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<document id="5CE0EE894E30BC79E09A666B52DE29B4" ID-DOI="10.1080/00222930310001613575" ID-ISSN="1464-5262" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629850176377" checkinUser="felipe" docAuthor="Clausen, Martha Weis" docDate="2004" docId="038587CDFF939A18CF8EFD4B419AFF2E" docLanguage="en" docName="JNatHist.38.15.1851-1880.pdf" docOrigin="Journal of Natural History 38 (15)" docSource="http://dx.doi.org/10.1080/00222930310001613575" docStyle="DocumentStyle:443144A73DAE3E092C51A378E4CD0138.2:JNatHist.2003-2004.journal_article.1cover" docStyleId="443144A73DAE3E092C51A378E4CD0138" docStyleName="JNatHist.2003-2004.journal_article.1cover" docStyleVersion="2" docTitle="Libyodrilus platybrachyoncus Clausen, 2004, n. sp." docType="treatment" docVersion="2" lastPageNumber="1864" masterDocId="FFBCFFB5FF9D9A17CE4FFFAF4372FFC9" masterDocTitle="Description of seven new species of Libyodrilus Beddard from Cameroon, with a key to the species of the genus (Oligochaeta: Eudrilidae)" masterLastPageNumber="1880" masterPageNumber="1851" pageNumber="1864" updateTime="1699120809012" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="6C35AF5EC4E0D640214A8FF39A709523" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="25C52AF3AB1A29259E59B93C1429831F">
<mods:title id="6A83F1C46BAB54C89470695A2A7C1ED1">Description of seven new species of Libyodrilus Beddard from Cameroon, with a key to the species of the genus (Oligochaeta: Eudrilidae)</mods:title>
</mods:titleInfo>
<mods:name id="B9B8CF85C1A1BBCDF7C37C5734673138" type="personal">
<mods:role id="51827E38DD6A6CD6ABDBA1EC5CF504A0">
<mods:roleTerm id="C78451D47F9D111C936B95AC6821239E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5D7A98744FD325C9939CC8ADDDB93CAF">Clausen, Martha Weis</mods:namePart>
</mods:name>
<mods:typeOfResource id="7438ACE37A747775FE62EC24A13F8D2F">text</mods:typeOfResource>
<mods:relatedItem id="A7FEDFAC839AA7F5A0661BBF401E70BD" type="host">
<mods:titleInfo id="F43FAA4F86A6D0582787F4DF65BEC4FD">
<mods:title id="974FB41647DD675284EAE0767A2AA935">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="11F5A02CDAE83A0B68BCAC46F3D21647">
<mods:date id="CC77494BA5CF739A1376427D2C602DDE">2004</mods:date>
<mods:detail id="EDD5AF782B7F8AF506961F575563751C" type="pubDate">
<mods:number id="2F11356359BDA0690112408FA75051B8">2004-08-31</mods:number>
</mods:detail>
<mods:detail id="A703D20298063F317F9C61EDB0AF3DE7" type="volume">
<mods:number id="5650F49740F30615D0DF00A42E5D8586">38</mods:number>
</mods:detail>
<mods:detail id="A8A20ACE6961277DB80588DF6D15A827" type="issue">
<mods:number id="CA776BF4C05E258B1A558B1299DF8CF8">15</mods:number>
</mods:detail>
<mods:extent id="0FFF9FCF45F8B51261D7973514A7E884" unit="page">
<mods:start id="CB6CC70534051A98AA4337F57C0B92E5">1851</mods:start>
<mods:end id="8367250D7CA6383B130F8EE7C54113B1">1880</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="6AD6D01D295B659380B9F8444FB45D9F">
<mods:url id="9CFA9E26085A33118F1D54534CDB5022">http://dx.doi.org/10.1080/00222930310001613575</mods:url>
</mods:location>
<mods:classification id="91B4F622DCBB9545D2376227D93C5BE3">journal article</mods:classification>
<mods:identifier id="5A5244431FCA077CB4054B836F2F5914" type="DOI">10.1080/00222930310001613575</mods:identifier>
<mods:identifier id="B1860866607CD2E304A275C9F0AC14DA" type="ISSN">1464-5262</mods:identifier>
</mods:mods>
<treatment id="038587CDFF939A18CF8EFD4B419AFF2E" LSID="urn:lsid:plazi:treatment:038587CDFF939A18CF8EFD4B419AFF2E" httpUri="http://treatment.plazi.org/id/038587CDFF939A18CF8EFD4B419AFF2E" lastPageId="15" lastPageNumber="1864" pageId="14" pageNumber="1864">
<subSubSection id="C3366550FF939A19CF8EFD4B4060FCD5" pageId="14" pageNumber="1864" type="nomenclature">
<paragraph id="8B9336DBFF939A19CF8EFD4B4025FD35" blockId="14.[449,855,740,764]" box="[449,855,740,764]" pageId="14" pageNumber="1864">
<heading id="D0DB81B7FF939A19CF8EFD4B4025FD35" box="[449,855,740,764]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="1864" reason="5">
<taxonomicName id="4C2C4D58FF939A19CF8EFD4B407EFD35" ID-CoL="8LDDY" authorityName="Clausen" authorityYear="2004" box="[449,780,740,764]" class="Clitellata" family="Eudrilidae" genus="Libyodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="14" pageNumber="1864" phylum="Annelida" rank="species" species="platybrachyoncus" status="sp. nov.">
<emphasis id="B958EAC9FF939A19CF8EFD4B407EFD35" bold="true" box="[449,780,740,764]" italics="true" pageId="14" pageNumber="1864">Libyodrilus platybrachyoncus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B57B2FF939A19CD59FD4B4025FD35" box="[790,855,740,764]" pageId="14" pageNumber="1864" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8B9336DBFF939A19CC48FCAB4060FCD5" blockId="14.[519,786,772,796]" box="[519,786,772,796]" pageId="14" pageNumber="1864">(figures 6E, 7B, 9C, D)</paragraph>
</subSubSection>
<subSubSection id="C3366550FF939A18CED5FCFF419AFF2E" lastPageId="15" lastPageNumber="1865" pageId="14" pageNumber="1864" type="materials_examined">
<paragraph id="8B9336DBFF939A19CED5FCFF4203FCA1" blockId="14.[154,1111,848,1097]" box="[154,369,848,872]" pageId="14" pageNumber="1864">
<emphasis id="B958EAC9FF939A19CED5FCFF4203FCA1" box="[154,369,848,872]" italics="true" pageId="14" pageNumber="1864">Material examined</emphasis>
</paragraph>
<paragraph id="8B9336DBFF939A19CE8DFCDF4195FC21" blockId="14.[154,1111,848,1097]" pageId="14" pageNumber="1864">
<emphasis id="B958EAC9FF939A19CE8DFCDF4219FC41" box="[194,363,880,904]" italics="true" pageId="14" pageNumber="1864">Type material.</emphasis>
<materialsCitation id="3B443C86FF939A19CFEFFCC04191FC21" collectingDate="1966-01-11" collectionCode="ZMUC" collectorName="Akom II &amp; M. W. Clausen" country="Cameroon" location="Kribi" pageId="14" pageNumber="1864" specimenCode="OLI-7, OLI-8" specimenCount="1" specimenCount-juvenile="1" typeStatus="holotype">
<collectingCountry id="F33B764BFF939A19CFEFFCC04169FC4E" box="[416,539,879,903]" name="Cameroon" pageId="14" pageNumber="1864">Cameroon</collectingCountry>
,
<quantity id="4CD49B3EFF939A19CC62FCC041FDFC4E" box="[557,655,879,903]" metricMagnitude="4" metricUnit="m" metricValue="2.0921472" pageId="14" pageNumber="1864" unit="mi" value="13.0">13 miles</quantity>
E of
<location id="8EF36000FF939A19CCAEFCC04053FC4E" LSID="urn:lsid:plazi:treatment:038587CDFF939A18CF8EFD4B419AFF2E:8EF36000FF939A19CCAEFCC04053FC4E" box="[737,801,879,903]" country="Cameroon" name="Kribi" pageId="14" pageNumber="1864">Kribi</location>
, 2
<emphasis id="B958EAC9FF939A19CD0FFCDF4038FC41" bold="true" box="[832,842,880,904]" pageId="14" pageNumber="1864"></emphasis>
57
<emphasis id="B958EAC9FF939A19CD2AFCC04019FC41" bold="true" box="[869,875,879,904]" pageId="14" pageNumber="1864"></emphasis>
N, 9
<emphasis id="B958EAC9FF939A19CDECFCDF40DFFC41" bold="true" box="[931,941,880,904]" pageId="14" pageNumber="1864"></emphasis>
55
<emphasis id="B958EAC9FF939A19CD87FCC040BCFC41" bold="true" box="[968,974,879,904]" pageId="14" pageNumber="1864"></emphasis>
E at Kribi-
<collectorName id="26D9530DFF939A19CED5FC204274FC6E" box="[154,262,911,935]" pageId="14" pageNumber="1864">Akom II</collectorName>
road, in wet soil on bank of small stream,
<date id="FF92101BFF939A19CD0BFC204760FC6E" box="[836,1042,911,935]" pageId="14" pageNumber="1864" value="1966-01-11">
<collectingDate id="EFD6E9F3FF939A19CD0BFC204760FC6E" box="[836,1042,911,935]" pageId="14" pageNumber="1864" value="1966-01-11">11 January 1966</collectingDate>
</date>
, coll.
<collectorName id="26D9530DFF939A19CED5FC004223FC0E" box="[154,337,943,967]" pageId="14" pageNumber="1864">M. W. Clausen</collectorName>
[
<typeStatus id="54978879FF939A19CF2BFC0042BEFC0E" box="[356,460,943,967]" pageId="14" pageNumber="1864" type="holotype">holotype</typeStatus>
: one clitellate,
<collectionCode id="ED3DAE1EFF939A19CCCBFC0041ABFC0E" box="[644,729,943,967]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="14" pageNumber="1864">ZMUC</collectionCode>
<specimenCode id="DB8A9EA0FF939A19CCA9FC00405DFC0E" box="[742,815,943,967]" collectionCode="OLI-" pageId="14" pageNumber="1864">OLI-7</specimenCode>
;
<typeStatus id="54978879FF939A19CD0FFC0040C0FC0E" box="[832,946,943,967]" pageId="14" pageNumber="1864" type="paratype">paratypes</typeStatus>
: one clitellate (badly damaged),
<specimenCount id="9D2AFD52FF939A19CF21FC7F428FFC21" box="[366,509,976,1000]" pageId="14" pageNumber="1864" type="juvenile">one juvenile</specimenCount>
, all
<collectionCode id="ED3DAE1EFF939A19CC7DFC7F41F5FC21" box="[562,647,976,1000]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="14" pageNumber="1864">ZMUC</collectionCode>
<specimenCode id="DB8A9EA0FF939A19CCDEFC7F41AEFC21" box="[657,732,976,1000]" collectionCode="OLI-" pageId="14" pageNumber="1864">OLI-8</specimenCode>
]
</materialsCitation>
.
</paragraph>
<paragraph id="8B9336DBFF939A19CE8DFC5E406CFB80" blockId="14.[154,1111,848,1097]" pageId="14" pageNumber="1864">
<emphasis id="B958EAC9FF939A19CE8DFC5E42D8FBC0" box="[194,426,1009,1033]" italics="true" pageId="14" pageNumber="1864">Additional material.</emphasis>
<materialsCitation id="3B443C86FF939A19CFADFC5E4068FB80" collectingDate="1966-01-11" collectionCode="ZMUC" collectorName="Kribi-Akom II &amp; M. W. Clausen" country="Cameroon" location="Kribi" pageId="14" pageNumber="1864" specimenCode="OLI-9" specimenCount="1">
<collectingCountry id="F33B764BFF939A19CFADFC5E412FFBC0" box="[482,605,1009,1033]" name="Cameroon" pageId="14" pageNumber="1864">Cameroon</collectingCountry>
,
<quantity id="4CD49B3EFF939A19CC3EFC5E41A4FBC0" box="[625,726,1009,1033]" metricMagnitude="4" metricUnit="m" metricValue="2.2530816" pageId="14" pageNumber="1864" unit="mi" value="14.0">14 miles</quantity>
E of
<location id="8EF36000FF939A19CD7FFC5E401FFBC0" LSID="urn:lsid:plazi:treatment:038587CDFF939A18CF8EFD4B419AFF2E:8EF36000FF939A19CD7FFC5E401FFBC0" box="[816,877,1009,1033]" country="Cameroon" name="Kribi" pageId="14" pageNumber="1864">Kribi</location>
at
<collectorName id="26D9530DFF939A19CDEBFC5E4724FBC0" box="[932,1110,1009,1033]" pageId="14" pageNumber="1864">Kribi-Akom II</collectorName>
road, wet soil on bank of small stream,
<date id="FF92101BFF939A19CCC6FBBE4023FBE0" box="[649,849,1041,1065]" pageId="14" pageNumber="1864" value="1966-01-11">
<collectingDate id="EFD6E9F3FF939A19CCC6FBBE4023FBE0" box="[649,849,1041,1065]" pageId="14" pageNumber="1864" value="1966-01-11">11 January 1966</collectingDate>
</date>
, coll.
<collectorName id="26D9530DFF939A19CDD0FBBE4724FBE0" box="[927,1110,1041,1065]" pageId="14" pageNumber="1864">M. W. Clausen</collectorName>
(one clitellate with hind end damaged,
<collectionCode id="ED3DAE1EFF939A19CC28FB9E41CEFB80" box="[615,700,1073,1097]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="14" pageNumber="1864">ZMUC</collectionCode>
<specimenCode id="DB8A9EA0FF939A19CC88FB9E4060FB80" box="[711,786,1073,1097]" collectionCode="OLI-" pageId="14" pageNumber="1864">OLI-9</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9336DBFF939A19CED5FB23428AFB6A" blockId="14.[154,1111,1163,1732]" box="[154,504,1163,1188]" pageId="14" pageNumber="1864">
<emphasis id="B958EAC9FF939A19CED5FB23420AFB6D" box="[154,376,1164,1188]" italics="true" pageId="14" pageNumber="1864">External characters</emphasis>
(figure 7B)
</paragraph>
<paragraph id="8B9336DBFF939A19CE8DFB044075FA0D" blockId="14.[154,1111,1163,1732]" pageId="14" pageNumber="1864">
<typeStatus id="54978879FF939A19CE8DFB044248FB0B" box="[194,314,1195,1219]" pageId="14" pageNumber="1864" type="holotype">HOLOTYPE</typeStatus>
: length
<quantity id="4CD49B3EFF939A19CFD3FB044285FB0A" box="[412,503,1195,1219]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" pageId="14" pageNumber="1864" unit="mm" value="125.0">125 mm</quantity>
, maximum diameter of all three clitellate specimens
<quantity id="4CD49B3EFF939A19CED5FB64439DFB2A" box="[154,239,1227,1251]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="1864" unit="mm" value="4.5">4.5 mm</quantity>
, number of segments of type 166. Annulation: segment 9 bi-annulate, 10 tri-annulate, 1113 and 1820 multi-annulate, annulation posterior to 20 gradually fading away. Prostomium prolobous. Clitellum orange, genital swellings yellowish, rest of body without pigment. Setae uniform and closely paired. Setal formula:
<emphasis id="B958EAC9FF939A19CF0BFAE3422DFAAD" box="[324,351,1356,1380]" italics="true" pageId="14" pageNumber="1864">aa</emphasis>
:
<emphasis id="B958EAC9FF939A19CF29FAE342F3FAAD" box="[358,385,1356,1380]" italics="true" pageId="14" pageNumber="1864">ab</emphasis>
:
<emphasis id="B958EAC9FF939A19CFC8FAE342D2FAAD" box="[391,416,1356,1380]" italics="true" pageId="14" pageNumber="1864">bc</emphasis>
:
<emphasis id="B958EAC9FF939A19CFE8FAE342B2FAAD" box="[423,448,1356,1380]" italics="true" pageId="14" pageNumber="1864">cd</emphasis>
<emphasis id="B958EAC9FF939A19CF8FFAE342A9FAAD" bold="true" box="[448,475,1356,1380]" pageId="14" pageNumber="1864">~</emphasis>
8.0:1.0:5.5:1.0,
<emphasis id="B958EAC9FF939A19CCC8FAE341D0FAAD" box="[647,674,1356,1380]" italics="true" pageId="14" pageNumber="1864">dd</emphasis>
<emphasis id="B958EAC9FF939A19CCEDFAE341CFFAAD" bold="true" box="[674,701,1356,1380]" pageId="14" pageNumber="1864">~</emphasis>
two-thirds of body circumference at segment 10; 5.5:1.0:4.0:1.0,
<emphasis id="B958EAC9FF939A19CFAFFAC34289FA4D" box="[480,507,1388,1412]" italics="true" pageId="14" pageNumber="1864">dd</emphasis>
<emphasis id="B958EAC9FF939A19CFB4FAC34164FA4D" bold="true" box="[507,534,1388,1412]" pageId="14" pageNumber="1864">~</emphasis>
two-thirds of body circumference at segment 30. Clitellum annular, with 3
<emphasis id="B958EAC9FF939A19CF89FA234293FA6D" bold="true" box="[454,481,1420,1444]" pageId="14" pageNumber="1864">K</emphasis>
segments in segments 141/2 17, replaced by genital swelling ventrally in posterior half of 17 (see below).
</paragraph>
<paragraph id="8B9336DBFF939A19CE8DFA634073FA2D" blockId="14.[154,1111,1163,1732]" box="[194,769,1484,1508]" pageId="14" pageNumber="1864">No dorsal pores. Nephridiopores not discernible.</paragraph>
<paragraph id="8B9336DBFF939A19CE8DFA434262F98D" blockId="14.[154,1111,1163,1732]" pageId="14" pageNumber="1864">
Spermarial pore unpaired, transverse slit (length about 1 1/2
<emphasis id="B958EAC9FF939A19CDC6FA4340D6F9CD" box="[905,932,1516,1540]" italics="true" pageId="14" pageNumber="1864">ab</emphasis>
) m/v in 13 just behind setal zone. Slit surrounded by low swelling (width about 1/2
<emphasis id="B958EAC9FF939A19CD85F9A34097F9ED" box="[970,997,1548,1572]" italics="true" pageId="14" pageNumber="1864">aa</emphasis>
, length a little less).
</paragraph>
<paragraph id="8B9336DBFF939A19CE8DF9E3410CF94D" blockId="14.[154,1111,1163,1732]" pageId="14" pageNumber="1864">
Female pores paired, minute openings on segment 15 between setal zone and 15/16, dorsal to
<emphasis id="B958EAC9FF939A19CF13F9C2421BF94C" box="[348,361,1645,1669]" italics="true" pageId="14" pageNumber="1864">d</emphasis>
, distance to
<emphasis id="B958EAC9FF939A19CC4BF9C24163F94C" box="[516,529,1645,1669]" italics="true" pageId="14" pageNumber="1864">d</emphasis>
<emphasis id="B958EAC9FF939A19CC5EF9C2415EF94C" bold="true" box="[529,556,1645,1669]" pageId="14" pageNumber="1864">~</emphasis>
14
<emphasis id="B958EAC9FF939A19CC10F9C2410AF94C" box="[607,632,1645,1669]" italics="true" pageId="14" pageNumber="1864">cd</emphasis>
.
</paragraph>
<paragraph id="8B9336DBFF939A18CE8DF923419AFF2E" blockId="14.[154,1111,1163,1732]" lastBlockId="15.[159,1115,173,231]" lastPageId="15" lastPageNumber="1865" pageId="14" pageNumber="1864">
Male pores closely paired, m/v in 17 between eq and 17/
<quantity id="4CD49B3EFF939A19CD72F9234007F96D" box="[829,885,1676,1700]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="14" pageNumber="1864" unit="in" value="18.0">18 in</quantity>
<typeStatus id="54978879FF939A19CD33F92340C2F96D" box="[892,944,1676,1700]" pageId="14" pageNumber="1864">type</typeStatus>
. The pores are situated on yellow glandular swelling (clearly distinctive from clitellar tissue) covering or almost covering area between setal lines
<emphasis id="B958EAC9FF929A18CD49FF014061FF0F" box="[774,787,174,198]" italics="true" pageId="15" pageNumber="1865">d</emphasis>
on anterior half of segment 17. Penial sacs opening together with euprostates.
</paragraph>
</subSubSection>
</treatment>
</document>