt. Trigonium stout, a little inflated near midlength, ending in short lateral points and a median spoon-shaped appendage with downcurved blunt tip. The parameroids are longer than the trigonium. Their slender bases curve away from the trigonium but their wide pore-bearing flat tips overlap dorsally from the trigonium (
<figureCitation id="01A02A3E9B52FFD7FDE3FD43FD0EFD3F" box="[615,699,692,717]" captionStart="FIGURES 29 − 37" captionStartId="10.[151,264,1707,1730]" captionTargetBox="[246,1345,203,1671]" captionTargetId="figure@10.[228,1358,197,1686]" captionTargetPageId="10" captionText="FIGURES 29 − 37. Calvarium (Calvariellum) spp., males. C. cochlearifer, n. sp.: 29, T 8 and S 8, superimposed; 30, penis; 31, T 9, S 9 and tegmen (tn) with parameres (p) superimposed. C. bellendenker, n. sp.: 32, T 8; 33, S 8; 34, T 9; 35, S 9; 36, penis; 37, tegmen and parameres. Figures for a given species to the same scale." httpUri="https://zenodo.org/record/286799/files/figure.png" pageId="11" pageNumber="12">Fig. 30</figureCitation>
).
</paragraph>
<paragraph id="992436BB9B52FFD7FF43FD20FD9DFCE7" blockId="11.[151,1437,151,968]" pageId="11" pageNumber="12">
<emphasis id="ABEFEAA99B52FFD7FF43FD20FEABFD03" bold="true" box="[199,286,727,752]" pageId="11" pageNumber="12">Female</emphasis>
. A flat structure in the gonoduct consisting of two lateral sclerites with wide-lobed overlapping tips seems to be the prehensor (
<figureCitation id="01A02A3E9B52FFD7FE43FD0BFDA9FCE7" box="[455,540,764,789]" captionStart="FIGURES 26 − 28" captionStartId="9.[151,264,1931,1954]" captionTargetBox="[253,1329,906,1914]" captionTargetId="figure@9.[245,1339,900,1920]" captionTargetPageId="9" captionText="FIGURES 26 − 28. Calvarium (Calvariellum) spp., females: 26, C. hamifer, n. sp. (paratype from Station Ck), abdomen with protracted ovipositor and prehensor in situ; 27, prehensor enlarged, diagrammatic. 28, C. cochlearifer, n. sp. (from Bamboo Ck), prehensor. Abbreviation: cf, circular fold. 27 and 28 to the same scale." httpUri="https://zenodo.org/record/286798/files/figure.png" pageId="11" pageNumber="12">Fig. 28</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D18165309B52FFD7FF43FCE8FAD2FC3B" pageId="11" pageNumber="12" type="etymology">
<paragraph id="992436BB9B52FFD7FF43FCE8FE78FCAF" blockId="11.[151,1437,151,968]" pageId="11" pageNumber="12">
<emphasis id="ABEFEAA99B52FFD7FF43FCE8FEF2FCCB" bold="true" box="[199,327,799,824]" pageId="11" pageNumber="12">Etymology</emphasis>
. The Latin noun in apposition translates as spoon-carrier (
<emphasis id="ABEFEAA99B52FFD7FC58FCD7FB8BFCCB" box="[988,1086,800,824]" italics="true" pageId="11" pageNumber="12">cochlear</emphasis>
= spoon, Latin), a reference to the shape of the penis apex.
</paragraph>
<paragraph id="992436BB9B52FFD7FF43FC90FAD2FC3B" blockId="11.[151,1437,151,968]" pageId="11" pageNumber="12">
<emphasis id="ABEFEAA99B52FFD7FF43FC90FEB3FC73" bold="true" box="[199,262,871,896]" pageId="11" pageNumber="12">Note.</emphasis>
Known only from tropical North Queensland. The outgrowth from the ventral side of T8 and the strongly curved appendage at the penis tip suggest
<taxonomicName id="5E9B4D389B52FFD7FD51FC7BFCFBFC57" box="[725,846,908,932]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="hamifer">
<emphasis id="ABEFEAA99B52FFD7FD51FC7BFCFBFC57" box="[725,846,908,932]" italics="true" pageId="11" pageNumber="12">C. hamifer</emphasis>
</taxonomicName>
and
<taxonomicName id="5E9B4D389B52FFD7FC01FC7BFB9AFC57" box="[901,1071,908,932]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cochlearifer">
<emphasis id="ABEFEAA99B52FFD7FC01FC7BFB9AFC57" box="[901,1071,908,932]" italics="true" pageId="11" pageNumber="12">C. cochlearifer</emphasis>
</taxonomicName>
are sister taxa. They were taken together at Bamboo Creek, the association of sexes based on common occurrence seems nevertheless reliable.
</paragraph>
</subSubSection>
</treatment>
</document>
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