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<document id="EFE60ADDD144BDE226D0D840646B0D2F" ID-CLB-Dataset="4593" ID-DOI="10.5281/zenodo.5353860" ID-GBIF-Dataset="600feca3-8ebf-4e20-bdd8-8eb2c2d454a5" ID-ISSN="1942-1354" ID-Zenodo-Dep="5353860" ID-ZooBank="CD47CC7D-0D74-42CA-902B-72889B0E8528" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630212701397" checkinUser="felipe" docAuthor="Rifkind, Jacques" docDate="2020" docId="03F49E67FFCFFFC77BDCFA152507F8AB" docLanguage="en" docName="0831.pdf" docOrigin="Insecta Mundi 2020 (831)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Aphelocerus thomasi Rifkind 2020, new species" docType="treatment" docVersion="7" lastPageNumber="2" masterDocId="FFCDE61FFFCDFFC47B1CFFD3201AFFA2" masterDocTitle="Aphelocerus thomasi, a new species of checkered beetle (Coleoptera: Cleridae: Clerinae) from Mexico" masterLastPageNumber="3" masterPageNumber="1" pageNumber="1" updateTime="1699211380402" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="B0649BB3C851314E9B219992AB7A2DB0">Aphelocerus thomasi, a new species of checkered beetle (Coleoptera: Cleridae: Clerinae) from Mexico</mods:title>
</mods:titleInfo>
<mods:name id="06A1C32BC33AD6919B1374B08AF10E82" type="personal">
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<mods:namePart id="7DAECCF224E19FAF79B4F52B93360782">Rifkind, Jacques</mods:namePart>
<mods:affiliation id="0A97CF476CD38853056C677759B4C063">Research Associate, California State Collection of Arthropods, 3294 Meadowview Road Sacramento, CA 95832 U. S. A.</mods:affiliation>
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<mods:title id="B26823BA04EE7A2229DAE51AC9D0E376">Insecta Mundi</mods:title>
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<mods:part id="416F62102D8B4CB33A2910602B5CCD7E">
<mods:date id="0A14FE05003B6B5586A95FEA3182A41D">2020</mods:date>
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<mods:identifier id="950DFCA49F0FBDA452B855E2991FD002" type="DOI">10.5281/zenodo.5353860</mods:identifier>
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<treatment id="03F49E67FFCFFFC77BDCFA152507F8AB" ID-DOI="http://doi.org/10.5281/zenodo.5451825" ID-GBIF-Taxon="185260834" ID-Zenodo-Dep="5451825" LSID="urn:lsid:plazi:treatment:03F49E67FFCFFFC77BDCFA152507F8AB" httpUri="http://treatment.plazi.org/id/03F49E67FFCFFFC77BDCFA152507F8AB" lastPageId="3" lastPageNumber="2" pageId="2" pageNumber="1">
<subSubSection id="C3477CFAFFCFFFC67BDCFA1522CBFA41" box="[192,721,1478,1507]" pageId="2" pageNumber="1" type="nomenclature">
<paragraph id="8BE22F71FFCFFFC67BDCFA1522CBFA41" blockId="2.[192,721,1478,1540]" box="[192,721,1478,1507]" pageId="2" pageNumber="1">
<heading id="D0AA981DFFCFFFC67BDCFA1522CBFA41" box="[192,721,1478,1507]" fontSize="12" level="2" pageId="2" pageNumber="1" reason="2">
<emphasis id="B929F363FFCFFFC67BDCFA1522CBFA41" bold="true" box="[192,721,1478,1507]" pageId="2" pageNumber="1">
<taxonomicName id="4C5D54F2FFCFFFC67BDCFA15222BFA41" authority="Rifkind" authorityName="Rifkind" authorityYear="2020" box="[192,561,1478,1507]" class="Insecta" family="Cleridae" genus="Aphelocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="thomasi" status="sp. nov.">
<emphasis id="B929F363FFCFFFC67BDCFA1521DFFA41" bold="true" box="[192,453,1478,1507]" italics="true" pageId="2" pageNumber="1">Aphelocerus thomasi</emphasis>
Rifkind
</taxonomicName>
,
<taxonomicNameLabel id="A21A4E18FFCFFFC67921FA1422CBFA41" box="[573,721,1478,1507]" pageId="2" pageNumber="1" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3477CFAFFCFFFC77BDCFA392260FAFC" lastPageId="3" lastPageNumber="2" pageId="2" pageNumber="1" type="description">
<paragraph id="8BE22F71FFCFFFC67BDCFA392137F9A6" blockId="2.[192,721,1478,1540]" box="[192,301,1514,1540]" pageId="2" pageNumber="1">
(
<figureCitation id="136633F4FFCFFFC67BD4FA39213EF9A6" box="[200,292,1514,1540]" captionStart="Figures 12" captionStartId="3.[285,372,958,984]" captionTargetBox="[285,1346,213,926]" captionTargetId="figure-351@3.[285,1347,213,927]" captionTargetPageId="3" captionText="Figures 12. Aphelocerus thomasi, holotype. 1) Habitus. 2) Head and prothorax." figureDoi="http://doi.org/10.5281/zenodo.5353862" httpUri="https://zenodo.org/record/5353862/files/figure.png" pageId="2" pageNumber="1">Fig. 12</figureCitation>
)
</paragraph>
<paragraph id="8BE22F71FFCFFFC67BDCF9C0213AF9ED" blockId="2.[192,1439,1555,1615]" pageId="2" pageNumber="1">
<emphasis id="B929F363FFCFFFC67BDCF9C021ACF98C" bold="true" box="[192,438,1555,1582]" pageId="2" pageNumber="1">Specimen examined.</emphasis>
<materialsCitation id="3B35252CFFCFFFC67ADDF9C7213AF9ED" ID-GBIF-Occurrence="3340069301" collectingDate="1962-06-21" collectionCode="CSCA" collectorName="J. M. Campbell" country="Mexico" location="Huichihuayan" pageId="2" pageNumber="1" specimenCount="1" stateProvince="San Luis Potosi" typeStatus="holotype">
<emphasis id="B929F363FFCFFFC67ADDF9C7226BF98C" bold="true" box="[449,625,1556,1582]" pageId="2" pageNumber="1">
<typeStatus id="54E691D3FFCFFFC67ADDF9C72234F98C" box="[449,558,1556,1582]" pageId="2" pageNumber="1" type="holotype">Holotype</typeStatus>
male
</emphasis>
:
<collectingCountry id="F34A6FE1FFCFFFC6799FF9C722C2F98C" box="[643,728,1556,1582]" name="Mexico" pageId="2" pageNumber="1">México</collectingCountry>
,
<collectingRegion id="4999E193FFCFFFC679FBF9C72386F98C" box="[743,924,1556,1582]" country="Mexico" name="San Luis Potosi" pageId="2" pageNumber="1">San Luis Potosí</collectingRegion>
,
<location id="8E8279AAFFCFFFC678B0F9C72457F98C" LSID="urn:lsid:plazi:treatment:03F49E67FFCFFFC77BDCFA152507F8AB:8E8279AAFFCFFFC678B0F9C72457F98C" box="[940,1101,1556,1582]" country="Mexico" name="Huichihuayan" pageId="2" pageNumber="1" stateProvince="San Luis Potosi">Huichihuayán</location>
,
<date id="FFE309B1FFCFFFC67F40F9C724C5F98C" box="[1116,1247,1556,1582]" pageId="2" pageNumber="1" value="1962-06-21">
<collectingDate id="EFA7F059FFCFFFC67F40F9C724C5F98C" box="[1116,1247,1556,1582]" pageId="2" pageNumber="1" value="1962-06-21">VI-21-1962</collectingDate>
</date>
,
<collectorName id="26A84AA7FFCFFFC67FF3F9C72585F98C" box="[1263,1439,1556,1582]" pageId="2" pageNumber="1">J. M. Campbell</collectorName>
(
<collectionCode id="ED4CB7B4FFCFFFC67BD4F9E6210FF9ED" box="[200,277,1589,1615]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="2" pageNumber="1" type="Museum">CSCA</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BE22F71FFCFFFC77BDCF98C2422FA96" blockId="2.[191,1441,1631,1927]" lastBlockId="3.[191,1442,1037,1333]" lastPageId="3" lastPageNumber="2" pageId="2" pageNumber="1">
<emphasis id="B929F363FFCFFFC67BDCF98C21AEF9DB" bold="true" box="[192,436,1631,1657]" pageId="2" pageNumber="1">Description. Length:</emphasis>
6.0 mm. Color: orange testaceous; head, prothorax, abdomen, black; posterior 1/3 of elytra piceous; mouthparts testaceous, antennae reddish brown; piceous elytral markings arcuately emarginate on anterior border, slightly produced anteriorly at suture (
<figureCitation id="136633F4FFCFFFC67824F9712360F91E" box="[824,890,1698,1724]" captionStart="Figures 12" captionStartId="3.[285,372,958,984]" captionTargetBox="[285,1346,213,926]" captionTargetId="figure-351@3.[285,1347,213,927]" captionTargetPageId="3" captionText="Figures 12. Aphelocerus thomasi, holotype. 1) Habitus. 2) Head and prothorax." figureDoi="http://doi.org/10.5281/zenodo.5353862" httpUri="https://zenodo.org/record/5353862/files/figure.png" pageId="2" pageNumber="1">Fig. 1</figureCitation>
).
<emphasis id="B929F363FFCFFFC6788EF97123C2F91E" bold="true" box="[914,984,1698,1724]" pageId="2" pageNumber="1">Head:</emphasis>
shining, shallowly, finely punctate; surface inconspicuously set with fine, short, silvery, adpressed setae, and a few longer, more robust, black setae; antennae moderate in length; club composed of distal 4 antennomeres, gradually enlarged.
<emphasis id="B929F363FFCFFFC67FD5F9352553F8A2" bold="true" box="[1225,1353,1766,1792]" pageId="2" pageNumber="1">Pronotum:</emphasis>
slightly broader than long, subflattened on disk above; anterior depression distinct; surface shining, moderately densely but inconspicuously vested with fine, medium length, reclinate silvery setae, intermixed with longer, more robust suberect black setae (
<figureCitation id="136633F4FFCFFFC67AB0F89921F4F8C7" box="[428,494,1866,1893]" captionStart="Figures 12" captionStartId="3.[285,372,958,984]" captionTargetBox="[285,1346,213,926]" captionTargetId="figure-351@3.[285,1347,213,927]" captionTargetPageId="3" captionText="Figures 12. Aphelocerus thomasi, holotype. 1) Habitus. 2) Head and prothorax." figureDoi="http://doi.org/10.5281/zenodo.5353862" httpUri="https://zenodo.org/record/5353862/files/figure.png" pageId="2" pageNumber="1">Fig. 2</figureCitation>
). Scutellum small, densely covered with white setae.
<emphasis id="B929F363FFCFFFC67F21F8982497F8C7" bold="true" box="[1085,1165,1867,1893]" pageId="2" pageNumber="1">Elytra:</emphasis>
elongate (ratio of length to width 13:7); umbones distinct; sides subparallel, arcuately, but rather gradually convergent posteriorly to separately rounded, dehiscent apices; dorsum slightly subflattened; integument shining, finely, sparsely, shallowly punctate, anterior 2/3 sparsely and inconspicuously set with short, erect pale setae, and fewer, somewhat longer and more robust erect, tawny setae; posterior margin of testaceous area with an irregular transverse band formed by a concentration of moderately long, reclinate, posteriorly oriented white setae, most conspicuous internally at suture; elytral posterior 1/3 rather densely covered with suberect and longer, erect black setae.
<emphasis id="B929F363FFCEFFC77FE1FB4725B8FB0C" bold="true" box="[1277,1442,1172,1198]" pageId="3" pageNumber="2">Metasternum:</emphasis>
shining, moderately densely clothed with medium length pale setae.
<emphasis id="B929F363FFCEFFC778A0FB66242DFB6D" bold="true" box="[956,1079,1205,1231]" pageId="3" pageNumber="2">Abdomen:</emphasis>
ventrites shining, coarsely, shallowly, punctate and moderately densely clothed with fine, pale setae, ventrite 5 with hind margin broadly, feebly emarginate; ventrite 6 smaller, hind margin broadly, feebly emarginate; tergite 6 obliquely rounded laterally, subtruncate apically, somewhat excavate ventrally, surpassing ventrite 6 posteriorly.
</paragraph>
<caption id="DF227FF9FFCEFFC77A01FC6D2480FC7A" ID-DOI="http://doi.org/10.5281/zenodo.5353862" ID-Zenodo-Dep="5353862" box="[285,1178,958,985]" httpUri="https://zenodo.org/record/5353862/files/figure.png" pageId="3" pageNumber="2" startId="3.[285,372,958,984]" targetBox="[285,1346,213,926]" targetPageId="3">
<paragraph id="8BE22F71FFCEFFC77A01FC6D2480FC7A" blockId="3.[285,1178,958,985]" box="[285,1178,958,985]" pageId="3" pageNumber="2">
<emphasis id="B929F363FFCEFFC77A01FC6D21B6FC7A" bold="true" box="[285,428,958,985]" pageId="3" pageNumber="2">Figures 12.</emphasis>
<taxonomicName id="4C5D54F2FFCEFFC77AAEFC6D228FFC7A" authorityName="Rifkind" authorityYear="2020" box="[434,661,958,984]" class="Insecta" family="Cleridae" genus="Aphelocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="thomasi">
<emphasis id="B929F363FFCEFFC77AAEFC6D228FFC7A" box="[434,661,958,984]" italics="true" pageId="3" pageNumber="2">Aphelocerus thomasi</emphasis>
</taxonomicName>
, holotype.
<emphasis id="B929F363FFCEFFC7780CFC6D2333FC7A" bold="true" box="[784,809,958,984]" pageId="3" pageNumber="2">1)</emphasis>
Habitus.
<emphasis id="B929F363FFCEFFC7788FFC6D23B1FC7A" bold="true" box="[915,939,958,984]" pageId="3" pageNumber="2">2)</emphasis>
Head and prothorax.
</paragraph>
</caption>
<paragraph id="8BE22F71FFCEFFC77BDCFA902260FAFC" blockId="3.[192,634,1347,1374]" box="[192,634,1347,1374]" pageId="3" pageNumber="2">
<emphasis id="B929F363FFCEFFC77BDCFA90212CFAFF" bold="true" box="[192,310,1347,1373]" pageId="3" pageNumber="2">Variation.</emphasis>
Only the
<typeStatus id="54E691D3FFCEFFC77AB8FA97221CFAFC" box="[420,518,1348,1374]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
is known.
</paragraph>
</subSubSection>
<subSubSection id="C3477CFAFFCEFFC77BDCFABE21FEFA0B" pageId="3" pageNumber="2" type="etymology">
<paragraph id="8BE22F71FFCEFFC77BDCFABE21FEFA0B" blockId="3.[192,1440,1389,1449]" pageId="3" pageNumber="2">
<emphasis id="B929F363FFCEFFC77BDCFABE215EFA25" bold="true" box="[192,324,1389,1415]" pageId="3" pageNumber="2">Etymology.</emphasis>
I dedicate this species to the late Michael C. Thomas, in recognition of his many contributions to the systematics of
<taxonomicName id="4C5D54F2FFCEFFC77A7DFA5C21FAFA0B" authorityName="Linnaeus" authorityYear="1758" box="[353,480,1423,1449]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3477CFAFFCEFFC77BDCFA6B2250FA57" pageId="3" pageNumber="2" type="distribution">
<paragraph id="8BE22F71FFCEFFC77BDCFA6B2250FA57" blockId="3.[192,1441,1464,1525]" pageId="3" pageNumber="2">
<emphasis id="B929F363FFCEFFC77BDCFA6B2140FA70" bold="true" box="[192,346,1464,1490]" pageId="3" pageNumber="2">Distribution.</emphasis>
The new species is known from
<collectingRegion id="4999E193FFCEFFC779D0FA6A2360FA71" box="[716,890,1464,1491]" country="Mexico" name="San Luis Potosi" pageId="3" pageNumber="2">San Luis Potosí</collectingRegion>
, on the Atlantic versant of north central
<collectingCountry id="F34A6FE1FFCEFFC77E5BFA6B2586FA70" box="[1351,1436,1464,1490]" name="Mexico" pageId="3" pageNumber="2">Mexico</collectingCountry>
. The
<typeStatus id="54E691D3FFCEFFC77BEDFA092149FA56" box="[241,339,1498,1524]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
was collected in June.
</paragraph>
</subSubSection>
<subSubSection id="C3477CFAFFCEFFC77BDCF9D02507F8AB" pageId="3" pageNumber="2" type="diagnosis">
<paragraph id="8BE22F71FFCEFFC77BDCF9D02507F8AB" blockId="3.[192,1441,1539,1801]" pageId="3" pageNumber="2">
<emphasis id="B929F363FFCEFFC77BDCF9D02126F9BF" bold="true" box="[192,316,1539,1565]" pageId="3" pageNumber="2">Diagnosis.</emphasis>
<taxonomicName id="4C5D54F2FFCEFFC77A5FF9D7223CF9BC" authorityName="Rifkind" authorityYear="2020" box="[323,550,1540,1566]" class="Insecta" family="Cleridae" genus="Aphelocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="thomasi">
<emphasis id="B929F363FFCEFFC77A5FF9D7223CF9BC" box="[323,550,1540,1566]" italics="true" pageId="3" pageNumber="2">Aphelocerus thomasi</emphasis>
</taxonomicName>
can be distinguished from congeners by a unique combination of color pattern and elytral shape. Most
<taxonomicName id="4C5D54F2FFCEFFC77AD5F9F622B9F99D" box="[457,675,1573,1599]" class="Insecta" family="Cleridae" genus="Aphelocerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B929F363FFCEFFC77AD5F9F62254F99D" box="[457,590,1573,1599]" italics="true" pageId="3" pageNumber="2">Aphelocerus</emphasis>
species
</taxonomicName>
are uniformly black. Of the few bicolorous species,
<taxonomicName id="4C5D54F2FFCEFFC77FF8F9F52541F99D" authorityName="Rifkind" authorityYear="2020" box="[1252,1371,1573,1600]" class="Insecta" family="Cleridae" genus="Aphelocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="thomasi">
<emphasis id="B929F363FFCEFFC77FF8F9F524ECF9E2" box="[1252,1270,1574,1600]" italics="true" pageId="3" pageNumber="2">A</emphasis>
.
<emphasis id="B929F363FFCEFFC77E1FF9F62541F99D" box="[1283,1371,1573,1599]" italics="true" pageId="3" pageNumber="2">thomasi</emphasis>
</taxonomicName>
could only be confused with the similarly sized
<taxonomicName id="4C5D54F2FFCEFFC77997F9942376F9C3" authority="(Barr)" baseAuthorityName="Barr" box="[651,876,1607,1633]" class="Insecta" family="Cleridae" genus="Aphelocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="delicatulus">
<emphasis id="B929F363FFCEFFC77997F9942284F9C3" box="[651,670,1607,1633]" italics="true" pageId="3" pageNumber="2">A</emphasis>
.
<emphasis id="B929F363FFCEFFC779B7F994233AF9C3" box="[683,800,1607,1633]" italics="true" pageId="3" pageNumber="2">delicatulus</emphasis>
(Barr)
</taxonomicName>
, but the latter has the head and prothorax partially testaceous, the anterior 2/3 of the elytra with a broad longitudinal dark band at the suture, and the elytral apices distinctly more rounded than the former (
<figureCitation id="136633F4FFCEFFC77986F95922C1F906" box="[666,731,1674,1700]" captionStart="Figure 3" captionStartId="4.[574,650,921,947]" captionTargetBox="[575,1058,213,891]" captionTargetId="figure-92@4.[574,1058,211,891]" captionTargetPageId="4" captionText="Figure 3. Aphelocerus delicatulus, habitus." figureDoi="http://doi.org/10.5281/zenodo.5353864" httpUri="https://zenodo.org/record/5353864/files/figure.png" pageId="3" pageNumber="2">Fig. 3</figureCitation>
). Furthermore,
<taxonomicName id="4C5D54F2FFCEFFC77892F959241FF906" authorityName="Rifkind" authorityYear="2020" box="[910,1029,1674,1701]" class="Insecta" family="Cleridae" genus="Aphelocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="thomasi">
<emphasis id="B929F363FFCEFFC77892F95923BAF906" box="[910,928,1674,1700]" italics="true" pageId="3" pageNumber="2">A</emphasis>
.
<emphasis id="B929F363FFCEFFC778B1F959241FF906" box="[941,1029,1674,1700]" italics="true" pageId="3" pageNumber="2">thomasi</emphasis>
</taxonomicName>
is known from a specimen collected in June on Mexicos Atlantic side, whereas
<taxonomicName id="4C5D54F2FFCEFFC779B2F97F235FF964" box="[686,837,1708,1734]" class="Insecta" family="Cleridae" genus="Aphelocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="delicatulus">
<emphasis id="B929F363FFCEFFC779B2F97F22DBF964" box="[686,705,1708,1734]" italics="true" pageId="3" pageNumber="2">A</emphasis>
.
<emphasis id="B929F363FFCEFFC779CCF97F235FF964" box="[720,837,1708,1734]" italics="true" pageId="3" pageNumber="2">delicatulus</emphasis>
</taxonomicName>
has only been recorded from the Pacific versant (in the states of
<collectingRegion id="4999E193FFCEFFC77A52F91E21B8F945" box="[334,418,1741,1767]" country="Mexico" name="Sinaloa" pageId="3" pageNumber="2">Sinaloa</collectingRegion>
,
<collectingRegion id="4999E193FFCEFFC77AB3F91E21E3F945" box="[431,505,1741,1767]" country="Mexico" name="Jalisco" pageId="3" pageNumber="2">Jalisco</collectingRegion>
,
<collectingRegion id="4999E193FFCEFFC77918F91E2243F945" box="[516,601,1741,1767]" country="Mexico" name="Colima" pageId="3" pageNumber="2">Colima</collectingRegion>
,
<collectingRegion id="4999E193FFCEFFC77979F91E22D9F945" box="[613,707,1741,1767]" country="Mexico" name="Morelos" pageId="3" pageNumber="2">Morelos</collectingRegion>
,
<collectingRegion id="4999E193FFCEFFC779D3F91E2301F945" box="[719,795,1741,1767]" country="Mexico" name="Puebla" pageId="3" pageNumber="2">Puebla</collectingRegion>
and
<collectingRegion id="4999E193FFCEFFC77849F91E23B6F945" box="[853,940,1741,1767]" country="Mexico" name="Oaxaca" pageId="3" pageNumber="2">Oaxaca</collectingRegion>
), with all collections made from late August through October. The two species therefore appear to be both allopatric and phenologically isolated.
</paragraph>
</subSubSection>
</treatment>
</document>