518 lines
66 KiB
XML
518 lines
66 KiB
XML
<document id="B9AF789E77BF4B263FEDB733ECF9F360" ID-DOI="10.11606/1807-0205/2020.60.special-issue.19" ID-ISSN="1807-0205" ID-Zenodo-Dep="4985175" ID-ZooBank="B723DFDD-1F03-4110-903A-A2B8A4CA7083" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624028473964" checkinUser="carolina" docAuthor="Bevilaqua, Marcus & Fonseca, Claudio Ruy Vasconcelos da" docDate="2020" docId="039E3733FFA67D08ECC09C60FB75FD4A" docLanguage="en" docName="PapAvZool.60.special-issue.e202060.19.pdf" docOrigin="Papéis Avulsos de Zoologia (Pap. Avulsos Zool., S. Paulo) 60 (19)" docSource="http://dx.doi.org/10.11606/1807-0205/2020.60.special-issue.19" docStyle="DocumentStyle:6844D5473582683DD13A9A75B001759F.4:PapAvZool.2018-.journal_article" docStyleId="6844D5473582683DD13A9A75B001759F" docStyleName="PapAvZool.2018-.journal_article" docStyleVersion="4" docTitle="Passalus (Passalus) cleidecostae Bevilaqua & Fonseca 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="8" masterDocId="FFA74F4BFFA57D0FEC6F9F5AFFF5FFD9" masterDocTitle="Two new species of Passalus Fabricius (Coleoptera: Passalidae) from the western Brazilian Amazon with comments on the taxonomic limits of the subgenera" masterLastPageNumber="13" masterPageNumber="1" pageNumber="4" updateTime="1698969348260" updateUser="plazi" zenodo-license-document="CC-BY-NC-4.0" zenodo-license-figures="CC-BY-NC-4.0">
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<mods:title id="336ED4F2CC73950B9039872FE83FB26E">Two new species of Passalus Fabricius (Coleoptera: Passalidae) from the western Brazilian Amazon with comments on the taxonomic limits of the subgenera</mods:title>
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<mods:name id="C2FA2B4C1AD3214A8A291A8E923B2C4C" type="personal">
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<mods:namePart id="920FF3E3B9B4F031D00B3E2A152EB5BE">Bevilaqua, Marcus</mods:namePart>
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<mods:roleTerm id="F5EF134182F3C5AB78028BF773D09524">Author</mods:roleTerm>
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<mods:namePart id="F7C38693A513356B0136C87267903730">Fonseca, Claudio Ruy Vasconcelos da</mods:namePart>
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<mods:title id="1C9FF4E8DF31A16C07F56327B5BE5860">Papéis Avulsos de Zoologia</mods:title>
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<mods:part id="7DD064E4DD966B7B3758E811DEB92F14">
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<mods:date id="4F66DF1A92AD5ABD4BCE63C9535644A9">2020</mods:date>
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<mods:title id="6BCCA9EE7DCCA18B6373026944A3B68F">Pap. Avulsos Zool., S. Paulo</mods:title>
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<mods:number id="AE322DE00BBE002116E7B84A5E2569DB">2020-03-04</mods:number>
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<mods:number id="15138796F15EE24F8DF5FF37DCF09055">60</mods:number>
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<mods:url id="9248BF7D0674A35B2CCA4159D4BFEF9A">http://dx.doi.org/10.11606/1807-0205/2020.60.special-issue.19</mods:url>
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<mods:classification id="D0690C9EB390C24B6C983023E1C280E2">journal article</mods:classification>
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<subSubSection id="C32DD5AEFFA67D0CECC09C60FD88FC8A" box="[175,637,826,851]" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="8B888625FFA67D0CECC09C60FD88FC8A" blockId="3.[94,719,826,1523]" box="[175,637,826,851]" pageId="3" pageNumber="4">
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<heading id="D0C03149FFA67D0CECC09C60FD88FC8A" box="[175,637,826,851]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="4" reason="2">
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<emphasis id="B9435A37FFA67D0CECC09C60FD88FC8A" bold="true" box="[175,637,826,851]" pageId="3" pageNumber="4">
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<taxonomicName id="4C37FDA6FFA67D0CECC09C60FDEBFC8A" authority="Bevilaqua & Fonseca, 2020" authorityName="Bevilaqua & Fonseca" authorityYear="2020" box="[175,542,826,851]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov." subGenus="Passalus">
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<emphasis id="B9435A37FFA67D0CECC09C60FDEBFC8A" bold="true" box="[175,542,826,851]" italics="true" pageId="3" pageNumber="4">Passalus (Passalus) cleidecostae</emphasis>
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<taxonomicNameLabel id="A270E74CFFA67D0CEE4B9C60FD88FC8A" box="[548,637,826,851]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C32DD5AEFFA67D0CED579C00FE01FCAA" box="[312,500,858,883]" pageId="3" pageNumber="4" type="description">
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<paragraph id="8B888625FFA67D0CED579C00FE01FCAA" blockId="3.[94,719,826,1523]" box="[312,500,858,883]" pageId="3" pageNumber="4">
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<emphasis id="B9435A37FFA67D0CED579C00FE01FCAA" bold="true" box="[312,500,858,883]" pageId="3" pageNumber="4">
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(
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<figureCitation id="130C9AA0FFA67D0CED509C00FE7BFCAA" box="[319,398,858,883]" captionStart="Figure 4" captionStartId="8.[170,229,2106,2131]" captionTargetBox="[170,1493,338,2077]" captionTargetId="figure-55@8.[170,1493,338,2077]" captionTargetPageId="8" captionText="Figure 4. Passalus (Passalus) cleidecostaesp.nov., habitus.(A) dorsal;(B) ventral;(C) lateral." figureDoi="http://doi.org/10.5281/zenodo.4985191" httpUri="https://zenodo.org/record/4985191/files/figure.png" pageId="3" pageNumber="4">Figs. 4</figureCitation>
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,
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<figureCitation id="130C9AA0FFA67D0CEDF79C00FE5CFCAA" box="[408,425,858,883]" captionStart="Figure 5" captionStartId="9.[94,153,2074,2099]" captionTargetBox="[94,1417,494,2044]" captionTargetId="figure-135@9.[94,1417,494,2044]" captionTargetPageId="9" captionText="Figure 5. Passalus (Passalus) cleidecostae sp.nov.(A) head and prothorax,dorsal;(B) head,ventral (C) head and prothorax,dorsolateral;(D) posterior region of the prosternum and anterior region of the mesosternum;(E) metasternum (left side);(F) aedeagus (ventral,dorsal and lateral view)." figureDoi="http://doi.org/10.5281/zenodo.4985199" httpUri="https://zenodo.org/record/4985199/files/figure.png" pageId="3" pageNumber="4">5</figureCitation>
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,
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<figureCitation id="130C9AA0FFA67D0CEDDC9C00FE35FCAA" box="[435,448,858,883]" captionStart="Figure 6" captionStartId="10.[171,230,2074,2099]" captionTargetBox="[195,1460,666,2031]" captionTargetId="figure-193@10.[170,1493,646,2044]" captionTargetPageId="10" captionText="Figure 6. Passalus (Passalus) cleidecostae sp. nov. (A) cephalic capsule, dorsal; (B) pronotum, lateral; (C) meso- and metasternum. Passalus (Passalus) bucki Luederwaldt,1931:(D) cephalic capsule,dorsal v.(E) pronotum,lateral.(F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985201" httpUri="https://zenodo.org/record/4985201/files/figure.png" pageId="3" pageNumber="4">6</figureCitation>
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A-C)
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</emphasis>
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<emphasis id="B9435A37FFA67D0CEC309CC0FEE7FC6A" bold="true" box="[95,274,922,947]" pageId="3" pageNumber="4">Type material:</emphasis>
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:
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23n – praia redonda /
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col”
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:“Sena Madureira / Tr23n – praia redonda /
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/ FBP Gouveia,
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<collectionCode id="ED261EE0FFA67D0CECA49B60FF19FB8A" box="[203,236,1082,1107]" country="0" name="Goucher College" pageId="3" pageNumber="4">GC</collectionCode>
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de Sá col. (
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). Sena Madureira / Tr22n – praia redonda /
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/
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<date id="FF89A0E5FFA67D0CEC309B20FF07FB4A" box="[95,242,1146,1171]" pageId="3" pageNumber="4" value="2010-11-15" valueMax="2010-11-19" valueMin="2010-11-15">15-19.xi.2010</date>
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/ FBP Gouveia,
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<collectionCode id="ED261EE0FFA67D0CEDF69B20FE4FFB4A" box="[409,442,1146,1171]" country="0" name="Goucher College" pageId="3" pageNumber="4">GC</collectionCode>
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de Sá col. (
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<specimenCount id="9D314DACFFA67D0CEE5B9B20FDA2FB4B" box="[564,599,1143,1171]" pageId="3" pageNumber="4" type="male">1 ♂</specimenCount>
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,
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<specimenCount id="9D314DACFFA67D0CEE0E9B20FD76FB4B" box="[609,643,1143,1171]" pageId="3" pageNumber="4" type="female">2 ♀</specimenCount>
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,
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<collectionCode id="ED261EE0FFA67D0CEEE29B20FD33FB4A" box="[653,710,1146,1171]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="3" pageNumber="4">INPA</collectionCode>
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). Sena Madureira / Tr21n – praia redonda /
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,
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/
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/ FBP Gouveia,
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<collectionCode id="ED261EE0FFA67D0CEE099BE0FD72FB0A" box="[614,647,1210,1235]" country="0" name="Goucher College" pageId="3" pageNumber="4">GC</collectionCode>
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de Sá col. (
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,
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<specimenCount id="9D314DACFFA67D0CECAF9B80FF17FB2B" box="[192,226,1239,1267]" pageId="3" pageNumber="4" type="female">1 ♀</specimenCount>
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,
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).Sena Madureira / Tr27n – praia redonda /
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||
,
|
||
<geoCoordinate id="EE03E0E2FFA67D0CED549BA0FE2BFACA" box="[315,478,1274,1300]" degrees="68" direction="west" minutes="45" orientation="longitude" pageId="3" pageNumber="4" precision="1" seconds="44.30" value="-68.76231">68°45′44.30″W</geoCoordinate>
|
||
/
|
||
<date id="FF89A0E5FFA67D0CED989BA0FD7EFACA" box="[503,651,1274,1299]" pageId="3" pageNumber="4" value="2010-11-15" valueMax="2010-11-19" valueMin="2010-11-15">15-19.xi.2010</date>
|
||
/ FBP Gouveia,
|
||
<collectionCode id="ED261EE0FFA67D0CECA79A40FF1CFAEA" box="[200,233,1306,1331]" country="0" name="Goucher College" pageId="3" pageNumber="4">GC</collectionCode>
|
||
de Sá col. (
|
||
<specimenCount id="9D314DACFFA67D0CED009A40FE63FAEB" box="[367,406,1303,1331]" pageId="3" pageNumber="4" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D314DACFFA67D0CEDCC9A40FE3DFAEB" box="[419,456,1303,1331]" pageId="3" pageNumber="4" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED261EE0FFA67D0CEDBA9A40FDFBFAEA" box="[469,526,1306,1331]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="3" pageNumber="4">INPA</collectionCode>
|
||
). Sena Madureira / Tr30n – praia redonda /
|
||
<geoCoordinate id="EE03E0E2FFA67D0CEDEB9A60FDD5FA8A" box="[388,544,1338,1364]" degrees="09" direction="south" minutes="09" orientation="latitude" pageId="3" pageNumber="4" precision="1" seconds="34.70" value="-9.159638">09°09′34.70″S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE03E0E2FFA67D0CEE449A60FD3BFA8A" box="[555,718,1338,1364]" degrees="68" direction="west" minutes="45" orientation="longitude" pageId="3" pageNumber="4" precision="1" seconds="41.34" value="-68.76148">68°45′41.34″W</geoCoordinate>
|
||
/
|
||
<date id="FF89A0E5FFA67D0CEC1F9A00FEF6FAAA" box="[112,259,1370,1395]" pageId="3" pageNumber="4" value="2010-11-15" valueMax="2010-11-19" valueMin="2010-11-15">15-19.xi.2010</date>
|
||
/ FBP Gouveia,
|
||
<collectionCode id="ED261EE0FFA67D0CEDD89A00FE2DFAAA" box="[439,472,1370,1395]" country="0" name="Goucher College" pageId="3" pageNumber="4">GC</collectionCode>
|
||
de Sá col. (
|
||
<specimenCount id="9D314DACFFA67D0CEE359A00FD75FAAB" box="[602,640,1367,1395]" pageId="3" pageNumber="4" type="female">2 ♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED261EE0FFA67D0CEEE29A00FD33FAAA" box="[653,710,1370,1395]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="3" pageNumber="4">INPA</collectionCode>
|
||
). Sena Madureira / Tr29n – praia redonda /
|
||
<geoCoordinate id="EE03E0E2FFA67D0CEE409A20FD3EFA4A" box="[559,715,1402,1428]" degrees="09" direction="south" minutes="09" orientation="latitude" pageId="3" pageNumber="4" precision="1" seconds="34.70" value="-9.159638">09°09′34.70″S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE03E0E2FFA67D0CEC319AC0FEF4FA6A" box="[94,257,1434,1460]" degrees="68" direction="west" minutes="45" orientation="longitude" pageId="3" pageNumber="4" precision="1" seconds="41.34" value="-68.76148">68°45′41.34″W</geoCoordinate>
|
||
/
|
||
<date id="FF89A0E5FFA67D0CED739AC0FE5AFA6A" box="[284,431,1434,1459]" pageId="3" pageNumber="4" value="2010-11-15" valueMax="2010-11-19" valueMin="2010-11-15">15-19.xi.2010</date>
|
||
/ FBP Gouveia,
|
||
<collectionCode id="ED261EE0FFA67D0CEE099AC0FD72FA6A" box="[614,647,1434,1459]" country="0" name="Goucher College" pageId="3" pageNumber="4">GC</collectionCode>
|
||
de Sá col. (
|
||
<specimenCount id="9D314DACFFA67D0CECFC9AE0FF4CFA0B" box="[147,185,1463,1491]" pageId="3" pageNumber="4" type="female">2 ♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED261EE0FFA67D0CECA99AE0FF0AFA0A" box="[198,255,1466,1491]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="3" pageNumber="4">INPA</collectionCode>
|
||
). Sena Madureira / roÇa D. [ona] Marina / (
|
||
<specimenCount id="9D314DACFFA67D0CEC0A9A80FF79FA2B" box="[101,140,1495,1523]" pageId="3" pageNumber="4" type="male">2 ♂</specimenCount>
|
||
,
|
||
<collectionCode id="ED261EE0FFA67D0CECF89A80FF3AFA2A" box="[151,207,1498,1523]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="3" pageNumber="4">INPA</collectionCode>
|
||
).”
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32DD5AEFFA67D0CEC319940FABDFB6A" pageId="3" pageNumber="4" type="diagnosis">
|
||
<paragraph id="8B888625FFA67D0CEC319940FABDFB6A" blockId="3.[94,719,1562,2131]" lastBlockId="3.[794,1418,826,1203]" pageId="3" pageNumber="4">
|
||
<emphasis id="B9435A37FFA67D0CEC319940FF2EF9EA" bold="true" box="[94,219,1562,1587]" pageId="3" pageNumber="4">Diagnosis:</emphasis>
|
||
medium sized specimens (
|
||
<quantity id="4CCF2BC0FFA67D0CEE4D9940FD6BF9EA" box="[546,670,1562,1587]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" pageId="3" pageNumber="4" unit="mm" value="29.5" valueMax="31.0" valueMin="28.0">28-31 mm</quantity>
|
||
, total length); body flat; anterior frontal edge of the front straight; anterior mediofrontal area with coarse punctures,posterior mediofrontal area with large rounded cephalic nodule; laterofrontal tubercles large, conspicuous, with acute apices; mediofrontal tubercles with round- ed apices, smaller than laterofrontal tubercles, located externally to these; secondary mediofrontal tubercles smaller than laterofrontal, with acute apices, distance between them less than distance to laterofrontal tubercles; inner tubercles small, but slightly detached from the anterofrontal ridges; posterofrontal ridges elevated, strong and slightly straight, starting at apex of central tubercle; anterofrontal ridges arched, larger, lower and weaker than posterofrontal ridges; central tubercle flat, with not free apex; conspicuous and slightly rounded lateroposterior tubercles; lateropostfrontal areas with a few small punctures; mentum with non-dilated mediobasal area, large and deep lateral scars; pronotum with anterior angles not protruded and slightly acute; pronotal punctures only on marginal grooves and near lateral fossae; mesosternal scars deep and elongated; metasternal disc barely delimited, without carina, with large group of punctures on posterior region; metasternal grooves deep and pubescent; in ventral view, aedeagus ventrally with basal piece fused to parameres, and with V-shaped emargination in middle region; median lobe globular occupying almost half of all aedeagus, with two longitudinal sclerotized plates located near outer edges; in dorsal view, apices of parameres very close to each other.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32DD5AEFFA67D08EF759B80FED1FD2A" lastPageId="7" lastPageNumber="8" pageId="3" pageNumber="4" type="description">
|
||
<paragraph id="8B888625FFA67D0CEF759B80FC51FB2A" blockId="3.[794,932,1242,1267]" box="[794,932,1242,1267]" pageId="3" pageNumber="4">
|
||
<emphasis id="B9435A37FFA67D0CEF759B80FC51FB2A" bold="true" box="[794,932,1242,1267]" pageId="3" pageNumber="4">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA67D0CEF759A40FC84FA8A" blockId="3.[794,1417,1306,1363]" pageId="3" pageNumber="4">
|
||
<emphasis id="B9435A37FFA67D0CEF759A40FC17FAEA" bold="true" box="[794,994,1306,1331]" pageId="3" pageNumber="4">
|
||
Body (
|
||
<figureCitation id="130C9AA0FFA67D0CEF059A40FC21FAEA" box="[874,980,1306,1331]" captionStart-0="Figure 4" captionStart-1="Figure 5" captionStart-2="Figure 6" captionStartId-0="8.[170,229,2106,2131]" captionStartId-1="9.[94,153,2074,2099]" captionStartId-2="10.[171,230,2074,2099]" captionTargetBox-0="[170,1493,338,2077]" captionTargetBox-1="[94,1417,494,2044]" captionTargetBox-2="[195,1460,666,2031]" captionTargetId-0="figure-55@8.[170,1493,338,2077]" captionTargetId-1="figure-135@9.[94,1417,494,2044]" captionTargetId-2="figure-193@10.[170,1493,646,2044]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="Figure 4. Passalus (Passalus) cleidecostaesp.nov., habitus.(A) dorsal;(B) ventral;(C) lateral." captionText-1="Figure 5. Passalus (Passalus) cleidecostae sp.nov.(A) head and prothorax,dorsal;(B) head,ventral (C) head and prothorax,dorsolateral;(D) posterior region of the prosternum and anterior region of the mesosternum;(E) metasternum (left side);(F) aedeagus (ventral,dorsal and lateral view)." captionText-2="Figure 6. Passalus (Passalus) cleidecostae sp. nov. (A) cephalic capsule, dorsal; (B) pronotum, lateral; (C) meso- and metasternum. Passalus (Passalus) bucki Luederwaldt,1931:(D) cephalic capsule,dorsal v.(E) pronotum,lateral.(F)meso- and metasternum." figureDoi-0="http://doi.org/10.5281/zenodo.4985191" figureDoi-1="http://doi.org/10.5281/zenodo.4985199" figureDoi-2="http://doi.org/10.5281/zenodo.4985201" httpUri-0="https://zenodo.org/record/4985191/files/figure.png" httpUri-1="https://zenodo.org/record/4985199/files/figure.png" httpUri-2="https://zenodo.org/record/4985201/files/figure.png" pageId="3" pageNumber="4">Figs. 4-6</figureCitation>
|
||
):
|
||
</emphasis>
|
||
flat;
|
||
<emphasis id="B9435A37FFA67D0CE84C9A40FBA5FAEA" box="[1059,1104,1306,1331]" italics="true" pageId="3" pageNumber="4">size:</emphasis>
|
||
medium (
|
||
<quantity id="4CCF2BC0FFA67D0CE8A39A40FAB2FAEA" box="[1228,1351,1306,1331]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" pageId="3" pageNumber="4" unit="mm" value="29.5" valueMax="31.0" valueMin="28.0">28-31 mm</quantity>
|
||
, total length).
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA67D0AEF759A20FDA2FEAA" blockId="3.[794,1419,1402,2131]" lastBlockId="5.[94,719,218,371]" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
|
||
<emphasis id="B9435A37FFA67D0CEF759A20FB96FA4A" bold="true" box="[794,1123,1402,1427]" pageId="3" pageNumber="4">
|
||
Head (
|
||
<figureCitation id="130C9AA0FFA67D0CEF099A20FC46FA4A" box="[870,947,1402,1427]" captionStart="Figure 4" captionStartId="8.[170,229,2106,2131]" captionTargetBox="[170,1493,338,2077]" captionTargetId="figure-55@8.[170,1493,338,2077]" captionTargetPageId="8" captionText="Figure 4. Passalus (Passalus) cleidecostaesp.nov., habitus.(A) dorsal;(B) ventral;(C) lateral." figureDoi="http://doi.org/10.5281/zenodo.4985191" httpUri="https://zenodo.org/record/4985191/files/figure.png" pageId="3" pageNumber="4">Figs. 4</figureCitation>
|
||
A-C, 5A-C, 6A):
|
||
</emphasis>
|
||
<emphasis id="B9435A37FFA67D0CE8049A20FB4BFA4A" box="[1131,1214,1402,1427]" italics="true" pageId="3" pageNumber="4">Labrum</emphasis>
|
||
concave on anterior border.
|
||
<emphasis id="B9435A37FFA67D0CEFCB9AC0FC09FA6A" box="[932,1020,1434,1459]" italics="true" pageId="3" pageNumber="4">Clypeus:</emphasis>
|
||
hidden under frons, with anterior angles small and obtuse and positioned below laterofrontal tubercles.
|
||
<emphasis id="B9435A37FFA67D0CEFBD9A80FB46FA2A" box="[978,1203,1498,1523]" italics="true" pageId="3" pageNumber="4">Anterior frontal edge:</emphasis>
|
||
straight.
|
||
<emphasis id="B9435A37FFA67D0CE9769A80FBE6F9CA" italics="true" pageId="3" pageNumber="4">Secondary mediofrontal tubercles:</emphasis>
|
||
small, smaller than laterofrontal tubercles, with acute apices, near to each other than distance to laterofrontal tubercles.
|
||
<emphasis id="B9435A37FFA67D0CE8E09960FA7CF98A" box="[1167,1417,1594,1619]" italics="true" pageId="3" pageNumber="4">Laterofrontal tubercles:</emphasis>
|
||
large, conspicuous, with acute apices projected forward.
|
||
<emphasis id="B9435A37FFA67D0CEF759920FBFBF94A" box="[794,1038,1658,1683]" italics="true" pageId="3" pageNumber="4">Mediofrontal tubercles:</emphasis>
|
||
small, with rounded apices, small- er than laterofrontal tubercles, located near and externally to this.
|
||
<emphasis id="B9435A37FFA67D0CEFD599E0FADEF90A" box="[954,1323,1722,1747]" italics="true" pageId="3" pageNumber="4">Secondary laterofrontal tubercles:</emphasis>
|
||
absent.
|
||
<emphasis id="B9435A37FFA67D0CEF759980FC14F92A" box="[794,993,1754,1779]" italics="true" pageId="3" pageNumber="4">Mediofrontal area:</emphasis>
|
||
caliciform 2.1× wider than long, with anterior region deeper than posterior; with coarse punctures scattered across anterior region.
|
||
<emphasis id="B9435A37FFA67D0CE8BA9840FA7FF8EA" box="[1237,1418,1818,1843]" italics="true" pageId="3" pageNumber="4">Cephalic nodule:</emphasis>
|
||
large, well-marked, rounded.
|
||
<emphasis id="B9435A37FFA67D0CE8069860FAFBF88A" box="[1129,1294,1850,1875]" italics="true" pageId="3" pageNumber="4">Inner tubercles:</emphasis>
|
||
small, but detached from the anterofrontal ridges, with rounded apices, a little closer to the central tubercle.
|
||
<emphasis id="B9435A37FFA67D0CE8959820FCAAF86A" italics="true" pageId="3" pageNumber="4">Anterofrontal ridges:</emphasis>
|
||
arched, larger, lower and weaker than the posterofrontal ridges.
|
||
<emphasis id="B9435A37FFA67D0CEF8598E0FB38F80A" box="[1002,1229,1978,2003]" italics="true" pageId="3" pageNumber="4">Posterofrontal ridges:</emphasis>
|
||
elevated, strong and slightly straight, starting in apex of central tubercle.
|
||
<emphasis id="B9435A37FFA67D0CEF7598A0FC3CF7CA" box="[794,969,2042,2067]" italics="true" pageId="3" pageNumber="4">Central tubercle:</emphasis>
|
||
conical, flat, with apex not free, dilated posteriorly.
|
||
<emphasis id="B9435A37FFA67D0CEFCF9740FB44F7EA" box="[928,1201,2074,2099]" italics="true" pageId="3" pageNumber="4">Lateroposterior tubercles:</emphasis>
|
||
conspicuous, with evident apices,slightly rounded,slightly distant from cen- tral tubercle.
|
||
<emphasis id="B9435A37FFA17D0BED559F80FDFFFF2A" box="[314,522,218,243]" italics="true" pageId="4" pageNumber="5">Laterofrontal areas:</emphasis>
|
||
slightly flat and smooth, without punctures or wrinkles.
|
||
<emphasis id="B9435A37FFA17D0BEE789FA0FCECFECA" box="[535,793,250,275]" italics="true" pageId="4" pageNumber="5">Lateropostfrontal areas:</emphasis>
|
||
deep, with smooth surface containing some punctures.
|
||
<emphasis id="B9435A37FFA17D0BEF0A9F80FBEBFF2A" box="[869,1054,218,243]" italics="true" pageId="4" pageNumber="5">Postfrontal area:</emphasis>
|
||
shallow, smooth surface.
|
||
<emphasis id="B9435A37FFA17D0BE90F9F80FC41FECA" italics="true" pageId="4" pageNumber="5">Postfrontal groove:</emphasis>
|
||
well-marked, lacking medial notch.
|
||
<emphasis id="B9435A37FFA17D0BE9299FA0FC55FEEA" italics="true" pageId="4" pageNumber="5">Epicranial sutures:</emphasis>
|
||
well-marked.
|
||
<emphasis id="B9435A37FFA17D0BE8579E40FB26FEEA" box="[1080,1235,282,307]" italics="true" pageId="4" pageNumber="5">Epicranial pits:</emphasis>
|
||
deep.
|
||
<emphasis id="B9435A37FFA17D0AE9759E40FF6FFF2A" italics="true" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">Anterior angles of head:</emphasis>
|
||
well-developed, with acute apices, slightly smaller than laterofrontal tubercles.
|
||
<emphasis id="B9435A37FFA07D0AEDFE9FA0FDC3FECA" box="[401,566,250,275]" italics="true" pageId="5" pageNumber="6">Canthus ocular:</emphasis>
|
||
apices almost straight, not reaching half of eye.
|
||
<emphasis id="B9435A37FFA07D0AEDA59E40FDCDFEEA" box="[458,568,282,307]" italics="true" pageId="5" pageNumber="6">Antennae:</emphasis>
|
||
trilamellated, with large and robust lamellae not folded inward, distal lamella almost twice as wide as the other two.
|
||
</paragraph>
|
||
<caption id="DF48D6ADFFA17D0BECC59760FC6BF78A" ID-DOI="http://doi.org/10.5281/zenodo.4985179" ID-Zenodo-Dep="4985179" box="[170,926,2106,2131]" httpUri="https://zenodo.org/record/4985179/files/figure.png" pageId="4" pageNumber="5" startId="4.[170,229,2106,2131]" targetBox="[170,1493,338,2077]" targetPageId="4">
|
||
<paragraph id="8B888625FFA17D0BECC59760FC6BF78A" blockId="4.[170,926,2106,2131]" box="[170,926,2106,2131]" pageId="4" pageNumber="5">
|
||
Figure 1.
|
||
<taxonomicName id="4C37FDA6FFA17D0BEC909760FDF4F78A" authority="Bevilaqua & Fonseca, 2020" authorityName="Bevilaqua & Fonseca" authorityYear="2020" box="[255,513,2106,2131]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="deuterocerus" status="sp. nov." subGenus="Pertinax">
|
||
<emphasis id="B9435A37FFA17D0BEC909760FDF4F78A" bold="true" box="[255,513,2106,2131]" italics="true" pageId="4" pageNumber="5">Passalus (Pertinax) deuterocerus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A270E74CFFA17D0BEE6B9760FDB0F78A" box="[516,581,2106,2131]" pageId="4" pageNumber="5" rank="species">sp.nov.</taxonomicNameLabel>
|
||
,habitus.(A) dorsal;(B) ventral;(C) lateral.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B888625FFA07D0AEC319EC0FD3AFE0A" blockId="5.[94,719,410,467]" pageId="5" pageNumber="6">
|
||
<emphasis id="B9435A37FFA07D0AEC319EC0FE97FE6A" bold="true" box="[94,354,410,435]" pageId="5" pageNumber="6">
|
||
Mouthparts (
|
||
<figureCitation id="130C9AA0FFA07D0AEC929EC0FEA0FE6A" box="[253,341,410,435]" captionStart="Figure 4" captionStartId="8.[170,229,2106,2131]" captionTargetBox="[170,1493,338,2077]" captionTargetId="figure-55@8.[170,1493,338,2077]" captionTargetPageId="8" captionText="Figure 4. Passalus (Passalus) cleidecostaesp.nov., habitus.(A) dorsal;(B) ventral;(C) lateral." figureDoi="http://doi.org/10.5281/zenodo.4985191" httpUri="https://zenodo.org/record/4985191/files/figure.png" pageId="5" pageNumber="6">Fig. 4B</figureCitation>
|
||
):
|
||
</emphasis>
|
||
<emphasis id="B9435A37FFA07D0AED039EC0FE41FE6A" box="[364,436,410,435]" italics="true" pageId="5" pageNumber="6">
|
||
<taxonomicName id="4C37FDA6FFA07D0AED039EC0FE5BFE6A" box="[364,430,410,435]" class="Cestoda" family="Diphyllobothriidae" genus="Ligula" kingdom="Animalia" order="Diphyllobothriidea" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="genus">Ligula</taxonomicName>
|
||
:
|
||
</emphasis>
|
||
tridentate, with median tooth larger than lateral teeth.
|
||
<emphasis id="B9435A37FFA07D0AEDDE9EE0FD78FE0A" box="[433,653,442,467]" italics="true" pageId="5" pageNumber="6">Hypostomal process:</emphasis>
|
||
wide,
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA07D0AEF759F80FC16FE0A" blockId="5.[794,1418,218,467]" pageId="5" pageNumber="6">
|
||
glabrous, slightly close to mentum. Mentum:
|
||
<emphasis id="B9435A37FFA07D0AE9779F80FC93FECA" italics="true" pageId="5" pageNumber="6">mediobasal area</emphasis>
|
||
not dilated, smooth and glabrous, with slightly protruded anterior region, and median notch;
|
||
<emphasis id="B9435A37FFA07D0AE92A9E40FCA7FE8A" italics="true" pageId="5" pageNumber="6">lateral lobes</emphasis>
|
||
with rounded apices;
|
||
<emphasis id="B9435A37FFA07D0AE8229E60FB38FE8A" box="[1101,1229,314,339]" italics="true" pageId="5" pageNumber="6">lateral scars</emphasis>
|
||
large, deep, glabrous, unpunctate and rounded.
|
||
<emphasis id="B9435A37FFA07D0AE8E39E00FAF4FEAA" box="[1164,1281,346,371]" italics="true" pageId="5" pageNumber="6">Mandibles:</emphasis>
|
||
incisor lobe with three well-formed apical teeth; suprainternal teeth robust; infrabasal pits inconspicuous.
|
||
<emphasis id="B9435A37FFA07D0AE8B09EC0FAC6FE6A" box="[1247,1331,410,435]" italics="true" pageId="5" pageNumber="6">Maxilla:</emphasis>
|
||
lacinia bidentate at apex.
|
||
</paragraph>
|
||
<caption id="DF48D6ADFFA07D0AEC319740FB81F78A" ID-DOI="http://doi.org/10.5281/zenodo.4985183" ID-Zenodo-Dep="4985183" httpUri="https://zenodo.org/record/4985183/files/figure.png" pageId="5" pageNumber="6" startId="5.[94,153,2074,2099]" targetBox="[94,1417,494,2044]" targetPageId="5">
|
||
<paragraph id="8B888625FFA07D0AEC319740FB81F78A" blockId="5.[94,1417,2074,2131]" pageId="5" pageNumber="6">
|
||
Figure 2.
|
||
<taxonomicName id="4C37FDA6FFA07D0AECDC9740FE41F7EA" authority="Bevilaqua & Fonseca, 2020" authorityName="Bevilaqua & Fonseca" authorityYear="2020" box="[179,436,2074,2099]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="deuterocerus" status="sp. nov." subGenus="Pertinax">
|
||
<emphasis id="B9435A37FFA07D0AECDC9740FE41F7EA" bold="true" box="[179,436,2074,2099]" italics="true" pageId="5" pageNumber="6">Passalus (Pertinax) deuterocerus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A270E74CFFA07D0AEDD79740FE03F7EA" box="[440,502,2074,2099]" pageId="5" pageNumber="6" rank="species">sp.nov.</taxonomicNameLabel>
|
||
(A) head and prothorax,dorsal;(B) head,ventral;(C) head and prothorax,dorsolateral;(D)posterior region of the prosternum and anterior region of the mesosternum;(E) metasternum (left side);(F) aedeagus (ventral,dorsal and lateral view).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B888625FFA37D09ECC59F80FDA6FDCA" blockId="6.[170,795,218,531]" pageId="6" pageNumber="7">
|
||
<emphasis id="B9435A37FFA37D09ECC59F80FD71FF2A" bold="true" box="[170,644,218,243]" pageId="6" pageNumber="7">
|
||
Prothorax:Pronotum(
|
||
<figureCitation id="130C9AA0FFA37D09EDC29F80FDFFFF2A" box="[429,522,218,243]" captionStart="Figure 4" captionStartId="8.[170,229,2106,2131]" captionTargetBox="[170,1493,338,2077]" captionTargetId="figure-55@8.[170,1493,338,2077]" captionTargetPageId="8" captionText="Figure 4. Passalus (Passalus) cleidecostaesp.nov., habitus.(A) dorsal;(B) ventral;(C) lateral." figureDoi="http://doi.org/10.5281/zenodo.4985191" httpUri="https://zenodo.org/record/4985191/files/figure.png" pageId="6" pageNumber="7">Figs.4A</figureCitation>
|
||
,
|
||
<figureCitation id="130C9AA0FFA37D09EE7E9F80FDB8FF2A" box="[529,589,218,243]" captionStart="Figure 5" captionStartId="9.[94,153,2074,2099]" captionTargetBox="[94,1417,494,2044]" captionTargetId="figure-135@9.[94,1417,494,2044]" captionTargetPageId="9" captionText="Figure 5. Passalus (Passalus) cleidecostae sp.nov.(A) head and prothorax,dorsal;(B) head,ventral (C) head and prothorax,dorsolateral;(D) posterior region of the prosternum and anterior region of the mesosternum;(E) metasternum (left side);(F) aedeagus (ventral,dorsal and lateral view)." figureDoi="http://doi.org/10.5281/zenodo.4985199" httpUri="https://zenodo.org/record/4985199/files/figure.png" pageId="6" pageNumber="7">5A,C</figureCitation>
|
||
,
|
||
<figureCitation id="130C9AA0FFA37D09EE3A9F80FD83FF2A" box="[597,630,218,243]" captionStart="Figure 6" captionStartId="10.[171,230,2074,2099]" captionTargetBox="[195,1460,666,2031]" captionTargetId="figure-193@10.[170,1493,646,2044]" captionTargetPageId="10" captionText="Figure 6. Passalus (Passalus) cleidecostae sp. nov. (A) cephalic capsule, dorsal; (B) pronotum, lateral; (C) meso- and metasternum. Passalus (Passalus) bucki Luederwaldt,1931:(D) cephalic capsule,dorsal v.(E) pronotum,lateral.(F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985201" httpUri="https://zenodo.org/record/4985201/files/figure.png" pageId="6" pageNumber="7">6B</figureCitation>
|
||
):
|
||
</emphasis>
|
||
anterior edge slightly straight; anterior angles slightly acute and not protruding; marginal groove well-marked, deep, narrow and punctate throughout, with apex dilated, not reaching ⅓ of pronotal width; lateral fossae large, well-marked, deep and rounded; with group of coarse punctures only in the region near the lateral fossae. Prosternum (
|
||
<figureCitation id="130C9AA0FFA37D09EEAB9EC0FCFAFE6A" box="[708,783,410,435]" captionStart="Figure 4" captionStartId="8.[170,229,2106,2131]" captionTargetBox="[170,1493,338,2077]" captionTargetId="figure-55@8.[170,1493,338,2077]" captionTargetPageId="8" captionText="Figure 4. Passalus (Passalus) cleidecostaesp.nov., habitus.(A) dorsal;(B) ventral;(C) lateral." figureDoi="http://doi.org/10.5281/zenodo.4985191" httpUri="https://zenodo.org/record/4985191/files/figure.png" pageId="6" pageNumber="7">Fig. 4B</figureCitation>
|
||
):
|
||
<emphasis id="B9435A37FFA37D09ECC59EE0FECCFE0A" box="[170,313,442,467]" italics="true" pageId="6" pageNumber="7">prepisternum</emphasis>
|
||
with a few long, sparse setae on outer face;
|
||
<emphasis id="B9435A37FFA37D09ECC59E80FEDBFE2A" box="[170,302,474,499]" italics="true" pageId="6" pageNumber="7">prepimerum</emphasis>
|
||
with a few sparse bristled punctures;
|
||
<emphasis id="B9435A37FFA37D09EE8C9E80FEFEFDCA" italics="true" pageId="6" pageNumber="7">prosternelum</emphasis>
|
||
rhomboidal, with acute base.
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA37D09ECC59D60FD76FD4A" blockId="6.[170,795,570,659]" pageId="6" pageNumber="7">
|
||
<emphasis id="B9435A37FFA37D09ECC59D60FDEAFD8A" bold="true" box="[170,543,570,595]" pageId="6" pageNumber="7">
|
||
Mesosternum (
|
||
<figureCitation id="130C9AA0FFA37D09ED359D60FE4FFD8A" box="[346,442,570,595]" captionStart="Figure 4" captionStartId="8.[170,229,2106,2131]" captionTargetBox="[170,1493,338,2077]" captionTargetId="figure-55@8.[170,1493,338,2077]" captionTargetPageId="8" captionText="Figure 4. Passalus (Passalus) cleidecostaesp.nov., habitus.(A) dorsal;(B) ventral;(C) lateral." figureDoi="http://doi.org/10.5281/zenodo.4985191" httpUri="https://zenodo.org/record/4985191/files/figure.png" pageId="6" pageNumber="7">Figs. 4B</figureCitation>
|
||
,
|
||
<figureCitation id="130C9AA0FFA37D09EDAB9D60FE13FD8A" box="[452,486,570,595]" captionStart="Figure 5" captionStartId="9.[94,153,2074,2099]" captionTargetBox="[94,1417,494,2044]" captionTargetId="figure-135@9.[94,1417,494,2044]" captionTargetPageId="9" captionText="Figure 5. Passalus (Passalus) cleidecostae sp.nov.(A) head and prothorax,dorsal;(B) head,ventral (C) head and prothorax,dorsolateral;(D) posterior region of the prosternum and anterior region of the mesosternum;(E) metasternum (left side);(F) aedeagus (ventral,dorsal and lateral view)." figureDoi="http://doi.org/10.5281/zenodo.4985199" httpUri="https://zenodo.org/record/4985199/files/figure.png" pageId="6" pageNumber="7">5D</figureCitation>
|
||
,
|
||
<figureCitation id="130C9AA0FFA37D09ED9E9D60FDE6FD8A" box="[497,531,570,595]" captionStart="Figure 6" captionStartId="10.[171,230,2074,2099]" captionTargetBox="[195,1460,666,2031]" captionTargetId="figure-193@10.[170,1493,646,2044]" captionTargetPageId="10" captionText="Figure 6. Passalus (Passalus) cleidecostae sp. nov. (A) cephalic capsule, dorsal; (B) pronotum, lateral; (C) meso- and metasternum. Passalus (Passalus) bucki Luederwaldt,1931:(D) cephalic capsule,dorsal v.(E) pronotum,lateral.(F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985201" httpUri="https://zenodo.org/record/4985201/files/figure.png" pageId="6" pageNumber="7">6C</figureCitation>
|
||
):
|
||
</emphasis>
|
||
smooth and glabrous;
|
||
<emphasis id="B9435A37FFA37D09ECC59D00FE9EFDAA" box="[170,363,602,627]" italics="true" pageId="6" pageNumber="7">mesosternal scars</emphasis>
|
||
elongated, deep, with slightly rough surface, without punctures or pubescence.
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA37D09EF0A9F80FC2AFE6A" blockId="6.[869,1494,218,435]" pageId="6" pageNumber="7">
|
||
<emphasis id="B9435A37FFA37D09EF0A9F80FB1AFF2A" bold="true" box="[869,1263,218,243]" pageId="6" pageNumber="7">
|
||
Metasternum (
|
||
<figureCitation id="130C9AA0FFA37D09E8749F80FB74FF2A" box="[1051,1153,218,243]" captionStart="Figure 4" captionStartId="8.[170,229,2106,2131]" captionTargetBox="[170,1493,338,2077]" captionTargetId="figure-55@8.[170,1493,338,2077]" captionTargetPageId="8" captionText="Figure 4. Passalus (Passalus) cleidecostaesp.nov., habitus.(A) dorsal;(B) ventral;(C) lateral." figureDoi="http://doi.org/10.5281/zenodo.4985191" httpUri="https://zenodo.org/record/4985191/files/figure.png" pageId="6" pageNumber="7">Figs. 4B</figureCitation>
|
||
,
|
||
<figureCitation id="130C9AA0FFA37D09E8FF9F80FB45FF2A" box="[1168,1200,218,243]" captionStart="Figure 5" captionStartId="9.[94,153,2074,2099]" captionTargetBox="[94,1417,494,2044]" captionTargetId="figure-135@9.[94,1417,494,2044]" captionTargetPageId="9" captionText="Figure 5. Passalus (Passalus) cleidecostae sp.nov.(A) head and prothorax,dorsal;(B) head,ventral (C) head and prothorax,dorsolateral;(D) posterior region of the prosternum and anterior region of the mesosternum;(E) metasternum (left side);(F) aedeagus (ventral,dorsal and lateral view)." figureDoi="http://doi.org/10.5281/zenodo.4985199" httpUri="https://zenodo.org/record/4985199/files/figure.png" pageId="6" pageNumber="7">5E</figureCitation>
|
||
,
|
||
<figureCitation id="130C9AA0FFA37D09E8AF9F80FB16FF2A" box="[1216,1251,218,243]" captionStart="Figure 6" captionStartId="10.[171,230,2074,2099]" captionTargetBox="[195,1460,666,2031]" captionTargetId="figure-193@10.[170,1493,646,2044]" captionTargetPageId="10" captionText="Figure 6. Passalus (Passalus) cleidecostae sp. nov. (A) cephalic capsule, dorsal; (B) pronotum, lateral; (C) meso- and metasternum. Passalus (Passalus) bucki Luederwaldt,1931:(D) cephalic capsule,dorsal v.(E) pronotum,lateral.(F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985201" httpUri="https://zenodo.org/record/4985201/files/figure.png" pageId="6" pageNumber="7">6C</figureCitation>
|
||
):
|
||
</emphasis>
|
||
<emphasis id="B9435A37FFA37D09E8959F80FAD1FF2A" box="[1274,1316,218,243]" italics="true" pageId="6" pageNumber="7">disc</emphasis>
|
||
well-delimited by weak lateral carina formed by large group of punctures on lateroposterior region of metasternal disc, almost reaching metasternal lateral groove; with bristled punctures on lateroanterior region;
|
||
<emphasis id="B9435A37FFA37D09E9549E00FA20FEAA" box="[1339,1493,346,371]" italics="true" pageId="6" pageNumber="7">lateral groove</emphasis>
|
||
narrower than mesotibiae, deep, with dense bristled punctures.
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA37D09EF0A9E80FC5DFD4A" blockId="6.[869,1494,474,659]" pageId="6" pageNumber="7">
|
||
<emphasis id="B9435A37FFA37D09EF0A9E80FBD1FE2A" bold="true" box="[869,1060,474,499]" pageId="6" pageNumber="7">
|
||
Elytra (
|
||
<figureCitation id="130C9AA0FFA37D09EFD29E80FBE3FE2A" box="[957,1046,474,499]" captionStart="Figure 4" captionStartId="8.[170,229,2106,2131]" captionTargetBox="[170,1493,338,2077]" captionTargetId="figure-55@8.[170,1493,338,2077]" captionTargetPageId="8" captionText="Figure 4. Passalus (Passalus) cleidecostaesp.nov., habitus.(A) dorsal;(B) ventral;(C) lateral." figureDoi="http://doi.org/10.5281/zenodo.4985191" httpUri="https://zenodo.org/record/4985191/files/figure.png" pageId="6" pageNumber="7">Fig. 4A</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Approximately 2.5× longer and 1.1× wider than pronotum;
|
||
<emphasis id="B9435A37FFA37D09E8069EA0FB5EFDCA" box="[1129,1195,506,531]" italics="true" pageId="6" pageNumber="7">striae:</emphasis>
|
||
narrower than interstriae, marked by circular punctures, on dorsal striae smaller than that on lateral striae;
|
||
<emphasis id="B9435A37FFA37D09E8F19D60FAF0FD8A" box="[1182,1285,570,595]" italics="true" pageId="6" pageNumber="7">epipleura</emphasis>
|
||
with setae on region closest to the humeri which have a tuft of long setae.
|
||
</paragraph>
|
||
<caption id="DF48D6ADFFA37D09ECC59740FCD4F78A" ID-DOI="http://doi.org/10.5281/zenodo.4985187" ID-Zenodo-Dep="4985187" httpUri="https://zenodo.org/record/4985187/files/figure.png" pageId="6" pageNumber="7" startId="6.[170,229,2074,2099]" targetBox="[177,1482,696,2032]" targetPageId="6">
|
||
<paragraph id="8B888625FFA37D09ECC59740FCD4F78A" blockId="6.[170,1494,2074,2131]" pageId="6" pageNumber="7">
|
||
Figure 3.
|
||
<taxonomicName id="4C37FDA6FFA37D09EC909740FDF5F7EA" authority="Bevilaqua & Fonseca, 2020" authorityName="Bevilaqua & Fonseca" authorityYear="2020" box="[255,512,2074,2099]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="deuterocerus" status="sp. nov." subGenus="Pertinax">
|
||
<emphasis id="B9435A37FFA37D09EC909740FDF5F7EA" bold="true" box="[255,512,2074,2099]" italics="true" pageId="6" pageNumber="7">Passalus(Pertinax) deuterocerus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A270E74CFFA37D09EE6B9740FDB7F7EA" box="[516,578,2074,2099]" pageId="6" pageNumber="7" rank="species">sp.nov.</taxonomicNameLabel>
|
||
(A) cephalic capsule,dorsal;(B)mentum;(C) meso- and metasternum.
|
||
<taxonomicName id="4C37FDA6FFA37D09E8EB9740FF29F78A" authority="(Kuwert, 1891)" baseAuthorityName="Kuwert" baseAuthorityYear="1891" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="epiphanoides" subGenus="Pertinax">
|
||
<emphasis id="B9435A37FFA37D09E8EB9740FA7FF7EA" bold="true" box="[1156,1418,2074,2099]" italics="true" pageId="6" pageNumber="7">Passalus (Pertinax) epiphanoides</emphasis>
|
||
(Kuwert, 1891)
|
||
</taxonomicName>
|
||
:(D) cephalic capsule,dorsal.(E)mentum. (F)meso- and metasternum.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B888625FFA27D08EC319F80FECCFEAA" blockId="7.[94,718,218,371]" pageId="7" pageNumber="8">
|
||
<emphasis id="B9435A37FFA27D08EC319F80FE62FF2A" bold="true" box="[94,407,218,243]" pageId="7" pageNumber="8">
|
||
Legs (
|
||
<figureCitation id="130C9AA0FFA27D08ECCB9F80FF07FF2A" box="[164,242,218,243]" captionStart="Figure 4" captionStartId="8.[170,229,2106,2131]" captionTargetBox="[170,1493,338,2077]" captionTargetId="figure-55@8.[170,1493,338,2077]" captionTargetPageId="8" captionText="Figure 4. Passalus (Passalus) cleidecostaesp.nov., habitus.(A) dorsal;(B) ventral;(C) lateral." figureDoi="http://doi.org/10.5281/zenodo.4985191" httpUri="https://zenodo.org/record/4985191/files/figure.png" pageId="7" pageNumber="8">Figs. 4</figureCitation>
|
||
A-C, 5A, C, E):
|
||
</emphasis>
|
||
<emphasis id="B9435A37FFA27D08EDF19F80FDF7FF2A" box="[414,514,218,243]" italics="true" pageId="7" pageNumber="8">profemur</emphasis>
|
||
with well-marked groove on anteroventral border; posteroventral border with a few setae near apex;
|
||
<emphasis id="B9435A37FFA27D08EDE39E40FE14FEEA" box="[396,481,282,307]" italics="true" pageId="7" pageNumber="8">protibia</emphasis>
|
||
not dilated;
|
||
<emphasis id="B9435A37FFA27D08EE059E40FF8DFE8A" italics="true" pageId="7" pageNumber="8">mesotibiae</emphasis>
|
||
with one small spine on outer face;
|
||
<emphasis id="B9435A37FFA27D08ED929E60FD84FE8A" box="[509,625,314,339]" italics="true" pageId="7" pageNumber="8">metatibiae</emphasis>
|
||
without spine on outer face.
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA27D08EC319EC0FE6FFE0A" blockId="7.[94,718,410,467]" pageId="7" pageNumber="8">
|
||
<emphasis id="B9435A37FFA27D08EC319EC0FEB4FE6A" bold="true" box="[94,321,410,435]" pageId="7" pageNumber="8">
|
||
Abdomen (
|
||
<figureCitation id="130C9AA0FFA27D08EC8E9EC0FEC1FE6A" box="[225,308,410,435]" captionStart="Figure 4" captionStartId="8.[170,229,2106,2131]" captionTargetBox="[170,1493,338,2077]" captionTargetId="figure-55@8.[170,1493,338,2077]" captionTargetPageId="8" captionText="Figure 4. Passalus (Passalus) cleidecostaesp.nov., habitus.(A) dorsal;(B) ventral;(C) lateral." figureDoi="http://doi.org/10.5281/zenodo.4985191" httpUri="https://zenodo.org/record/4985191/files/figure.png" pageId="7" pageNumber="8">Fig. 4B</figureCitation>
|
||
):
|
||
</emphasis>
|
||
sternite VII with complete and wellmarked groove; sides rough.
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA27D08EC319EA0FED1FD2A" blockId="7.[94,719,506,755]" pageId="7" pageNumber="8">
|
||
<emphasis id="B9435A37FFA27D08EC319EA0FEB4FDCA" bold="true" box="[94,321,506,531]" pageId="7" pageNumber="8">
|
||
Aedeagus (
|
||
<figureCitation id="130C9AA0FFA27D08EC8D9EA0FEC1FDCA" box="[226,308,506,531]" captionStart="Figure 5" captionStartId="9.[94,153,2074,2099]" captionTargetBox="[94,1417,494,2044]" captionTargetId="figure-135@9.[94,1417,494,2044]" captionTargetPageId="9" captionText="Figure 5. Passalus (Passalus) cleidecostae sp.nov.(A) head and prothorax,dorsal;(B) head,ventral (C) head and prothorax,dorsolateral;(D) posterior region of the prosternum and anterior region of the mesosternum;(E) metasternum (left side);(F) aedeagus (ventral,dorsal and lateral view)." figureDoi="http://doi.org/10.5281/zenodo.4985199" httpUri="https://zenodo.org/record/4985199/files/figure.png" pageId="7" pageNumber="8">Fig. 5F</figureCitation>
|
||
):
|
||
</emphasis>
|
||
<emphasis id="B9435A37FFA27D08ED289EA0FE27FDCA" box="[327,466,506,531]" italics="true" pageId="7" pageNumber="8">Median lobe:</emphasis>
|
||
narrower than parameres and basal piece together; ventrally with two sclerotized plates lacking at median region. Basal piece attached to the parameres forming a tegmen.
|
||
<emphasis id="B9435A37FFA27D08EE2C9D00FD6DFDAA" box="[579,664,602,627]" italics="true" pageId="7" pageNumber="8">Tegmen</emphasis>
|
||
ventrally with median anterior region with V-shaped median notch; laterally, with slightly straight projections almost reaching half-length of median lobe; dorsally, with projections attached.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32DD5AEFFA27D08EC319C40FF61FBEA" pageId="7" pageNumber="8" type="etymology">
|
||
<paragraph id="8B888625FFA27D08EC319C40FF18FC6A" blockId="7.[94,718,794,947]" pageId="7" pageNumber="8">
|
||
<emphasis id="B9435A37FFA27D08EC319C40FF13FCEA" bold="true" box="[94,230,794,819]" pageId="7" pageNumber="8">Etymology:</emphasis>
|
||
We dedicate this new species to Dr. Cleide Costa, a researcher with extensive career in
|
||
<taxonomicName id="4C37FDA6FFA27D08EE3C9C60FD3BFC8A" box="[595,718,826,851]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
|
||
systematics, working especially with Neotropical immature insects and for her dedication on training taxonomists.
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA27D08EC319C80FF61FBEA" blockId="7.[94,718,986,1075]" pageId="7" pageNumber="8">
|
||
<emphasis id="B9435A37FFA27D08EC319C80FEEAFC2A" bold="true" box="[94,287,986,1011]" pageId="7" pageNumber="8">Polymorphisms:</emphasis>
|
||
The size of the inner tubercles, and the density of punctures on the lateropostfrontal areas may vary.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32DD5AEFFA27D08EC319B00FB75FD4A" pageId="7" pageNumber="8" type="discussion">
|
||
<paragraph id="8B888625FFA27D08EC319B00FBC4FECA" blockId="7.[94,719,1114,2131]" lastBlockId="7.[794,1419,218,659]" pageId="7" pageNumber="8">
|
||
<emphasis id="B9435A37FFA27D08EC319B00FF38FBAA" bold="true" box="[94,205,1114,1139]" pageId="7" pageNumber="8">Remarks:</emphasis>
|
||
<taxonomicName id="4C37FDA6FFA27D08ECBC9B00FE42FBAA" authority="Bevilaqua & Fonseca, 2020" authorityName="Bevilaqua & Fonseca" authorityYear="2020" box="[211,439,1114,1139]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
|
||
<emphasis id="B9435A37FFA27D08ECBC9B00FE42FBAA" box="[211,439,1114,1139]" italics="true" pageId="7" pageNumber="8">Passalus cleidecostae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A270E74CFFA27D08EDD19B00FDE1FBAA" box="[446,532,1114,1139]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be assigned to subgenus
|
||
<emphasis id="B9435A37FFA27D08EC809B20FE4AFB4A" box="[239,447,1146,1171]" italics="true" pageId="7" pageNumber="8">
|
||
<taxonomicName id="4C37FDA6FFA27D08EC809B20FE49FB4A" authority="(Passalus)" box="[239,444,1146,1171]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subGenus" subGenus="Passalus">Passalus (Passalus)</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
precisely in the “Neleus” section by the following characters: mediofrontal area short and transverse; presence of two well-developed secondary mediofrontal tubercles; mediofrontal and laterofrontal tubercles not fused, but close together; inner tubercles small; metasternal disc well-delimit- ed, and humeri, epipleura, lateroanterior region of the metasternum and metasternal groove pubescent. It is more similar to species whose apex of the central tubercle is not free, such as
|
||
<taxonomicName id="4C37FDA6FFA27D08ED059AC0FD3BFA6A" authority="Eschscholtz, 1829" authorityName="Eschscholtz" authorityYear="1829" box="[362,718,1434,1459]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="interstitialis">
|
||
<emphasis id="B9435A37FFA27D08ED059AC0FE0BFA6A" box="[362,510,1434,1459]" italics="true" pageId="7" pageNumber="8">P. interstitialis</emphasis>
|
||
Eschscholtz, 1829
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C37FDA6FFA27D08ECFE9AE0FE48FA0A" authority="Luederwaldt, 1931" authorityName="Luederwaldt" authorityYear="1931" box="[145,445,1466,1491]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bucki">
|
||
<emphasis id="B9435A37FFA27D08ECFE9AE0FF17FA0A" box="[145,226,1466,1491]" italics="true" pageId="7" pageNumber="8">P. bucki</emphasis>
|
||
<bibRefCitation id="EFA6FBD4FFA27D08EC859AE0FE48FA0A" author="Luederwaldt, H." box="[234,445,1466,1491]" pageId="7" pageNumber="8" pagination="1 - 262" refId="ref8431" refString="Luederwaldt, H. 1931. Monografia dos passalideos do Brasil (Col.). Revista do Museu Paulista, 17: 1 - 262." type="journal article" year="1931">Luederwaldt, 1931</bibRefCitation>
|
||
</taxonomicName>
|
||
. The new species differs from
|
||
<taxonomicName id="4C37FDA6FFA27D08ECF39A80FEDAFA2A" authorityName="Eschscholtz" authorityYear="1829" box="[156,303,1498,1523]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="interstitialis">
|
||
<emphasis id="B9435A37FFA27D08ECF39A80FEDAFA2A" box="[156,303,1498,1523]" italics="true" pageId="7" pageNumber="8">P. interstitialis</emphasis>
|
||
</taxonomicName>
|
||
by presenting smaller and more obtuse secondary mediofrontal tubercles; smaller mediofrontal tubercles and not overlapping laterofrontal tubercles; less developed inner tubercles; less straight frontal ridges; shorter and thicker antennal lamellae, not turned inwards; weaker carina on metasternal disc, and sparser pubescence on the humeri, epipleura, and metasternal anterolateral region (
|
||
<tableCitation id="C6B5B39EFFA27D08ED2A99E0FE6CF90A" box="[325,409,1722,1747]" captionStart="Table 2" captionStartId="11.[94,144,218,243]" captionText="Table 2. Differential characters of Passalus cleidecostaesp.nov.,P.interstitialis Eschscholtz,1829 and P.bucki Luederwaldt,1931." pageId="7" pageNumber="8">Table 2</tableCitation>
|
||
). The similarities with
|
||
<taxonomicName id="4C37FDA6FFA27D08EEE599E0FF85F92A" authorityName="Luederwaldt" authorityYear="1931" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bucki">
|
||
<emphasis id="B9435A37FFA27D08EEE599E0FF85F92A" italics="true" pageId="7" pageNumber="8">P. bucki</emphasis>
|
||
</taxonomicName>
|
||
are more pronounced, since both species have secondary mediofrontal tubercles smaller than laterofrontal tubercles,and with obtuse apices, presence of punctures on anterior mediofrontal and lateropostfrontal areas. However, they can be differentiated since
|
||
<taxonomicName id="4C37FDA6FFA27D08EE5A9800FD3BF8AA" authority="Bevilaqua & Fonseca, 2020" authorityName="Bevilaqua & Fonseca" authorityYear="2020" box="[565,718,1882,1907]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
|
||
<emphasis id="B9435A37FFA27D08EE5A9800FD3BF8AA" box="[565,718,1882,1907]" italics="true" pageId="7" pageNumber="8">P. cleidecostae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A270E74CFFA27D08EC319820FF46F84A" box="[94,179,1914,1939]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
presents secondary mediofrontal tubercles with more acute apices (
|
||
<figureCitation id="130C9AA0FFA27D08ED5E98C0FE50F86A" box="[305,421,1946,1971]" captionStart="Figure 6" captionStartId="10.[171,230,2074,2099]" captionTargetBox="[195,1460,666,2031]" captionTargetId="figure-193@10.[170,1493,646,2044]" captionTargetPageId="10" captionText="Figure 6. Passalus (Passalus) cleidecostae sp. nov. (A) cephalic capsule, dorsal; (B) pronotum, lateral; (C) meso- and metasternum. Passalus (Passalus) bucki Luederwaldt,1931:(D) cephalic capsule,dorsal v.(E) pronotum,lateral.(F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985201" httpUri="https://zenodo.org/record/4985201/files/figure.png" pageId="7" pageNumber="8">Figs.6A, D</figureCitation>
|
||
), inner tubercles larger and more conspicuous (
|
||
<figureCitation id="130C9AA0FFA27D08ED2F98E0FE35F80A" box="[320,448,1978,2003]" captionStart="Figure 6" captionStartId="10.[171,230,2074,2099]" captionTargetBox="[195,1460,666,2031]" captionTargetId="figure-193@10.[170,1493,646,2044]" captionTargetPageId="10" captionText="Figure 6. Passalus (Passalus) cleidecostae sp. nov. (A) cephalic capsule, dorsal; (B) pronotum, lateral; (C) meso- and metasternum. Passalus (Passalus) bucki Luederwaldt,1931:(D) cephalic capsule,dorsal v.(E) pronotum,lateral.(F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985201" httpUri="https://zenodo.org/record/4985201/files/figure.png" pageId="7" pageNumber="8">Figs. 6A, D</figureCitation>
|
||
), frontal ridges smooth (
|
||
<figureCitation id="130C9AA0FFA27D08EC0B9880FF2EF82A" box="[100,219,2010,2035]" captionStart="Figure 6" captionStartId="10.[171,230,2074,2099]" captionTargetBox="[195,1460,666,2031]" captionTargetId="figure-193@10.[170,1493,646,2044]" captionTargetPageId="10" captionText="Figure 6. Passalus (Passalus) cleidecostae sp. nov. (A) cephalic capsule, dorsal; (B) pronotum, lateral; (C) meso- and metasternum. Passalus (Passalus) bucki Luederwaldt,1931:(D) cephalic capsule,dorsal v.(E) pronotum,lateral.(F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985201" httpUri="https://zenodo.org/record/4985201/files/figure.png" pageId="7" pageNumber="8">Figs. 6A, D</figureCitation>
|
||
), anterior mediofrontal and lateropostfrontal areas with smaller and sparser punctures (
|
||
<figureCitation id="130C9AA0FFA27D08EE2998A0FD36F7CA" box="[582,707,2042,2067]" captionStart="Figure 6" captionStartId="10.[171,230,2074,2099]" captionTargetBox="[195,1460,666,2031]" captionTargetId="figure-193@10.[170,1493,646,2044]" captionTargetPageId="10" captionText="Figure 6. Passalus (Passalus) cleidecostae sp. nov. (A) cephalic capsule, dorsal; (B) pronotum, lateral; (C) meso- and metasternum. Passalus (Passalus) bucki Luederwaldt,1931:(D) cephalic capsule,dorsal v.(E) pronotum,lateral.(F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985201" httpUri="https://zenodo.org/record/4985201/files/figure.png" pageId="7" pageNumber="8">Figs. 6A, D</figureCitation>
|
||
), pronotal punctures concentrated only near the lateral fossae (
|
||
<figureCitation id="130C9AA0FFA27D08ECDB9760FEDEF78A" box="[180,299,2106,2131]" captionStart="Figure 6" captionStartId="10.[171,230,2074,2099]" captionTargetBox="[195,1460,666,2031]" captionTargetId="figure-193@10.[170,1493,646,2044]" captionTargetPageId="10" captionText="Figure 6. Passalus (Passalus) cleidecostae sp. nov. (A) cephalic capsule, dorsal; (B) pronotum, lateral; (C) meso- and metasternum. Passalus (Passalus) bucki Luederwaldt,1931:(D) cephalic capsule,dorsal v.(E) pronotum,lateral.(F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985201" httpUri="https://zenodo.org/record/4985201/files/figure.png" pageId="7" pageNumber="8">Figs. 6B, E</figureCitation>
|
||
), and metasternal punctures not ex- ceeding the lateroposterior region of the metasternal disc (
|
||
<figureCitation id="130C9AA0FFA27D08EF3E9FA0FC35FECA" box="[849,960,250,275]" captionStart="Figure 6" captionStartId="10.[171,230,2074,2099]" captionTargetBox="[195,1460,666,2031]" captionTargetId="figure-193@10.[170,1493,646,2044]" captionTargetPageId="10" captionText="Figure 6. Passalus (Passalus) cleidecostae sp. nov. (A) cephalic capsule, dorsal; (B) pronotum, lateral; (C) meso- and metasternum. Passalus (Passalus) bucki Luederwaldt,1931:(D) cephalic capsule,dorsal v.(E) pronotum,lateral.(F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985201" httpUri="https://zenodo.org/record/4985201/files/figure.png" pageId="7" pageNumber="8">Figs. 6C, F</figureCitation>
|
||
) (
|
||
<tableCitation id="C6B5B39EFFA27D08EFBD9FA0FBD3FECA" box="[978,1062,250,275]" captionStart="Table 2" captionStartId="11.[94,144,218,243]" captionText="Table 2. Differential characters of Passalus cleidecostaesp.nov.,P.interstitialis Eschscholtz,1829 and P.bucki Luederwaldt,1931." pageId="7" pageNumber="8">Table 2</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA27D08EF509E40FB75FD4A" blockId="7.[794,1419,218,659]" pageId="7" pageNumber="8">
|
||
Although
|
||
<taxonomicName id="4C37FDA6FFA27D08EFC09E40FBB2FEEA" authority="Bevilaqua & Fonseca, 2020" authorityName="Bevilaqua & Fonseca" authorityYear="2020" box="[943,1095,282,307]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cleidecostae" status="sp. nov.">
|
||
<emphasis id="B9435A37FFA27D08EFC09E40FBB2FEEA" box="[943,1095,282,307]" italics="true" pageId="7" pageNumber="8">P. cleidecostae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A270E74CFFA27D08E8229E40FB54FEEA" box="[1101,1185,282,307]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C37FDA6FFA27D08E8B99E40FAD3FEEA" authorityName="Luederwaldt" authorityYear="1931" box="[1238,1318,282,307]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bucki">
|
||
<emphasis id="B9435A37FFA27D08E8B99E40FAD3FEEA" box="[1238,1318,282,307]" italics="true" pageId="7" pageNumber="8">P. bucki</emphasis>
|
||
</taxonomicName>
|
||
are similar species, they have enough distinctive characteristics, which was possible to see after comparison with the
|
||
<taxonomicName id="4C37FDA6FFA27D08EF759E20FC9FFE4A" authorityName="Luederwaldt" authorityYear="1931" box="[794,874,378,403]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bucki">
|
||
<emphasis id="B9435A37FFA27D08EF759E20FC9FFE4A" box="[794,874,378,403]" italics="true" pageId="7" pageNumber="8">P. bucki</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="548C3887FFA27D08EF1E9E20FC56FE4A" box="[881,931,378,403]" pageId="7" pageNumber="8">type</typeStatus>
|
||
series deposited at the Museu de Zoologia da Universidade de
|
||
<collectingRegion id="49F348C7FFA27D08EF979EC0FB90FE6A" box="[1016,1125,410,435]" country="Brazil" name="Sao Paulo" pageId="7" pageNumber="8">São Paulo</collectingRegion>
|
||
(MZUSP), as well as specimens from other localities and species that also belong to the Neleus section. Both species may be more similar, since the presence of flattened central tubercle flatted, and absence of free apex are not common characteristics in this group. It is possible that within the Neleus section these species form a group which the main characteristic is related to the central tubercle.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |