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<document id="BF2814D220C2E4E60666259B3CFE6094" ID-DOI="10.37828/em.2020.31.2" ID-ISSN="2336-9744" ID-Zenodo-Dep="13231643" ID-ZooBank="urn:lsid:zoobank.org:pub:665E8674-ABFF-457A-A923-4F6B5D728718" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722913104154" checkinUser="felipe" docAuthor="Spitsyn, Vitaly M., ., .., ., ., Rmbh, N., . &amp; . V, Axio" docDate="2020" docId="038587FE603B5804A2F9F8C799F6F8CF" docLanguage="en" docName="EcolMontenegrina.31.6-9.pdf" docOrigin="Ecologica Montenegrina 31" docSource="http://dx.doi.org/10.37828/em.2020.31.2" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Cyana pseudojavanica Spitsyn, ., ., ., Rmbh, N., . &amp; V, 2020, sp. nov." docType="treatment" docVersion="2" lastPageNumber="8" masterDocId="FFBCFF86603B5806A26EFFD99A46FF85" masterDocTitle="Cyana pseudojavanica sp. nov. from Thailand and Cambodia (Lepidoptera: Erebidae: Arctiinae)" masterLastPageNumber="9" masterPageNumber="6" pageNumber="6" updateTime="1722914306330" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D716E3223BB3D1749179DA5E967789EA">Cyana pseudojavanica sp. nov. from Thailand and Cambodia (Lepidoptera: Erebidae: Arctiinae)</mods:title>
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<mods:namePart id="4CBF2C89D38076E058DA5FD79ECA8607">Spitsyn, Vitaly M.</mods:namePart>
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<mods:namePart id="4988123AB24269F8EEF4D2486FB10472">., ..</mods:namePart>
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<mods:namePart id="47E5E8F1626AC405F5FD50EC56FFF9D9">. V, Axio</mods:namePart>
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<mods:date id="E65D1FACCE8878FEEE45840DE34942C8">2020</mods:date>
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<treatment id="038587FE603B5804A2F9F8C799F6F8CF" ID-DOI="http://doi.org/10.5281/zenodo.13231649" ID-Zenodo-Dep="13231649" LSID="urn:lsid:plazi:treatment:038587FE603B5804A2F9F8C799F6F8CF" httpUri="http://treatment.plazi.org/id/038587FE603B5804A2F9F8C799F6F8CF" lastPageId="2" lastPageNumber="8" pageId="0" pageNumber="6">
<subSubSection id="C3366563603B5806A2F9F8C79849F8BD" box="[151,527,1822,1848]" pageId="0" pageNumber="6" type="nomenclature">
<paragraph id="8B9336E8603B5806A2F9F8C79849F8BD" blockId="0.[151,1159,1822,1912]" box="[151,527,1822,1848]" pageId="0" pageNumber="6">
<heading id="D0DB8184603B5806A2F9F8C79849F8BD" box="[151,527,1822,1848]" fontSize="11" level="2" pageId="0" pageNumber="6" reason="2">
<emphasis id="B958EAFA603B5806A2F9F8C79849F8BD" bold="true" box="[151,527,1822,1848]" pageId="0" pageNumber="6">
<taxonomicName id="4C2C4D6B603B5806A2F9F8C79BEFF8BD" authorityName="Spitsyn &amp; . &amp; . &amp; . &amp; Rmbh &amp; N. &amp; . &amp; .V" authorityYear="2020" box="[151,425,1822,1848]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="6" phylum="Arthropoda" rank="species" species="pseudojavanica" status="sp. nov.">
<emphasis id="B958EAFA603B5806A2F9F8C79BEFF8BD" bold="true" box="[151,425,1822,1848]" italics="true" pageId="0" pageNumber="6">Cyana pseudojavanica</emphasis>
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<taxonomicNameLabel id="A26B5781603B5806A3DEF8C79849F8BD" box="[432,527,1822,1848]" pageId="0" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3366563603B5806A2F9F8E69EC1F8FD" pageId="0" pageNumber="6" type="description">
<paragraph id="8B9336E8603B5806A2F9F8E69ABEF8DF" blockId="0.[151,1159,1822,1912]" box="[151,248,1855,1882]" pageId="0" pageNumber="6">
<figureCitation id="13172A6D603B5806A2F9F8E69ABEF8DF" box="[151,248,1855,1882]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStartId-0="1.[151,226,1699,1723]" captionStartId-1="2.[151,226,1407,1431]" captionTargetBox-0="[151,1435,189,1680]" captionTargetBox-1="[220,1356,200,1374]" captionTargetId-0="figure-16@1.[151,1435,189,1680]" captionTargetId-1="figure-10@2.[195,1392,189,1400]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figure 1. Cyana spp.: A) Holotype male of C. pseudojavanica sp. nov., Thailand; B) male of C. javanica, Flores Island, Indonesia. Scale bar = 5 mm. Photos: Vitaly M. Spitsyn." captionText-1="Figure 2. Male genitalia and aedeagus of Cyana spp.: A-B) Holotype of C. pseudojavanica sp. nov., Thailand; C-D) C. javanica, Flores Island, Indonesia. Photos: Vitaly M. Spitsyn." figureDoi-0="http://doi.org/10.5281/zenodo.13231645" figureDoi-1="http://doi.org/10.5281/zenodo.13231647" httpUri-0="https://zenodo.org/record/13231645/files/figure.png" httpUri-1="https://zenodo.org/record/13231647/files/figure.png" pageId="0" pageNumber="6">Figs 1-2</figureCitation>
</paragraph>
<paragraph id="8B9336E8603B5806A2F9F8B99EC1F8FD" blockId="0.[151,1159,1822,1912]" box="[151,1159,1888,1912]" pageId="0" pageNumber="6">
https://zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection id="C3366563603B5806A2F9F8789F36F85B" pageId="0" pageNumber="6" type="materials_examined">
<paragraph id="8B9336E8603B5806A2F9F8789F36F85B" blockId="0.[151,1435,1953,2014]" pageId="0" pageNumber="6">
<emphasis id="B958EAFA603B5806A2F9F8789B14F83E" bold="true" box="[151,338,1953,1979]" pageId="0" pageNumber="6">Type material.</emphasis>
<materialsCitation id="3B443CB5603B5806A333F8789F36F85B" collectingDate="2016-05-30" collectingDateMax="2016-06-19" collectingDateMin="2016-05-30" collectionCode="V, RMBH" collectorName="Spitsyn" country="Thailand" latitude="19.572" location="Mountain" longLatPrecision="1" longitude="98.280785" municipality="Cave Lodge" pageId="0" pageNumber="6" specimenCount="1" stateProvince="Mae Hong Son" typeStatus="holotype">
<emphasis id="B958EAFA603B5806A333F8789850F83E" bold="true" box="[349,534,1953,1979]" pageId="0" pageNumber="6">
<typeStatus id="5497884A603B5806A333F8789B89F83E" box="[349,463,1953,1979]" pageId="0" pageNumber="6" type="holotype">Holotype</typeStatus>
male
</emphasis>
:
<collectingCountry id="F33B7678603B5806A047F8789882F839" box="[553,708,1953,1980]" name="Thailand" pageId="0" pageNumber="6">THAILAND</collectingCountry>
,
<collectingRegion id="49E8F80A603B5806A0BDF87B9E47F839" box="[723,1025,1953,1980]" country="Thailand" name="Mae Hong Son" pageId="0" pageNumber="6">Mae Hong Son Province</collectingRegion>
,
<collectingMunicipality id="6BF7AC92603B5806A661F8789EE2F839" box="[1039,1188,1953,1980]" pageId="0" pageNumber="6">Cave Lodge</collectingMunicipality>
,
<location id="8EF36033603B5806A6DDF8789F63F839" LSID="urn:lsid:plazi:treatment:038587FE603B5804A2F9F8C799F6F8CF:8EF36033603B5806A6DDF8789F63F839" box="[1203,1317,1953,1980]" country="Thailand" latitude="19.572" longLatPrecision="1" longitude="98.280785" municipality="Cave Lodge" name="Mountain" pageId="0" pageNumber="6" stateProvince="Mae Hong Son">Mountain</location>
monsoon forest in the river valley,
<geoCoordinate id="EE18502F603B5806A3D1F81A981DF85B" box="[447,603,1987,2014]" degrees="19" direction="north" minutes="34.320" orientation="latitude" pageId="0" pageNumber="6" precision="1" value="19.572">19°34.320N</geoCoordinate>
,
<geoCoordinate id="EE18502F603B5806A008F81A98B9F85B" box="[614,767,1987,2014]" degrees="98" direction="east" minutes="16.847" orientation="longitude" pageId="0" pageNumber="6" precision="1" value="98.280785">98°16.847E</geoCoordinate>
,
<date id="FF921028603B5806A164F81A998BF85B" box="[778,973,1987,2014]" pageId="0" pageNumber="6" value="2016-05-30" valueMax="2016-06-19" valueMin="2016-05-30">
<collectingDate id="EFD6E9C0603B5806A164F81A998BF85B" box="[778,973,1987,2014]" pageId="0" pageNumber="6" value="2016-05-30" valueMax="2016-06-19" valueMin="2016-05-30">30.v19.vi.2016</collectingDate>
</date>
,
<collectionCode id="ED3DAE2D603B5806A1B7F81D99B4F85B" box="[985,1010,1988,2014]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="0" pageNumber="6" type="Museum">V</collectionCode>
.
<collectorName id="26D9533E603B5806A193F81A9E12F85B" box="[1021,1108,1987,2014]" pageId="0" pageNumber="6">Spitsyn</collectorName>
leg. (
<collectionCode id="ED3DAE2D603B5806A6F8F81A9EA9F85B" box="[1174,1263,1987,2014]" pageId="0" pageNumber="6">RMBH</collectionCode>
Sph0848).
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</paragraph>
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<caption id="DF536660603A5807A2F9F97A9979F95C" ID-DOI="http://doi.org/10.5281/zenodo.13231645" ID-Zenodo-Dep="13231645" httpUri="https://zenodo.org/record/13231645/files/figure.png" pageId="1" pageNumber="7" startId="1.[151,226,1699,1723]" targetBox="[151,1435,189,1680]" targetPageId="1" targetType="figure">
<paragraph id="8B9336E8603A5807A2F9F97A9979F95C" blockId="1.[151,1436,1698,1753]" pageId="1" pageNumber="7">
<emphasis id="B958EAFA603A5807A2F9F97A9B47F93E" bold="true" box="[151,257,1699,1723]" pageId="1" pageNumber="7">Figure 1.</emphasis>
<taxonomicName id="4C2C4D6B603A5807A362F97A9BCAF93F" box="[268,396,1699,1722]" class="Insecta" family="Erebidae" genus="Cyana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="7" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B958EAFA603A5807A362F97A9B14F93F" box="[268,338,1699,1722]" italics="true" pageId="1" pageNumber="7">Cyana</emphasis>
spp.
</taxonomicName>
:
<emphasis id="B958EAFA603A5807A3F4F97A9BF1F93E" bold="true" box="[410,439,1699,1723]" pageId="1" pageNumber="7">A)</emphasis>
Holotype male of
<taxonomicName id="4C2C4D6B603A5807A0FEF97A991CF93F" authorityName="Spitsyn &amp; . &amp; . &amp; . &amp; Rmbh &amp; N. &amp; . &amp; .V" authorityYear="2020" box="[656,858,1698,1722]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="7" phylum="Arthropoda" rank="species" species="pseudojavanica" status="sp. nov.">
<emphasis id="B958EAFA603A5807A0FEF97A991CF93F" box="[656,858,1698,1722]" italics="true" pageId="1" pageNumber="7">C. pseudojavanica</emphasis>
</taxonomicName>
<emphasis id="B958EAFA603A5807A108F97A9987F93E" bold="true" box="[870,961,1699,1723]" pageId="1" pageNumber="7">
<taxonomicNameLabel id="A26B5781603A5807A108F97A9987F93E" box="[870,961,1699,1723]" pageId="1" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Thailand;
<emphasis id="B958EAFA603A5807A62DF97A9E18F93E" bold="true" box="[1091,1118,1699,1723]" pageId="1" pageNumber="7">B)</emphasis>
male of
<taxonomicName id="4C2C4D6B603A5807A6A7F97A9F0EF93F" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2016" baseAuthorityName="Butler" baseAuthorityYear="1877" box="[1225,1352,1699,1722]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="7" phylum="Arthropoda" rank="species" species="javanica">
<emphasis id="B958EAFA603A5807A6A7F97A9F0EF93F" box="[1225,1352,1699,1722]" italics="true" pageId="1" pageNumber="7">C. javanica</emphasis>
</taxonomicName>
, Flores Island, Indonesia. Scale bar = 5 mm. Photos: Vitaly M. Spitsyn.
</paragraph>
</caption>
<subSubSection id="C3366563603A5807A2F9F8FD99CDF821" pageId="1" pageNumber="7" type="diagnosis">
<paragraph id="8B9336E8603A5807A2F9F8FD99CDF821" blockId="1.[151,1436,1828,1956]" pageId="1" pageNumber="7">
<emphasis id="B958EAFA603A5807A2F9F8FD9B51F8BB" bold="true" box="[151,279,1828,1854]" pageId="1" pageNumber="7">Diagnosis.</emphasis>
The novel species is morphologically similar to
<taxonomicName id="4C2C4D6B603A5807A1E2F8FD9F4FF8BA" authority="(Butler, 1877)" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2016" baseAuthorityName="Butler" baseAuthorityYear="1877" box="[908,1289,1828,1855]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="7" phylum="Arthropoda" rank="species" species="javanica">
<emphasis id="B958EAFA603A5807A1E2F8FD9E0AF8BB" box="[908,1100,1828,1854]" italics="true" pageId="1" pageNumber="7">Cyana javanica</emphasis>
(Butler, 1877)
</taxonomicName>
and
<taxonomicName id="4C2C4D6B603A5807A73EF8FD9B86F8E5" authority="(Walker, 1862)" baseAuthorityName="Walker" baseAuthorityYear="1862" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="7" phylum="Arthropoda" rank="species" species="conclusa">
<emphasis id="B958EAFA603A5807A73EF8FD9AB9F8E5" italics="true" pageId="1" pageNumber="7">Cyana conclusa</emphasis>
(Walker, 1862)
</taxonomicName>
, but can be distinguished from those taxa by a broad distal section of the valva (vs narrow and long distal section of the valva in
<taxonomicName id="4C2C4D6B603A5807A0B6F8B19922F804" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2016" baseAuthorityName="Butler" baseAuthorityYear="1877" box="[728,868,1896,1922]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="7" phylum="Arthropoda" rank="species" species="javanica">
<emphasis id="B958EAFA603A5807A0B6F8B19922F804" box="[728,868,1896,1922]" italics="true" pageId="1" pageNumber="7">C. javanica</emphasis>
</taxonomicName>
) and by the lack of a cluster of spines in the aedeagus vesica (vs the presence of such cluster in
<taxonomicName id="4C2C4D6B603A5807A081F853993CF821" baseAuthorityName="Walker" baseAuthorityYear="1862" box="[751,890,1929,1956]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="7" phylum="Arthropoda" rank="species" species="conclusa">
<emphasis id="B958EAFA603A5807A081F853993CF821" box="[751,890,1929,1956]" italics="true" pageId="1" pageNumber="7">C. conclusa</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="DF53666060395804A2F9FAA6996EFA30" ID-DOI="http://doi.org/10.5281/zenodo.13231647" ID-Zenodo-Dep="13231647" httpUri="https://zenodo.org/record/13231647/files/figure.png" pageId="2" pageNumber="8" startId="2.[151,226,1407,1431]" targetBox="[220,1356,200,1374]" targetPageId="2" targetType="figure">
<paragraph id="8B9336E860395804A2F9FAA6996EFA30" blockId="2.[151,1436,1406,1461]" pageId="2" pageNumber="8">
<emphasis id="B958EAFA60395804A2F9FAA69ABBFA12" bold="true" box="[151,253,1407,1431]" pageId="2" pageNumber="8">Figure 2.</emphasis>
Male genitalia and aedeagus of
<taxonomicName id="4C2C4D6B60395804A036FAA69893FA13" box="[600,725,1407,1430]" class="Insecta" family="Erebidae" genus="Cyana" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B958EAFA60395804A036FAA698D8FA13" box="[600,670,1407,1430]" italics="true" pageId="2" pageNumber="8">Cyana</emphasis>
spp.
</taxonomicName>
:
<emphasis id="B958EAFA60395804A08EFAA69950FA12" bold="true" box="[736,790,1407,1431]" pageId="2" pageNumber="8">A-B)</emphasis>
Holotype of
<taxonomicName id="4C2C4D6B60395804A1CBFAA69E2DFA13" authorityName="Spitsyn &amp; . &amp; . &amp; . &amp; Rmbh &amp; N. &amp; . &amp; .V" authorityYear="2020" box="[933,1131,1406,1430]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="8" phylum="Arthropoda" rank="species" species="pseudojavanica" status="sp. nov.">
<emphasis id="B958EAFA60395804A1CBFAA69E2DFA13" box="[933,1131,1406,1430]" italics="true" pageId="2" pageNumber="8">C. pseudojavanica</emphasis>
</taxonomicName>
<emphasis id="B958EAFA60395804A61CFAA69E8CFA12" bold="true" box="[1138,1226,1407,1431]" pageId="2" pageNumber="8">
<taxonomicNameLabel id="A26B578160395804A61CFAA69E8CFA12" box="[1138,1226,1407,1431]" pageId="2" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Thailand;
<emphasis id="B958EAFA60395804A72AFAA69F3AFA12" bold="true" box="[1348,1404,1407,1431]" pageId="2" pageNumber="8">C-D)</emphasis>
<taxonomicName id="4C2C4D6B60395804A7EAFAA69AB5FA30" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2016" baseAuthorityName="Butler" baseAuthorityYear="1877" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="8" phylum="Arthropoda" rank="species" species="javanica">
<emphasis id="B958EAFA60395804A7EAFAA69AB5FA30" italics="true" pageId="2" pageNumber="8">C. javanica</emphasis>
</taxonomicName>
, Flores Island, Indonesia. Photos: Vitaly M. Spitsyn.
</paragraph>
</caption>
<subSubSection id="C336656360395804A2F9F9D99806F882" pageId="2" pageNumber="8" type="description">
<paragraph id="8B9336E860395804A2F9F9D99EC3F927" blockId="2.[151,1436,1536,1799]" pageId="2" pageNumber="8">
<emphasis id="B958EAFA60395804A2F9F9D99B69F99F" bold="true" box="[151,303,1536,1562]" pageId="2" pageNumber="8">Description.</emphasis>
<emphasis id="B958EAFA60395804A356F9D99854F99E" box="[312,530,1536,1563]" italics="true" pageId="2" pageNumber="8">Male morphology:</emphasis>
Wingspan
<quantity id="4CD49B0D60395804A0F7F9D998B6F99E" box="[665,752,1536,1563]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="2" pageNumber="8" unit="mm" value="36.0">36 mm</quantity>
, forewing length
<quantity id="4CD49B0D60395804A1D3F9D99E53F99E" box="[957,1045,1536,1563]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="8" unit="mm" value="18.0">18 mm</quantity>
. Eye gray. Antenna brown. Head white. Labial palpus short, brown. Thorax white. Patagium, and tegula white, with pink apex. Legs white with pink spots. Forewing upperside white; basal, antemedial and postmedial lines red; outer margin orange; apex of the central cell with two black spots. A line of androconial scales is located along the costal margin. Hindwings pink. Underside of forewings white, hindwings light pink. Abdomen pink.
</paragraph>
<paragraph id="8B9336E860395804A29BF9719806F882" blockId="2.[151,1436,1536,1799]" pageId="2" pageNumber="8">
<emphasis id="B958EAFA60395804A29BF9719BE0F946" box="[245,422,1704,1731]" italics="true" pageId="2" pageNumber="8">Male genitalia</emphasis>
: Uncus long; tegumen long, u-shaped; saccus broad. Valva long, distal section of valva broad; the ampulla is long, wide and curved; distal saccular process long, curved. Aedeagus straight, without a cluster of spines in vesica.
</paragraph>
</subSubSection>
<subSubSection id="C336656360395804A2F9F8E999F6F8CF" box="[151,944,1839,1866]" pageId="2" pageNumber="8" type="distribution">
<paragraph id="8B9336E860395804A2F9F8E999F6F8CF" blockId="2.[151,944,1839,1866]" box="[151,944,1839,1866]" pageId="2" pageNumber="8">
<emphasis id="B958EAFA60395804A2F9F8E99B71F8CF" bold="true" box="[151,311,1840,1866]" pageId="2" pageNumber="8">Distribution.</emphasis>
<collectingCountry id="F33B767860395804A351F8F69BE1F8CF" box="[319,423,1839,1866]" name="Thailand" pageId="2" pageNumber="8">Thailand</collectingCountry>
and
<collectingCountry id="F33B767860395804A38EF8F6981FF8CF" box="[480,601,1839,1866]" name="Cambodia" pageId="2" pageNumber="8">Cambodia</collectingCountry>
(
<bibRefCitation id="EFBD4B1960395804A006F8F699E5F8CF" author="Bayarsaikhan, U. &amp; Bae, Y. S." box="[616,931,1839,1866]" pageId="2" pageNumber="8" pagination="447 - 463" refId="ref840" refString="Bayarsaikhan, U. &amp; Bae, Y. S. (2016) A review of the genus Cyana Walker, 1854 (Lepidoptera, Erebidae, Arctiinae) from Cambodia, with description of new species. Zootaxa, 4114 (4), 447 - 463. http: // doi. org / 10.11646 / zootaxa. 4114.4.5" type="journal article" year="2016">Bayarsaikhan &amp; Bae 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>