123 lines
15 KiB
XML
123 lines
15 KiB
XML
<document id="C69F597F75D90EEAB4B7093EB8D7F734" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE345FFD2FF75F96DBA8AFD9D" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena (Monomadraena) tibiodentipes Perkins 2017" docType="treatment" docVersion="9" lastPageNumber="68" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="67" updateTime="1698480684898" updateUser="ExternalLinkService">
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<mods:titleInfo id="7A47DB58F3FE4CEF959DC01BDACC484A">
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<mods:title id="166BEB7BE038498F2E4460D58C5EEFAF">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
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<mods:name id="19B6C813D76F0101344E3B09FD63DE35" type="personal">
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<mods:namePart id="300FEBE8FCDB4F90C05E17C4FE54F900">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="45797D427DE96E3F738B6BA5BB8CD5DC">text</mods:typeOfResource>
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<mods:title id="295C0291959B50376D957185AA78B7A2">Zootaxa</mods:title>
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<mods:part id="9016F447F154D350E2AA04B57582A562">
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<mods:date id="D30CBBB1D93F7B93CA17980F2223FDA3">2017</mods:date>
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<mods:number id="6712AF1E813E7CD23555BB6BBB3DE1A9">2017-11-03</mods:number>
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<mods:number id="FC31BC2014E9F65029277BE3C3098D0B">4342</mods:number>
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<mods:identifier id="CA147F93E59C450F7B786B6FF50CAB57" type="DOI">10.11646/zootaxa.4342.1.1</mods:identifier>
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<mods:identifier id="87A386B581404C88202A152EBA1249B5" type="ISSN">1175-5326</mods:identifier>
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<treatment id="038587BBE345FFD2FF75F96DBA8AFD9D" ID-DOI="http://doi.org/10.5281/zenodo.5323528" ID-GBIF-Taxon="137233472" ID-Zenodo-Dep="5323528" LSID="urn:lsid:plazi:treatment:038587BBE345FFD2FF75F96DBA8AFD9D" httpUri="http://treatment.plazi.org/id/038587BBE345FFD2FF75F96DBA8AFD9D" lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67">
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<subSubSection id="C3366526E345FFD3FF75F96DB8E5F8EA" pageId="66" pageNumber="67" type="nomenclature">
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<paragraph id="8B9336ADE345FFD3FF75F96DBA00F967" blockId="66.[151,828,1736,1762]" box="[151,828,1736,1762]" pageId="66" pageNumber="67">
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<heading id="D0DB81C1E345FFD3FF75F96DBA00F967" bold="true" box="[151,828,1736,1762]" fontSize="11" level="1" pageId="66" pageNumber="67" reason="1">
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<emphasis id="B958EABFE345FFD3FF75F96DBA00F967" bold="true" box="[151,828,1736,1762]" pageId="66" pageNumber="67">
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<taxonomicName id="4C2C4D2EE345FFD3FF75F96DBBB0F967" ID-CoL="6MJ3H" authorityName="Perkins" authorityYear="2017" box="[151,652,1736,1762]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="tibiodentipes" subGenus="Monomadraena">
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<emphasis id="B958EABFE345FFD3FF75F96DB82CF967" bold="true" box="[151,272,1736,1762]" italics="true" pageId="66" pageNumber="67">Hydraena</emphasis>
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(
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<emphasis id="B958EABFE345FFD3FEC2F96DB8DEF967" bold="true" box="[288,482,1736,1762]" italics="true" pageId="66" pageNumber="67">Monomadraena</emphasis>
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)
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<emphasis id="B958EABFE345FFD3FE11F96DBBB0F967" bold="true" box="[499,652,1736,1762]" italics="true" pageId="66" pageNumber="67">tibiodentipes</emphasis>
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</taxonomicName>
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species group
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9336ADE345FFD3FF75F8B5B8E5F8EA" blockId="66.[151,1437,1807,2012]" pageId="66" pageNumber="67">
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Fourteen species are included in this group:
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<emphasis id="B958EABFE345FFD3FD60F8B5B834F8F5" italics="true" pageId="66" pageNumber="67">
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<taxonomicName id="4C2C4D2EE345FFD3FD60F8B5BA0BF8AD" authorityName="Perkins" authorityYear="2017" box="[642,823,1807,1832]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="tibiodentipes">H. tibiodentipes</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE345FFD3FCA0F8B5BAF2F8AD" authorityName="Perkins" authorityYear="2017" box="[834,974,1807,1832]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="bisinuata">H. bisinuata</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE345FFD3FC3BF8B5BD9AF8AD" authorityName="Perkins" authorityYear="2017" box="[985,1190,1807,1832]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="contracolorata">H. contracolorata</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE345FFD3FB52F8B5BC4BF8AD" authorityName="Perkins" authorityYear="2017" box="[1200,1399,1807,1832]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="amplipunctata">H. amplipunctata</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE345FFD3FA63F8B5B817F8CE" authorityName="Perkins" authorityYear="2017" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="parvispinosa">H. parvispinosa</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE345FFD3FED7F891B8EFF8C9" authorityName="Perkins" authorityYear="2017" box="[309,467,1843,1868]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="triaequalis">H. triaequalis</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE345FFD3FE3CF891BBB6F8C9" authorityName="Perkins" authorityYear="2017" box="[478,650,1843,1868]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="rubrifurcata">H. rubrifurcata</taxonomicName>
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<taxonomicName id="4C2C4D2EE345FFD3FD72F891BA24F8CE" authorityName="Perkins" authorityYear="2017" box="[656,792,1844,1867]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="inseriata">H. inseriata</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE345FFD3FCC6F891BAFDF8C9" authorityName="Perkins" authorityYear="2017" box="[804,961,1843,1868]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="levifurcata">H. levifurcata</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE345FFD3FC2EF891BD62F8CE" authorityName="Perkins" authorityYear="2017" box="[972,1118,1844,1867]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="compacta">H. compacta</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE345FFD3FB8AF891BC3DF8C9" authorityName="Perkins" authorityYear="2017" box="[1128,1281,1843,1868]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="upsilonica">H. upsilonica</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE345FFD3FAEFF891BCA4F8CE" authorityName="Perkins" authorityYear="2017" box="[1293,1432,1844,1867]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="tripartita">H. tripartita</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE345FFD3FF75F8FDB838F8F5" authorityName="Perkins" authorityYear="2017" box="[151,260,1879,1904]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="jubata">H. jubata</taxonomicName>
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,
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</emphasis>
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and
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<taxonomicName id="4C2C4D2EE345FFD3FEDCF8FDB8EDF8F5" authorityName="Perkins" authorityYear="2017" box="[318,465,1879,1904]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="sculponea">
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<emphasis id="B958EABFE345FFD3FEDCF8FDB8EDF8F5" box="[318,465,1879,1904]" italics="true" pageId="66" pageNumber="67">H. sculponea</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E345FFD2FF25F8DEBA8AFD9D" lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67" type="description">
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<paragraph id="8B9336ADE345FFD2FF25F8DEBD36FEC5" blockId="66.[151,1437,1807,2012]" lastBlockId="67.[151,1437,151,536]" lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67">
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Members of this group are of moderate size (ca.
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<quantity id="4CD49B48E345FFD3FD00F8D9BAB4F811" box="[738,904,1916,1940]" metricMagnitude="-3" metricUnit="m" metricValue="1.845" metricValueMax="2.21" metricValueMin="1.48" pageId="66" pageNumber="67" unit="mm" value="1.845" valueMax="2.21" valueMin="1.48">1.48–2.21 mm</quantity>
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), usually dark brown or reddish brown dorsally, infrequently with a pronotal macula, and have coarse and dense dorsal punctation. The genae have a strong posterior ridge. The metaventrite has a strong anteromedian carina that is, in ten of the species, contiguous with the plaques, together forming an inverted Y-shape (e.g.,
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<figureCitation id="13172A28E344FFD2FCFBFF32BA49FF35" box="[793,885,151,176]" captionStart="FIGURE 74" captionStartId="85.[151,250,1404,1426]" captionText="FIGURE 74. Hydraena upsilonica, holotype, habitus dorsal, lateral, and ventral." pageId="67" pageNumber="68">Fig. 74</figureCitation>
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). The metaventrite ridges extending from the mesocoxal cavities are also quite strong. The intercoxal processes are relatively wide, with P1 and P2 approximately equal in width. Mesotibiae of males have a row of spines on the inner margin, quite prominent in all but one species, number varying from 2–3 to 4–5, and the margin is usually notched between the spines (e.g.,
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<figureCitation id="13172A28E344FFD2FA92FEA1B98AFEC5" captionStart="FIGURE 57" captionStartId="68.[151,250,1215,1237]" captionText="FIGURE 57. Hydraena tibiodentipes, holotype, habitus dorsal, lateral, and ventral." pageId="67" pageNumber="68">Fig. 57</figureCitation>
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; in
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<taxonomicName id="4C2C4D2EE344FFD2FF3CFE8CB8ADFEC5" authorityName="Perkins" authorityYear="2017" box="[222,401,297,320]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="parvispinosa">
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<emphasis id="B958EABFE344FFD2FF3CFE8CB8ADFEC5" box="[222,401,297,320]" italics="true" pageId="67" pageNumber="68">H. parvispinosa</emphasis>
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</taxonomicName>
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the spines are very short, and the margin is not notched.
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</paragraph>
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<paragraph id="8B9336ADE344FFD2FF25FEE9BA8AFD9D" blockId="67.[151,1437,151,536]" pageId="67" pageNumber="68">
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The aedeagi have a similar general plan: the main piece is slender in profile, the single paramere (right side) is large in dorsal aspect, originates near the basal orifice, and extends to or slightly beyond the main piece. In all but one species the gonopore is located at the end of a moderately long to very long, usually coiled, flagellum (e.g.,
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<figureCitation id="13172A28E344FFD2FF75FE12B81FFE55" box="[151,291,439,464]" captionStart="FIGURES 67 – 68" captionStartId="78.[151,264,1879,1902]" captionText="FIGURES 67 – 68. Hydraena, aedeagi of holotypes." pageId="67" pageNumber="68">Figs. 67, 68</figureCitation>
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). The aedeagus of
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<taxonomicName id="4C2C4D2EE344FFD2FE1CFE1CBB56FE55" authorityName="Perkins" authorityYear="2017" box="[510,618,439,464]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="jubata">
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<emphasis id="B958EABFE344FFD2FE1CFE1CBB56FE55" box="[510,618,439,464]" italics="true" pageId="67" pageNumber="68">H. jubata</emphasis>
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</taxonomicName>
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does not appear to have a flagellum, but only one male is known; it is possible that the flagellum is broken off; the aedeagus is also unusual in the size and shape of the distal piece. Consequently
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<taxonomicName id="4C2C4D2EE344FFD2FEDAFDA4B89EFD9D" authorityName="Perkins" authorityYear="2017" box="[312,418,511,536]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="jubata">
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<emphasis id="B958EABFE344FFD2FEDAFDA4B89EFD9D" box="[312,418,511,536]" italics="true" pageId="67" pageNumber="68">H. jubata</emphasis>
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</taxonomicName>
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is only tentatively placed in this species group.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |