treatments-xml/data/03/85/87/038587F46803FFC04E8D6665FD01FC01.xml

<document id="C79EBE80E099E44626AFEA41D8B6EF3C" ID-CLB-Dataset="7213" ID-DOI="10.11646/zootaxa.4958.1.31" ID-GBIF-Dataset="a1df68e3-0e4c-4010-9556-cb15d51a5499" ID-ISSN="1175-5326" ID-Zenodo-Dep="4692369" ID-ZooBank="9E63F8E2-1CCA-4F58-9019-39AE7A3D0640" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1618471284689" checkinUser="plazi" docAuthor="Salini, S. &amp; Kment, Petr" docDate="2021" docId="038587F46803FFC04E8D6665FD01FC01" docLanguage="en" docName="zootaxa.4958.1.31.pdf" docOrigin="Zootaxa 4958 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Agathocles normalis Salini &amp; Kment 2021, comb. nov." docType="treatment" docVersion="7" lastPageNumber="543" masterDocId="FFBCFF8C6823FFE14E1A672FFFFDFF80" masterDocTitle="The genera Agathocles and Surenus (Hemiptera: Heteroptera: Pentatomidae) tribal reassessment, redescription, new synonyms, and description of two new species" masterLastPageNumber="559" masterPageNumber="510" pageNumber="542" updateTime="1698936917073" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="42D307FD4DC01C68BEA33442A22A2E83">The genera Agathocles and Surenus (Hemiptera: Heteroptera: Pentatomidae) tribal reassessment, redescription, new synonyms, and description of two new species</mods:title>
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<mods:namePart id="F0CE92AE9C15EF6F81FE28586FD87C66">Salini, S.</mods:namePart>
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<mods:namePart id="6B7805710F82756939F0D74D977CD211">Kment, Petr</mods:namePart>
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<mods:affiliation id="D6541FD544E685BD3660B69C2C3737B0">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic sigara @ post. cz; https: // orcid. org / 0000 - 0002 - 7026 - 5691</mods:affiliation>
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<taxonomicName id="4C2C4D616803FFC14E8D6665FDADFEE4" authority="(Distant, 1901)" authorityName="Salini &amp; Kment" authorityYear="2021" baseAuthorityName="Distant" baseAuthorityYear="1901" box="[151,592,330,357]" class="Insecta" family="Pentatomidae" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="542" phylum="Arthropoda" rank="species" species="normalis" status="comb. nov.">Agathocles normalis (Distant, 1901)</taxonomicName>
, 
<taxonomicNameLabel id="A26B578B6803FFC14C446664FD19FEE5" box="[606,740,331,357]" pageId="32" pageNumber="542" rank="species">comb. nov.</taxonomicNameLabel>
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<subSubSection id="C33665696803FFC14E8D6640FDE8FE09" box="[151,533,367,393]" pageId="32" pageNumber="542" type="description">
<paragraph id="8B9336E26803FFC14E8D6640FDE8FE09" blockId="32.[151,740,330,393]" box="[151,533,367,393]" pageId="32" pageNumber="542">
(
<figureCitation id="13172A676803FFC14E856640FF15FE09" box="[159,232,367,393]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="32" pageNumber="542">Figs 9</figureCitation>
, 
<figureCitation id="13172A676803FFC14EEE6640FE9BFE09" box="[244,358,367,393]" captionStart="FIGURES 14–22" captionStartId="9.[151,264,1779,1803]" captionTargetBox="[151,1436,181,1755]" captionTargetId="figure-29@9.[151,1436,181,1755]" captionTargetPageId="9" captionText="FIGURES 14–22. External scent efferent system of metathoracic scent gland. 14—Agathocles normalis (Distant, 1901), male, Laos: Phou Pane Mt. (23×). 15—A. joceliae sp. nov., female, Malaysia: Kelantan: Kampong Timor. 16–19—ostiole and peritreme: 16—A. limbatus Stål, 1876 (90×); 17—A. flavipes sp. nov. (100×); 18—A. joceliae sp. nov. (75×); 19—A. normalis (Distant, 1901) (85×). 20—A. joceliae sp. nov., evaporatorium laterally on mesopleuron (75×). 21—A. normalis, metathoracic spiracle (42×). 22—A. limbatus, detail of metathoracic spiracle with reticulate filter processes (320×). Lettering: ev—evaporatorium, mf—median furrow, o—ostiole, pe—peritreme, sms—smooth spot on mesopleuron, sp—spiracle. Scale bars: 1 mm (14, 15, 21), 0.5 mm (16–20), 0.1 mm (22). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692379" httpUri="https://zenodo.org/record/4692379/files/figure.png" pageId="32" pageNumber="542">14, 19, 21</figureCitation>
, 
<figureCitation id="13172A676803FFC14F686640FE2AFE09" box="[370,471,367,393]" captionStart-0="FIGURES 102–107" captionStart-1="FIGURES 108–112" captionStart-2="FIGURES 113–117" captionStart-3="FIGURES 118–124" captionStartId-0="28.[151,264,1763,1787]" captionStartId-1="30.[151,264,1652,1676]" captionStartId-2="31.[151,264,896,920]" captionStartId-3="31.[151,264,1924,1948]" captionTargetBox-0="[161,1424,189,1732]" captionTargetBox-1="[153,1428,193,1628]" captionTargetBox-2="[306,1282,181,873]" captionTargetBox-3="[311,1277,1053,1901]" captionTargetId-0="figure-19@28.[151,1436,181,1740]" captionTargetId-1="figure-19@30.[151,1436,181,1628]" captionTargetId-2="figure-29@31.[306,1282,181,873]" captionTargetId-3="figure-85@31.[311,1277,1053,1901]" captionTargetPageId-0="28" captionTargetPageId-1="30" captionTargetPageId-2="31" captionTargetPageId-3="31" captionText-0="FIGURES 102–107. Penis of Agathocles species. 102–105—A. joceliae sp. nov. (102–104—not inflated, 105—inflated): 102—dorsal view, 103—lateral view, 104—ventral view, 105—apical view. 106–107—A. normalis (Distant, 1901), inflated: 106—dorsal view, 107—ventral view. Lettering: ad—aedeagus, dlc—dorsolateral conjunctival processes, mpp—median penial plate, pt—phallotheca. Scale bars: 1 mm. Orig. P. Kment." captionText-1="FIGURES 108–112. Agathocles normalis (Distant, 1901), habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of Surenus normalis Distant, 1901, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112)." captionText-2="FIGURES 113–117. Male genital capsule of Agathocles normalis (Distant, 1901): 113—caudal view (27×); 114—caudal view, dissected (32×); 115—ventral view (21×); 116—lateral view (27×); 117—superior process of dorsal rim (150×). Lettering: plp—posterolateral projection of genital capsule, sp—superior process of dorsal rim. Scale bars: 1 mm (113–116), 0.1 mm (117). Orig. P. Kment." captionText-3="FIGURES 118–124. Paramere of Agathocles normalis (Distant, 1901), five different views (118–123, 100×) and detail of microsculpture (124, 150×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 1 mm (118–123), 0.25 mm (124). Orig. P. Kment." figureDoi-0="http://doi.org/10.5281/zenodo.4692409" figureDoi-1="http://doi.org/10.5281/zenodo.4692411" figureDoi-2="http://doi.org/10.5281/zenodo.4692413" figureDoi-3="http://doi.org/10.5281/zenodo.4692415" httpUri-0="https://zenodo.org/record/4692409/files/figure.png" httpUri-1="https://zenodo.org/record/4692411/files/figure.png" httpUri-2="https://zenodo.org/record/4692413/files/figure.png" httpUri-3="https://zenodo.org/record/4692415/files/figure.png" pageId="32" pageNumber="542">106–124</figureCitation>
, 
<figureCitation id="13172A676803FFC14FF86640FDF0FE09" box="[482,525,367,393]" captionStart="FIGURE 139" captionStartId="39.[151,250,1769,1793]" captionTargetBox="[151,1435,1007,1745]" captionTargetId="figure-340@39.[151,1435,1007,1745]" captionTargetPageId="39" captionText="FIGURE 139. Distribution of the Agathocles species. T—type locality;?—doubtful record. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692423" httpUri="https://zenodo.org/record/4692423/files/figure.png" pageId="32" pageNumber="542">139</figureCitation>
)
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<treatmentCitation id="0A8D10F36803FFC14E8D6681FDE1FE46" author="Distant, W. L." box="[151,540,430,454]" page="107" pageId="32" pageNumber="542" year="1901">
<taxonomicName id="4C2C4D616803FFC14E8D6681FDE1FE46" authority="Distant, 1901 a: 107" authorityName="Distant. We" authorityPageNumber="107" authorityYear="1901" baseAuthorityName="Distant" baseAuthorityYear="1901" box="[151,540,430,454]" class="Insecta" family="Pentatomidae" genus="Surenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="542" phylum="Arthropoda" rank="species" species="normalis" status="comb. nov.">
<emphasis id="B958EAF06803FFC14E8D6681FEB7FE46" box="[151,330,430,454]" italics="true" pageId="32" pageNumber="542">Surenus normalis</emphasis>
<materialsCitation id="3B443CBF6803FFC14F486681FDE1FE46" ID-GBIF-Occurrence="3083795359" box="[338,540,430,454]" collectionCode="MCSN" country="Myanmar" location="Carin Cheba" pageId="32" pageNumber="542" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<bibRefCitation id="EFBD4B136803FFC14F486681FDE1FE46" author="Distant, W. L." box="[338,540,430,454]" pageId="32" pageNumber="542" pagination="99 - 114" refId="ref29459" refString="Distant, W. L. (1901 a) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant, 1901a: 107</bibRefCitation>
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(original description). 
<typeStatus id="549788406803FFC14D106681FC92FE46" box="[778,879,430,454]" pageId="32" pageNumber="542" type="lectotype">Lectotype</typeStatus>
(here designated) (see 
<figureCitation id="13172A676803FFC14A7A6681FB40FE46" box="[1120,1213,430,454]" captionStart="FIGURES 1–5" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[203,1384,183,1912]" captionTargetId="figure-29@7.[196,1392,179,1917]" captionTargetPageId="7" captionText="FIGURES 1–5. Agathocles limbatus Stål, 1876. 1–2—habitus of male from Laos: Ban Saluey → Phou Pane Mt. (22.2 mm). 3–5—habitus and labels of holotype of A. yunnanensis Zhang &amp; Lin, 1984, syn. nov., female, China: Yunnan: Cang-yuan (scale bar 1 mm). Orig. P. Kment (1–2), D. Rédei (3–5)." figureDoi="http://doi.org/10.5281/zenodo.4692373" httpUri="https://zenodo.org/record/4692373/files/figure.png" pageId="32" pageNumber="542">Figs 1–2</figureCitation>
): 
<specimenCount id="9D2AFD6B6803FFC14ACB6681FB19FE45" box="[1233,1252,430,453]" pageId="32" pageNumber="542" type="female">♀</specimenCount>
, 
<collectingCountry id="F33B76726803FFC14AE96681FAA7FE46" box="[1267,1370,430,454]" name="Myanmar" pageId="32" pageNumber="542">Myanmar</collectingCountry>
: 
<location id="8EF360396803FFC14B7E6681FEF5FE64" LSID="urn:lsid:plazi:treatment:038587F46803FFC04E8D6665FD01FC01:8EF360396803FFC14B7E6681FEF5FE64" country="Myanmar" name="Carin Cheba" pageId="32" pageNumber="542">Carin Cheba</location>
(MCSN).
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<treatmentCitationGroup id="AB3C11CC6803FFC14E8D66C4FEDDFDC0" pageId="32" pageNumber="542">
<treatmentCitation id="0A8D10F36803FFC14E8D66C4FDE4FD83" author="Distant, W. L." box="[151,537,491,515]" page="117" pageId="32" pageNumber="542" year="1902">
<taxonomicName id="4C2C4D616803FFC14E8D66C4FE16FD83" authority=": Distant (1902)" authorityName="Distant" authorityYear="1902" baseAuthorityName="Distant" baseAuthorityYear="1901" box="[151,491,491,515]" class="Insecta" family="Pentatomidae" genus="Surenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="542" phylum="Arthropoda" rank="species" species="normalis" status="comb. nov.">
<emphasis id="B958EAF06803FFC14E8D66C4FEB5FD83" box="[151,328,491,515]" italics="true" pageId="32" pageNumber="542">Surenus normalis</emphasis>
: 
<bibRefCitation id="EFBD4B136803FFC14F4E66C4FE16FD83" author="Distant, W. L." box="[340,491,491,515]" pageId="32" pageNumber="542" pagination="1 - 438" refId="ref29587" refString="Distant, W. L. (1902) Rhynchota. Vol. I. Heteroptera. In: Blanford, W. T. (Ed.), The Fauna of British India Series Including Ceylon and Burma. Taylor &amp; Francis, London, pp. i - xxxvii + 1 - 438." type="book chapter" year="1902">Distant (1902)</bibRefCitation>
</taxonomicName>
: 117
</treatmentCitation>
(description repeated, figures of dorsal habitus and thorax and abdomen in ventral view); 
<treatmentCitation id="0A8D10F36803FFC14EDD6525FE5FFDA2" author="Bergroth, E." box="[199,418,522,546]" page="154" pageId="32" pageNumber="542" year="1908">
<bibRefCitation id="EFBD4B136803FFC14EDD6525FE8CFDA2" author="Bergroth, E." box="[199,369,522,546]" pageId="32" pageNumber="542" pagination="131 - 200" refId="ref28911" refString="Bergroth, E. (1908) Enumeratio Pentatomidarum post Catalogum bruxellensem descriptarum. Memoires de la Societe Entomologique de Belgique, 15, 131 - 200." type="journal article" year="1908">Bergroth (1908)</bibRefCitation>
: 154
</treatmentCitation>
(catalogue); 
<treatmentCitation id="0A8D10F36803FFC14C316525FCFAFDA2" author="Kirkaldy, G. W." box="[555,775,522,546]" page="195" pageId="32" pageNumber="542" year="1909">
<bibRefCitation id="EFBD4B136803FFC14C316525FD28FDA2" author="Kirkaldy, G. W." box="[555,725,522,546]" pageId="32" pageNumber="542" refId="ref31156" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of foodplants and parasites, etc. Prefaced by a discussion on Nomenclature and an analytical table of families. Vol. I. Cimicidae. Felix L. Dames, Berlin, xl + 392 pp. https: // doi. org / 10.5962 / bhl. title. 15205" type="book" year="1909">Kirkaldy (1909)</bibRefCitation>
: 195
</treatmentCitation>
(world catalogue); 
<treatmentCitation id="0A8D10F36803FFC14DCE6525FB11FDA2" author="Memon, N. &amp; Gilbert, F. &amp; Ahmad, I." box="[980,1260,522,546]" page="1153" pageId="32" pageNumber="542" year="2011">
<bibRefCitation id="EFBD4B136803FFC14DCE6525FB52FDA2" author="Memon, N. &amp; Gilbert, F. &amp; Ahmad, I." box="[980,1199,522,546]" pageId="32" pageNumber="542" pagination="1149 - 1169" refId="ref32198" refString="Memon, N., Gilbert, F. &amp; Ahmad, I. (2011) Phylogeny of the South Asian Halyine stink bugs (Hemiptera: Pentatomidae: Halyini) based on morphological characters. Annals of Entomological Society of America, 104 (6), 1149 - 1169. https: // doi. org / 10.1603 / AN 10109" type="journal article" year="2011">
Memon 
<emphasis id="B958EAF06803FFC14A316524FB9DFDA2" box="[1067,1120,522,546]" italics="true" pageId="32" pageNumber="542">et al.</emphasis>
(2011)
</bibRefCitation>
: 1153
</treatmentCitation>
(figure of dorsal habitus).
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</paragraph>
<paragraph id="8B9336E26803FFC14E8D6568FD0EFDDF" blockId="32.[151,1436,430,607]" box="[151,755,583,607]" pageId="32" pageNumber="542">
<treatmentCitationGroup id="AB3C11CC6803FFC14E8D6568FD0EFDDF" box="[151,755,583,607]" pageId="32" pageNumber="542">
<treatmentCitation id="0A8D10F36803FFC14E8D6568FD6EFDDF" author="Arnold, K. &amp; Arnold, M. - L." box="[151,659,583,607]" page="13" pageId="32" pageNumber="542" year="2017">
<taxonomicName id="4C2C4D616803FFC14E8D6568FD8CFDDF" authority="(Distant, 1901)" baseAuthorityName="Distant" baseAuthorityYear="1901" box="[151,625,583,607]" class="Insecta" family="Pentatomidae" genus="Surenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="542" phylum="Arthropoda" rank="species" species="normalis" status="comb. nov.">
<emphasis id="B958EAF06803FFC14E8D6568FE94FDDF" box="[151,361,583,607]" italics="true" pageId="32" pageNumber="542">Surenus cf. normalis</emphasis>
: 
<bibRefCitation id="EFBD4B136803FFC14F6F6568FD8CFDDF" author="Arnold, K. &amp; Arnold, M. - L." box="[373,625,583,607]" pageId="32" pageNumber="542" pagination="9 - 21" refId="ref28681" refString="Arnold, K. &amp; Arnold, M. - L. (2017) 7. Beitrag zur Heteropterenfauna der Inselwelt Indonesiens, speziell der Insel Sumatra so wie weiterer ostlicher bzw. sudostlicher Bereiche unter Einbeziehung der sudostlichen Orientalischen Region (Insecta: Hemiptera: Heteroptera). Edessana, 7, 9 - 21." type="journal article" year="2017">Arnold &amp; Arnold (2017)</bibRefCitation>
</taxonomicName>
: 13
</treatmentCitation>
(record).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33665696803FFC14E8D65A7FD1BFC66" pageId="32" pageNumber="542" type="materials_examined">
<paragraph id="8B9336E26803FFC14E8D65A7FF0FFD46" blockId="32.[151,1437,648,2043]" pageId="32" pageNumber="542">
<materialsCitation id="3B443CBF6803FFC14E8D65A7FF13FD46" ID-GBIF-Occurrence="3083795333" collectionCode="R" country="Myanmar" latitude="19.216667" location="Carin Cheba'" longLatPrecision="1272" longitude="-96.583336" pageId="32" pageNumber="542" specimenCount="1" typeStatus="holotype">
<emphasis id="B958EAF06803FFC14E8D65A7FECAFD22" bold="true" box="[151,311,648,674]" pageId="32" pageNumber="542">
<typeStatus id="549788406803FFC14E8D65A7FF2FFD22" box="[151,210,648,674]" pageId="32" pageNumber="542">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F33B76726803FFC14F5A65A6FE4CFD23" box="[320,433,649,675]" name="Myanmar" pageId="32" pageNumber="542">Myanmar</collectingCountry>
: ‘ 
<location id="8EF360396803FFC14FDB65A7FDA1FD22" LSID="urn:lsid:plazi:treatment:038587F46803FFC04E8D6665FD01FC01:8EF360396803FFC14FDB65A7FDA1FD22" box="[449,604,648,674]" country="Myanmar" latitude="19.216667" longLatPrecision="1272" longitude="-96.583336" name="Carin Cheba'" pageId="32" pageNumber="542">Carin Chebà’</location>
[= Karen Hills, ca. 
<geoCoordinate id="EE1850256803FFC14D5865A6FC5DFD24" box="[834,928,649,676]" degrees="19" direction="north" minutes="13" orientation="latitude" pageId="32" pageNumber="542" precision="925" value="19.216667">19°13′N</geoCoordinate>
<geoCoordinate id="EE1850256803FFC14DB365A6FBF1FD24" box="[937,1036,649,676]" degrees="96" direction="west" minutes="35" orientation="longitude" pageId="32" pageNumber="542" precision="925" value="-96.583336">96°35′W</geoCoordinate>
— 
<collectionCode id="ED3DAE276803FFC14A3165A6FBC2FD24" box="[1067,1087,649,676]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="32" pageNumber="542">R</collectionCode>
. Poggi, pers. comm. in 
<bibRefCitation id="EFBD4B136803FFC14B5565A7FF15FD46" author="Jendek, E." pageId="32" pageNumber="542" pagination="179 - 190" refId="ref31031" refString="Jendek, E. (2003) Studies in the Palaearctic and Oriental Agrilus (Coleoptera, Buprestidae). III. Biologia (Bratislava), 58 (2), 179 - 190." type="journal article" year="2003">Jendek (2003)</bibRefCitation>
]
</materialsCitation>
.
</paragraph>
<paragraph id="8B9336E26803FFC14EDD65FFFBC6FCFA" blockId="32.[151,1437,648,2043]" pageId="32" pageNumber="542">
<emphasis id="B958EAF06803FFC14EDD65FFFE8AFD6A" bold="true" box="[199,375,720,746]" pageId="32" pageNumber="542">Type material.</emphasis>
<typeStatus id="549788406803FFC14F6565FEFE12FD6B" box="[383,495,721,747]" pageId="32" pageNumber="542" type="lectotype">Lectotype</typeStatus>
(here designated) (see 
<figureCitation id="13172A676803FFC14CE365FFFC65FD6A" box="[761,920,720,746]" captionStart="FIGURES 108–112" captionStartId="30.[151,264,1652,1676]" captionTargetBox="[153,1428,193,1628]" captionTargetId="figure-19@30.[151,1436,181,1628]" captionTargetPageId="30" captionText="FIGURES 108–112. Agathocles normalis (Distant, 1901), habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of Surenus normalis Distant, 1901, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112)." figureDoi="http://doi.org/10.5281/zenodo.4692411" httpUri="https://zenodo.org/record/4692411/files/figure.png" pageId="32" pageNumber="542">Figs 110–112</figureCitation>
): 
<specimenCount id="9D2AFD6B6803FFC14DB665FEFC3CFD6B" box="[940,961,721,747]" pageId="32" pageNumber="542" type="female">♀</specimenCount>
, 
<collectingCountry id="F33B76726803FFC14DCA65FEFBBCFD6B" box="[976,1089,721,747]" name="Myanmar" pageId="32" pageNumber="542">Myanmar</collectingCountry>
: ‘Carin Chebà / 
<quantity id="4CD49B076803FFC14AE465FFFA71FD6B" box="[1278,1420,720,747]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="32" pageNumber="542" unit="m" value="1000.0" valueMax="1100.0" valueMin="900.0">900–1100 m</quantity>
/ L.Fea [p] 
<collectionCode id="ED3DAE276803FFC14F1865DAFEEBFC8F" box="[258,278,757,783]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="32" pageNumber="542" type="Museum">V</collectionCode>
.XII 88 [hw, white label with black submarginal frame] // Typus [p in red, white label with red submarginal frame] // 
<taxonomicName id="4C2C4D616803FFC14F1C6437FE9DFCB2" authorityName=": Distant" authorityYear="1902" box="[262,352,792,818]" class="Insecta" family="Pentatomidae" genus="Surenus" kingdom="Plantae" order="Hemiptera" pageId="32" pageNumber="553" phylum="Arthropoda" rank="genus">Surenus</taxonomicName>
/ normalis / Dist. [hw, white label] // 
<taxonomicName id="4C2C4D616803FFC14D3E6437FC83FCB2" authorityName=": Distant" authorityYear="1902" box="[804,894,792,818]" class="Insecta" family="Pentatomidae" genus="Surenus" kingdom="Plantae" order="Hemiptera" pageId="32" pageNumber="553" phylum="Arthropoda" rank="genus">Surenus</taxonomicName>
/ normalis / Typus! Dist. [hw, orange label] // normalis / Dist. [hw, white label]’ (
<collectionCode id="ED3DAE276803FFC14C056413FD88FCD6" box="[543,629,828,854]" country="Italy" httpUri="http://grbio.org/cool/353t-cj9k" name="Museo Civico di Storia Naturale, Verona" pageId="32" pageNumber="542">MCSN</collectionCode>
). We provided the specimen with the following label: ‘ 
<typeStatus id="549788406803FFC14ACC6413FA73FCD6" box="[1238,1422,828,854]" pageId="32" pageNumber="542" type="lectotype">LECTOTYPUS</typeStatus>
/ 
<taxonomicName id="4C2C4D616803FFC14E8D644FFEEDFCFA" authorityName=": Distant" authorityYear="1902" box="[151,272,864,890]" class="Insecta" family="Pentatomidae" genus="Surenus" kingdom="Plantae" order="Hemiptera" pageId="32" pageNumber="542" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF06803FFC14E8D644FFEEDFCFA" box="[151,272,864,890]" italics="true" pageId="32" pageNumber="542">SURENUS</emphasis>
</taxonomicName>
/ 
<emphasis id="B958EAF06803FFC14F3C644FFE50FCFA" box="[294,429,864,890]" italics="true" pageId="32" pageNumber="542">NORMALIS</emphasis>
/ Distant, 1901 / des. Salini &amp; Kment 2020 [p, red label]’.
</paragraph>
<paragraph id="8B9336E26803FFC14EDD64ABFD1BFC66" blockId="32.[151,1437,648,2043]" pageId="32" pageNumber="542">
<emphasis id="B958EAF06803FFC14EDD64ABFDC8FC1E" bold="true" box="[199,565,900,926]" pageId="32" pageNumber="542">
<collectingRegion id="49E8F8006803FFC14EDD64ABFEBAFC1E" box="[199,327,900,926]" pageId="32" pageNumber="542">Additional</collectingRegion>
material examined.
</emphasis>
<materialsCitation id="3B443CBF6803FFC14C2664ABFD1FFC66" ID-GBIF-Occurrence="3083795313" collectingDate="2012-06-01" collectingDateMax="2012-06-24" collectingDateMin="2012-06-01" collectionCode="NMPC" collectorName="Vit Kuban &amp; Lao" country="Laos" determinerName="P. Kment" elevation="1605" location="Phou Pane Mt." longitude="104.01667" municipality="Ban Saluey" pageId="32" pageNumber="542" specimenCount="1" specimenCount-male="1" stateProvince="Houa Phan Province">
<emphasis id="B958EAF06803FFC14C2664ABFC64FC1E" bold="true" box="[572,921,900,926]" pageId="32" pageNumber="542">
<collectingCountry id="F33B76726803FFC14C2664ABFD71FC1E" box="[572,652,900,926]" name="Laos" pageId="32" pageNumber="542">LAOS</collectingCountry>
: 
<collectingRegion id="49E8F8006803FFC14C8264ABFC69FC1E" box="[664,916,900,926]" pageId="32" pageNumber="542">Houa Phan Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BF7AC986803FFC14DBA64AAFBDCFC1E" box="[928,1057,900,927]" pageId="32" pageNumber="542">Ban Saluey</collectingMunicipality>
→ 
<location id="8EF360396803FFC14A5164ABFB09FC1F" LSID="urn:lsid:plazi:treatment:038587F46803FFC04E8D6665FD01FC01:8EF360396803FFC14A5164ABFB09FC1F" box="[1099,1268,900,927]" country="Laos" longitude="104.01667" municipality="Ban Saluey" name="Phou Pane Mt." pageId="32" pageNumber="542" stateProvince="Houa Phan Province">Phou Pane Mt.</location>
, 
<geoCoordinate id="EE1850256803FFC14B1A64AAFABCFC20" box="[1280,1345,901,928]" pageId="32" pageNumber="542">20°12</geoCoordinate>
–13.5′N 103°59.5′– 
<geoCoordinate id="EE1850256803FFC14F0E6486FE81FC44" box="[276,380,937,964]" degrees="104" direction="east" minutes="01" orientation="longitude" pageId="32" pageNumber="542" precision="925" value="104.01667">104°01′E</geoCoordinate>
, 
<elevation id="0001D1D16803FFC14F906487FD9BFC42" box="[394,614,936,963]" metricMagnitude="3" metricUnit="m" metricValue="1.605" metricValueMax="1.8699999999999999" metricValueMin="1.34" pageId="32" pageNumber="542" unit="m" value="1605.0" valueMax="1870.0" valueMin="1340.0">
<quantity id="4CD49B076803FFC14F906487FDDAFC43" box="[394,551,936,963]" metricMagnitude="3" metricUnit="m" metricValue="1.605" metricValueMax="1.8699999999999999" metricValueMin="1.34" pageId="32" pageNumber="542" unit="m" value="1605.0" valueMax="1870.0" valueMin="1340.0">1340–1870 m</quantity>
a.s.l.
</elevation>
, primary mountain forest, individual collecting, 
<date id="FF9210226803FFC14A886487FAD2FC42" box="[1170,1327,936,962]" pageId="32" pageNumber="542" value="2012-06-01" valueMax="2012-06-24" valueMin="2012-06-01">
<collectingDate id="EFD6E9CA6803FFC14A886487FAD2FC42" box="[1170,1327,936,962]" pageId="32" pageNumber="542" value="2012-06-01" valueMax="2012-06-24" valueMin="2012-06-01">1.–24.vi.2012</collectingDate>
</date>
, 
<specimenCount id="9D2AFD6B6803FFC14B276487FA96FC43" box="[1341,1387,936,963]" pageId="32" pageNumber="542" type="male">1 ♂</specimenCount>
, 
<collectorName id="26D953346803FFC14B626487FF1FFC66" pageId="32" pageNumber="542">Vít Kubáň</collectorName>
&amp; 
<collectorName id="26D953346803FFC14F1C64E2FECFFC67" box="[262,306,973,999]" pageId="32" pageNumber="542">Lao</collectorName>
collectors lgt., 
<determinerName id="7E4FDA266803FFC14FC564E2FDB8FC67" box="[479,581,973,999]" pageId="32" pageNumber="542">P. Kment</determinerName>
det. (
<collectionCode id="ED3DAE276803FFC14C9E64E3FD27FC66" box="[644,730,972,998]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="32" pageNumber="542">NMPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33665696803FFC04EDD64DFFE7AFE75" lastPageId="33" lastPageNumber="543" pageId="32" pageNumber="542" type="description">
<paragraph id="8B9336E26803FFC14EDD64DFFC8BFB3E" blockId="32.[151,1437,648,2043]" pageId="32" pageNumber="542">
<emphasis id="B958EAF06803FFC14EDD64DFFE90FB8A" bold="true" box="[199,365,1008,1034]" pageId="32" pageNumber="542">Redescription</emphasis>
(male). 
<emphasis id="B958EAF06803FFC14FCF64C0FD9CFB8A" bold="true" box="[469,609,1007,1034]" italics="true" pageId="32" pageNumber="542">Colouration</emphasis>
(
<figureCitation id="13172A676803FFC14C6E64DFFCEFFB8A" box="[628,786,1008,1034]" captionStart="FIGURES 108–112" captionStartId="30.[151,264,1652,1676]" captionTargetBox="[153,1428,193,1628]" captionTargetId="figure-19@30.[151,1436,181,1628]" captionTargetPageId="30" captionText="FIGURES 108–112. Agathocles normalis (Distant, 1901), habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of Surenus normalis Distant, 1901, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112)." figureDoi="http://doi.org/10.5281/zenodo.4692411" httpUri="https://zenodo.org/record/4692411/files/figure.png" pageId="32" pageNumber="542">Figs 108–111</figureCitation>
). Body black, except clavus, corium, acetabula, genital capsule posteroventrally, antennomeres I–III, and legs blackish brown; labium and distiflagellum (IV) pale brown. Apices of humeri, apex of scutellum, propleuron sublaterally, lateral margin of abdominal ventrite II, antero- and posterolateral angles of connexival segments III to VII, and exocorium with small to very small, yellowish ochraceous callosities. Peritremal surface and small round fields surrounding abdominal trichobothria black. Membrane dark brown, translucent. Remaining parts of genital capsule yellowish.
</paragraph>
<paragraph id="8B9336E26803FFC14EDD63E7FC0DFAF3" blockId="32.[151,1437,648,2043]" pageId="32" pageNumber="542">
<emphasis id="B958EAF06803FFC14EDD63E7FE09FB62" bold="true" box="[199,500,1223,1250]" italics="true" pageId="32" pageNumber="542">Integument and vestiture.</emphasis>
Body shining, clavus, corium and evaporatorium matt. Body above covered with dense, coarse, black punctures, including apex of scutellum; only endocorium with punctures distinctly smaller and sparser than on exocorium; disc of pronotum (posteromedially) and scutellum (anteriorly) with prominent transverse wrinkles (
<figureCitation id="13172A676803FFC14F1C621BFE95FACE" box="[262,360,1332,1358]" captionStart="FIGURES 108–112" captionStartId="30.[151,264,1652,1676]" captionTargetBox="[153,1428,193,1628]" captionTargetId="figure-19@30.[151,1436,181,1628]" captionTargetPageId="30" captionText="FIGURES 108–112. Agathocles normalis (Distant, 1901), habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of Surenus normalis Distant, 1901, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112)." figureDoi="http://doi.org/10.5281/zenodo.4692411" httpUri="https://zenodo.org/record/4692411/files/figure.png" pageId="32" pageNumber="542">Fig. 108</figureCitation>
); head underside and thoracic sterna with dense, coarse punctures; surface of abdominal sternites smooth with fine punctures. Body mostly glabrous (see generic description).
</paragraph>
<paragraph id="8B9336E26803FFC14EDD6253FDBDF9EF" blockId="32.[151,1437,648,2043]" pageId="32" pageNumber="542">
<emphasis id="B958EAF06803FFC14EDD6253FE7CFA16" box="[199,385,1404,1430]" italics="true" pageId="32" pageNumber="542">
<emphasis id="B958EAF06803FFC14EDD6253FEC6FA16" bold="true" box="[199,315,1404,1430]" italics="true" pageId="32" pageNumber="542">Structure.</emphasis>
Head
</emphasis>
(
<figureCitation id="13172A676803FFC14F886253FE25FA16" box="[402,472,1404,1430]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="32" pageNumber="542">Fig. 9</figureCitation>
). Head slightly shorter than wide (length: width ratio 0.88). Mandibular plates with lateral margins distinctly sinuate in front of eyes above the small antenniferous tubercle, parallel-sided about their midlength, and regularly rounded anteriorly; mandibular plates not meeting anteriorly, apex of clypeus free, inner margins of apical incisure parallel. Antennomeres from shortest to longest: I &lt;IIa &lt;IV &lt;IIb &lt;III. Scape (I) reaching about apex of head, basipedicellite (IIa) distinctly (1.54×) longer than scape and shorter than IIb (length of IIa: IIb ratio: 1: 1.70). Labiomeres from shortest to longest (n = 1): I = IV (
<quantity id="4CD49B076803FFC14DA5611FFBDBF9CB" box="[959,1062,1584,1611]" metricMagnitude="-3" metricUnit="m" metricValue="1.47" pageId="32" pageNumber="542" unit="mm" value="1.47">1.47 mm</quantity>
) &lt;III (
<quantity id="4CD49B076803FFC14A62611FFB23F9CB" box="[1144,1246,1584,1611]" metricMagnitude="-3" metricUnit="m" metricValue="2.11" pageId="32" pageNumber="542" unit="mm" value="2.11">2.11 mm</quantity>
) &lt;II (
<quantity id="4CD49B076803FFC14B3C611FFA6DF9CB" box="[1318,1424,1584,1611]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="32" pageNumber="542" unit="mm" value="2.3">2.30 mm</quantity>
). Labium reaching between metacoxae.
</paragraph>
<paragraph id="8B9336E26803FFC14EDD6156FDEAF95B" blockId="32.[151,1437,648,2043]" pageId="32" pageNumber="542">
<emphasis id="B958EAF06803FFC14EDD6156FECBF912" box="[199,310,1657,1682]" italics="true" pageId="32" pageNumber="542">Pronotum</emphasis>
(
<figureCitation id="13172A676803FFC14F5E6157FE1CF912" box="[324,481,1656,1682]" captionStart="FIGURES 108–112" captionStartId="30.[151,264,1652,1676]" captionTargetBox="[153,1428,193,1628]" captionTargetId="figure-19@30.[151,1436,181,1628]" captionTargetPageId="30" captionText="FIGURES 108–112. Agathocles normalis (Distant, 1901), habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of Surenus normalis Distant, 1901, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112)." figureDoi="http://doi.org/10.5281/zenodo.4692411" httpUri="https://zenodo.org/record/4692411/files/figure.png" pageId="32" pageNumber="542">Figs 108, 110</figureCitation>
). Lateral tooth on anterolateral angles small, directed laterad (
<figureCitation id="13172A676803FFC14A8B6157FB28F912" box="[1169,1237,1656,1682]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="32" pageNumber="542">Fig. 9</figureCitation>
). Lateral pronotal margin anteriorly only minutely crenulate. Humeri rectangular, slightly surpassing laterad over base of corium. Posterior margin slightly concave.
</paragraph>
<paragraph id="8B9336E26803FFC14EDD61CBFE9AF8A2" blockId="32.[151,1437,648,2043]" pageId="32" pageNumber="542">
<emphasis id="B958EAF06803FFC14EDD61CBFEC8F97E" box="[199,309,1764,1790]" italics="true" pageId="32" pageNumber="542">Scutellum</emphasis>
(
<figureCitation id="13172A676803FFC14F5F61CBFE19F97E" box="[325,484,1764,1790]" captionStart="FIGURES 108–112" captionStartId="30.[151,264,1652,1676]" captionTargetBox="[153,1428,193,1628]" captionTargetId="figure-19@30.[151,1436,181,1628]" captionTargetPageId="30" captionText="FIGURES 108–112. Agathocles normalis (Distant, 1901), habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of Surenus normalis Distant, 1901, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112)." figureDoi="http://doi.org/10.5281/zenodo.4692411" httpUri="https://zenodo.org/record/4692411/files/figure.png" pageId="32" pageNumber="542">Figs 108, 110</figureCitation>
). Disc of scutellum convex in prefrenal portion, only its postfrenal portion flat; apex narrowly rounded.
</paragraph>
<paragraph id="8B9336E26803FFC14EDD6003FBBDF8C6" blockId="32.[151,1437,648,2043]" box="[199,1088,1836,1863]" pageId="32" pageNumber="542">
<emphasis id="B958EAF06803FFC14EDD6003FEC0F8C6" box="[199,317,1836,1862]" italics="true" pageId="32" pageNumber="542">Hemelytra</emphasis>
. Costal margin of corium only slightly insinuated basally (
<figureCitation id="13172A676803FFC14DCB6003FBCEF8C6" box="[977,1075,1836,1862]" captionStart="FIGURES 108–112" captionStartId="30.[151,264,1652,1676]" captionTargetBox="[153,1428,193,1628]" captionTargetId="figure-19@30.[151,1436,181,1628]" captionTargetPageId="30" captionText="FIGURES 108–112. Agathocles normalis (Distant, 1901), habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of Surenus normalis Distant, 1901, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112)." figureDoi="http://doi.org/10.5281/zenodo.4692411" httpUri="https://zenodo.org/record/4692411/files/figure.png" pageId="32" pageNumber="542">Fig. 108</figureCitation>
).
</paragraph>
<paragraph id="8B9336E26803FFC14EDD607FFDFBF80E" blockId="32.[151,1437,648,2043]" pageId="32" pageNumber="542">
<emphasis id="B958EAF06803FFC14EDD607FFEE5F8EA" box="[199,280,1872,1898]" italics="true" pageId="32" pageNumber="542">Thorax</emphasis>
. Ostiole large, oval, opening posteriad; median furrow deep, reaching nearly apex of peritreme (
<figureCitation id="13172A676803FFC14B75607FFF4BF80E" captionStart="FIGURES 14–22" captionStartId="9.[151,264,1779,1803]" captionTargetBox="[151,1436,181,1755]" captionTargetId="figure-29@9.[151,1436,181,1755]" captionTargetPageId="9" captionText="FIGURES 14–22. External scent efferent system of metathoracic scent gland. 14—Agathocles normalis (Distant, 1901), male, Laos: Phou Pane Mt. (23×). 15—A. joceliae sp. nov., female, Malaysia: Kelantan: Kampong Timor. 16–19—ostiole and peritreme: 16—A. limbatus Stål, 1876 (90×); 17—A. flavipes sp. nov. (100×); 18—A. joceliae sp. nov. (75×); 19—A. normalis (Distant, 1901) (85×). 20—A. joceliae sp. nov., evaporatorium laterally on mesopleuron (75×). 21—A. normalis, metathoracic spiracle (42×). 22—A. limbatus, detail of metathoracic spiracle with reticulate filter processes (320×). Lettering: ev—evaporatorium, mf—median furrow, o—ostiole, pe—peritreme, sms—smooth spot on mesopleuron, sp—spiracle. Scale bars: 1 mm (14, 15, 21), 0.5 mm (16–20), 0.1 mm (22). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692379" httpUri="https://zenodo.org/record/4692379/files/figure.png" pageId="32" pageNumber="542">Fig. 19</figureCitation>
). Metasternum medium wide.
</paragraph>
<paragraph id="8B9336E26803FFC04EDD60B7FBEFFEE6" blockId="32.[151,1437,648,2043]" lastBlockId="33.[151,1437,151,897]" lastPageId="33" lastPageNumber="543" pageId="32" pageNumber="542">
<emphasis id="B958EAF06803FFC14EDD60B7FE90F832" box="[199,365,1944,1970]" italics="true" pageId="32" pageNumber="542">Male genitalia</emphasis>
. Genital capsule (
<figureCitation id="13172A676803FFC14C2160B7FD2AF833" box="[571,727,1944,1971]" captionStart="FIGURES 113–117" captionStartId="31.[151,264,896,920]" captionTargetBox="[306,1282,181,873]" captionTargetId="figure-29@31.[306,1282,181,873]" captionTargetPageId="31" captionText="FIGURES 113–117. Male genital capsule of Agathocles normalis (Distant, 1901): 113—caudal view (27×); 114—caudal view, dissected (32×); 115—ventral view (21×); 116—lateral view (27×); 117—superior process of dorsal rim (150×). Lettering: plp—posterolateral projection of genital capsule, sp—superior process of dorsal rim. Scale bars: 1 mm (113–116), 0.1 mm (117). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692413" httpUri="https://zenodo.org/record/4692413/files/figure.png" pageId="32" pageNumber="542">Figs 113–116</figureCitation>
) nearly quadrangular in dorsal/ventral view, longer than wide. Ventral rim deeply incised medially (
<figureCitation id="13172A676803FFC14C346093FD70F857" box="[558,653,1980,2007]" captionStart="FIGURES 113–117" captionStartId="31.[151,264,896,920]" captionTargetBox="[306,1282,181,873]" captionTargetId="figure-29@31.[306,1282,181,873]" captionTargetPageId="31" captionText="FIGURES 113–117. Male genital capsule of Agathocles normalis (Distant, 1901): 113—caudal view (27×); 114—caudal view, dissected (32×); 115—ventral view (21×); 116—lateral view (27×); 117—superior process of dorsal rim (150×). Lettering: plp—posterolateral projection of genital capsule, sp—superior process of dorsal rim. Scale bars: 1 mm (113–116), 0.1 mm (117). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692413" httpUri="https://zenodo.org/record/4692413/files/figure.png" pageId="32" pageNumber="542">Fig. 115</figureCitation>
); ventral rim infolding wide, medially with crescent-shaped depression (
<figureCitation id="13172A676803FFC14E8560CFFEBCF87A" box="[159,321,2016,2042]" captionStart="FIGURES 113–117" captionStartId="31.[151,264,896,920]" captionTargetBox="[306,1282,181,873]" captionTargetId="figure-29@31.[306,1282,181,873]" captionTargetPageId="31" captionText="FIGURES 113–117. Male genital capsule of Agathocles normalis (Distant, 1901): 113—caudal view (27×); 114—caudal view, dissected (32×); 115—ventral view (21×); 116—lateral view (27×); 117—superior process of dorsal rim (150×). Lettering: plp—posterolateral projection of genital capsule, sp—superior process of dorsal rim. Scale bars: 1 mm (113–116), 0.1 mm (117). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692413" httpUri="https://zenodo.org/record/4692413/files/figure.png" pageId="32" pageNumber="542">Figs 113–114</figureCitation>
). Superior processes of dorsal rim present, small, subtriangular (
<figureCitation id="13172A676803FFC14A2360CFFAE9F87A" box="[1081,1300,2016,2042]" captionStart="FIGURES 113–117" captionStartId="31.[151,264,896,920]" captionTargetBox="[306,1282,181,873]" captionTargetId="figure-29@31.[306,1282,181,873]" captionTargetPageId="31" captionText="FIGURES 113–117. Male genital capsule of Agathocles normalis (Distant, 1901): 113—caudal view (27×); 114—caudal view, dissected (32×); 115—ventral view (21×); 116—lateral view (27×); 117—superior process of dorsal rim (150×). Lettering: plp—posterolateral projection of genital capsule, sp—superior process of dorsal rim. Scale bars: 1 mm (113–116), 0.1 mm (117). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692413" httpUri="https://zenodo.org/record/4692413/files/figure.png" pageId="32" pageNumber="542">Figs 113–114, 117</figureCitation>
), trapezoid, laterally constricted about midlength (
<figureCitation id="13172A676802FFC04C5C67B8FD54FF31" box="[582,681,151,177]" captionStart="FIGURES 113–117" captionStartId="31.[151,264,896,920]" captionTargetBox="[306,1282,181,873]" captionTargetId="figure-29@31.[306,1282,181,873]" captionTargetPageId="31" captionText="FIGURES 113–117. Male genital capsule of Agathocles normalis (Distant, 1901): 113—caudal view (27×); 114—caudal view, dissected (32×); 115—ventral view (21×); 116—lateral view (27×); 117—superior process of dorsal rim (150×). Lettering: plp—posterolateral projection of genital capsule, sp—superior process of dorsal rim. Scale bars: 1 mm (113–116), 0.1 mm (117). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692413" httpUri="https://zenodo.org/record/4692413/files/figure.png" pageId="33" pageNumber="543">Fig. 113</figureCitation>
). Parameral socket open (
<figureCitation id="13172A676802FFC04DD467B8FBCDFF31" box="[974,1072,151,177]" captionStart="FIGURES 113–117" captionStartId="31.[151,264,896,920]" captionTargetBox="[306,1282,181,873]" captionTargetId="figure-29@31.[306,1282,181,873]" captionTargetPageId="31" captionText="FIGURES 113–117. Male genital capsule of Agathocles normalis (Distant, 1901): 113—caudal view (27×); 114—caudal view, dissected (32×); 115—ventral view (21×); 116—lateral view (27×); 117—superior process of dorsal rim (150×). Lettering: plp—posterolateral projection of genital capsule, sp—superior process of dorsal rim. Scale bars: 1 mm (113–116), 0.1 mm (117). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692413" httpUri="https://zenodo.org/record/4692413/files/figure.png" pageId="33" pageNumber="543">Fig. 114</figureCitation>
). Paramere (
<figureCitation id="13172A676802FFC04AA467B8FAA6FF31" box="[1214,1371,151,177]" captionStart="FIGURES 118–124" captionStartId="31.[151,264,1924,1948]" captionTargetBox="[311,1277,1053,1901]" captionTargetId="figure-85@31.[311,1277,1053,1901]" captionTargetPageId="31" captionText="FIGURES 118–124. Paramere of Agathocles normalis (Distant, 1901), five different views (118–123, 100×) and detail of microsculpture (124, 150×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 1 mm (118–123), 0.25 mm (124). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692415" httpUri="https://zenodo.org/record/4692415/files/figure.png" pageId="33" pageNumber="543">Figs 118–124</figureCitation>
) with basal process with a foliaceous, roughly semicircular lobe positioned rather vertically, bearing row of many long setae (
<figureCitation id="13172A676802FFC04EC567F0FE0CFF79" box="[223,497,223,249]" captionStart="FIGURES 118–124" captionStartId="31.[151,264,1924,1948]" captionTargetBox="[311,1277,1053,1901]" captionTargetId="figure-85@31.[311,1277,1053,1901]" captionTargetPageId="31" captionText="FIGURES 118–124. Paramere of Agathocles normalis (Distant, 1901), five different views (118–123, 100×) and detail of microsculpture (124, 150×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 1 mm (118–123), 0.25 mm (124). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692415" httpUri="https://zenodo.org/record/4692415/files/figure.png" pageId="33" pageNumber="543">Figs 118, 121–122, 124</figureCitation>
); apical process hook-shaped, apically obtuse (
<figureCitation id="13172A676802FFC04A1D67F0FB1CFF79" box="[1031,1249,223,249]" captionStart="FIGURES 118–124" captionStartId="31.[151,264,1924,1948]" captionTargetBox="[311,1277,1053,1901]" captionTargetId="figure-85@31.[311,1277,1053,1901]" captionTargetPageId="31" captionText="FIGURES 118–124. Paramere of Agathocles normalis (Distant, 1901), five different views (118–123, 100×) and detail of microsculpture (124, 150×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 1 mm (118–123), 0.25 mm (124). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692415" httpUri="https://zenodo.org/record/4692415/files/figure.png" pageId="33" pageNumber="543">Figs 118, 122, 124</figureCitation>
); mesal angle of stem without additional projection (
<figureCitation id="13172A676802FFC04C2A662CFCF1FE9D" box="[560,780,259,285]" captionStart="FIGURES 118–124" captionStartId="31.[151,264,1924,1948]" captionTargetBox="[311,1277,1053,1901]" captionTargetId="figure-85@31.[311,1277,1053,1901]" captionTargetPageId="31" captionText="FIGURES 118–124. Paramere of Agathocles normalis (Distant, 1901), five different views (118–123, 100×) and detail of microsculpture (124, 150×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 1 mm (118–123), 0.25 mm (124). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692415" httpUri="https://zenodo.org/record/4692415/files/figure.png" pageId="33" pageNumber="543">Figs 118, 122, 124</figureCitation>
). Phallus (
<figureCitation id="13172A676802FFC04D9F662CFBDEFE9E" box="[901,1059,259,286]" captionStart="FIGURES 102–107" captionStartId="28.[151,264,1763,1787]" captionTargetBox="[161,1424,189,1732]" captionTargetId="figure-19@28.[151,1436,181,1740]" captionTargetPageId="28" captionText="FIGURES 102–107. Penis of Agathocles species. 102–105—A. joceliae sp. nov. (102–104—not inflated, 105—inflated): 102—dorsal view, 103—lateral view, 104—ventral view, 105—apical view. 106–107—A. normalis (Distant, 1901), inflated: 106—dorsal view, 107—ventral view. Lettering: ad—aedeagus, dlc—dorsolateral conjunctival processes, mpp—median penial plate, pt—phallotheca. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692409" httpUri="https://zenodo.org/record/4692409/files/figure.png" pageId="33" pageNumber="543">Figs 106–107</figureCitation>
) with pair of dorsal conjunctival processes transversely placed and ending in sclerotised processes, median penial plates (mpp) fused basally, outer margins broadly and thickly sclerotized, encircling short, curved aedeagus (ad).
</paragraph>
<paragraph id="8B9336E26802FFC04EDD6640FDF7FE09" blockId="33.[151,1437,151,897]" box="[199,522,367,393]" pageId="33" pageNumber="543">
<emphasis id="B958EAF06802FFC04EDD6640FE7BFE09" box="[199,390,367,393]" italics="true" pageId="33" pageNumber="543">Female genitalia</emphasis>
. Unknown.
</paragraph>
<paragraph id="8B9336E26802FFC04EDD66BCFAE9FE2D" blockId="33.[151,1437,151,897]" box="[199,1300,403,429]" pageId="33" pageNumber="543">
<emphasis id="B958EAF06802FFC04EDD66BCFE94FE2C" box="[199,361,403,428]" italics="true" pageId="33" pageNumber="543">Measurements</emphasis>
(mm). See 
<tableCitation id="C6AE03596802FFC04FF666BCFDBCFE2D" box="[492,577,403,429]" captionStart="TABLE 2" captionStartId="34.[151,239,507,533]" captionText="TABLE 2. Measurements of the Agathocles species presented as median, with minimum and maximum values given in parentheses." pageId="33" pageNumber="543">Table 2</tableCitation>
. 
<bibRefCitation id="EFBD4B136802FFC04C5666BCFD06FE2D" author="Distant, W. L." box="[588,763,403,429]" pageId="33" pageNumber="543" pagination="99 - 114" refId="ref29459" refString="Distant, W. L. (1901 a) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant (1901a)</bibRefCitation>
give the length of female 
<typeStatus id="549788406802FFC04A3B66BCFB7EFE2D" box="[1057,1155,403,429]" pageId="33" pageNumber="543" type="holotype">holotype</typeStatus>
as 20.0 mm.
</paragraph>
<paragraph id="8B9336E26802FFC04EDD6699FE7AFE75" blockId="33.[151,1437,151,897]" pageId="33" pageNumber="543">
<emphasis id="B958EAF06802FFC04EDD6699FEC2FE50" bold="true" box="[199,319,438,464]" pageId="33" pageNumber="543">Variation.</emphasis>
The 
<typeStatus id="549788406802FFC04F6D6698FE23FE51" box="[375,478,439,465]" pageId="33" pageNumber="543" type="lectotype">lectotype</typeStatus>
female differs from the examined male by presence of small yellowish callosities also on corium (
<figureCitation id="13172A676802FFC04F0366F4FE87FE75" box="[281,378,475,501]" captionStart="FIGURES 108–112" captionStartId="30.[151,264,1652,1676]" captionTargetBox="[153,1428,193,1628]" captionTargetId="figure-19@30.[151,1436,181,1628]" captionTargetPageId="30" captionText="FIGURES 108–112. Agathocles normalis (Distant, 1901), habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of Surenus normalis Distant, 1901, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112)." figureDoi="http://doi.org/10.5281/zenodo.4692411" httpUri="https://zenodo.org/record/4692411/files/figure.png" pageId="33" pageNumber="543">Fig. 110</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33665696802FFC04EDD66D1FC8BFDE1" pageId="33" pageNumber="543" type="diagnosis">
<paragraph id="8B9336E26802FFC04EDD66D1FC8BFDE1" blockId="33.[151,1437,151,897]" pageId="33" pageNumber="543">
<emphasis id="B958EAF06802FFC04EDD66D1FE2CFD98" bold="true" box="[199,465,510,536]" pageId="33" pageNumber="543">Differential diagnosis.</emphasis>
See key and 
<tableCitation id="C6AE03596802FFC04C6E66D0FD31FD99" box="[628,716,511,537]" captionStart="TABLE 3" captionStartId="40.[151,239,479,505]" captionText="TABLE 3. Summary of main morphological differences of Agathocles species." pageId="33" pageNumber="543">Table 3</tableCitation>
. This species could be recognized by combination of apically narrowly free clypeus, scape about reaching apex of head, and blackish-brown colouration with less conspicuous whitish spot with concolorous punctures at the apex of scutellum.
</paragraph>
</subSubSection>
<subSubSection id="C33665696802FFC04EDD6545FD01FC01" pageId="33" pageNumber="543" type="distribution">
<paragraph id="8B9336E26802FFC04EDD6545FC88FD29" blockId="33.[151,1437,151,897]" pageId="33" pageNumber="543">
<emphasis id="B958EAF06802FFC04EDD6545FEA4FD04" bold="true" box="[199,345,618,644]" pageId="33" pageNumber="543">Distribution</emphasis>
(
<figureCitation id="13172A676802FFC04F766544FE2FFD05" box="[364,466,619,645]" captionStart="FIGURE 139" captionStartId="39.[151,250,1769,1793]" captionTargetBox="[151,1435,1007,1745]" captionTargetId="figure-340@39.[151,1435,1007,1745]" captionTargetPageId="39" captionText="FIGURE 139. Distribution of the Agathocles species. T—type locality;?—doubtful record. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692423" httpUri="https://zenodo.org/record/4692423/files/figure.png" pageId="33" pageNumber="543">Fig. 139</figureCitation>
). 
<collectingCountry id="F33B76726802FFC04FF26544FDE3FD05" box="[488,542,619,645]" name="Laos" pageId="33" pageNumber="543">Laos</collectingCountry>
(new record), 
<collectingCountry id="F33B76726802FFC04CD46544FCC6FD05" box="[718,827,619,645]" name="Myanmar" pageId="33" pageNumber="543">Myanmar</collectingCountry>
(
<bibRefCitation id="EFBD4B136802FFC04D566544FC0DFD05" author="Distant, W. L." box="[844,1008,619,645]" pageId="33" pageNumber="543" pagination="99 - 114" refId="ref29459" refString="Distant, W. L. (1901 a) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant 1901a</bibRefCitation>
, 
<bibRefCitation id="EFBD4B136802FFC04DE56544FBC6FD05" author="Distant, W. L." box="[1023,1083,619,645]" pageId="33" pageNumber="543" pagination="1 - 438" refId="ref29587" refString="Distant, W. L. (1902) Rhynchota. Vol. I. Heteroptera. In: Blanford, W. T. (Ed.), The Fauna of British India Series Including Ceylon and Burma. Taylor &amp; Francis, London, pp. i - xxxvii + 1 - 438." type="book chapter" year="1902">1902</bibRefCitation>
). The record from 
<collectingCountry id="F33B76726802FFC04B006544FAA5FD05" box="[1306,1368,619,645]" name="India" pageId="33" pageNumber="543">India</collectingCountry>
: 
<collectingRegion id="49E8F8006802FFC04B7F6544FF16FD29" country="India" name="West Bengal" pageId="33" pageNumber="543">West Bengal</collectingRegion>
: Darjiling (
<bibRefCitation id="EFBD4B136802FFC04F7365A0FD90FD29" author="Arnold, K. &amp; Arnold, M. - L." box="[361,621,655,681]" pageId="33" pageNumber="543" pagination="9 - 21" refId="ref28681" refString="Arnold, K. &amp; Arnold, M. - L. (2017) 7. Beitrag zur Heteropterenfauna der Inselwelt Indonesiens, speziell der Insel Sumatra so wie weiterer ostlicher bzw. sudostlicher Bereiche unter Einbeziehung der sudostlichen Orientalischen Region (Insecta: Hemiptera: Heteroptera). Edessana, 7, 9 - 21." type="journal article" year="2017">Arnold &amp; Arnold 2017</bibRefCitation>
) requires confirmation.
</paragraph>
<paragraph id="8B9336E26802FFC04EDD659CFD01FC01" blockId="33.[151,1437,151,897]" pageId="33" pageNumber="543">
<emphasis id="B958EAF06802FFC04EDD659CFE8AFD4D" bold="true" box="[199,375,691,717]" pageId="33" pageNumber="543">Nomenclature.</emphasis>
<bibRefCitation id="EFBD4B136802FFC04F9A659CFDCDFD4D" author="Distant, W. L." box="[384,560,691,717]" pageId="33" pageNumber="543" pagination="99 - 114" refId="ref29459" refString="Distant, W. L. (1901 a) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant (1901a)</bibRefCitation>
did not indicate the number and sex of the specimen(s) he studied, though a single measurement given suggests only one specimen. The ventral illustration of abdomen in 
<bibRefCitation id="EFBD4B136802FFC04AB365F8FAB7FD71" author="Distant, W. L." box="[1193,1354,727,753]" pageId="33" pageNumber="543" pagination="1 - 438" refId="ref29587" refString="Distant, W. L. (1902) Rhynchota. Vol. I. Heteroptera. In: Blanford, W. T. (Ed.), The Fauna of British India Series Including Ceylon and Burma. Taylor &amp; Francis, London, pp. i - xxxvii + 1 - 438." type="book chapter" year="1902">Distant (1902)</bibRefCitation>
clearly shows a female. According to Maria Tavano (pers. comm.), there is only a single female labelled as ‘typus’ in the collection of the Museum of Natural History in Genoa, most probably the only specimen Distant studied. According to the Recommendation 73F of the 
<bibRefCitation id="EFBD4B136802FFC04C04646CFD4DFCDD" author="ICZN" box="[542,688,835,861]" pageId="33" pageNumber="543" refId="ref31005" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN (1999)</bibRefCitation>
we designate here the single type specimen present in MCSN (
<figureCitation id="13172A676802FFC04B70646CFF04FC01" captionStart="FIGURES 108–112" captionStartId="30.[151,264,1652,1676]" captionTargetBox="[153,1428,193,1628]" captionTargetId="figure-19@30.[151,1436,181,1628]" captionTargetPageId="30" captionText="FIGURES 108–112. Agathocles normalis (Distant, 1901), habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of Surenus normalis Distant, 1901, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112)." figureDoi="http://doi.org/10.5281/zenodo.4692411" httpUri="https://zenodo.org/record/4692411/files/figure.png" pageId="33" pageNumber="543">Figs 110–112</figureCitation>
) as 
<typeStatus id="549788406802FFC04F3C6448FE70FC01" box="[294,397,871,897]" pageId="33" pageNumber="543" type="lectotype">lectotype</typeStatus>
to fix the identity of the species.
</paragraph>
</subSubSection>
</treatment>
</document>