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<document id="6394912D8E6CDE8E4063C2D2BDE2CC3D" ID-CLB-Dataset="7213" ID-DOI="10.11646/zootaxa.4958.1.31" ID-GBIF-Dataset="a1df68e3-0e4c-4010-9556-cb15d51a5499" ID-ISSN="1175-5326" ID-Zenodo-Dep="4692369" ID-ZooBank="9E63F8E2-1CCA-4F58-9019-39AE7A3D0640" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1618471284689" checkinUser="plazi" docAuthor="Salini, S. &amp; Kment, Petr" docDate="2021" docId="038587F46806FFC64E8D61F2FE5EFEC1" docLanguage="en" docName="zootaxa.4958.1.31.pdf" docOrigin="Zootaxa 4958 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Riazocoris Ahmad &amp; Afzal 1979" docType="treatment" docVersion="6" lastPageNumber="549" masterDocId="FFBCFF8C6823FFE14E1A672FFFFDFF80" masterDocTitle="The genera Agathocles and Surenus (Hemiptera: Heteroptera: Pentatomidae) tribal reassessment, redescription, new synonyms, and description of two new species" masterLastPageNumber="559" masterPageNumber="510" pageNumber="547" updateTime="1698936917073" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2EEBA2FE731B313E3860D0767E99D764">The genera Agathocles and Surenus (Hemiptera: Heteroptera: Pentatomidae) tribal reassessment, redescription, new synonyms, and description of two new species</mods:title>
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<mods:namePart id="C390B4C76D445B19ACDE5D10AE19A0F4">Salini, S.</mods:namePart>
<mods:affiliation id="0BFFC1B8CA26A70B174B070794D222DF">National Bureau of Agricultural Insect Resources, Bangalore, 560024, India</mods:affiliation>
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<mods:namePart id="4F27F8B31CE995AD3725CF9A62CFA70A">Kment, Petr</mods:namePart>
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<mods:affiliation id="CFD7CAD645206EAF4A4E4417A9B121AA">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic sigara @ post. cz; https: // orcid. org / 0000 - 0002 - 7026 - 5691</mods:affiliation>
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<mods:number id="1FA1C7FCECDA0BD7C85CA2358999774F">2021-04-14</mods:number>
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<subSubSection id="C33665696806FFC44E8D61F2FDD1F977" box="[151,556,1757,1783]" pageId="37" pageNumber="547" type="nomenclature">
<paragraph id="8B9336E26806FFC44E8D61F2FDD1F977" blockId="37.[151,556,1757,1783]" box="[151,556,1757,1783]" pageId="37" pageNumber="547">
<heading id="D0DB818E6806FFC44E8D61F2FDD1F977" bold="true" box="[151,556,1757,1783]" fontSize="11" level="1" pageId="37" pageNumber="547" reason="1">
<taxonomicName id="4C2C4D616806FFC44E8D61F2FDD1F977" authority="Ahmad &amp; Afzal, 1979" authorityName="Ahmad &amp; Afzal" authorityYear="1979" box="[151,556,1757,1783]" class="Insecta" family="Pentatomidae" genus="Riazocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF06806FFC44E8D61F2FDD1F977" bold="true" box="[151,556,1757,1783]" pageId="37" pageNumber="547">
<emphasis id="B958EAF06806FFC44E8D61F2FEEBF977" bold="true" box="[151,278,1757,1783]" italics="true" pageId="37" pageNumber="547">Riazocoris</emphasis>
<bibRefCitation id="EFBD4B136806FFC44F0461F2FDD1F977" author="Ahmad, I. &amp; Afzal, M." box="[286,556,1757,1783]" pageId="37" pageNumber="547" pagination="1 - 14" refId="ref27835" refString="Ahmad, I. &amp; Afzal, M. (1979) Resurrection of the tribe Caystrini Stal (Heteroptera, Pentatomidae, Pentatominae) with description of two new genera from Oriental region. Annotationes Zoologicae et Botanicae, 133, 1 - 14." type="journal article" year="1979">Ahmad &amp; Afzal, 1979</bibRefCitation>
</emphasis>
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</paragraph>
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<paragraph id="8B9336E26806FFC44E8D600DFCC2F8D8" blockId="37.[151,1437,1826,1972]" pageId="37" pageNumber="547">
<treatmentCitationGroup id="AB3C11CC6806FFC44E8D600DFCC2F8D8" pageId="37" pageNumber="547">
<taxonomicName id="4C2C4D616806FFC44E8D600DFEFEF8B9" authorityName="Ahmad &amp; Afzal" authorityYear="1979" box="[151,259,1826,1849]" class="Insecta" family="Pentatomidae" genus="Riazocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF06806FFC44E8D600DFEFEF8B9" box="[151,259,1826,1849]" italics="true" pageId="37" pageNumber="547">Riazocoris</emphasis>
</taxonomicName>
(unavailable):
<treatmentCitation id="0A8D10F36806FFC44FB8600DFD50F8BA" author="Ahmad, I. &amp; Afzal, M. &amp; Siddiqui, M. I." box="[418,685,1826,1850]" page="161" pageId="37" pageNumber="547" year="1977">
<bibRefCitation id="EFBD4B136806FFC44FB8600DFD81F8BA" author="Ahmad, I. &amp; Afzal, M. &amp; Siddiqui, M. I." box="[418,636,1826,1850]" pageId="37" pageNumber="547" pagination="161 - 165" refId="ref27925" refString="Ahmad, I., Afzal, M. &amp; Siddiqui, M. I. (1977) Aspects of morphology of Riazocoris nigra Ahmad and Afzal (Pentatominae: Caystrini) with phylogenetic considerations. Pakistan Journal of Zoology, 9 (2), 161 - 165." type="journal article" year="1977">
Ahmad
<emphasis id="B958EAF06806FFC44FEF600CFDD1F8BA" box="[501,556,1826,1850]" italics="true" pageId="37" pageNumber="547">et al.</emphasis>
(1977)
</bibRefCitation>
: 161
</treatmentCitation>
–164,
<figureCitation id="13172A676806FFC44CEB600DFCB1F8BA" box="[753,844,1826,1850]" captionStart="FIGURES 1–5" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[203,1384,183,1912]" captionTargetId="figure-29@7.[196,1392,179,1917]" captionTargetPageId="7" captionText="FIGURES 1–5. Agathocles limbatus Stål, 1876. 1–2—habitus of male from Laos: Ban Saluey → Phou Pane Mt. (22.2 mm). 3–5—habitus and labels of holotype of A. yunnanensis Zhang &amp; Lin, 1984, syn. nov., female, China: Yunnan: Cang-yuan (scale bar 1 mm). Orig. P. Kment (1–2), D. Rédei (3–5)." figureDoi="http://doi.org/10.5281/zenodo.4692373" httpUri="https://zenodo.org/record/4692373/files/figure.png" pageId="37" pageNumber="547">Figs 1–2</figureCitation>
(description of morphology of alimentary and male and female reproductive organs, salivary glands and scent glands).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9336E26806FFC44E8D604FFEA5F834" blockId="37.[151,1437,1826,1972]" pageId="37" pageNumber="547">
<treatmentCitationGroup id="AB3C11CC6806FFC44E8D604FFEA5F834" pageId="37" pageNumber="547">
<treatmentCitation id="0A8D10F36806FFC44E8D604FFDDFF8F7" author="Ahmad, I. &amp; Afzal, M." box="[151,546,1887,1911]" page="7" pageId="37" pageNumber="547" year="1979">
<taxonomicName id="4C2C4D616806FFC44E8D604FFDDFF8F7" authority="Ahmad &amp; Afzal, 1979: 4 - 7" authorityName="Ahmad &amp; Afzal" authorityPageNumber="4 - 7" authorityYear="1979" box="[151,546,1887,1911]" class="Insecta" family="Pentatomidae" genus="Riazocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF06806FFC44E8D604FFEFEF8F7" box="[151,259,1888,1911]" italics="true" pageId="37" pageNumber="547">Riazocoris</emphasis>
<bibRefCitation id="EFBD4B136806FFC44F116070FDDFF8F7" author="Ahmad, I. &amp; Afzal, M." box="[267,546,1887,1911]" pageId="37" pageNumber="547" pagination="1 - 14" refId="ref27835" refString="Ahmad, I. &amp; Afzal, M. (1979) Resurrection of the tribe Caystrini Stal (Heteroptera, Pentatomidae, Pentatominae) with description of two new genera from Oriental region. Annotationes Zoologicae et Botanicae, 133, 1 - 14." type="journal article" year="1979">Ahmad &amp; Afzal, 1979: 4–7</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(description, illustrations);
<treatmentCitation id="0A8D10F36806FFC44D266070FBFAF8F7" author="Ahmad, I." box="[828,1031,1887,1911]" page="125" pageId="37" pageNumber="547" year="1980">
<bibRefCitation id="EFBD4B136806FFC44D266070FC28F8F7" author="Ahmad, I." box="[828,981,1887,1911]" pageId="37" pageNumber="547" pagination="115 - 155" refId="ref27587" refString="Ahmad, I. (1980) Insect fauna of Pakistan and Azad Kashmir - some groups within the order Hemiptera. Proceedings of the Pakistan Congress of Zoology, 1, 115 - 155." type="journal article" year="1980">Ahmad (1980)</bibRefCitation>
: 125
</treatmentCitation>
, 137 (list);
<treatmentCitation id="0A8D10F36806FFC44A666070FAC1F8F7" author="Ahmad, I." box="[1148,1340,1887,1911]" page="52" pageId="37" pageNumber="547" year="1981">
<bibRefCitation id="EFBD4B136806FFC44A666070FAE8F8F7" author="Ahmad, I." box="[1148,1301,1887,1911]" pageId="37" pageNumber="547" pagination="1 - 113" refId="ref27623" refString="Ahmad, I. (1981) A revision of the superfamilies Coreoidea and Pentatomoidea (Heteroptera: Pentatomomorpha) from Pakistan, Azad Kashmir and Bangladesh. Part 1: Additions and corrections of coreid and pentatomid fauna with phylogenetic considerations. Supplement of the Entomological Society of Karachi, 4 (1), 1 - 113. [1979]" type="journal article" year="1981">Ahmad (1981)</bibRefCitation>
: 52
</treatmentCitation>
, 105, fig. 8 (checklist, illustration);
<treatmentCitation id="0A8D10F36806FFC44FD46051FCC4F816" author="Ahmad, I. &amp; Kamaluddin, S." box="[462,825,1918,1942]" page="170" pageId="37" pageNumber="547" year="1989">
<bibRefCitation id="EFBD4B136806FFC44FD46051FCFAF816" author="Ahmad, I. &amp; Kamaluddin, S." box="[462,775,1918,1942]" pageId="37" pageNumber="547" pagination="169 - 183" refId="ref28067" refString="Ahmad, I. &amp; Kamaluddin, S. (1989) A revision of the tribe Caystrini (Hemiptera: Pentatomidae: Pentatominae) from Indo- Pakistan subcontinent with description of two new species from Pakistan and their cladistic analysis. Proceedings of the Pakistan Congress of Zoology, 9, 169 - 183." type="journal article" year="1989">Ahmad &amp; Kamaluddin (1989)</bibRefCitation>
: 170
</treatmentCitation>
, 174, 176, 181–182 (key to genera, diagnosis, phylogenetic relationships).
</treatmentCitationGroup>
</paragraph>
<caption id="DF53666A6805FFC74E8D606DFB41F86A" ID-DOI="http://doi.org/10.5281/zenodo.4692421" ID-Zenodo-Dep="4692421" httpUri="https://zenodo.org/record/4692421/files/figure.png" pageId="38" pageNumber="548" startId="38.[151,264,1858,1882]" targetBox="[221,1366,207,1834]" targetPageId="38">
<paragraph id="8B9336E26805FFC74E8D606DFB41F86A" blockId="38.[151,1437,1858,2026]" pageId="38" pageNumber="548">
<emphasis id="B958EAF06805FFC74E8D606DFE86F8DA" bold="true" box="[151,379,1858,1882]" pageId="38" pageNumber="548">FIGURES 130–-138.</emphasis>
Out-group comparison. 130–132—parameres (lateral view):130—
<taxonomicName id="4C2C4D616805FFC74A21606DFA64F8DA" authority="Distant, 1901" authorityName="Distant" authorityYear="1901" box="[1083,1433,1858,1882]" class="Insecta" family="Pentatomidae" genus="Caystrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="548" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B958EAF06805FFC74A21606DFB01F8DA" box="[1083,1276,1858,1882]" italics="true" pageId="38" pageNumber="548">Caystrus obscurus</emphasis>
Distant, 1901
</taxonomicName>
; 131—
<taxonomicName id="4C2C4D616805FFC74ECD6048FEDBF8FE" authorityName="Stal" authorityYear="1861" box="[215,294,1895,1918]" class="Insecta" family="Pentatomidae" genus="Laprius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="548" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF06805FFC74ECD6048FEDBF8FE" box="[215,294,1895,1918]" italics="true" pageId="38" pageNumber="548">Laprius</emphasis>
</taxonomicName>
sp.; 132—
<taxonomicName id="4C2C4D616805FFC74F876049FD01F8FE" authority="Westwood, 1837" authorityName="Westwood" authorityYear="1837" box="[413,764,1894,1918]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="38" pageNumber="548" phylum="Chordata" rank="species" species="serrigera">
<emphasis id="B958EAF06805FFC74F876049FDBCF8FE" box="[413,577,1894,1918]" italics="true" pageId="38" pageNumber="548">Halys serrigera</emphasis>
Westwood, 1837
</taxonomicName>
. 133–136—phallus: 133—
<taxonomicName id="4C2C4D616805FFC74A0C6048FB6AF8FE" authorityName="Westwood" authorityYear="1837" box="[1046,1175,1895,1918]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="38" pageNumber="548" phylum="Chordata" rank="species" species="serrigera">
<emphasis id="B958EAF06805FFC74A0C6048FB6AF8FE" box="[1046,1175,1895,1918]" italics="true" pageId="38" pageNumber="548">H. serrigera</emphasis>
</taxonomicName>
(ventral view); 134—
<taxonomicName id="4C2C4D616805FFC74B9F6049FF0FF822" authorityName="Distant" authorityYear="1901" class="Insecta" family="Pentatomidae" genus="Caystrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="548" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B958EAF06805FFC74B9F6049FF0FF822" italics="true" pageId="38" pageNumber="548">C. obscurus</emphasis>
</taxonomicName>
(ventral view); 135–136—
<taxonomicName id="4C2C4D616805FFC74C0E60A4FD9EF822" authorityName="Stal" authorityYear="1861" box="[532,611,1931,1954]" class="Insecta" family="Pentatomidae" genus="Laprius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="548" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF06805FFC74C0E60A4FD9EF822" box="[532,611,1931,1954]" italics="true" pageId="38" pageNumber="548">Laprius</emphasis>
</taxonomicName>
sp. (135—ventral view, 136—dorsal view). 137–138—external scent efferent system of the metathoracic scent gland: 137—
<taxonomicName id="4C2C4D616805FFC74C926080FCF4F846" authorityName="Westwood" authorityYear="1837" box="[648,777,1967,1990]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="38" pageNumber="548" phylum="Chordata" rank="species" species="serrigera">
<emphasis id="B958EAF06805FFC74C926080FCF4F846" box="[648,777,1967,1990]" italics="true" pageId="38" pageNumber="548">H. serrigera</emphasis>
</taxonomicName>
, 138—
<taxonomicName id="4C2C4D616805FFC74D406081FC39F846" authorityName="Distant" authorityYear="1902" box="[858,964,1966,1990]" class="Insecta" genus="Exithemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="548" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF06805FFC74D406081FC39F846" box="[858,964,1966,1990]" italics="true" pageId="38" pageNumber="548">Exithemus</emphasis>
</taxonomicName>
sp. Lettering: dlc—dorsolateral conjunctival process, pe—peritreme, vlc—ventrolateral conjunctical process. Orig. Salini (130–137), P. Kment (138).
</paragraph>
</caption>
<paragraph id="8B9336E26804FFC64E8D67B8FE5EFEC1" blockId="39.[151,1436,151,321]" pageId="39" pageNumber="549">
<emphasis id="B958EAF06804FFC64E8D67B8FEBAFF31" bold="true" box="[151,327,151,177]" pageId="39" pageNumber="549">Nomenclature.</emphasis>
The generic name
<taxonomicName id="4C2C4D616804FFC64C3967B8FCAAFF31" authority="Ahmad &amp; Afzal" authorityName="Ahmad &amp; Afzal" authorityYear="1979" box="[547,855,151,177]" class="Insecta" family="Pentatomidae" genus="Riazocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="549" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF06804FFC64C3967B8FD66FF30" box="[547,667,151,176]" italics="true" pageId="39" pageNumber="549">Riazocoris</emphasis>
Ahmad &amp; Afzal
</taxonomicName>
first appeared in the paper by
<bibRefCitation id="EFBD4B136804FFC64AAF67B8FA61FF31" author="Ahmad, I. &amp; Afzal, M. &amp; Siddiqui, M. I." box="[1205,1436,151,177]" pageId="39" pageNumber="549" pagination="161 - 165" refId="ref27925" refString="Ahmad, I., Afzal, M. &amp; Siddiqui, M. I. (1977) Aspects of morphology of Riazocoris nigra Ahmad and Afzal (Pentatominae: Caystrini) with phylogenetic considerations. Pakistan Journal of Zoology, 9 (2), 161 - 165." type="journal article" year="1977">
Ahmad
<emphasis id="B958EAF06804FFC64B0A67B8FAB4FF31" box="[1296,1353,151,177]" italics="true" pageId="39" pageNumber="549">et al.</emphasis>
(1977)
</bibRefCitation>
describing the internal organs of
<taxonomicName id="4C2C4D616804FFC64C0F6794FD2CFF54" authorityName="Ahmad &amp; Afzal" authorityYear="1977" box="[533,721,187,212]" class="Insecta" family="Pentatomidae" genus="Riazocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="549" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B958EAF06804FFC64C0F6794FD2CFF54" box="[533,721,187,212]" italics="true" pageId="39" pageNumber="549">Riazocoris niger</emphasis>
</taxonomicName>
. In that paper neither the
<typeStatus id="549788406804FFC64A186794FBCFFF55" box="[1026,1074,187,213]" pageId="39" pageNumber="549">type</typeStatus>
species of
<taxonomicName id="4C2C4D616804FFC64AAD6794FAD2FF54" authorityName="Ahmad &amp; Afzal" authorityYear="1979" box="[1207,1327,187,212]" class="Insecta" family="Pentatomidae" genus="Riazocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="549" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF06804FFC64AAD6794FAD2FF54" box="[1207,1327,187,212]" italics="true" pageId="39" pageNumber="549">Riazocoris</emphasis>
</taxonomicName>
has been designated (
<bibRefCitation id="EFBD4B136804FFC64F3A67F0FE55FF79" author="ICZN" box="[288,424,223,249]" pageId="39" pageNumber="549" refId="ref31005" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
: Art. 13.3) nor it became available as a combined description of new genus and species because none of the taxa was marked as new (
<bibRefCitation id="EFBD4B136804FFC64C86662CFCDEFE9D" author="ICZN" box="[668,803,259,285]" pageId="39" pageNumber="549" refId="ref31005" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
: Art. 13.4). The name
<taxonomicName id="4C2C4D616804FFC64A04662CFB6BFE9C" authorityName="Ahmad &amp; Afzal" authorityYear="1979" box="[1054,1174,259,284]" class="Insecta" family="Pentatomidae" genus="Riazocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="549" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF06804FFC64A04662CFB6BFE9C" box="[1054,1174,259,284]" italics="true" pageId="39" pageNumber="549">Riazocoris</emphasis>
</taxonomicName>
was made available by
<bibRefCitation id="EFBD4B136804FFC64E8D6608FE62FEC1" author="Ahmad, I. &amp; Afzal, M." box="[151,415,295,321]" pageId="39" pageNumber="549" pagination="1 - 14" refId="ref27835" refString="Ahmad, I. &amp; Afzal, M. (1979) Resurrection of the tribe Caystrini Stal (Heteroptera, Pentatomidae, Pentatominae) with description of two new genera from Oriental region. Annotationes Zoologicae et Botanicae, 133, 1 - 14." type="journal article" year="1979">Ahmad &amp; Afzal (1979)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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