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<document id="75EC1D93DEBBAD262EACEC05CE61D0A2" ID-DOI="10.11646/zootaxa.3693.2.6" ID-GBIF-Dataset="d0232cdf-3ee1-4dcf-983a-9b3898741072" ID-ISSN="1175-5326" ID-Zenodo-Dep="222022" ID-ZooBank="0EB0518C-BDFC-4EFC-9FEE-35B535579E90" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460654806266" checkinUser="plazi" docAuthor="Loeb, Marina Vianna &amp; Alcântara, Ayda Vera" docDate="2013" docId="038587DC5E59AC41FF23FBB6FDA47B9E" docLanguage="en" docName="zt03693p206.pdf" docOrigin="Zootaxa 3693 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lycengraulis figueiredoi Loeb &amp; Alcântara, 2013, sp. n." docType="treatment" docVersion="8" lastPageNumber="204" masterDocId="FFBCFFA45E58AC45FFB4FFA1FFAA783A" masterDocTitle="A new species of Lycengraulis Günther, 1868 (Clupeiformes: Engraulinae) from the Amazon basin, Brazil, with comments on Lycengraulis batesii (Günther, 1868)" masterLastPageNumber="206" masterPageNumber="200" pageNumber="201" updateTime="1698344490041" updateUser="plazi">
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<mods:title id="BE22780D74442BA0CA6102A580433136">A new species of Lycengraulis Günther, 1868 (Clupeiformes: Engraulinae) from the Amazon basin, Brazil, with comments on Lycengraulis batesii (Günther, 1868)</mods:title>
</mods:titleInfo>
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<mods:namePart id="8B7C823C3AAE11E79018A3A65D46C563">Loeb, Marina Vianna</mods:namePart>
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<mods:namePart id="17750817944DC44C0CA95C5B75A6FF79">Alcântara, Ayda Vera</mods:namePart>
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<mods:title id="D8B92C7BC9C4ACD398EBE024E10E5F5A">Zootaxa</mods:title>
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<mods:date id="942CC6DD2F425068D856555EC7861624">2013</mods:date>
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<mods:number id="3985C225202FBCFB42A1AE75A7608852">3693</mods:number>
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<treatment id="038587DC5E59AC41FF23FBB6FDA47B9E" ID-DOI="http://doi.org/10.5281/zenodo.5689053" ID-GBIF-Taxon="119573469" ID-Zenodo-Dep="5689053" LSID="urn:lsid:plazi:treatment:038587DC5E59AC41FF23FBB6FDA47B9E" httpUri="http://treatment.plazi.org/id/038587DC5E59AC41FF23FBB6FDA47B9E" lastPageId="4" lastPageNumber="204" pageId="1" pageNumber="201">
<subSubSection id="C33665415E59AC44FF23FBB6FB1B7C8D" pageId="1" pageNumber="201" type="nomenclature">
<paragraph id="8B9336CA5E59AC44FF23FBB6FDBA7C0B" blockId="1.[151,528,1047,1106]" box="[151,528,1047,1073]" pageId="1" pageNumber="201">
<heading id="D0DB81A65E59AC44FF23FBB6FDBA7C0B" bold="true" box="[151,528,1047,1073]" fontSize="11" level="1" pageId="1" pageNumber="201" reason="1">
<emphasis id="B958EAD85E59AC44FF23FBB6FDBA7C0B" bold="true" box="[151,528,1047,1073]" pageId="1" pageNumber="201">
<emphasis id="B958EAD85E59AC44FF23FBB6FE6C7C0B" bold="true" box="[151,454,1047,1073]" italics="true" pageId="1" pageNumber="201">
<taxonomicName id="4C2C4D495E59AC44FF23FBB6FE6A7C0B" ID-CoL="6R24H" box="[151,448,1047,1073]" class="Actinopterygii" family="Engraulidae" genus="Lycengraulis" kingdom="Animalia" order="Clupeiformes" pageId="1" pageNumber="201" phylum="Chordata" rank="species" species="figueiredoi" status="sp. nov.">Lycengraulis figueiredoi</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A26B57A35E59AC44FE79FBB6FDBA7C0B" box="[461,528,1047,1073]" pageId="1" pageNumber="201" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9336CA5E59AC44FF23FB9BFE947C68" blockId="1.[151,528,1047,1106]" box="[151,318,1082,1106]" pageId="1" pageNumber="201">
(
<figureCitation id="13172A4F5E59AC44FF2BFB9BFF487C68" box="[159,226,1082,1106]" captionStart="FIGURES 1" captionStartId="3.[151,264,601,624]" captionTargetBox="[221,1355,195,565]" captionTargetId="figure@3.[221,1355,195,566]" captionTargetPageId="3" captionText="FIGURES 1. Lycengraulis figueiredoi, holotype, MZUSP 6032, 67.6 mm SL, Amazonas, lago Beruri at rio Purus, Brazil." httpUri="https://zenodo.org/record/222023/files/figure.png" pageId="1" pageNumber="201">Fig. 1</figureCitation>
,
<tableCitation id="C6AE03715E59AC44FF5BFB9BFE997C68" box="[239,307,1082,1106]" captionStart="TABLE 1" captionStartId="2.[151,239,331,355]" captionTargetBox="[159,1353,414,1891]" captionTargetPageId="2" captionText="TABLE 1. Morphometrics and meristics of Lycengraulis figueiredoi, new species. SD = Standard Deviation, frequency in parenthesis." httpUri="http://table.plazi.org/id/DF5366425E5AAC47FF23FEEAFE9A79B9" pageId="1" pageNumber="201" tableUuid="DF5366425E5AAC47FF23FEEAFE9A79B9">Tab. 1</tableCitation>
)
</paragraph>
<paragraph id="8B9336CA5E59AC44FF23FB23FB1B7C8D" blockId="1.[151,1436,1154,1207]" pageId="1" pageNumber="201">
<taxonomicName id="4C2C4D495E59AC44FF23FB23FDC07CA2" authority="Alcantara, 1972:19" authorityName="Alcantara" authorityPageNumber="19" authorityYear="1972" box="[151,618,1154,1176]" class="Actinopterygii" family="Engraulidae" genus="Lycengraulis" kingdom="Animalia" order="Clupeiformes" pageId="1" pageNumber="201" phylum="Chordata" rank="species" species="amazonensis">
<emphasis id="B958EAD85E59AC44FF23FB23FE0B7CA2" box="[151,417,1154,1176]" italics="true" pageId="1" pageNumber="201">Lycengraulis amazonensis</emphasis>
Alcântara, 1972:19
</taxonomicName>
[not a valid name, described as new in Alcântara's unpublished master thesis].— Whitehead
<emphasis id="B958EAD85E59AC44FE8FFB03FEC47C8D" box="[315,366,1185,1207]" italics="true" pageId="1" pageNumber="201">et al.</emphasis>
, 1985:391 [erroneously considered a junior synonym of
<taxonomicName id="4C2C4D495E59AC44FC19FB00FB087C8D" box="[941,1186,1185,1207]" class="Actinopterygii" family="Engraulidae" genus="Lycengraulis" kingdom="Animalia" order="Clupeiformes" pageId="1" pageNumber="201" phylum="Chordata" rank="species" species="grossidens">
<emphasis id="B958EAD85E59AC44FC19FB00FB087C8D" box="[941,1186,1185,1207]" italics="true" pageId="1" pageNumber="201">Lycengraulis grossidens</emphasis>
</taxonomicName>
].
</paragraph>
</subSubSection>
<subSubSection id="C33665415E59AC44FF23FB45FD477EB2" pageId="1" pageNumber="201" type="materials_examined">
<paragraph id="8B9336CA5E59AC44FF23FB45FD2B7D25" blockId="1.[151,1436,1252,2034]" pageId="1" pageNumber="201">
<typeStatus id="549788685E59AC44FF23FB45FEAE7CC6" box="[151,260,1252,1276]" pageId="1" pageNumber="201" type="holotype">
<emphasis id="B958EAD85E59AC44FF23FB45FEAE7CC6" bold="true" box="[151,260,1252,1276]" pageId="1" pageNumber="201">Holotype</emphasis>
</typeStatus>
. MZUSP
<quantity id="4CD49B2F5E59AC44FEC8FB45FDED7CC6" box="[380,583,1252,1276]" metricMagnitude="3" metricUnit="m" metricValue="6.0321298" pageId="1" pageNumber="201" unit="mm" value="6032129.8">6032, 129.80 mm</quantity>
SL.
<collectingCountry id="F33B765A5E59AC44FD35FB45FD627CC6" box="[641,712,1252,1276]" name="Brazil" pageId="1" pageNumber="201">Brazil</collectingCountry>
, Amazonas, lago Beruri, rio Purus,
<geoCoordinate id="EE18500D5E59AC44FBDDFB45FB197CC6" box="[1129,1203,1252,1276]" direction="south" orientation="latitude" pageId="1" pageNumber="201" precision="925" value="-3.8333333">3°50'S</geoCoordinate>
<geoCoordinate id="EE18500D5E59AC44FB09FB45FA8A7CC6" box="[1213,1312,1252,1276]" direction="west" orientation="longitude" pageId="1" pageNumber="201" precision="925" value="-61.333332">61°20'W</geoCoordinate>
,
<date id="FF92100A5E59AC44FA9AFB45FF787D25" pageId="1" pageNumber="201" value="1967-04-07" valueMax="1967-04-08" valueMin="1967-04-07">78 April 1967</date>
, Expedição Permanente da Amazônia.
</paragraph>
<paragraph id="8B9336CA5E59AC44FF73FA8DFD477EB2" blockId="1.[151,1436,1252,2034]" pageId="1" pageNumber="201">
<emphasis id="B958EAD85E59AC44FF73FA8DFEEC7D7E" bold="true" box="[199,326,1324,1348]" pageId="1" pageNumber="201">
<typeStatus id="549788685E59AC44FF73FA8DFEEB7D7E" box="[199,321,1324,1348]" pageId="1" pageNumber="201" type="paratype">Paratypes</typeStatus>
.
</emphasis>
All from
<collectingCountry id="F33B765A5E59AC44FE01FA8DFE577D7E" box="[437,509,1324,1348]" name="Brazil" pageId="1" pageNumber="201">Brazil</collectingCountry>
. ANSP 193934 (2,
<quantity id="4CD49B2F5E59AC44FD62FA8DFCD37D7E" box="[726,889,1324,1348]" metricMagnitude="-2" metricUnit="m" metricValue="7.185" metricValueMax="7.88" metricValueMin="6.49" pageId="1" pageNumber="201" unit="mm" value="71.85" valueMax="78.8" valueMin="64.9">64.978.8 mm</quantity>
SL), INPA 39522 (2, 63.677.0 mm SL), MNRJ 40719 (2,
<quantity id="4CD49B2F5E59AC44FEBDFAEEFE627D52" box="[265,456,1359,1384]" metricMagnitude="-2" metricUnit="m" metricValue="7.409" metricValueMax="8.186" metricValueMin="6.632" pageId="1" pageNumber="201" unit="mm" value="74.09" valueMax="81.86" valueMin="66.32">66.3281.86 mm</quantity>
SL): Pará, rio Trombetas, Oriximiná,
<geoCoordinate id="EE18500D5E59AC44FCC7FAEEFC167D5D" box="[883,956,1359,1383]" direction="south" orientation="latitude" pageId="1" pageNumber="201" precision="925" value="-1.7666667">1°46'S</geoCoordinate>
<geoCoordinate id="EE18500D5E59AC44FC77FAEEFB8C7D5D" box="[963,1062,1359,1383]" direction="west" orientation="longitude" pageId="1" pageNumber="201" precision="925" value="-55.866665">55°52'W</geoCoordinate>
,
<date id="FF92100A5E59AC44FB85FAEEFAB07D5D" box="[1073,1306,1359,1383]" pageId="1" pageNumber="201" value="1967-10-12" valueMax="1967-10-15" valueMin="1967-10-12">1215 October 1967</date>
, Expedição Permanente da Amazônia. MZUSP 6363 (
<quantity id="4CD49B2F5E59AC44FDC0FAD5FCAA7DB6" box="[628,768,1396,1420]" metricMagnitude="0" metricUnit="m" metricValue="1.1012" pageId="1" pageNumber="201" unit="mm" value="1101.2">1, 101.2 mm</quantity>
SL), Amazonas, lago Castro, rio Purus at mouth,
<date id="FF92100A5E59AC44FA84FAD5FF787D95" pageId="1" pageNumber="201" value="1967-04-07" valueMax="1967-04-08" valueMin="1967-04-07">78 April 1967</date>
, Expedição Permanente da Amazônia. MZUSP 9815-21 (5, 112.11
<quantity id="4CD49B2F5E59AC44FC66FA36FBF87D8A" box="[978,1106,1431,1456]" metricMagnitude="-1" metricUnit="m" metricValue="1.2324" pageId="1" pageNumber="201" unit="mm" value="123.24">123.24 mm</quantity>
SL), Amazonas, lago Beruri, rio Purus,
<date id="FF92100A5E59AC44FEB9FA1DFE177DEE" box="[269,445,1468,1492]" pageId="1" pageNumber="201" value="1967-04-07" valueMax="1967-04-08" valueMin="1967-04-07">78 April 1967</date>
, Expedição Permanente da Amazônia. MZUSP 6749 (
<quantity id="4CD49B2F5E59AC44FB9EFA1DFB1D7DEE" box="[1066,1207,1468,1492]" metricMagnitude="0" metricUnit="m" metricValue="1.1185999999999998" pageId="1" pageNumber="201" unit="mm" value="1118.6">1, 118.6 mm</quantity>
SL), Amazonas, rio Negro at Manaus neighborhood,
<date id="FF92100A5E59AC44FDBBFA7EFCB27DCD" box="[527,792,1503,1527]" pageId="1" pageNumber="201" value="1967-11-15" valueMax="1967-11-23" valueMin="1967-11-15">1523 November 1967</date>
, Expedição Permanente da Amazônia. MZUSP 7658 (
<quantity id="4CD49B2F5E59AC44FA3DFA7EFEA17E26" metricMagnitude="0" metricUnit="m" metricValue="1.1524" pageId="1" pageNumber="201" unit="mm" value="1152.4">1, 152.4 mm</quantity>
SL), Amazonas, lago José Açu at mouth, Parintins,
<date id="FF92100A5E59AC44FCD1F9A5FBC37E26" box="[869,1129,1540,1564]" pageId="1" pageNumber="201" value="1967-12-11" valueMax="1967-12-12" valueMin="1967-12-11">1112 December 1967</date>
, Expedição Permanente da Amazônia. MZUSP 9822 (15,
<quantity id="4CD49B2F5E59AC44FE59F986FD3A7E7A" box="[493,656,1575,1600]" metricMagnitude="-2" metricUnit="m" metricValue="6.540000000000001" metricValueMax="7.859999999999999" metricValueMin="5.220000000000001" pageId="1" pageNumber="201" unit="mm" value="65.4" valueMax="78.6" valueMin="52.2">52.278.6 mm</quantity>
SL), Pará, rio Trombetas, Oriximiná,
<geoCoordinate id="EE18500D5E59AC44FB83F986FB2B7E05" box="[1079,1153,1575,1599]" direction="south" orientation="latitude" pageId="1" pageNumber="201" precision="925" value="-1.7666667">1°46'S</geoCoordinate>
<geoCoordinate id="EE18500D5E59AC44FB3CF986FB407E05" box="[1160,1258,1575,1599]" direction="west" orientation="longitude" pageId="1" pageNumber="201" precision="925" value="-55.866665">55°52'W</geoCoordinate>
,
<date id="FF92100A5E59AC44FB41F986FF787E5E" pageId="1" pageNumber="201" value="1967-10-12" valueMax="1967-10-15" valueMin="1967-10-12">1215 October 1967</date>
, Expedição Permanente da Amazônia. MZUSP 5537 (
<quantity id="4CD49B2F5E59AC44FCEDF9EDFC457E5E" box="[857,1007,1612,1636]" metricMagnitude="0" metricUnit="m" metricValue="1.1696" pageId="1" pageNumber="201" unit="mm" value="1169.6">1, 169.6 mm</quantity>
SL), Pará, Lago Jacupá, Oriximiná,
<date id="FF92100A5E59AC44FF23F9D1FE947EB2" box="[151,318,1648,1672]" pageId="1" pageNumber="201" value="1967-02">February 1967</date>
, Expedição Permanente da Amazônia.
</paragraph>
</subSubSection>
<subSubSection id="C33665415E59AC44FF73F932FDB07FBD" pageId="1" pageNumber="201" type="diagnosis">
<paragraph id="8B9336CA5E59AC44FF73F932FDB07FBD" blockId="1.[151,1436,1252,2034]" pageId="1" pageNumber="201">
<emphasis id="B958EAD85E59AC44FF73F932FEEB7E96" bold="true" box="[199,321,1683,1708]" pageId="1" pageNumber="201">Diagnosis.</emphasis>
<taxonomicName id="4C2C4D495E59AC44FEF8F935FDC97E96" box="[332,611,1684,1708]" class="Actinopterygii" family="Engraulidae" genus="Lycengraulis" kingdom="Animalia" order="Clupeiformes" pageId="1" pageNumber="201" phylum="Chordata" rank="species" species="figueiredoi">
<emphasis id="B958EAD85E59AC44FEF8F935FDC97E96" box="[332,611,1684,1708]" italics="true" pageId="1" pageNumber="201">Lycengraulis figueiredoi</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C2C4D495E59AC44FC1DF934FBE87E96" box="[937,1090,1684,1708]" class="Actinopterygii" family="Engraulidae" genus="Lycengraulis" kingdom="Animalia" order="Clupeiformes" pageId="1" pageNumber="201" phylum="Chordata" rank="species" species="grossidens">
<emphasis id="B958EAD85E59AC44FC1DF934FBE87E96" box="[937,1090,1684,1708]" italics="true" pageId="1" pageNumber="201">L. grossidens</emphasis>
</taxonomicName>
by a short upper maxilla, its posterior margin not reaching the lower maxilla joint (vs. upper maxilla longer, its posterior margin reaching to or a little past of mandible joint).
<taxonomicName id="4C2C4D495E59AC44FE5AF97DFCAA7ECE" box="[494,768,1756,1780]" class="Actinopterygii" family="Engraulidae" genus="Lycengraulis" kingdom="Animalia" order="Clupeiformes" pageId="1" pageNumber="201" phylum="Chordata" rank="species" species="figueiredoi">
<emphasis id="B958EAD85E59AC44FE5AF97DFCAA7ECE" box="[494,768,1756,1780]" italics="true" pageId="1" pageNumber="201">Lycengraulis figueiredoi</emphasis>
</taxonomicName>
can be distinguished also from
<taxonomicName id="4C2C4D495E59AC44FBD5F97CFB117ECE" box="[1121,1211,1757,1780]" class="Actinopterygii" family="Engraulidae" genus="Lycengraulis" kingdom="Animalia" order="Clupeiformes" pageId="1" pageNumber="201" phylum="Chordata" rank="species" species="poeyi">
<emphasis id="B958EAD85E59AC44FBD5F97CFB117ECE" box="[1121,1211,1757,1780]" italics="true" pageId="1" pageNumber="201">L. poeyi</emphasis>
</taxonomicName>
by 26 to 31 anal-fin rays (
<emphasis id="B958EAD85E59AC44FF61F8A0FF5E7F22" box="[213,244,1793,1816]" italics="true" pageId="1" pageNumber="201">vs.</emphasis>
21 to 23 anal-fin rays) and by 44 to 46 vertebrae (
<emphasis id="B958EAD85E59AC44FC90F8A0FCE97F22" box="[804,835,1793,1816]" italics="true" pageId="1" pageNumber="201">vs.</emphasis>
42 vertebrae), and from
<taxonomicName id="4C2C4D495E59AC44FBE3F8A0FB157F22" box="[1111,1215,1792,1816]" class="Actinopterygii" family="Engraulidae" genus="Lycengraulis" kingdom="Animalia" order="Clupeiformes" pageId="1" pageNumber="201" phylum="Chordata" rank="species" species="batesii">
<emphasis id="B958EAD85E59AC44FBE3F8A0FB157F22" box="[1111,1215,1792,1816]" italics="true" pageId="1" pageNumber="201">L. batesii</emphasis>
</taxonomicName>
by having the analfin origin at vertical through base of second to fifth dorsal-fin ray (
<emphasis id="B958EAD85E59AC44FC28F884FC107F06" box="[924,954,1829,1852]" italics="true" pageId="1" pageNumber="201">vs.</emphasis>
anal-fin origin at vertical through base of sixth to 10th dorsal-fin ray) and by 1721 gill rakers on the lower branch of first gill arch (
<emphasis id="B958EAD85E59AC44FB27F8EDFB1B7F59" box="[1171,1201,1868,1891]" italics="true" pageId="1" pageNumber="201">vs.</emphasis>
1215 gill rakers on the lower branch of first gill arch).
</paragraph>
</subSubSection>
<subSubSection id="C33665415E59AC41FF73F833FD937AF6" lastPageId="4" lastPageNumber="204" pageId="1" pageNumber="201" type="description">
<paragraph id="8B9336CA5E59AC47FF73F833FC5D7926" blockId="1.[151,1436,1252,2034]" lastBlockId="2.[151,1436,151,284]" lastPageId="2" lastPageNumber="202" pageId="1" pageNumber="201">
<emphasis id="B958EAD85E59AC44FF73F833FEF37F91" bold="true" box="[199,345,1938,1963]" pageId="1" pageNumber="201">Description.</emphasis>
Morphometric data of the
<typeStatus id="549788685E59AC44FD25F833FD597F90" box="[657,755,1938,1962]" pageId="1" pageNumber="201" type="holotype">holotype</typeStatus>
and 30
<typeStatus id="549788685E59AC44FCE1F832FC6A7F91" box="[853,960,1939,1963]" pageId="1" pageNumber="201" type="paratype">paratypes</typeStatus>
are presented in
<tableCitation id="C6AE03715E59AC44FB3EF833FB4A7F90" box="[1162,1248,1938,1962]" captionStart="TABLE 1" captionStartId="2.[151,239,331,355]" captionTargetBox="[159,1353,414,1891]" captionTargetPageId="2" captionText="TABLE 1. Morphometrics and meristics of Lycengraulis figueiredoi, new species. SD = Standard Deviation, frequency in parenthesis." httpUri="http://table.plazi.org/id/DF5366425E5AAC47FF23FEEAFE9A79B9" pageId="1" pageNumber="201" tableUuid="DF5366425E5AAC47FF23FEEAFE9A79B9">Table 1</tableCitation>
. Body elongate, laterally compressed; greatest body depth at vertical through dorsal-fin origin. Dorsal body profile gently convex from tip of snout to vertical through posterior margin of posterior nostril, convex from latter point to the top of the head, approximately straight from the top of the head to dorsal-fin origin, straight and downward along dorsal-fin base, approximately straight along caudal peduncle. Ventral body profile gently convex from anterior tip of lower jaw to posterior margin of orbit, convex from latter point to the posterior end of anal-fin base, nearly straight along caudal peduncle. Caudal peduncle longer than deep. Vertebrae 4446 (n = 8).
</paragraph>
<caption id="DF5366425E5AAC47FF23FEEAFE9A79B9" ID-Table-UUID="DF5366425E5AAC47FF23FEEAFE9A79B9" httpUri="http://table.plazi.org/id/DF5366425E5AAC47FF23FEEAFE9A79B9" pageId="2" pageNumber="202" targetBox="[159,1353,414,1891]" targetIsTable="true" targetPageId="2">
<paragraph id="8B9336CA5E5AAC47FF23FEEAFE9A79B9" blockId="2.[151,1435,331,387]" pageId="2" pageNumber="202">
<emphasis id="B958EAD85E5AAC47FF23FEEAFEA37959" bold="true" box="[151,265,331,355]" pageId="2" pageNumber="202">TABLE 1.</emphasis>
Morphometrics and meristics of
<taxonomicName id="4C2C4D495E5AAC47FDC5FEEDFCDD7959" box="[625,887,332,355]" class="Actinopterygii" family="Engraulidae" genus="Lycengraulis" kingdom="Animalia" order="Clupeiformes" pageId="2" pageNumber="202" phylum="Chordata" rank="species" species="figueiredoi" status="sp. nov.">
<emphasis id="B958EAD85E5AAC47FDC5FEEDFCDD7959" box="[625,887,332,355]" italics="true" pageId="2" pageNumber="202">Lycengraulis figueiredoi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26B57A35E5AAC47FC31FEEDFBA37959" box="[901,1033,332,355]" pageId="2" pageNumber="202" rank="species">new species</taxonomicNameLabel>
. SD = Standard Deviation, frequency in parenthesis.
</paragraph>
</caption>
<paragraph id="8B9336CA5E5AAC47FF2BFE3FFBD67F59" pageId="2" pageNumber="202">
<table id="F92CC46A5E5A53BAFF2BFE3FFAE37F59" box="[159,1353,414,1891]" gridcols="4" gridrows="28" pageId="2" pageNumber="202">
<tr id="351C34885E5A53BAFF2BFE3FFAE3798E" box="[159,1353,414,436]" gridrow="0" pageId="2" pageNumber="202">
<th id="76CD5DF45E5A53BAFF2BFE3FFE7E798E" box="[159,468,414,436]" gridcol="0" gridrow="0" pageId="2" pageNumber="202">Morphometrics</th>
<th id="76CD5DF45E5A53BAFDAEFE3FFDD5798E" box="[538,639,414,436]" gridcol="1" gridrow="0" pageId="2" pageNumber="202">Holotype</th>
<th id="76CD5DF45E5A53BAFD7FFE3FFB01798E" box="[715,1195,414,436]" gridcol="2" gridrow="0" pageId="2" pageNumber="202">n Range Mean</th>
<th id="76CD5DF45E5A53BAFAA9FE3FFAE3798E" box="[1309,1353,414,436]" gridcol="3" gridrow="0" pageId="2" pageNumber="202">SD</th>
</tr>
<tr id="351C34885E5A53BAFF2BFE6DFAE37A04" box="[159,1353,460,574]" gridrow="1" pageId="2" pageNumber="202">
<th id="76CD5DF45E5A53BAFF2BFE6DFE7E7A04" box="[159,468,460,574]" gridcol="0" gridrow="1" pageId="2" pageNumber="202">Standard length (mm) Head length (mm) Percentages of standard length</th>
<td id="76CD5DF45E5A53BAFDAEFE6DFDD57A04" box="[538,639,460,574]" gridcol="1" gridrow="1" pageId="2" pageNumber="202">129.8 30.2</td>
<td id="76CD5DF45E5A53BAFD7FFE6DFB017A04" box="[715,1195,460,574]" gridcol="2" gridrow="1" pageId="2" pageNumber="202">30 52.2 169.6 85.6 30 12.5 41.1 20.9</td>
<td id="76CD5DF45E5A53BAFAA9FE6DFAE37A04" box="[1309,1353,460,574]" gridcol="3" gridrow="1" pageId="2" pageNumber="202">29.5 6.8</td>
</tr>
<tr id="351C34885E5A53BAFF2BFDF7FAE37A56" box="[159,1353,598,620]" gridrow="2" pageId="2" pageNumber="202">
<th id="76CD5DF45E5A53BAFF2BFDF7FE7E7A56" box="[159,468,598,620]" gridcol="0" gridrow="2" pageId="2" pageNumber="202">Body depth</th>
<td id="76CD5DF45E5A53BAFDAEFDF7FDD57A56" box="[538,639,598,620]" gridcol="1" gridrow="2" pageId="2" pageNumber="202">20.6</td>
<td id="76CD5DF45E5A53BAFD7FFDF7FB017A56" box="[715,1195,598,620]" gridcol="2" gridrow="2" pageId="2" pageNumber="202">30 19.1 24.4 21.7</td>
<td id="76CD5DF45E5A53BAFAA9FDF7FAE37A56" box="[1309,1353,598,620]" gridcol="3" gridrow="2" pageId="2" pageNumber="202">1.3</td>
</tr>
<tr id="351C34885E5A53BAFF2BFD25FAE37AA0" box="[159,1353,644,666]" gridrow="3" pageId="2" pageNumber="202">
<th id="76CD5DF45E5A53BAFF2BFD25FE7E7AA0" box="[159,468,644,666]" gridcol="0" gridrow="3" pageId="2" pageNumber="202">Caudal peduncle depth</th>
<td id="76CD5DF45E5A53BAFDAEFD25FDD57AA0" box="[538,639,644,666]" gridcol="1" gridrow="3" pageId="2" pageNumber="202">8.8</td>
<td id="76CD5DF45E5A53BAFD7FFD25FB017AA0" box="[715,1195,644,666]" gridcol="2" gridrow="3" pageId="2" pageNumber="202">30 8.2 11.2 9.9</td>
<td id="76CD5DF45E5A53BAFAA9FD25FAE37AA0" box="[1309,1353,644,666]" gridcol="3" gridrow="3" pageId="2" pageNumber="202">0.6</td>
</tr>
<tr id="351C34885E5A53BAFF2BFD13FAE37AF2" box="[159,1353,690,712]" gridrow="4" pageId="2" pageNumber="202">
<th id="76CD5DF45E5A53BAFF2BFD13FE7E7AF2" box="[159,468,690,712]" gridcol="0" gridrow="4" pageId="2" pageNumber="202">Dorsal-fin base length</th>
<td id="76CD5DF45E5A53BAFDAEFD13FDD57AF2" box="[538,639,690,712]" gridcol="1" gridrow="4" pageId="2" pageNumber="202">11.2</td>
<td id="76CD5DF45E5A53BAFD7FFD13FB017AF2" box="[715,1195,690,712]" gridcol="2" gridrow="4" pageId="2" pageNumber="202">30 10.0 13.9 12.0</td>
<td id="76CD5DF45E5A53BAFAA9FD13FAE37AF2" box="[1309,1353,690,712]" gridcol="3" gridrow="4" pageId="2" pageNumber="202">0.8</td>
</tr>
<tr id="351C34885E5A53BAFF2BFD41FAE37ACC" box="[159,1353,736,758]" gridrow="5" pageId="2" pageNumber="202">
<th id="76CD5DF45E5A53BAFF2BFD41FE7E7ACC" box="[159,468,736,758]" gridcol="0" gridrow="5" pageId="2" pageNumber="202">Anal-fin base length</th>
<td id="76CD5DF45E5A53BAFDAEFD41FDD57ACC" box="[538,639,736,758]" gridcol="1" gridrow="5" pageId="2" pageNumber="202">22.8</td>
<td id="76CD5DF45E5A53BAFD7FFD41FB017ACC" box="[715,1195,736,758]" gridcol="2" gridrow="5" pageId="2" pageNumber="202">30 25.6 31.0 27.3</td>
<td id="76CD5DF45E5A53BAFAA9FD41FAE37ACC" box="[1309,1353,736,758]" gridcol="3" gridrow="5" pageId="2" pageNumber="202">1.1</td>
</tr>
<tr id="351C34885E5A53BAFF2BFCAFFAE37B1E" box="[159,1353,782,804]" gridrow="6" pageId="2" pageNumber="202">
<th id="76CD5DF45E5A53BAFF2BFCAFFE7E7B1E" box="[159,468,782,804]" gridcol="0" gridrow="6" pageId="2" pageNumber="202">Pectoral fin length</th>
<td id="76CD5DF45E5A53BAFDAEFCAFFDD57B1E" box="[538,639,782,804]" gridcol="1" gridrow="6" pageId="2" pageNumber="202">19.7</td>
<td id="76CD5DF45E5A53BAFD7FFCAFFB017B1E" box="[715,1195,782,804]" gridcol="2" gridrow="6" pageId="2" pageNumber="202">30 13.4 21.6 18.2</td>
<td id="76CD5DF45E5A53BAFAA9FCAFFAE37B1E" box="[1309,1353,782,804]" gridcol="3" gridrow="6" pageId="2" pageNumber="202">2.3</td>
</tr>
<tr id="351C34885E5A53BAFF2BFC9DFAE37B68" box="[159,1353,828,850]" gridrow="7" pageId="2" pageNumber="202">
<th id="76CD5DF45E5A53BAFF2BFC9DFE7E7B68" box="[159,468,828,850]" gridcol="0" gridrow="7" pageId="2" pageNumber="202">Pelvic fin length</th>
<td id="76CD5DF45E5A53BAFDAEFC9DFDD57B68" box="[538,639,828,850]" gridcol="1" gridrow="7" pageId="2" pageNumber="202">10.5</td>
<td id="76CD5DF45E5A53BAFD7FFC9DFB017B68" box="[715,1195,828,850]" gridcol="2" gridrow="7" pageId="2" pageNumber="202">30 9.0 13.5 10.7</td>
<td id="76CD5DF45E5A53BAFAA9FC9DFAE37B68" box="[1309,1353,828,850]" gridcol="3" gridrow="7" pageId="2" pageNumber="202">1.2</td>
</tr>
<tr id="351C34885E5A53BAFF2BFCCBFAE37BBA" box="[159,1353,874,896]" gridrow="8" pageId="2" pageNumber="202">
<th id="76CD5DF45E5A53BAFF2BFCCBFE7E7BBA" box="[159,468,874,896]" gridcol="0" gridrow="8" pageId="2" pageNumber="202">Prepelvic length</th>
<td id="76CD5DF45E5A53BAFDAEFCCBFDD57BBA" box="[538,639,874,896]" gridcol="1" gridrow="8" pageId="2" pageNumber="202">40.9</td>
<td id="76CD5DF45E5A53BAFD7FFCCBFB017BBA" box="[715,1195,874,896]" gridcol="2" gridrow="8" pageId="2" pageNumber="202">30 38.0 45.2 42.4</td>
<td id="76CD5DF45E5A53BAFAA9FCCBFAE37BBA" box="[1309,1353,874,896]" gridcol="3" gridrow="8" pageId="2" pageNumber="202">1.7</td>
</tr>
<tr id="351C34885E5A53BAFF2BFC39FAE37B94" box="[159,1353,920,942]" gridrow="9" pageId="2" pageNumber="202">
<th id="76CD5DF45E5A53BAFF2BFC39FE7E7B94" box="[159,468,920,942]" gridcol="0" gridrow="9" pageId="2" pageNumber="202">Prepectoral length</th>
<td id="76CD5DF45E5A53BAFDAEFC39FDD57B94" box="[538,639,920,942]" gridcol="1" gridrow="9" pageId="2" pageNumber="202">24.3</td>
<td id="76CD5DF45E5A53BAFD7FFC39FB017B94" box="[715,1195,920,942]" gridcol="2" gridrow="9" pageId="2" pageNumber="202">30 21.7 26.5 24.7</td>
<td id="76CD5DF45E5A53BAFAA9FC39FAE37B94" box="[1309,1353,920,942]" gridcol="3" gridrow="9" pageId="2" pageNumber="202">1.2</td>
</tr>
<tr id="351C34885E5A53BAFF2BFC67FAE37BE6" box="[159,1353,966,988]" gridrow="10" pageId="2" pageNumber="202">
<th id="76CD5DF45E5A53BAFF2BFC67FE7E7BE6" box="[159,468,966,988]" gridcol="0" gridrow="10" pageId="2" pageNumber="202">Predorsal length</th>
<td id="76CD5DF45E5A53BAFDAEFC67FDD57BE6" box="[538,639,966,988]" gridcol="1" gridrow="10" pageId="2" pageNumber="202">55.3</td>
<td id="76CD5DF45E5A53BAFD7FFC67FB017BE6" box="[715,1195,966,988]" gridcol="2" gridrow="10" pageId="2" pageNumber="202">30 55.2 86.2 59.0</td>
<td id="76CD5DF45E5A53BAFAA9FC67FAE37BE6" box="[1309,1353,966,988]" gridcol="3" gridrow="10" pageId="2" pageNumber="202">5.3</td>
</tr>
<tr id="351C34885E5A53BAFF2BFC55FAE37C30" box="[159,1353,1012,1034]" gridrow="11" pageId="2" pageNumber="202">
<th id="76CD5DF45E5A53BAFF2BFC55FE7E7C30" box="[159,468,1012,1034]" gridcol="0" gridrow="11" pageId="2" pageNumber="202">Preanal length</th>
<td id="76CD5DF45E5A53BAFDAEFC55FDD57C30" box="[538,639,1012,1034]" gridcol="1" gridrow="11" pageId="2" pageNumber="202">56.8</td>
<td id="76CD5DF45E5A53BAFD7FFC55FB017C30" box="[715,1195,1012,1034]" gridcol="2" gridrow="11" pageId="2" pageNumber="202">30 57.5 62.6 60.5</td>
<td id="76CD5DF45E5A53BAFAA9FC55FAE37C30" box="[1309,1353,1012,1034]" gridcol="3" gridrow="11" pageId="2" pageNumber="202">1.5</td>
</tr>
<tr id="351C34885E5A53BAFF2BFB83FAE37C02" box="[159,1353,1058,1080]" gridrow="12" pageId="2" pageNumber="202">
<th id="76CD5DF45E5A53BAFF2BFB83FE7E7C02" box="[159,468,1058,1080]" gridcol="0" gridrow="12" pageId="2" pageNumber="202">Head length</th>
<td id="76CD5DF45E5A53BAFDAEFB83FDD57C02" box="[538,639,1058,1080]" gridcol="1" gridrow="12" pageId="2" pageNumber="202">23.3</td>
<td id="76CD5DF45E5A53BAFD7FFB83FB017C02" box="[715,1195,1058,1080]" gridcol="2" gridrow="12" pageId="2" pageNumber="202">30 22.5 26.8 24.6</td>
<td id="76CD5DF45E5A53BAFAA9FB83FAE37C02" box="[1309,1353,1058,1080]" gridcol="3" gridrow="12" pageId="2" pageNumber="202">0.9</td>
</tr>
<tr id="351C34885E5A53BAFF2BFBF1FAE37C5C" box="[159,1353,1104,1126]" gridrow="13" pageId="2" pageNumber="202" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76CD5DF45E5A53BAFF2BFBF1FE7E7C5C" box="[159,468,1104,1126]" gridcol="0" gridrow="13" pageId="2" pageNumber="202">Percentages of head length</th>
</tr>
<tr id="351C34885E5A53BAFF2BFBDFFAE37CAE" box="[159,1353,1150,1172]" gridrow="14" pageId="2" pageNumber="202">
<th id="76CD5DF45E5A53BAFF2BFBDFFE7E7CAE" box="[159,468,1150,1172]" gridcol="0" gridrow="14" pageId="2" pageNumber="202">Cheek length</th>
<td id="76CD5DF45E5A53BAFDAEFBDFFDD57CAE" box="[538,639,1150,1172]" gridcol="1" gridrow="14" pageId="2" pageNumber="202">47.7</td>
<td id="76CD5DF45E5A53BAFD7FFBDFFB017CAE" box="[715,1195,1150,1172]" gridcol="2" gridrow="14" pageId="2" pageNumber="202">30 35.3 48.8 39.8</td>
<td id="76CD5DF45E5A53BAFAA9FBDFFAE37CAE" box="[1309,1353,1150,1172]" gridcol="3" gridrow="14" pageId="2" pageNumber="202">3.7</td>
</tr>
<tr id="351C34885E5A53BAFF2BFB0DFAE37CF8" box="[159,1353,1196,1218]" gridrow="15" pageId="2" pageNumber="202">
<th id="76CD5DF45E5A53BAFF2BFB0DFE7E7CF8" box="[159,468,1196,1218]" gridcol="0" gridrow="15" pageId="2" pageNumber="202">Snout length</th>
<td id="76CD5DF45E5A53BAFDAEFB0DFDD57CF8" box="[538,639,1196,1218]" gridcol="1" gridrow="15" pageId="2" pageNumber="202">14.6</td>
<td id="76CD5DF45E5A53BAFD7FFB0DFB017CF8" box="[715,1195,1196,1218]" gridcol="2" gridrow="15" pageId="2" pageNumber="202">30 11.0 15.5 12.7</td>
<td id="76CD5DF45E5A53BAFAA9FB0DFAE37CF8" box="[1309,1353,1196,1218]" gridcol="3" gridrow="15" pageId="2" pageNumber="202">1.2</td>
</tr>
<tr id="351C34885E5A53BAFF2BFB7BFAE37CCA" box="[159,1353,1242,1264]" gridrow="16" pageId="2" pageNumber="202">
<th id="76CD5DF45E5A53BAFF2BFB7BFE7E7CCA" box="[159,468,1242,1264]" gridcol="0" gridrow="16" pageId="2" pageNumber="202">Orbital diameter</th>
<td id="76CD5DF45E5A53BAFDAEFB7BFDD57CCA" box="[538,639,1242,1264]" gridcol="1" gridrow="16" pageId="2" pageNumber="202">27.8</td>
<td id="76CD5DF45E5A53BAFD7FFB7BFB017CCA" box="[715,1195,1242,1264]" gridcol="2" gridrow="16" pageId="2" pageNumber="202">30 19.2 27.9 24.6</td>
<td id="76CD5DF45E5A53BAFAA9FB7BFAE37CCA" box="[1309,1353,1242,1264]" gridcol="3" gridrow="16" pageId="2" pageNumber="202">2.5</td>
</tr>
<tr id="351C34885E5A53BAFF2BFAA9FAE37D24" box="[159,1353,1288,1310]" gridrow="17" pageId="2" pageNumber="202">
<th id="76CD5DF45E5A53BAFF2BFAA9FE7E7D24" box="[159,468,1288,1310]" gridcol="0" gridrow="17" pageId="2" pageNumber="202">Upper jaw length</th>
<td id="76CD5DF45E5A53BAFDAEFAA9FDD57D24" box="[538,639,1288,1310]" gridcol="1" gridrow="17" pageId="2" pageNumber="202">74.8</td>
<td id="76CD5DF45E5A53BAFD7FFAA9FB017D24" box="[715,1195,1288,1310]" gridcol="2" gridrow="17" pageId="2" pageNumber="202">30 67.7 78.2 71.3</td>
<td id="76CD5DF45E5A53BAFAA9FAA9FAE37D24" box="[1309,1353,1288,1310]" gridcol="3" gridrow="17" pageId="2" pageNumber="202">2.8</td>
</tr>
<tr id="351C34885E5A53BAFF2BFA97FAE37D76" box="[159,1353,1334,1356]" gridrow="18" pageId="2" pageNumber="202">
<th id="76CD5DF45E5A53BAFF2BFA97FE7E7D76" box="[159,468,1334,1356]" gridcol="0" gridrow="18" pageId="2" pageNumber="202">Postorbital distance</th>
<td id="76CD5DF45E5A53BAFDAEFA97FDD57D76" box="[538,639,1334,1356]" gridcol="1" gridrow="18" pageId="2" pageNumber="202">63.0</td>
<td id="76CD5DF45E5A53BAFD7FFA97FB017D76" box="[715,1195,1334,1356]" gridcol="2" gridrow="18" pageId="2" pageNumber="202">30 57.1 66.8 62.7</td>
<td id="76CD5DF45E5A53BAFAA9FA97FAE37D76" box="[1309,1353,1334,1356]" gridcol="3" gridrow="18" pageId="2" pageNumber="202">2.6</td>
</tr>
<tr id="351C34885E5A53BAFF2BFAC5FAE37DFF" box="[159,1353,1380,1477]" gridrow="19" pageId="2" pageNumber="202">
<th id="76CD5DF45E5A53BAFF2BFAC5FE7E7DFF" box="[159,468,1380,1477]" gridcol="0" gridrow="19" pageId="2" pageNumber="202">Gill raker angle on1st gill arch length Meristics</th>
<td id="76CD5DF45E5A53BAFDAEFAC5FDD57DFF" box="[538,639,1380,1477]" gridcol="1" gridrow="19" pageId="2" pageNumber="202">11.9 Holotype</td>
<td id="76CD5DF45E5A53BAFD7FFAC5FB017DFF" box="[715,1195,1380,1477]" gridcol="2" gridrow="19" pageId="2" pageNumber="202">30 9.0 15.7 11.9 n Range</td>
<td id="76CD5DF45E5A53BAFAA9FAC5FAE37DFF" box="[1309,1353,1380,1477]" gridcol="3" gridrow="19" pageId="2" pageNumber="202">2.0</td>
</tr>
<tr id="351C34885E5A53BAFF2BFA7CFAE37DC9" box="[159,1353,1501,1523]" gridrow="20" pageId="2" pageNumber="202" rowspan-3="1">
<th id="76CD5DF45E5A53BAFF2BFA7CFE7E7DC9" box="[159,468,1501,1523]" gridcol="0" gridrow="20" pageId="2" pageNumber="202">Dorsal-fin rays</th>
<td id="76CD5DF45E5A53BAFDAEFA7CFDD57DC9" box="[538,639,1501,1523]" gridcol="1" gridrow="20" pageId="2" pageNumber="202">ii+11</td>
<td id="76CD5DF45E5A53BAFD7FFA7CFB017DC9" box="[715,1195,1501,1523]" gridcol="2" gridrow="20" pageId="2" pageNumber="202">30 13(2),14(16),15(11),16(1)</td>
</tr>
<tr id="351C34885E5A53BAFF2BF9AAFAE37E1B" box="[159,1353,1547,1569]" gridrow="21" pageId="2" pageNumber="202" rowspan-3="1">
<th id="76CD5DF45E5A53BAFF2BF9AAFE7E7E1B" box="[159,468,1547,1569]" gridcol="0" gridrow="21" pageId="2" pageNumber="202">Anal-fin rays</th>
<td id="76CD5DF45E5A53BAFDAEF9AAFDD57E1B" box="[538,639,1547,1569]" gridcol="1" gridrow="21" pageId="2" pageNumber="202">iii+23</td>
<td id="76CD5DF45E5A53BAFD7FF9AAFB017E1B" box="[715,1195,1547,1569]" gridcol="2" gridrow="21" pageId="2" pageNumber="202">25 26(1),27(1),28(7),29(6),30(8),31(2)</td>
</tr>
<tr id="351C34885E5A53BAFF2BF998FAE37E75" box="[159,1353,1593,1615]" gridrow="22" pageId="2" pageNumber="202" rowspan-3="1">
<th id="76CD5DF45E5A53BAFF2BF998FE7E7E75" box="[159,468,1593,1615]" gridcol="0" gridrow="22" pageId="2" pageNumber="202">Pectoral-fin rays</th>
<td id="76CD5DF45E5A53BAFDAEF998FDD57E75" box="[538,639,1593,1615]" gridcol="1" gridrow="22" pageId="2" pageNumber="202">i+14</td>
<td id="76CD5DF45E5A53BAFD7FF998FB017E75" box="[715,1195,1593,1615]" gridcol="2" gridrow="22" pageId="2" pageNumber="202">30 14(22),15(8)</td>
</tr>
<tr id="351C34885E5A53BAFF2BF9C6FAE37E47" box="[159,1353,1639,1661]" gridrow="23" pageId="2" pageNumber="202" rowspan-3="1">
<th id="76CD5DF45E5A53BAFF2BF9C6FE7E7E47" box="[159,468,1639,1661]" gridcol="0" gridrow="23" pageId="2" pageNumber="202">Pelvic-fin rays</th>
<td id="76CD5DF45E5A53BAFDAEF9C6FDD57E47" box="[538,639,1639,1661]" gridcol="1" gridrow="23" pageId="2" pageNumber="202">i+6</td>
<td id="76CD5DF45E5A53BAFD7FF9C6FB017E47" box="[715,1195,1639,1661]" gridcol="2" gridrow="23" pageId="2" pageNumber="202">30 7(30)</td>
</tr>
<tr id="351C34885E5A53BAFF2BF934FAE37E91" box="[159,1353,1685,1707]" gridrow="24" pageId="2" pageNumber="202" rowspan-3="1">
<th id="76CD5DF45E5A53BAFF2BF934FE7E7E91" box="[159,468,1685,1707]" gridcol="0" gridrow="24" pageId="2" pageNumber="202">Vertebrae</th>
<td id="76CD5DF45E5A53BAFDAEF934FDD57E91" box="[538,639,1685,1707]" gridcol="1" gridrow="24" pageId="2" pageNumber="202">-</td>
<td id="76CD5DF45E5A53BAFD7FF934FB017E91" box="[715,1195,1685,1707]" gridcol="2" gridrow="24" pageId="2" pageNumber="202">8 44(1),45(6),46(1)</td>
</tr>
<tr id="351C34885E5A53BAFF2BF962FAE37F3D" box="[159,1353,1731,1799]" gridrow="25" pageId="2" pageNumber="202" rowspan-3="1">
<th id="76CD5DF45E5A53BAFF2BF962FE7E7F3D" box="[159,468,1731,1799]" gridcol="0" gridrow="25" pageId="2" pageNumber="202">Gill rackers on1st gill arch Upper branch</th>
<td id="76CD5DF45E5A53BAFDAEF962FDD57F3D" box="[538,639,1731,1799]" gridcol="1" gridrow="25" pageId="2" pageNumber="202">14</td>
<td id="76CD5DF45E5A53BAFD7FF962FB017F3D" box="[715,1195,1731,1799]" gridcol="2" gridrow="25" pageId="2" pageNumber="202">30 13(2),14(21),15(7),16(1)</td>
</tr>
<tr id="351C34885E5A53BAFF2BF8BEFAE37F0F" box="[159,1353,1823,1845]" gridrow="26" pageId="2" pageNumber="202" rowspan-3="1">
<th id="76CD5DF45E5A53BAFF2BF8BEFE7E7F0F" box="[159,468,1823,1845]" gridcol="0" gridrow="26" pageId="2" pageNumber="202">Lower branch</th>
<td id="76CD5DF45E5A53BAFDAEF8BEFDD57F0F" box="[538,639,1823,1845]" gridcol="1" gridrow="26" pageId="2" pageNumber="202">20</td>
<td id="76CD5DF45E5A53BAFD7FF8BEFB017F0F" box="[715,1195,1823,1845]" gridcol="2" gridrow="26" pageId="2" pageNumber="202">30 17(4),18(9),19(13),20(4),21(1)</td>
</tr>
<tr id="351C34885E5A53BAFF2BF8ECFAE37F59" box="[159,1353,1869,1891]" gridrow="27" pageId="2" pageNumber="202" rowspan-3="1">
<th id="76CD5DF45E5A53BAFF2BF8ECFE7E7F59" box="[159,468,1869,1891]" gridcol="0" gridrow="27" pageId="2" pageNumber="202">Total</th>
<td id="76CD5DF45E5A53BAFDAEF8ECFDD57F59" box="[538,639,1869,1891]" gridcol="1" gridrow="27" pageId="2" pageNumber="202">34</td>
<td id="76CD5DF45E5A53BAFD7FF8ECFB017F59" box="[715,1195,1869,1891]" gridcol="2" gridrow="27" pageId="2" pageNumber="202">30 31(5),32(4),33(14),34(6),35(1),36(1)</td>
</tr>
</table>
</paragraph>
<caption id="DF5366425E5BAC46FF23FDF8FAC67A4A" box="[151,1388,601,624]" httpUri="https://zenodo.org/record/222023/files/figure.png" pageId="3" pageNumber="203" targetBox="[221,1355,195,565]" targetPageId="3">
<paragraph id="8B9336CA5E5BAC46FF23FDF8FAC67A4A" blockId="3.[151,1388,601,624]" box="[151,1388,601,624]" pageId="3" pageNumber="203">
<emphasis id="B958EAD85E5BAC46FF23FDF8FE8B7A55" bold="true" box="[151,289,601,624]" pageId="3" pageNumber="203">FIGURES 1.</emphasis>
<taxonomicName id="4C2C4D495E5BAC46FE93FDFBFDB47A4A" box="[295,542,602,624]" class="Actinopterygii" family="Engraulidae" genus="Lycengraulis" kingdom="Animalia" order="Clupeiformes" pageId="3" pageNumber="203" phylum="Chordata" rank="species" species="figueiredoi">
<emphasis id="B958EAD85E5BAC46FE93FDFBFDB47A4A" box="[295,542,602,624]" italics="true" pageId="3" pageNumber="203">Lycengraulis figueiredoi</emphasis>
</taxonomicName>
, holotype, MZUSP 6032, 67.6 mm SL, Amazonas, lago Beruri at rio Purus, Brazil.
</paragraph>
</caption>
<caption id="DF5366425E5BAC46FF23F9AFFA3D7E1F" box="[151,1431,1550,1573]" httpUri="https://zenodo.org/record/222024/files/figure.png" pageId="3" pageNumber="203" targetBox="[200,1333,674,1508]" targetPageId="3">
<paragraph id="8B9336CA5E5BAC46FF23F9AFFA3D7E1F" blockId="3.[151,1431,1550,1573]" box="[151,1431,1550,1573]" pageId="3" pageNumber="203">
<emphasis id="B958EAD85E5BAC46FF23F9AFFE8A7E1E" bold="true" box="[151,288,1550,1573]" pageId="3" pageNumber="203">FIGURES 2.</emphasis>
<taxonomicName id="4C2C4D495E5BAC46FE92F9AEFE587E1F" box="[294,498,1551,1573]" class="Actinopterygii" family="Engraulidae" genus="Lycengraulis" kingdom="Animalia" order="Clupeiformes" pageId="3" pageNumber="203" phylum="Chordata" rank="species" species="batesii">
<emphasis id="B958EAD85E5BAC46FE92F9AEFE587E1F" box="[294,498,1551,1573]" italics="true" pageId="3" pageNumber="203">Lycengraulis batesii</emphasis>
</taxonomicName>
, syntype, BMNH 1849.11.8.73-75, approximately 120 to 190 mm SL, Pará, rio Pará, Brazil.
</paragraph>
</caption>
<paragraph id="8B9336CA5E5BAC46FF73F9F0FE477EC3" blockId="3.[151,1437,1617,2002]" pageId="3" pageNumber="203">Conical-shaped head. Snout short, conical in lateral view. Mouth sub-terminal, inclined downward relative to body axis. Upper maxilla short and thin, posterior margin rounded, extending beyond the vertical through posterior margin of orbit, not reaching the mandible joint. Premaxilla, maxilla and dentary bearing one row of teeth. Teeth enlarged, well-spaced canine-like and slightly recurved at premaxilla and maxilla. Dentary with larger teeth intercalated with smaller ones.</paragraph>
<paragraph id="8B9336CA5E5BAC46FF73F8A6FBF27F5C" blockId="3.[151,1437,1617,2002]" pageId="3" pageNumber="203">Eye lateral on head, located dorsal to horizontal through pectoral-fin insertion; eye visible in dorsal and ventral views. Two confluent nostrils on each side of the head, the anterior elliptical and the posterior crescent-sharped. Perforated scale on the lateral line absent. Cicloid scales higher than large, deciduous.</paragraph>
<paragraph id="8B9336CA5E5BAC41FF73F8D0FA2479BD" blockId="3.[151,1437,1617,2002]" lastBlockId="4.[151,1436,151,932]" lastPageId="4" lastPageNumber="204" pageId="3" pageNumber="203">Dorsal-fin rays iiiii+1113 (n = 31), its origin closer to caudal-fin base than to tip of snout; dorsal-fin distal margin approximately straight in juveniles, slightly convex in adults. Anal-fin rays iiiii+2428 (n = 26), its origin at vertical through base of second to fifth dorsal-fin rays, anterior rays longer. Pectoral-fin rays i+1314 (n = 31); distal tip of longest pectoral-fin ray reaching pelvic-fin origin; pectoral-fin distal margin slightly convex. Pectoralfin axillary scale longer than half of pectoral-fin length, absent in most specimens, probably lost during collecting or storage. Pelvic fin-rays i+6 (n = 31), its pelvic-fin origin in midpoint of distance from pectoral-fin origin to anus; distal tip of longest pelvic-fin ray not reaching anal-fin origin; pelvic-fin distal margin slightly convex. Pelvic-fin axillary scale smaller than pectoral fin, absent in most specimens, probably lost during collecting or storage. Pelvic-fin origin closer to the base of the anteriormost pectoral-fin ray than to anal-fin origin. Caudal-fin rays 8,10,9,7, forked, lobes similar in size. Dorsal and ventral procurrent caudal-fin rays 8,7 respectively (n = 3 C&amp;S).</paragraph>
<paragraph id="8B9336CA5E5CAC41FF73FE35FBDE79CE" blockId="4.[151,1436,151,932]" pageId="4" pageNumber="204">Pseudobranch present, shorter than eye. Gill rakers on the first branchial arch long and thin in juveniles, short and thick in adults, 1316 (n = 31) on the upper branch, 1721 (n = 31) on the lower branch, 3136 (n = 31) in total. Length of raker inserted in the angle of first gill arch smaller than orbital diameter.</paragraph>
<paragraph id="8B9336CA5E5CAC41FF73FE5EFD937AF6" blockId="4.[151,1436,151,932]" pageId="4" pageNumber="204">
<emphasis id="B958EAD85E5CAC41FF73FE5EFE7A7A22" bold="true" box="[199,464,511,536]" pageId="4" pageNumber="204">Coloration in alcohol.</emphasis>
Body coloration pale to light yellow, juveniles with lighter coloration. Longitudinal stripe brownish to silver, absent in most preserved specimens, when present extending from the posterior margin of head to caudal peduncle; width of the stripe of about 20% of body depth at vertical through pectoral-fin origin. Caudal-fin borders and gill cavity darkened. Dark spots dorsally on the head, along dorsal portion of body, along dorsal margin of longitudinal line and along base of dorsal and caudal fins. Orbit and middle portion of the postorbital region translucent to pale.
</paragraph>
</subSubSection>
<subSubSection id="C33665415E5CAC41FF73FD79FEAA7B0D" pageId="4" pageNumber="204" type="etymology">
<paragraph id="8B9336CA5E5CAC41FF73FD79FEAA7B0D" blockId="4.[151,1436,151,932]" pageId="4" pageNumber="204">
<emphasis id="B958EAD85E5CAC41FF73FD79FEE47ACA" bold="true" box="[199,334,728,752]" pageId="4" pageNumber="204">Etymology.</emphasis>
The specific name was given in honor of José Lima de Figueiredo, a Brazilian ichthyologist from the Museu de Zoologia da Universidade de São Paulo who contributed significantly to the study of fish systematics in
<collectingCountry id="F33B765A5E5CAC41FF00FCBEFF567B0D" box="[180,252,799,823]" name="Brazil" pageId="4" pageNumber="204">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33665415E5CAC41FF73FCE2FDA47B9E" pageId="4" pageNumber="204" type="distribution">
<paragraph id="8B9336CA5E5CAC41FF73FCE2FDA47B9E" blockId="4.[151,1436,151,932]" pageId="4" pageNumber="204">
<emphasis id="B958EAD85E5CAC41FF73FCE2FECA7B66" bold="true" box="[199,352,835,860]" pageId="4" pageNumber="204">Distribution.</emphasis>
<taxonomicName id="4C2C4D495E5CAC41FEDCFCE5FDD17B66" box="[360,635,836,860]" class="Actinopterygii" family="Engraulidae" genus="Lycengraulis" kingdom="Animalia" order="Clupeiformes" pageId="4" pageNumber="204" phylum="Chordata" rank="species" species="figueiredoi">
<emphasis id="B958EAD85E5CAC41FEDCFCE5FDD17B66" box="[360,635,836,860]" italics="true" pageId="4" pageNumber="204">Lycengraulis figueiredoi</emphasis>
</taxonomicName>
is known only from lago Castro and lago Beruri, in the drainage of rio Purus; rio Negro; lago José Açu at Parintins; lago Jacupá and rio Trombetas at Oriximiná, middle Amazon basin, Amazonas and Pará State,
<collectingCountry id="F33B765A5E5CAC41FE75FC2DFDA37B9E" box="[449,521,908,932]" name="Brazil" pageId="4" pageNumber="204">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>