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<document id="F39E28BF0B05E4729E0B7395D2CA64CD" ID-DOI="10.5281/zenodo.5351598" ID-ISSN="2345-7600" ID-Zenodo-Dep="5351598" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630206282112" checkinUser="tatiana" docAuthor="Jäch, Manfred A., Díaz, Juan A. &amp; Skale, Andre" docDate="2013" docId="03AF8781EA312930CCE4BE0418FBDBBB" docLanguage="en" docName="61rbz053-071.pdf" docOrigin="Raffles Bulletin of Zoology 61 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Hydraena (Hydraenopsis) michaelbalkei Jäch &amp; Díaz &amp; Skale 2013, new species" docType="treatment" docVersion="4" lastPageNumber="65" masterDocId="FF96FFF9EA3D293CCC79BD781C2BDF4E" masterDocTitle="The Hydraenidae (Coleoptera) Of The Republic Of Singapore" masterLastPageNumber="71" masterPageNumber="53" pageNumber="65" updateTime="1699184966419" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="6291646C37CBAD398A84881D18156DD0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8137FB5AB894217199477F73B76F9C95">
<mods:title id="746B5F135EFF88D2C95EC70BA26FE7D1">The Hydraenidae (Coleoptera) Of The Republic Of Singapore</mods:title>
</mods:titleInfo>
<mods:name id="D58C831F6CCB82C5391917195D6EA468" type="personal">
<mods:role id="206C3E7E8936693F602CC87589333F18">
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<mods:namePart id="24926327873162BA0A3BE509C006EFC5">Jäch, Manfred A.</mods:namePart>
</mods:name>
<mods:name id="19543BA3DABC84DA6DFF69D5FA1A1DE8" type="personal">
<mods:role id="48B411638ED23800E7468AD0339E06F6">
<mods:roleTerm id="B6D095E8AFE6C162A2ECA8BFADBA696B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="82CA8F652920B89784D28FBF2C26673D">Díaz, Juan A.</mods:namePart>
</mods:name>
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<mods:role id="0A5706C5F78E783633DB7C03460B547C">
<mods:roleTerm id="7E1C92D099630ED12B25FECEF932DE9E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="47D512888564704E67D945138647E81E">Skale, Andre</mods:namePart>
</mods:name>
<mods:typeOfResource id="4D530ABCB6215D846DC24A325C1AFE09">text</mods:typeOfResource>
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<mods:titleInfo id="7C7C0BF0964AC3E98D08007992BD9F8F">
<mods:title id="C4D81E47C3CFE4A57C53E5B13A47F01E">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="0CC065C44030B641C8B49B6004346997">
<mods:date id="827FED000814E05B4DD6BD031D97E2A7">2013</mods:date>
<mods:detail id="E4AC6221C5D816BCE52D792B9EFEF2F0" type="pubDate">
<mods:number id="25D6D143FB764D8CD5AB0C25BF124B23">2013-02-28</mods:number>
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<mods:detail id="96E41339A1366B0E5B5F60821AB4C929" type="volume">
<mods:number id="78EABBD7A94DC2783957E1E6FD7989F2">61</mods:number>
</mods:detail>
<mods:detail id="7467C747D81ED19A796F06A048AD0CD7" type="issue">
<mods:number id="792C01FB07BE715A96CD54E4724B56A6">1</mods:number>
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<mods:classification id="FC056ABCAA641F144C3F176490FD9E16">journal article</mods:classification>
<mods:identifier id="3995B3430CE5CC08A83C851963EEE658" type="DOI">10.5281/zenodo.5351598</mods:identifier>
<mods:identifier id="DBE2ECA5F4AFA2B61CBAFFDE8DD7918A" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="18D04A5156F91F0FC61FE843FB271D48" type="Zenodo-Dep">5351598</mods:identifier>
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<treatment id="03AF8781EA312930CCE4BE0418FBDBBB" ID-DOI="http://doi.org/10.5281/zenodo.5449815" ID-Zenodo-Dep="5449815" LSID="urn:lsid:plazi:treatment:03AF8781EA312930CCE4BE0418FBDBBB" httpUri="http://treatment.plazi.org/id/03AF8781EA312930CCE4BE0418FBDBBB" lastPageNumber="65" pageId="12" pageNumber="65">
<subSubSection id="C31C651CEA312930CCE4BE041EDFDCDB" box="[157,756,892,917]" pageId="12" pageNumber="65" type="nomenclature">
<paragraph id="8BB93697EA312930CCE4BE041EDFDCDB" blockId="12.[157,756,892,949]" box="[157,756,892,917]" pageId="12" pageNumber="65">
<heading id="D0F181FBEA312930CCE4BE041EDFDCDB" bold="true" box="[157,756,892,917]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="65" reason="2">
<emphasis id="B972EA85EA312930CCE4BE041EDFDCDB" bold="true" box="[157,756,892,917]" pageId="12" pageNumber="65">
<taxonomicName id="4C064D14EA312930CCE4BE041E74DCDB" ID-CoL="3N3DR" authority="Jäch &amp; Díaz &amp; Skale, 2013" authorityName="Jäch &amp; Díaz &amp; Skale" authorityYear="2013" box="[157,607,892,917]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="michaelbalkei" status="sp. nov." subGenus="Hydraenopsis">
<emphasis id="B972EA85EA312930CCE4BE041D20DCDB" bold="true" box="[157,267,892,917]" italics="true" pageId="12" pageNumber="65">Hydraena</emphasis>
(
<emphasis id="B972EA85EA312930CD65BE041D98DCDB" bold="true" box="[284,435,892,917]" italics="true" pageId="12" pageNumber="65">Hydraenopsis</emphasis>
)
<emphasis id="B972EA85EA312930CDBCBE041E74DCDB" bold="true" box="[453,607,892,917]" italics="true" pageId="12" pageNumber="65">michaelbalkei</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24157FEEA312930CE17BE051EDFDCDB" box="[622,756,893,917]" pageId="12" pageNumber="65" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA312930CDFCBEE51E20DCFB" box="[389,523,925,949]" pageId="12" pageNumber="65" type="description">
<paragraph id="8BB93697EA312930CDFCBEE51E20DCFB" blockId="12.[157,756,892,949]" box="[389,523,925,949]" pageId="12" pageNumber="65">
(
<figureCitation id="133D2A12EA312930CDF4BEE51DF1DCFB" box="[397,474,925,949]" captionStart="Fig" captionStartId="7.[113,148,2062,2084]" captionTargetBox="[231,1337,197,2010]" captionTargetId="figure-8@7.[221,1352,177,2030]" captionTargetPageId="7" captionText="Fig. 8. Hydraena michaelbalkei: a, aedeagus in lateral view; b, aedeagus, dorsal view; c, aedeagus, ventral view; d, male sternite X and spiculum." figureDoi="http://doi.org/10.5281/zenodo.5351608" httpUri="https://zenodo.org/record/5351608/files/figure.png" pageId="12" pageNumber="65">Figs. 8</figureCitation>
,
<figureCitation id="133D2A12EA312930CD91BEE51E28DCFB" box="[488,515,925,949]" captionStart="Fig" captionStartId="14.[137,172,2090,2112]" captionTargetBox="[137,1470,1208,2063]" captionTargetId="figure-8@14.[137,1478,1202,2067]" captionTargetPageId="14" captionText="Fig. 15. Geographical map of Singapore, showing distribution of Hydraenidae." figureDoi="http://doi.org/10.5281/zenodo.5351618" httpUri="https://zenodo.org/record/5351618/files/figure.png" pageId="12" pageNumber="65">15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA312930CCF0BEA41D47DBCC" pageId="12" pageNumber="65" type="materials_examined">
<paragraph id="8BB93697EA312930CCF0BEA41E55DB41" blockId="12.[137,776,987,1039]" pageId="12" pageNumber="65">
<materialsCitation id="3B6E3CCAEA312930CCF0BEA41E51DB41" country="Singapore" latitude="1.3498334" location="Sime Road" longLatPrecision="1" longitude="103.81129" municipality="MacRitchie Reservoir" pageId="12" pageNumber="65" specimenCount="1" typeStatus="holotype">
<emphasis id="B972EA85EA312930CCF0BEA41D23DCBF" bold="true" box="[137,264,987,1009]" italics="true" pageId="12" pageNumber="65">
<typeStatus id="54BD8835EA312930CCF0BEA41C93DCBF" box="[137,184,988,1009]" pageId="12" pageNumber="65">Type</typeStatus>
locality.
</emphasis>
— Shaded pool with grass,
<geoCoordinate id="EE325050EA312930CE7ABEA41E5EDCBF" box="[515,629,988,1009]" degrees="1" direction="north" minutes="20.990" orientation="latitude" pageId="12" pageNumber="65" precision="1" value="1.3498334">1°20.990’N</geoCoordinate>
,
<geoCoordinate id="EE325050EA312930CE05BEA41F2FDCBC" box="[636,772,988,1010]" degrees="103" direction="east" minutes="48.677" orientation="longitude" pageId="12" pageNumber="65" precision="1" value="103.81129">103°48.677’E</geoCoordinate>
, near
<location id="8ED9604CEA312930CCC3BE811D03DB41" LSID="urn:lsid:plazi:treatment:03AF8781EA312930CCE4BE0418FBDBBB:8ED9604CEA312930CCC3BE811D03DB41" box="[186,296,1017,1039]" country="Singapore" latitude="1.3498334" longLatPrecision="1" longitude="103.81129" municipality="MacRitchie Reservoir" name="Sime Road" pageId="12" pageNumber="65">Sime Road</location>
,
<collectingMunicipality id="6BDDACEDEA312930CD4DBE811E20DB41" box="[308,523,1017,1039]" pageId="12" pageNumber="65">MacRitchie Reservoir</collectingMunicipality>
,
<collectingCountry id="F3117607EA312930CE6FBE811E51DB41" box="[534,634,1017,1039]" name="Singapore" pageId="12" pageNumber="65">Singapore</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB93697EA312930CCF0B94B1D47DBCC" blockId="12.[137,775,1073,1154]" pageId="12" pageNumber="65">
<emphasis id="B972EA85EA312930CCF0B94B1D15DB06" bold="true" box="[137,318,1073,1096]" pageId="12" pageNumber="65">
<emphasis id="B972EA85EA312930CCF0B94B1D36DB09" bold="true" box="[137,285,1073,1096]" italics="true" pageId="12" pageNumber="65">Type material.</emphasis>
—
</emphasis>
<materialsCitation id="3B6E3CCAEA312930CD31B94A1D47DBCC" collectingDate="1997-04-24" collectionCode="ZRC" collectorName="Maxillary" country="Singapore" location="Mac Ritchie" municipality="Sime" pageId="12" pageNumber="65" specimenCount="1" typeStatus="holotype">
<emphasis id="B972EA85EA312930CD31B94A1D8CDB09" bold="true" box="[328,423,1074,1095]" pageId="12" pageNumber="65">
<typeStatus id="54BD8835EA312930CD31B94A1D8CDB09" box="[328,423,1074,1095]" pageId="12" pageNumber="65" type="holotype">Holotype</typeStatus>
</emphasis>
male (
<collectionCode id="ED17AE52EA312930CD89B94A1E0EDB06" box="[496,549,1074,1096]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="65">ZRC</collectionCode>
): “
<collectingCountry id="F3117607EA312930CE3BB94A1EE4DB09" box="[578,719,1074,1095]" name="Singapore" pageId="12" pageNumber="65">SINGAPORE</collectingCountry>
,
<location id="8ED9604CEA312930CEA5B94B1CFFDB2B" LSID="urn:lsid:plazi:treatment:03AF8781EA312930CCE4BE0418FBDBBB:8ED9604CEA312930CEA5B94B1CFFDB2B" country="Singapore" municipality="Sime" name="Mac Ritchie" pageId="12" pageNumber="65">Mac Ritchie</location>
, nr
<collectingMunicipality id="6BDDACEDEA312930CC86B9371D19DB2B" box="[255,306,1103,1125]" pageId="12" pageNumber="65">Sime</collectingMunicipality>
rd,
<date id="FFB81057EA312930CD19B9371DEEDB2A" box="[352,453,1103,1124]" pageId="12" pageNumber="65" value="1997-04-24">
<collectingDate id="EFFCE9BFEA312930CD19B9371DEEDB2A" box="[352,453,1103,1124]" pageId="12" pageNumber="65" value="1997-04-24">24IV1997</collectingDate>
</date>
Balke &amp; Hendrich”.
<collectorName id="26F35341EA312930CEDCB9371F2DDB2B" box="[677,774,1103,1125]" pageId="12" pageNumber="65">Maxillary</collectorName>
palpi and legs missing.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA312930CCF0B9D2194BDB5B" pageId="12" pageNumber="65" type="diagnosis">
<paragraph id="8BB93697EA312930CCF0B9D2199ADE1B" blockId="12.[137,776,1194,1315]" lastBlockId="12.[835,1474,189,341]" pageId="12" pageNumber="65">
<emphasis id="B972EA85EA312930CCF0B9D21D2DDB8C" bold="true" box="[137,262,1194,1218]" italics="true" pageId="12" pageNumber="65">Diagnosis.</emphasis>
— Body length:
<quantity id="4CFE9B72EA312930CDA2B9D21E63DB8C" box="[475,584,1194,1218]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="12" pageNumber="65" unit="mm" value="1.25">1.25 mm</quantity>
. This species is very closely related with
<taxonomicName id="4C064D14EA312930CDC9B9B31D60DA4C" authority="Freitag &amp; Jach. Colouration" authorityName="Freitag &amp; Jach. Colouration" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="manguao">
<emphasis id="B972EA85EA312930CDC9B9B31EA5DBAC" box="[432,654,1227,1250]" italics="true" pageId="12" pageNumber="65">Hydraena manguao</emphasis>
Freitag &amp; Jäch. Colouration
</taxonomicName>
, dorsal punctation and body form agree very well with the latter (see Freitag &amp; Jäch, 2007:
<figureCitation id="133D2A12EA312930CEA5B8721F4BDF9B" box="[732,864,189,1314]" captionStart="Fig" captionStartId="11.[113,148,2063,2085]" captionTargetBox="[153,1421,173,2029]" captionTargetId="figure-8@11.[141,1429,173,2044]" captionTargetPageId="11" captionText="Fig. 11. Hydraena paulmoritz: a, aedeagus in lateral view; b, aedeagus, dorsal view; aedeagus, ventral view; d, male sternite X and spiculum; e, gonocoxite; f, female tergite X; g, h, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351614" httpUri="https://zenodo.org/record/5351614/files/figure.png" pageId="12" pageNumber="65">Fig. 11</figureCitation>
). Unfortunately, the legs of the
<typeStatus id="54BD8835EA312930C8ACBDC5191EDF9B" box="[1237,1333,189,213]" pageId="12" pageNumber="65" type="holotype">holotype</typeStatus>
are missing and therefore we are (except for the sexually dimorphic male tergite X) at present not able to find any significant characters to distinguish
<taxonomicName id="4C064D14EA312930C829BC671923DE78" authorityName="Jäch &amp; Díaz &amp; Skale" authorityYear="2013" box="[1104,1288,286,310]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="michaelbalkei">
<emphasis id="B972EA85EA312930C829BC671848DE78" box="[1104,1123,287,310]" italics="true" pageId="12" pageNumber="65">H</emphasis>
.
<emphasis id="B972EA85EA312930C80BBC671923DE78" box="[1138,1288,287,310]" italics="true" pageId="12" pageNumber="65">michaelbalkei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C064D14EA312930C944BC6719E9DE78" box="[1341,1474,286,310]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="manguao">
<emphasis id="B972EA85EA312930C944BC67197BDE78" box="[1341,1360,287,310]" italics="true" pageId="12" pageNumber="65">H</emphasis>
.
<emphasis id="B972EA85EA312930C926BC6719E9DE78" box="[1375,1474,287,310]" italics="true" pageId="12" pageNumber="65">manguao</emphasis>
</taxonomicName>
externally. Male tergite VIII apically distinctly produced.
</paragraph>
<paragraph id="8BB93697EA312930CF3ABC051F89DEBB" blockId="12.[835,1474,381,501]" pageId="12" pageNumber="65">
Male terminal sternite and spiculum (
<figureCitation id="133D2A12EA312930C89FBC051912DEDB" box="[1254,1337,381,405]" captionStart="Fig" captionStartId="7.[113,148,2062,2084]" captionTargetBox="[231,1337,197,2010]" captionTargetId="figure-8@7.[221,1352,177,2030]" captionTargetPageId="7" captionText="Fig. 8. Hydraena michaelbalkei: a, aedeagus in lateral view; b, aedeagus, dorsal view; c, aedeagus, ventral view; d, male sternite X and spiculum." figureDoi="http://doi.org/10.5281/zenodo.5351608" httpUri="https://zenodo.org/record/5351608/files/figure.png" pageId="12" pageNumber="65">Fig. 8d</figureCitation>
): Somewhat resembling a contrabass, more elongate than in
<taxonomicName id="4C064D14EA312930C947BCE719E9DEF8" box="[1342,1474,415,438]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="manguao">
<emphasis id="B972EA85EA312930C947BCE719E9DEF8" box="[1342,1474,415,438]" italics="true" pageId="12" pageNumber="65">H. manguao</emphasis>
</taxonomicName>
(Freitag &amp; Jäch 2007: Fig. 25d); apex more widely rounded.
</paragraph>
<paragraph id="8BB93697EA312930CF3ABF651984DC1B" blockId="12.[835,1475,541,854]" pageId="12" pageNumber="65">
Aedeagus (
<figureCitation id="133D2A12EA312930CFC5BF65180CDD7B" box="[956,1063,541,565]" captionStart="Fig" captionStartId="7.[113,148,2062,2084]" captionTargetBox="[231,1337,197,2010]" captionTargetId="figure-8@7.[221,1352,177,2030]" captionTargetPageId="7" captionText="Fig. 8. Hydraena michaelbalkei: a, aedeagus in lateral view; b, aedeagus, dorsal view; c, aedeagus, ventral view; d, male sternite X and spiculum." figureDoi="http://doi.org/10.5281/zenodo.5351608" httpUri="https://zenodo.org/record/5351608/files/figure.png" pageId="12" pageNumber="65">Fig. 8a–c</figureCitation>
): In ventral and dorsal view distinctly Y-shaped as in
<taxonomicName id="4C064D14EA312930CF8CBF471854DD18" box="[1013,1151,575,598]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="manguao">
<emphasis id="B972EA85EA312930CF8CBF471854DD18" box="[1013,1151,575,598]" italics="true" pageId="12" pageNumber="65">H. manguao</emphasis>
</taxonomicName>
(Freitag &amp; Jäch, 2007: Fig. 25a–c),
<taxonomicName id="4C064D14EA312930CFE2BF271883DD3B" authority="Jach &amp; Diaz" authorityName="Jach &amp; Diaz" authorityYear="1999" box="[923,1192,605,630]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="sautakei">
<emphasis id="B972EA85EA312930CFE2BF27183CDD38" box="[923,1047,607,630]" italics="true" pageId="12" pageNumber="65">H. sautakei</emphasis>
Jäch &amp; Díaz
</taxonomicName>
(Jäch &amp; Díaz, 1999:
<figureCitation id="133D2A12EA312930C9E1BF251F7ADDDB" captionStart="Fig" captionStartId="6.[136,171,2060,2082]" captionTargetBox="[166,1446,190,2011]" captionTargetId="figure-8@6.[43,1493,170,2020]" captionTargetPageId="6" captionText="Fig. 7. Hydraena yangae: a, aedeagus in lateral view; b, aedeagus, dorsal view (setae of left paramere omitted in a, b); c, aedeagus, ventral view (setae of right paramere omitted); d, male sternite X and spiculum; e, gonocoxite; f, female tergite X; g, h, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351606" httpUri="https://zenodo.org/record/5351606/files/figure.png" pageId="12" pageNumber="65">Fig. 7</figureCitation>
) and
<taxonomicName id="4C064D14EA312930CFF4BF07183EDDD8" authorityName="Sato" authorityYear="1959" box="[909,1045,639,662]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="miyatakei">
<emphasis id="B972EA85EA312930CFF4BF07183EDDD8" box="[909,1045,639,662]" italics="true" pageId="12" pageNumber="65">H. miyatakei</emphasis>
</taxonomicName>
(Jäch &amp; Díaz, 1999:
<figureCitation id="133D2A12EA312930C88EBF05191CDDDB" box="[1271,1335,637,661]" captionStart="Fig" captionStartId="7.[113,148,2062,2084]" captionTargetBox="[231,1337,197,2010]" captionTargetId="figure-8@7.[221,1352,177,2030]" captionTargetPageId="7" captionText="Fig. 8. Hydraena michaelbalkei: a, aedeagus in lateral view; b, aedeagus, dorsal view; c, aedeagus, ventral view; d, male sternite X and spiculum." figureDoi="http://doi.org/10.5281/zenodo.5351608" httpUri="https://zenodo.org/record/5351608/files/figure.png" pageId="12" pageNumber="65">Fig. 8</figureCitation>
). Main piece rather straight in lateral view; with only one short dorsal seta near base of distal lobe. Left paramere distinctly shorter than in
<taxonomicName id="4C064D14EA312930CF27BFA71FC9DDB8" box="[862,994,735,758]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="manguao">
<emphasis id="B972EA85EA312930CF27BFA71FC9DDB8" box="[862,994,735,758]" italics="true" pageId="12" pageNumber="65">H. manguao</emphasis>
</taxonomicName>
, distinctly thinner than in
<taxonomicName id="4C064D14EA312930C882BFA7195FDDB8" authorityName="Jach &amp; Diaz" authorityYear="1999" box="[1275,1396,735,758]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="sautakei">
<emphasis id="B972EA85EA312930C882BFA7195FDDB8" box="[1275,1396,735,758]" italics="true" pageId="12" pageNumber="65">H. sautakei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C064D14EA312930C9D1BFA71F80DC58" authorityName="Sato" authorityYear="1959" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="miyatakei">
<emphasis id="B972EA85EA312930C9D1BFA71F80DC58" italics="true" pageId="12" pageNumber="65">H. miyatakei</emphasis>
</taxonomicName>
, with few apical setae; right paramere (as in
<taxonomicName id="4C064D14EA312930C9D1BF871FB7DC78" authorityName="Jach &amp; Diaz" authorityYear="1999" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="sautakei">
<emphasis id="B972EA85EA312930C9D1BF871FB7DC78" italics="true" pageId="12" pageNumber="65">H. sautakei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C064D14EA312930CFA8BE671870DC78" authorityName="Sato" authorityYear="1959" box="[977,1115,799,822]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="miyatakei">
<emphasis id="B972EA85EA312930CFA8BE671870DC78" box="[977,1115,799,822]" italics="true" pageId="12" pageNumber="65">H. miyatakei</emphasis>
</taxonomicName>
) completely fused to main piece, mainly indicated by the presence of two groups of setae.
</paragraph>
<paragraph id="8BB93697EA312930CF3ABE05194BDB5B" blockId="12.[835,1474,893,1045]" pageId="12" pageNumber="65">Secondary sexual characters: Characters of male legs unknown. Male metaventral plaques thin and short (confined to basal half), carinate. Male tergite X with subapical, asymmetrically heart-shaped excavation; apex slightly excised, right corner longer than left one.</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA312930CF3AB944180CDBDB" pageId="12" pageNumber="65" type="etymology">
<paragraph id="8BB93697EA312930CF3AB944180CDBDB" blockId="12.[835,1474,1084,1173]" pageId="12" pageNumber="65">
<emphasis id="B972EA85EA312930CF3AB9441F95DB1B" bold="true" box="[835,958,1084,1109]" italics="true" pageId="12" pageNumber="65">Etymology.</emphasis>
— Named for Michael Balke (ZSM), excellent water beetle specialist, who collected the
<typeStatus id="54BD8835EA312930C887B9251970DB3B" box="[1278,1371,1117,1141]" pageId="12" pageNumber="65" type="holotype">holotype</typeStatus>
(together with Lars Hendrich).
</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA312930CF3AB9C418FBDBBB" pageId="12" pageNumber="65" type="distribution">
<paragraph id="8BB93697EA312930CF3AB9C418FBDBBB" blockId="12.[835,1473,1212,1269]" pageId="12" pageNumber="65">
<emphasis id="B972EA85EA312930CF3AB9C41FCBDB9B" bold="true" box="[835,992,1212,1237]" italics="true" pageId="12" pageNumber="65">Distribution.</emphasis>
— So far known only from
<collectingCountry id="F3117607EA312930C932B9C519EADB9B" box="[1355,1473,1213,1237]" name="Singapore" pageId="12" pageNumber="65">Singapore</collectingCountry>
(MacRitchie Reservoir), see
<figureCitation id="133D2A12EA312930C805B9A518E7DBBB" box="[1148,1228,1245,1269]" captionStart="Fig" captionStartId="14.[137,172,2090,2112]" captionTargetBox="[137,1470,1208,2063]" captionTargetId="figure-8@14.[137,1478,1202,2067]" captionTargetPageId="14" captionText="Fig. 15. Geographical map of Singapore, showing distribution of Hydraenidae." figureDoi="http://doi.org/10.5281/zenodo.5351618" httpUri="https://zenodo.org/record/5351618/files/figure.png" pageId="12" pageNumber="65">Fig. 15</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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