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<document id="0392874C5DF2AF905CEB0A116607B6C4" ID-DOI="10.5281/zenodo.5392079" ID-ISSN="1638-9387" ID-Zenodo-Dep="5392079" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630316032044" checkinUser="marcus" docAuthor="Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul &amp; Spiegel, Didier Van Den" docDate="2007" docId="03F4C378A2363053FC800210FE2FF935" docLanguage="en" docName="Zoosystema.2007.29.1.7-49.pdf" docOrigin="Zoosystema 29 (1)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Glyphiulus deharvengi Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel 2007, n. sp." docType="treatment" docVersion="3" lastPageNumber="34" masterDocId="FFCDBB00A22F3048FF94075DFFE6FF91" masterDocTitle="Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group" masterLastPageNumber="49" masterPageNumber="7" pageNumber="32" updateTime="1699223690750" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="A57AF8D9863E4D85EF772B597317F806" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="667EB899C39DA08668AE11A49BEB0AE8">
<mods:title id="A156C10B687B9F12EB1EB957A2110152">Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group</mods:title>
</mods:titleInfo>
<mods:name id="651B352B550AFD48D0B29671391E2CDE" type="personal">
<mods:role id="234E376CB8578D668150DE9349965F82">
<mods:roleTerm id="45622D80F4626F3C9AAE79A4EDC55F05">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BFCA9BD4C00726029D8C5782DE616F46">Golovatch, Sergei I.</mods:namePart>
<mods:affiliation id="B6D384C769C76F40F6AF7E7F11308E9D">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru</mods:affiliation>
<mods:nameIdentifier id="CFC8E108AECFA0AAE43B23A2A0275EDD" type="email">sgol@orc.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="D72057ACF1B666226A5882E4ABB0BCCD" type="personal">
<mods:role id="FABCD5D1BCE8F440443CFAEACF0BB711">
<mods:roleTerm id="F57906F764AA271480FD24ECE9F7ECDB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2F8DF85D4D27900B45D33ECE77215BE6">Geoffroy, Jean-Jacques</mods:namePart>
<mods:affiliation id="057F62FAD105291301A6429862F94C8E">Muséum national d’Histoire naturelle, Département Écologie et Gestion de la Biodiversité, USM 306 Réseaux trophiques du sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="1DB730FE05623E7D380473D75AF259FF" type="email">geoffroy@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="F2DCD4D7A03D243BEBF90090D9508BD7" type="personal">
<mods:role id="7F99F13716A776609D4690CDC60215B8">
<mods:roleTerm id="4678A296A04DF81ABD0344F8612A989B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C6DD74202E289F8178A4B079505DB0EE">Mauriès, Jean-Paul</mods:namePart>
<mods:affiliation id="CE5EAB188A68FFF0BCE7D35E13B133BE">Muséum national d’Histoire naturelle, Département Systématique et Évolution, USM 602, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="FC081D1BC7F20FCA5C3AF2F8DC25BC85" type="email">collmill@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="393936D1DFF339E0567376B4B901145B" type="personal">
<mods:role id="E22F31E3DC6CC5D05A1E9BA256298A98">
<mods:roleTerm id="D679BA6136C0E61FB995E664119A8E33">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AFD06CE2219125DB38AAB8880277B663">Spiegel, Didier Van Den</mods:namePart>
<mods:affiliation id="EC23A7FC4D90830E49B45376A85EF050">Musée royal de l’Afrique centrale, Leuvensesteenweg 13, B- 3080 Tervuren (Belgium) didier. van. den. spiegel @ africamuseum. be</mods:affiliation>
</mods:name>
<mods:typeOfResource id="EDCD24F058F73E568B22E0815DFCB6DE">text</mods:typeOfResource>
<mods:relatedItem id="533BACA681D94B4E98F5CAC5560810DA" type="host">
<mods:titleInfo id="4D8A074F638E58C18FCDD4D0DE219110">
<mods:title id="4BD5F6457FCA9AC0F15BD954DF36BB22">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="8737A10C1434DC0DF467135B1773FF11">
<mods:date id="B9CD5F84459BDF85ACCB1FAD55E6F4E1">2007</mods:date>
<mods:detail id="913034ABF9FEC571E1F9A6761444A0C3" type="volume">
<mods:number id="3B239E8DD31EFBE8DCFCC15227864FA9">29</mods:number>
</mods:detail>
<mods:detail id="C190992C9B4A3CAFF115BF45E45A0B3D" type="issue">
<mods:number id="C8F4D3961E913E3D0444DD50007FBB8C">1</mods:number>
</mods:detail>
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<mods:start id="D421144227B853BBE04A7CD5EE458FF7">7</mods:start>
<mods:end id="C4E0BA957E25EA57601FD8DFCDD8B472">49</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="0E91D0D6423CDB43879FA4E568F0BFBF">journal article</mods:classification>
<mods:identifier id="3297E99213461FDEBE8A4370C1510413" type="DOI">10.5281/zenodo.5392079</mods:identifier>
<mods:identifier id="5220303FF0457D1D5B3778202D536D63" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="36571CEFA68F9A320EAD160DFAA5440E" type="Zenodo-Dep">5392079</mods:identifier>
</mods:mods>
<treatment id="03F4C378A2363053FC800210FE2FF935" LSID="urn:lsid:plazi:treatment:03F4C378A2363053FC800210FE2FF935" httpUri="http://treatment.plazi.org/id/03F4C378A2363053FC800210FE2FF935" lastPageId="27" lastPageNumber="34" pageId="25" pageNumber="32">
<subSubSection id="C34721E5A2363051FC800210FBACFAF6" box="[788,1098,1357,1383]" pageId="25" pageNumber="32" type="nomenclature">
<paragraph id="8BE2726EA2363051FC800210FBACFAF6" blockId="25.[788,1098,1357,1415]" box="[788,1098,1357,1383]" pageId="25" pageNumber="32">
<heading id="D0AAC502A2363051FC800210FBACFAF6" box="[788,1098,1357,1383]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="32" reason="2">
<taxonomicName id="4C5D09EDA2363051FC800210FBEEFAF6" ID-CoL="3GGJP" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[788,1032,1357,1383]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="32" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
<emphasis id="B929AE7CA2363051FC800210FBEEFAF6" bold="true" box="[788,1032,1357,1383]" italics="true" pageId="25" pageNumber="32">Glyphiulus deharvengi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A1307A2363051FB9B0210FBACFAF6" box="[1039,1098,1357,1383]" pageId="25" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34721E5A2363051FCFE0230FC12FA16" box="[874,1012,1389,1415]" pageId="25" pageNumber="32" type="description">
<paragraph id="8BE2726EA2363051FCFE0230FC12FA16" blockId="25.[788,1098,1357,1415]" box="[874,1012,1389,1415]" pageId="25" pageNumber="32">
(
<figureCitation id="13666EEBA2363051FCE50230FC25FA16" box="[881,963,1389,1415]" captionStart="FIG" captionStartId="24.[113,124,1635,1651]" captionTargetBox="[120,1199,216,1503]" captionTargetId="figure-44@24.[390,661,767,951]" captionTargetPageId="24" captionText="FIG. 19. — Glyphiulus deharvengi n. sp., ♀ paratype: A, anterior part of body, lateral view; B, same, dorsal view; C, middle part of body, dorsal view; D, telson, ventral view; E, midbody segment section, caudal view; F, claw of a midbody leg. Scale bars: A, C, D, 0.5 mm; B, E, 1.0 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392119" httpUri="https://zenodo.org/record/5392119/files/figure.png" pageId="25" pageNumber="32">Figs 19</figureCitation>
;
<figureCitation id="13666EEBA2363051FC5B0230FC0AFA16" box="[975,1004,1389,1415]" captionStart="FIG" captionStartId="25.[113,124,985,1001]" captionTargetBox="[190,1126,202,956]" captionTargetId="figure-14@25.[190,1126,202,956]" captionTargetPageId="25" captionText="FIG. 20. — Glyphiulus deharvengi n. sp., ♂ paratype: A, antenna; B, gnathochilarium, caudal view; C, legs 1, caudal view; D, legs 2, caudal view;E, leg 3; F, midbody leg; G, anterior gonopods, caudal view;H, posterior gonopod, subfrontal view.Scale bar: A-F, 0.6 mm; G, H, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.5392121" httpUri="https://zenodo.org/record/5392121/files/figure.png" pageId="25" pageNumber="32">20</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34721E5A2363052FD3102EFFEC0FBEA" lastPageId="26" lastPageNumber="33" pageId="25" pageNumber="32" type="materials_examined">
<paragraph id="8BE2726EA2363052FD3102EFFEC0FBEA" blockId="25.[675,1210,1458,1699]" lastBlockId="26.[113,647,1071,1147]" lastPageId="26" lastPageNumber="33" pageId="25" pageNumber="32">
<materialsCitation id="3B357833A2363051FD3102EFFBC0F98A" collectingDate="1995-08-11" collectionCode="MNHN" collectorName="A. Bedos &amp; L. Deharveng" country="China" county="Longshan" location="Feihu Dong Cave" municipality="Huoyan" pageId="25" pageNumber="32" specimenCode="GA 040" specimenCount="1" specimenCount-male="1" stateProvince="Hunan" typeStatus="holotype">
TYPE MATERIAL. —
<emphasis id="B929AE7CA2363051FCE402EFFC49FA58" bold="true" box="[880,943,1458,1481]" pageId="25" pageNumber="32">
<collectingCountry id="F34A32FEA2363051FCE402EFFC49FA58" box="[880,943,1458,1481]" name="China" pageId="25" pageNumber="32">China</collectingCountry>
</emphasis>
.
<collectingRegion id="4999BC8CA2363051FC2B02EFFBA3FA58" box="[959,1093,1458,1481]" country="China" name="Hunan" pageId="25" pageNumber="32">Hunan Prov.</collectingRegion>
,
<collectingCounty id="62830AE2A2363051FBC402EFFB50FA58" box="[1104,1206,1458,1481]" pageId="25" pageNumber="32">Longshan</collectingCounty>
:
<collectingMunicipality id="6B86E814A2363051FD330290FCE6FA75" box="[679,768,1485,1508]" pageId="25" pageNumber="32">Huoyan</collectingMunicipality>
,
<location id="8E8224B5A2363051FC9A0290FC34FA75" LSID="urn:lsid:plazi:treatment:03F4C378A2363053FC800210FE2FF935:8E8224B5A2363051FC9A0290FC34FA75" box="[782,978,1485,1508]" country="China" county="Longshan" municipality="Huoyan" name="Feihu Dong Cave" pageId="25" pageNumber="32" stateProvince="Hunan">Feihu Dong Cave</location>
(
<location id="8E8224B5A2363051FC770290FB49FA75" LSID="urn:lsid:plazi:treatment:03F4C378A2363053FC800210FE2FF935:8E8224B5A2363051FC770290FB49FA75" box="[995,1199,1485,1508]" country="China" county="Longshan" municipality="Huoyan" name="Flying Tiger Cave" pageId="25" pageNumber="32" stateProvince="Hunan">Flying Tiger Cave</location>
),
<date id="FFE354AEA2363051FD3102B5FCC8FA6E" box="[677,814,1512,1535]" pageId="25" pageNumber="32" value="1995-08-11">
<collectingDate id="EFA7AD46A2363051FD3102B5FCC8FA6E" box="[677,814,1512,1535]" pageId="25" pageNumber="32" value="1995-08-11">11.VIII.1995</collectingDate>
</date>
, leg.
<collectorName id="26A817B8A2363051FCF202B5FC27FA6E" box="[870,961,1512,1535]" pageId="25" pageNumber="32">A. Bedos</collectorName>
&amp;
<collectorName id="26A817B8A2363051FC7202B5FB97FA6E" box="[998,1137,1512,1535]" pageId="25" pageNumber="32">L. Deharveng</collectorName>
(CHI- 022),
<typeStatus id="54E6CCCCA2363051FD770159FCDAF98A" box="[739,828,1540,1563]" pageId="25" pageNumber="32" type="holotype">holotype</typeStatus>
<specimenCount id="9D5BB9E7A2363051FCD1015BFCBFF98B" box="[837,857,1542,1562]" pageId="25" pageNumber="32" type="male">♂</specimenCount>
(
<collectionCode id="ED4CEAABA2363051FCFD0159FC59F98A" box="[873,959,1540,1563]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="32">MNHN</collectionCode>
<specimenCode id="DBFBDA15A2363051FC5C0159FBF8F98A" box="[968,1054,1540,1563]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="32">GA 040</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B357833A2363051FBA60159FB53F916" collectingDate="1995-08-15" collectionCode="MNHN" collectorName="A. Bedos &amp; L. Deharveng" country="China" county="Longshan" location="Baiyan Dong Cave" municipality="Huoyan" pageId="25" pageNumber="32" specimenCode="GA 040" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Hunan" typeStatus="paratype">
<typeStatus id="54E6CCCCA2363051FBA60159FB6EF98A" box="[1074,1160,1540,1563]" pageId="25" pageNumber="32" type="paratype">paratype</typeStatus>
<specimenCount id="9D5BB9E7A2363051FB060159FB5FF98B" box="[1170,1209,1540,1563]" pageId="25" pageNumber="32" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED4CEAABA2363051FD380142FCE0F9A7" box="[684,774,1567,1590]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="32">MNHN</collectionCode>
<specimenCode id="DBFBDA15A2363051FC860142FC89F9A7" box="[786,879,1567,1590]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="32">GA 040</specimenCode>
). —
<collectingRegion id="4999BC8CA2363051FC3C0142FBDCF9A7" box="[936,1082,1567,1590]" country="China" name="Hunan" pageId="25" pageNumber="32">Hunan Prov.</collectingRegion>
,
<collectingCounty id="62830AE2A2363051FBDC0142FB52F9A7" box="[1096,1204,1567,1590]" pageId="25" pageNumber="32">Longshan</collectingCounty>
,
<collectingMunicipality id="6B86E814A2363051FD310167FD1AF9C0" box="[677,764,1594,1617]" pageId="25" pageNumber="32">Huoyan</collectingMunicipality>
,
<location id="8E8224B5A2363051FC9D0167FC37F9C0" LSID="urn:lsid:plazi:treatment:03F4C378A2363053FC800210FE2FF935:8E8224B5A2363051FC9D0167FC37F9C0" box="[777,977,1594,1617]" country="China" county="Longshan" municipality="Huoyan" name="Baiyan Dong Cave" pageId="25" pageNumber="32" stateProvince="Hunan">Baiyan Dong Cave</location>
(
<location id="8E8224B5A2363051FC750167FB48F9C0" LSID="urn:lsid:plazi:treatment:03F4C378A2363053FC800210FE2FF935:8E8224B5A2363051FC750167FB48F9C0" box="[993,1198,1594,1617]" country="China" county="Longshan" municipality="Huoyan" name="White Rocks Cave" pageId="25" pageNumber="32" stateProvince="Hunan">White Rocks Cave</location>
),
<date id="FFE354AEA2363051FD310108FCC8F9FD" box="[677,814,1621,1644]" pageId="25" pageNumber="32" value="1995-08-15">
<collectingDate id="EFA7AD46A2363051FD310108FCC8F9FD" box="[677,814,1621,1644]" pageId="25" pageNumber="32" value="1995-08-15">15.VIII.1995</collectingDate>
</date>
, leg.
<collectorName id="26A817B8A2363051FCF20108FC27F9FD" box="[870,961,1621,1644]" pageId="25" pageNumber="32">A. Bedos</collectorName>
&amp;
<collectorName id="26A817B8A2363051FC720108FB97F9FD" box="[998,1137,1621,1644]" pageId="25" pageNumber="32">L. Deharveng</collectorName>
(CHI- 032),
<typeStatus id="54E6CCCCA2363051FD74012DFCD8F916" box="[736,830,1648,1671]" pageId="25" pageNumber="32" type="paratype">paratypes</typeStatus>
<specimenCount id="9D5BB9E7A2363051FCD7012DFC8FF916" box="[835,873,1648,1671]" pageId="25" pageNumber="32" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D5BB9E7A2363051FCE1012DFC7CF916" box="[885,922,1648,1671]" pageId="25" pageNumber="32" type="female">1 ♀</specimenCount>
,
<specimenCount id="9D5BB9E7A2363051FC32012DFC2DF916" box="[934,971,1648,1671]" pageId="25" pageNumber="32" type="female">1 ♀</specimenCount>
juv. (
<collectionCode id="ED4CEAABA2363051FB95012DFBB0F916" box="[1025,1110,1648,1671]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="32">MNHN</collectionCode>
<specimenCode id="DBFBDA15A2363051FBCF012DFB48F916" box="[1115,1198,1648,1671]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="32">GA 040</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B357833A2363052FD3701D1FEBCFBF0" collectingDate="1995-08-19" collectionCode="ZMUM" collectorName="A. Bedos &amp; L. Deharveng" country="China" county="Longshan" lastPageId="26" lastPageNumber="33" location="Renmi Dong Cave" municipality="Huoyan" pageId="25" pageNumber="32" specimenCount="1" specimenCount-male="1" stateProvince="Hunan" typeStatus="paratype">
<specimenCount id="9D5BB9E7A2363051FD3701D1FD2AF933" box="[675,716,1676,1699]" pageId="25" pageNumber="32" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED4CEAABA2363051FD4F01D1FCD5F932" box="[731,819,1676,1699]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="25" pageNumber="32">ZMUM</collectionCode>
). —
<collectingRegion id="4999BC8CA2363051FCF101D1FC0EF932" box="[869,1000,1676,1699]" country="China" name="Hunan" pageId="25" pageNumber="32">Hunan Prov.</collectingRegion>
,
<collectingCounty id="62830AE2A2363051FC6601D1FBBEF932" box="[1010,1112,1676,1699]" pageId="25" pageNumber="32">Longshan</collectingCounty>
:
<collectingMunicipality id="6B86E814A2363051FBF601D1FB53F932" box="[1122,1205,1676,1699]" pageId="25" pageNumber="32">Huoyan</collectingMunicipality>
,
<location id="8E8224B5A2353052FFE50372FED6FBD7" LSID="urn:lsid:plazi:treatment:03F4C378A2363053FC800210FE2FF935:8E8224B5A2353052FFE50372FED6FBD7" box="[113,304,1071,1094]" country="China" county="Longshan" municipality="Huoyan" name="Renmi Dong Cave" pageId="26" pageNumber="33" stateProvince="Hunan">Renmi Dong Cave</location>
,
<date id="FFE354AEA2353052FEAE0372FE25FBD7" box="[314,451,1071,1094]" pageId="26" pageNumber="33" value="1995-08-19">
<collectingDate id="EFA7AD46A2353052FEAE0372FE25FBD7" box="[314,451,1071,1094]" pageId="26" pageNumber="33" value="1995-08-19">19.VIII.1995</collectingDate>
</date>
, leg.
<collectorName id="26A817B8A2353052FE630372FDB6FBD7" box="[503,592,1071,1094]" pageId="26" pageNumber="33">A. Bedos</collectorName>
&amp;
<collectorName id="26A817B8A2353052FDE60372FF39FBF0" pageId="26" pageNumber="33">L. Deharveng</collectorName>
(CHI-043)
</materialsCitation>
;
<materialsCitation id="3B357833A2353052FEF30317FF43FBEA" collectionCode="MNHN" country="China" pageId="26" pageNumber="33" specimenCode="GA 040" specimenCount="1" specimenCount-male="1" stateProvince="Hunan" typeStatus="paratype">
<typeStatus id="54E6CCCCA2353052FEF30317FE23FBF0" box="[359,453,1098,1121]" pageId="26" pageNumber="33" type="paratype">paratypes</typeStatus>
<specimenCount id="9D5BB9E7A2353052FE5A0317FE13FBF1" box="[462,501,1098,1121]" pageId="26" pageNumber="33" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED4CEAABA2353052FD910317FDBCFBF0" box="[517,602,1098,1121]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="26" pageNumber="33">MNHN</collectionCode>
<specimenCode id="DBFBDA15A2353052FDF70317FF7BFBEA" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="26" pageNumber="33">GA 040</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B357833A2353052FF240339FEC7FBEA" box="[176,289,1124,1147]" collectionCode="SEM" country="China" pageId="26" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Hunan" typeStatus="paratype">
<specimenCount id="9D5BB9E7A2353052FF240339FF35FBEA" box="[176,211,1124,1147]" pageId="26" pageNumber="33" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED4CEAABA2353052FF760339FEFDFBEA" box="[226,283,1124,1147]" pageId="26" pageNumber="33">SEM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF2222E6A2353052FFE5049CFD8AFC76" ID-DOI="http://doi.org/10.5281/zenodo.5392123" ID-Zenodo-Dep="5392123" httpUri="https://zenodo.org/record/5392123/files/figure.png" pageId="26" pageNumber="33" startId="26.[113,124,961,977]" targetBox="[119,1201,217,909]" targetPageId="26">
<paragraph id="8BE2726EA2353052FFE5049CFD8AFC76" blockId="26.[113,1209,960,1000]" pageId="26" pageNumber="33">
FIG. 21. —
<taxonomicName id="4C5D09EDA2353052FF58049DFE7CFC40" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[204,410,960,977]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="26" pageNumber="33" phylum="Arthropoda" rank="species" species="pergranulatus" status="sp. nov.">
<emphasis id="B929AE7CA2353052FF58049DFE7CFC40" box="[204,410,960,977]" italics="true" pageId="26" pageNumber="33">Glyphiulus pergranulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A1307A2353052FE0B049CFE36FC40" box="[415,464,961,977]" pageId="26" pageNumber="33" rank="species">n. sp.</taxonomicNameLabel>
, ♀ paratype:
<emphasis id="B929AE7CA2353052FDAB049DFDAAFC40" bold="true" box="[575,588,960,977]" pageId="26" pageNumber="33">A</emphasis>
, anterior part of body, lateral view;
<emphasis id="B929AE7CA2353052FCE7049DFC66FC41" bold="true" box="[883,896,960,976]" pageId="26" pageNumber="33">B</emphasis>
, same, dorsal view;
<emphasis id="B929AE7CA2353052FBB8049DFBDCFC40" bold="true" box="[1068,1082,960,977]" pageId="26" pageNumber="33">C</emphasis>
, middle part of body, lateral view;
<emphasis id="B929AE7CA2353052FE98048AFEFCFC76" bold="true" box="[268,282,983,999]" pageId="26" pageNumber="33">D</emphasis>
, same, dorsal view. Scale bars: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C34721E5A2353052FFE503C9FF33FB54" pageId="26" pageNumber="33" type="etymology">
<paragraph id="8BE2726EA2353052FFE503C9FF33FB54" blockId="26.[113,646,1172,1221]" pageId="26" pageNumber="33">ETYMOLOGY. — Honours Louis Deharveng, one of the collectors.</paragraph>
</subSubSection>
<subSubSection id="C34721E5A2353052FFE50383FEE0FA3F" pageId="26" pageNumber="33" type="diagnosis">
<paragraph id="8BE2726EA2353052FFE50383FEE0FA3F" blockId="26.[112,648,1246,1454]" pageId="26" pageNumber="33">DIAGNOSIS. — Differs by the relatively large size, the nearly completely smooth collum, the complete depigmentation (even of ocellaria), the elongated legs (with claws devoid of a spine at base from leg 4 onwards), the low, but evident, transverse pleural ridge behind the gonopod opening on male segment 7, the bare but flagelliform distal process of the posterior gonopods, etc.</paragraph>
</subSubSection>
<subSubSection id="C34721E5A2353053FFE502B6FF0BF9B4" lastPageId="27" lastPageNumber="34" pageId="26" pageNumber="33" type="description">
<paragraph id="8BE2726EA2353052FFE502B6FEE4F992" blockId="26.[111,648,1515,1700]" box="[113,258,1515,1541]" pageId="26" pageNumber="33">DESCRIPTION</paragraph>
<paragraph id="8BE2726EA2353052FFE50156FECBF935" blockId="26.[111,648,1515,1700]" pageId="26" pageNumber="33">
Length of adults of both sexes
<quantity id="4CA5DF8BA2353052FE570156FDDDF9B4" box="[451,571,1547,1573]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" metricValueMax="5.5" metricValueMin="4.0" pageId="26" pageNumber="33" unit="mm" value="47.5" valueMax="55.0" valueMin="40.0">40-55 mm</quantity>
, width 2.0-
<quantity id="4CA5DF8BA2353052FF0B0176FF1AF9D4" box="[159,252,1579,1605]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="26" pageNumber="33" unit="mm" value="2.3">2.3 mm</quantity>
. Coloration usually uniformly dark brown due to heavy sclerotization, tegument being coloured only due to chitin; sometimes ozoporiferous tubercles reddish.
</paragraph>
<paragraph id="8BE2726EA2353052FD2F036DFC5EFB58" blockId="26.[674,1211,1072,1700]" pageId="26" pageNumber="33">
Adults with 55-68p+4-1a+T; juvenile
<specimenCount id="9D5BB9E7A2353052FBFE036EFB9BFBD8" box="[1130,1149,1075,1097]" pageId="26" pageNumber="33" type="female">♀</specimenCount>
with 40p+5a+T, recognised by considerably reduced ozoporiferous tubercles on segments 7-33, plus 5a+T. Length of
<typeStatus id="54E6CCCCA2353052FCF203D2FC21FB38" box="[870,967,1167,1193]" pageId="26" pageNumber="33" type="holotype">holotype</typeStatus>
about
<quantity id="4CA5DF8BA2353052FB8C03D2FB88FB38" box="[1048,1134,1167,1193]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="26" pageNumber="33" unit="mm" value="55.0">55 mm</quantity>
, width
<quantity id="4CA5DF8BA2353052FD3203F2FD19FB58" box="[678,767,1199,1225]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="26" pageNumber="33" unit="mm" value="2.3">2.3 mm</quantity>
, with 67p+1a+T.
</paragraph>
<paragraph id="8BE2726EA2353052FD2F0393FCBAFAF6" blockId="26.[674,1211,1072,1700]" pageId="26" pageNumber="33">
Antennae slightly more elongate than usual (
<figureCitation id="13666EEBA2353052FB190393FD37FA99" captionStart="FIG" captionStartId="24.[113,124,1635,1651]" captionTargetBox="[120,1199,216,1503]" captionTargetId="figure-44@24.[390,661,767,951]" captionTargetPageId="24" captionText="FIG. 19. — Glyphiulus deharvengi n. sp., ♀ paratype: A, anterior part of body, lateral view; B, same, dorsal view; C, middle part of body, dorsal view; D, telson, ventral view; E, midbody segment section, caudal view; F, claw of a midbody leg. Scale bars: A, C, D, 0.5 mm; B, E, 1.0 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392119" httpUri="https://zenodo.org/record/5392119/files/figure.png" pageId="26" pageNumber="33">Figs 19A</figureCitation>
;
<figureCitation id="13666EEBA2353052FD4E03B3FCECFA99" box="[730,778,1262,1288]" captionStart="FIG" captionStartId="25.[113,124,985,1001]" captionTargetBox="[190,1126,202,956]" captionTargetId="figure-14@25.[190,1126,202,956]" captionTargetPageId="25" captionText="FIG. 20. — Glyphiulus deharvengi n. sp., ♂ paratype: A, antenna; B, gnathochilarium, caudal view; C, legs 1, caudal view; D, legs 2, caudal view;E, leg 3; F, midbody leg; G, anterior gonopods, caudal view;H, posterior gonopod, subfrontal view.Scale bar: A-F, 0.6 mm; G, H, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.5392121" httpUri="https://zenodo.org/record/5392121/files/figure.png" pageId="26" pageNumber="33">20A</figureCitation>
). Ocellaria unpigmented, barely traceable, with about 10 or 11 ocelli in largest specimens. Gnathochilarium (
<figureCitation id="13666EEBA2353052FCEC0270FC3DFAD6" box="[888,987,1325,1351]" captionStart="FIG" captionStartId="25.[113,124,985,1001]" captionTargetBox="[190,1126,202,956]" captionTargetId="figure-14@25.[190,1126,202,956]" captionTargetPageId="25" captionText="FIG. 20. — Glyphiulus deharvengi n. sp., ♂ paratype: A, antenna; B, gnathochilarium, caudal view; C, legs 1, caudal view; D, legs 2, caudal view;E, leg 3; F, midbody leg; G, anterior gonopods, caudal view;H, posterior gonopod, subfrontal view.Scale bar: A-F, 0.6 mm; G, H, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.5392121" httpUri="https://zenodo.org/record/5392121/files/figure.png" pageId="26" pageNumber="33">Fig. 20B</figureCitation>
) with an undivided mentum (n = 1).
</paragraph>
<paragraph id="8BE2726EA2353053FD2F0230FE4BFB58" blockId="26.[674,1211,1072,1700]" lastBlockId="27.[113,649,1167,1573]" lastPageId="27" lastPageNumber="34" pageId="26" pageNumber="33">
Carinotaxy of collum unusual in that all crests are nearly obliterated (
<figureCitation id="13666EEBA2353052FC0602D1FBF6FA37" box="[914,1040,1420,1446]" captionStart="FIG" captionStartId="24.[113,124,1635,1651]" captionTargetBox="[120,1199,216,1503]" captionTargetId="figure-44@24.[390,661,767,951]" captionTargetPageId="24" captionText="FIG. 19. — Glyphiulus deharvengi n. sp., ♀ paratype: A, anterior part of body, lateral view; B, same, dorsal view; C, middle part of body, dorsal view; D, telson, ventral view; E, midbody segment section, caudal view; F, claw of a midbody leg. Scale bars: A, C, D, 0.5 mm; B, E, 1.0 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392119" httpUri="https://zenodo.org/record/5392119/files/figure.png" pageId="26" pageNumber="33">Fig. 19A, B</figureCitation>
), though a typical pattern is traceable. Carinotaxy of subsequent metaterga typical (
<figureCitation id="13666EEBA2353052FCFA0291FC5FFA77" box="[878,953,1484,1510]" captionStart="FIG" captionStartId="24.[113,124,1635,1651]" captionTargetBox="[120,1199,216,1503]" captionTargetId="figure-44@24.[390,661,767,951]" captionTargetPageId="24" captionText="FIG. 19. — Glyphiulus deharvengi n. sp., ♀ paratype: A, anterior part of body, lateral view; B, same, dorsal view; C, middle part of body, dorsal view; D, telson, ventral view; E, midbody segment section, caudal view; F, claw of a midbody leg. Scale bars: A, C, D, 0.5 mm; B, E, 1.0 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392119" httpUri="https://zenodo.org/record/5392119/files/figure.png" pageId="26" pageNumber="33">Fig. 19</figureCitation>
A-C, E), but crests low, especially dorsal ones on segments 2-6; only two lateral crests below ozoporiferous ones, divided from segment 7 onwards, followed by a dozen striae. A low, but evident, transverse pleural ridge behind gonopod opening on male segment 7. Epiproct slightly elongate apically, more narrowly rounded than usual; hypoproct more narrowly emarginate at caudal margin (
<figureCitation id="13666EEBA2343053FEAC03F2FE46FB58" box="[312,416,1199,1225]" captionStart="FIG" captionStartId="24.[113,124,1635,1651]" captionTargetBox="[120,1199,216,1503]" captionTargetId="figure-44@24.[390,661,767,951]" captionTargetPageId="24" captionText="FIG. 19. — Glyphiulus deharvengi n. sp., ♀ paratype: A, anterior part of body, lateral view; B, same, dorsal view; C, middle part of body, dorsal view; D, telson, ventral view; E, midbody segment section, caudal view; F, claw of a midbody leg. Scale bars: A, C, D, 0.5 mm; B, E, 1.0 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392119" httpUri="https://zenodo.org/record/5392119/files/figure.png" pageId="27" pageNumber="34">Fig. 19D</figureCitation>
).
</paragraph>
<caption id="DF2222E6A2343053FFE50354FC78FBA1" ID-DOI="http://doi.org/10.5281/zenodo.5392125" ID-Zenodo-Dep="5392125" httpUri="https://zenodo.org/record/5392125/files/figure.png" pageId="27" pageNumber="34" startId="27.[113,124,1033,1049]" targetBox="[121,1197,215,984]" targetPageId="27">
<paragraph id="8BE2726EA2343053FFE50354FC78FBA1" blockId="27.[113,1209,1032,1073]" pageId="27" pageNumber="34">
FIG. 22. —
<taxonomicName id="4C5D09EDA2343053FF5C0355FE72FB88" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[200,404,1032,1049]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="27" pageNumber="34" phylum="Arthropoda" rank="species" species="pergranulatus" status="sp. nov.">
<emphasis id="B929AE7CA2343053FF5C0355FE72FB88" box="[200,404,1032,1049]" italics="true" pageId="27" pageNumber="34">Glyphiulus pergranulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A1307A2343053FE0C0354FE21FB88" box="[408,455,1033,1049]" pageId="27" pageNumber="34" rank="species">n. sp.</taxonomicNameLabel>
, ♀ paratype:
<emphasis id="B929AE7CA2343053FDA60355FDD9FB88" bold="true" box="[562,575,1032,1049]" pageId="27" pageNumber="34">A</emphasis>
, caudal part of body, ventral view;
<emphasis id="B929AE7CA2343053FCCE0354FC81FB88" bold="true" box="[858,871,1033,1049]" pageId="27" pageNumber="34">B</emphasis>
, same, lateral view;
<emphasis id="B929AE7CA2343053FB980355FBFCFB88" bold="true" box="[1036,1050,1032,1049]" pageId="27" pageNumber="34">C</emphasis>
, claw of a midbody leg;
<emphasis id="B929AE7CA2343053FF01037DFF45FBA1" bold="true" box="[149,163,1056,1072]" pageId="27" pageNumber="34">D</emphasis>
, midbody segment section, caudal view. Scale bars: A, 0.2 mm; B, D, 0.5 mm; C, 0.01 mm.
</paragraph>
</caption>
<paragraph id="8BE2726EA2343053FF1C0393FF03FAB9" blockId="27.[113,649,1167,1573]" pageId="27" pageNumber="34">
Legs elongate (
<figureCitation id="13666EEBA2343053FEBB0393FE75FB79" box="[303,403,1230,1256]" captionStart="FIG" captionStartId="25.[113,124,985,1001]" captionTargetBox="[190,1126,202,956]" captionTargetId="figure-14@25.[190,1126,202,956]" captionTargetPageId="25" captionText="FIG. 20. — Glyphiulus deharvengi n. sp., ♂ paratype: A, antenna; B, gnathochilarium, caudal view; C, legs 1, caudal view; D, legs 2, caudal view;E, leg 3; F, midbody leg; G, anterior gonopods, caudal view;H, posterior gonopod, subfrontal view.Scale bar: A-F, 0.6 mm; G, H, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.5392121" httpUri="https://zenodo.org/record/5392121/files/figure.png" pageId="27" pageNumber="34">Fig. 20F</figureCitation>
), claws strongly elongate, without a spine near base starting from leg 4 (
<figureCitation id="13666EEBA2343053FFED0253FF3FFAB9" box="[121,217,1294,1320]" captionStart="FIG" captionStartId="24.[113,124,1635,1651]" captionTargetBox="[120,1199,216,1503]" captionTargetId="figure-44@24.[390,661,767,951]" captionTargetPageId="24" captionText="FIG. 19. — Glyphiulus deharvengi n. sp., ♀ paratype: A, anterior part of body, lateral view; B, same, dorsal view; C, middle part of body, dorsal view; D, telson, ventral view; E, midbody segment section, caudal view; F, claw of a midbody leg. Scale bars: A, C, D, 0.5 mm; B, E, 1.0 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392119" httpUri="https://zenodo.org/record/5392119/files/figure.png" pageId="27" pageNumber="34">Fig. 19F</figureCitation>
).
</paragraph>
<paragraph id="8BE2726EA2343053FF1C0270FF0BF9B4" blockId="27.[113,649,1167,1573]" pageId="27" pageNumber="34">
Male legs 1 with vestigial tubercular coxites (
<figureCitation id="13666EEBA2343053FFEF0210FF06FAF6" box="[123,224,1357,1383]" captionStart="FIG" captionStartId="25.[113,124,985,1001]" captionTargetBox="[190,1126,202,956]" captionTargetId="figure-14@25.[190,1126,202,956]" captionTargetPageId="25" captionText="FIG. 20. — Glyphiulus deharvengi n. sp., ♂ paratype: A, antenna; B, gnathochilarium, caudal view; C, legs 1, caudal view; D, legs 2, caudal view;E, leg 3; F, midbody leg; G, anterior gonopods, caudal view;H, posterior gonopod, subfrontal view.Scale bar: A-F, 0.6 mm; G, H, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.5392121" httpUri="https://zenodo.org/record/5392121/files/figure.png" pageId="27" pageNumber="34">Fig. 20C</figureCitation>
). Penes with 4 or 5 long setae distolaterally (
<figureCitation id="13666EEBA2343053FF500230FECDFA16" box="[196,299,1389,1415]" captionStart="FIG" captionStartId="25.[113,124,985,1001]" captionTargetBox="[190,1126,202,956]" captionTargetId="figure-14@25.[190,1126,202,956]" captionTargetPageId="25" captionText="FIG. 20. — Glyphiulus deharvengi n. sp., ♂ paratype: A, antenna; B, gnathochilarium, caudal view; C, legs 1, caudal view; D, legs 2, caudal view;E, leg 3; F, midbody leg; G, anterior gonopods, caudal view;H, posterior gonopod, subfrontal view.Scale bar: A-F, 0.6 mm; G, H, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.5392121" httpUri="https://zenodo.org/record/5392121/files/figure.png" pageId="27" pageNumber="34">Fig. 20D</figureCitation>
), male legs 3 elongate, much as in
<taxonomicName id="4C5D09EDA2343053FF0502D1FEC2FA37" baseAuthorityName="Gervais" baseAuthorityYear="1847" box="[145,292,1420,1446]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="27" pageNumber="34" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B929AE7CA2343053FF0502D1FEC2FA37" box="[145,292,1420,1446]" italics="true" pageId="27" pageNumber="34">G. granulatus</emphasis>
</taxonomicName>
(
<figureCitation id="13666EEBA2343053FEA102D1FE7AFA37" box="[309,412,1420,1446]" captionStart="FIG" captionStartId="25.[113,124,985,1001]" captionTargetBox="[190,1126,202,956]" captionTargetId="figure-14@25.[190,1126,202,956]" captionTargetPageId="25" captionText="FIG. 20. — Glyphiulus deharvengi n. sp., ♂ paratype: A, antenna; B, gnathochilarium, caudal view; C, legs 1, caudal view; D, legs 2, caudal view;E, leg 3; F, midbody leg; G, anterior gonopods, caudal view;H, posterior gonopod, subfrontal view.Scale bar: A-F, 0.6 mm; G, H, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.5392121" httpUri="https://zenodo.org/record/5392121/files/figure.png" pageId="27" pageNumber="34">Fig. 20E</figureCitation>
). Anterior gonopods with small subunciform protuberances at median corners of coxosternum (
<figureCitation id="13666EEBA2343053FE150291FE01FA77" box="[385,487,1484,1510]" captionStart="FIG" captionStartId="25.[113,124,985,1001]" captionTargetBox="[190,1126,202,956]" captionTargetId="figure-14@25.[190,1126,202,956]" captionTargetPageId="25" captionText="FIG. 20. — Glyphiulus deharvengi n. sp., ♂ paratype: A, antenna; B, gnathochilarium, caudal view; C, legs 1, caudal view; D, legs 2, caudal view;E, leg 3; F, midbody leg; G, anterior gonopods, caudal view;H, posterior gonopod, subfrontal view.Scale bar: A-F, 0.6 mm; G, H, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.5392121" httpUri="https://zenodo.org/record/5392121/files/figure.png" pageId="27" pageNumber="34">Fig. 20G</figureCitation>
). Posterior gonopods with a flagelliform but bare distal process (
<figureCitation id="13666EEBA2343053FFED0156FF07F9B4" box="[121,225,1547,1573]" captionStart="FIG" captionStartId="25.[113,124,985,1001]" captionTargetBox="[190,1126,202,956]" captionTargetId="figure-14@25.[190,1126,202,956]" captionTargetPageId="25" captionText="FIG. 20. — Glyphiulus deharvengi n. sp., ♂ paratype: A, antenna; B, gnathochilarium, caudal view; C, legs 1, caudal view; D, legs 2, caudal view;E, leg 3; F, midbody leg; G, anterior gonopods, caudal view;H, posterior gonopod, subfrontal view.Scale bar: A-F, 0.6 mm; G, H, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.5392121" httpUri="https://zenodo.org/record/5392121/files/figure.png" pageId="27" pageNumber="34">Fig. 20H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34721E5A2343053FFE50117FE2FF935" pageId="27" pageNumber="34" type="discussion">
<paragraph id="8BE2726EA2343053FFE50117FF33F9F3" blockId="27.[113,646,1610,1700]" box="[113,213,1610,1636]" pageId="27" pageNumber="34">REMARKS</paragraph>
<paragraph id="8BE2726EA2343053FFE50137FE2FF935" blockId="27.[113,646,1610,1700]" pageId="27" pageNumber="34">Based on its evident troglomorphic traits, this species seems to be a troglobite.</paragraph>
</subSubSection>
</treatment>
</document>
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