157 lines
17 KiB
XML
157 lines
17 KiB
XML
<document id="8614469D1BB379D9584183FAB3FEFA82" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE33AFFAEFF75FF3DB843FD2D" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena multiarcuata Perkins 2017, new species" docType="treatment" docVersion="9" lastPageNumber="64" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="62" updateTime="1698480684898" updateUser="ExternalLinkService">
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<mods:mods id="15022C9FA366B9640BFD33100E0CD1E3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9E7DB2DDE8C17BA791F9168A687CE559">
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<mods:title id="BBB948703DCAF447740EAB0AC6FC90AA">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
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</mods:titleInfo>
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<mods:name id="DC2DD859A5A97830A5AF68C655D5AC64" type="personal">
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<mods:role id="485C058C0329DF16E07CAA47E33857EA">
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<mods:roleTerm id="E78962569707C296991DC2474A8DE858">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C0C970E28262404168FFEFFA56FA6C36">Perkins, Philip D.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="19F977D095AF831A17AEB2633ACB871A">text</mods:typeOfResource>
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<mods:titleInfo id="4A86B6B91DBE426411D72D9C4A8F1D8B">
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<mods:title id="233CEDED398F3486118F19A6EB41D07A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="3BD9BECD77A2208830221FDF0E246C35">
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<mods:date id="512A2FBE8244B7603ECE560A56EC891F">2017</mods:date>
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<mods:detail id="CCA358FA84350CB94AC8509C19E4BBFA" type="pubDate">
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<mods:number id="6BC0A215B0BB24BFEE131929262352BC">2017-11-03</mods:number>
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</mods:detail>
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<mods:detail id="4B65885BF4CF92AE5707B5AA3A36F7AC" type="volume">
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<mods:number id="DE27684FF80D8DEDD980E476C2FCF322">4342</mods:number>
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</mods:detail>
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<mods:detail id="8E9B11C8F86312C57B079208ECD0A7A4" type="issue">
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<mods:number id="110FA355B7AC7AF8116556EB3F4CEF3D">1</mods:number>
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<mods:start id="4357EF6CB4EA8E3DA20CA7E0FAB17AEE">1</mods:start>
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<mods:end id="9C112248B219CD88E5A3B1A81EC7146E">264</mods:end>
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<mods:classification id="118D3114F7F75F26729DCE3C205A5F64">journal article</mods:classification>
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<mods:identifier id="A648FFE52A5C22F2D18D3AE49949CCF8" type="CLB-Dataset">31639</mods:identifier>
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<mods:identifier id="2D619DFF3572C11EEE2710F475A55CDE" type="DOI">10.11646/zootaxa.4342.1.1</mods:identifier>
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<mods:identifier id="13F781D05732D31CE62AE7A767C2E609" type="GBIF-Dataset">15799d2a-1968-4874-8100-452597d4e26b</mods:identifier>
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<mods:identifier id="05EE2F1DED70EBB447D7FDE93269CCEA" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="2DDB6B3EF5C203D697A715BDDBD03B6C" type="Zenodo-Dep">1041066</mods:identifier>
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<mods:identifier id="61EDE8B7A91438F96B48040ADC1C8A11" type="ZooBank">2ACD54D2-3487-432D-9323-EEC131FE2E64</mods:identifier>
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</mods:mods>
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<treatment id="038587BBE33AFFAEFF75FF3DB843FD2D" ID-DOI="http://doi.org/10.5281/zenodo.5323518" ID-GBIF-Taxon="137233464" ID-Zenodo-Dep="5323518" LSID="urn:lsid:plazi:treatment:038587BBE33AFFAEFF75FF3DB843FD2D" httpUri="http://treatment.plazi.org/id/038587BBE33AFFAEFF75FF3DB843FD2D" lastPageId="63" lastPageNumber="64" pageId="61" pageNumber="62">
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<subSubSection id="C3366526E33AFFACFF75FF3DBBB4FF56" pageId="61" pageNumber="62" type="nomenclature">
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<paragraph id="8B9336ADE33AFFACFF75FF3DBB6DFF37" blockId="61.[151,648,152,211]" box="[151,593,152,178]" pageId="61" pageNumber="62">
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<heading id="D0DB81C1E33AFFACFF75FF3DBB6DFF37" bold="true" box="[151,593,152,178]" fontSize="11" level="1" pageId="61" pageNumber="62" reason="1">
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<emphasis id="B958EABFE33AFFACFF75FF3DBB6DFF37" bold="true" box="[151,593,152,178]" pageId="61" pageNumber="62">
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<taxonomicName id="4C2C4D2EE33AFFACFF75FF3DB88EFF37" ID-CoL="3N3M4" authority="Perkins, 2017" authorityName="Perkins" authorityYear="2017" box="[151,434,152,178]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="multiarcuata" status="sp. nov.">
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<emphasis id="B958EABFE33AFFACFF75FF3DB88EFF37" bold="true" box="[151,434,152,178]" italics="true" pageId="61" pageNumber="62">Hydraena multiarcuata</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26B57C4E33AFFACFE23FF3DBB6DFF37" box="[449,593,152,178]" pageId="61" pageNumber="62" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9336ADE33AFFACFF75FF1FBBB4FF56" blockId="61.[151,648,152,211]" box="[151,648,186,211]" pageId="61" pageNumber="62">
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<figureCitation id="13172A28E33AFFACFF75FF1FB9CDFF56" box="[151,241,186,211]" captionStart="FIGURE 50" captionStartId="61.[151,250,1984,2006]" captionText="FIGURE 50. Hydraena multiarcuata, holotype, habitus dorsal, lateral, and ventral." pageId="61" pageNumber="62">Figs. 50</figureCitation>
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(habitus), 52 (aedeagus), 251 (map)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E33AFFACFF75FEA7BBAAFEBA" pageId="61" pageNumber="62" type="materials_examined">
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<paragraph id="8B9336ADE33AFFACFF75FEA7BBAAFEBA" blockId="61.[151,1437,258,895]" pageId="61" pageNumber="62">
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<materialsCitation id="3B443CF0E33AFFACFF75FEA7BBAAFEBA" ID-GBIF-Occurrence="1697157543" collectingDate="2004-11-01" collectingDateMax="2004-12-30" collectingDateMin="2004-11-01" collectionCode="ZSMC" collectorName="M. Balke" elevation="1123" latitude="-21.266666" location="Sahamalaotra" longLatPrecision="1264" longitude="47.416668" municipality="Ranomafana" pageId="61" pageNumber="62" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
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<emphasis id="B958EABFE33AFFACFF75FEA7B887FE9E" bold="true" box="[151,443,258,283]" pageId="61" pageNumber="62">
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<typeStatus id="5497880FE33AFFACFF75FEA7B9EEFE9E" box="[151,210,258,283]" pageId="61" pageNumber="62">Type</typeStatus>
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Material.
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<typeStatus id="5497880FE33AFFACFEACFEA7B887FE9E" box="[334,443,258,283]" pageId="61" pageNumber="62" type="holotype">Holotype</typeStatus>
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</emphasis>
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(male):
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<collectingRegion id="49E8F84FE33AFFACFDF8FEA7BB85FE9E" box="[538,697,258,283]" country="Madagascar" name="Fianarantsoa" pageId="61" pageNumber="62">
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<emphasis id="B958EABFE33AFFACFDF8FEA7BB85FE9E" bold="true" box="[538,697,258,283]" pageId="61" pageNumber="62">Fianarantsoa</emphasis>
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</collectingRegion>
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,
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<collectingMunicipality id="6BF7ACD7E33AFFACFD25FEA7BA64FE9E" box="[711,856,258,283]" pageId="61" pageNumber="62">Ranomafana</collectingMunicipality>
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,
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<location id="8EF36076E33AFFACFC80FEA7BAC1FE9E" LSID="urn:lsid:plazi:treatment:038587BBE33AFFAEFF75FF3DB843FD2D:8EF36076E33AFFACFC80FEA7BAC1FE9E" box="[866,1021,258,283]" latitude="-21.266666" longLatPrecision="1264" longitude="47.416668" municipality="Ranomafana" name="Sahamalaotra" pageId="61" pageNumber="62" stateProvince="Fianarantsoa">Sahamalaotra</location>
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, elev.
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<elevation id="0001D19EE33AFFACFBA1FEA6BDA5FE9F" box="[1091,1177,259,283]" metricMagnitude="3" metricUnit="m" metricValue="1.123" pageId="61" pageNumber="62" unit="m" value="1123.0">
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<quantity id="4CD49B48E33AFFACFBA1FEA6BDA5FE9F" box="[1091,1177,259,283]" metricMagnitude="3" metricUnit="m" metricValue="1.123" pageId="61" pageNumber="62" unit="m" value="1123.0">1123 m</quantity>
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</elevation>
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,
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<geoCoordinate id="EE18506AE33AFFACFB47FEA6BC30FE9E" box="[1189,1292,258,283]" direction="south" orientation="latitude" pageId="61" pageNumber="62" precision="925" value="-21.266666">21° 16' S</geoCoordinate>
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,
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<geoCoordinate id="EE18506AE33AFFACFAFBFEA6BCBFFE9F" box="[1305,1411,258,283]" direction="east" orientation="longitude" pageId="61" pageNumber="62" precision="925" value="47.416668">47° 25' E</geoCoordinate>
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,
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<collectingDate id="EFD6E985E33AFFACFA6CFEA6B801FEBA" pageId="61" pageNumber="62" value="2004-11-01" valueMax="2004-12-30" valueMin="2004-11-01">
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<date id="FF92106DE33AFFACFA6CFEA6B801FEBA" pageId="61" pageNumber="62" value="2004-11-01" valueMax="2004-12-30" valueMin="2004-11-01">1 xi–30 xii 2004</date>
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</collectingDate>
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,
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<collectorName id="26D9537BE33AFFACFEAAFE82B88CFEBA" box="[328,432,294,319]" pageId="61" pageNumber="62">M. Balke</collectorName>
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(MD 031) (
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<collectionCode id="ED3DAE68E33AFFACFDD5FE82BBB6FEBA" box="[567,650,295,319]" country="Germany" httpUri="http://grbio.org/cool/a6ms-py1z" name="Zoologische Staatssammlung" pageId="61" pageNumber="62">ZSMC</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E33AFFACFF25FEEFB8DCFE4A" pageId="61" pageNumber="62" type="diagnosis">
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<paragraph id="8B9336ADE33AFFACFF25FEEFB8DCFE4A" blockId="61.[151,1437,258,895]" pageId="61" pageNumber="62">
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<emphasis id="B958EABFE33AFFACFF25FEEFB8E9FEE6" bold="true" box="[199,469,330,355]" pageId="61" pageNumber="62">Differential Diagnosis.</emphasis>
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Differentiated from other members of the species group by the combination of the arcuate meso- and metatibiae, the arcuate metaventral plaques, and the arcuate elytra. The aedeagus perhaps has some very superficial resemblance to that of
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<taxonomicName id="4C2C4D2EE33AFFACFD74FE31BA7FFE2E" authorityName="Perkins" authorityYear="2017" box="[662,835,403,427]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="ranomafana">
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<emphasis id="B958EABFE33AFFACFD74FE31BA7FFE2E" box="[662,835,403,427]" italics="true" pageId="61" pageNumber="62">H. ranomafana</emphasis>
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</taxonomicName>
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, but differs in many respects of both distal and main piece, and size (
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<figureCitation id="13172A28E33AFFACFEA8FE13B894FE4A" box="[330,424,438,463]" captionStart="FIGURES 51 – 52" captionStartId="62.[151,264,1876,1899]" captionText="FIGURES 51 – 52. Hydraena, aedeagi of holotypes." pageId="61" pageNumber="62">Figs. 52</figureCitation>
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,
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<figureCitation id="13172A28E33AFFACFE56FE13B8EFFE4A" box="[436,467,438,463]" captionStart="FIGURES 55 – 56" captionStartId="66.[151,264,1638,1661]" captionText="FIGURES 55 – 56. Hydraena, aedeagi of holotypes." pageId="61" pageNumber="62">55</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E33AFFAEFF25FE7FBAD0FDE5" lastPageId="63" lastPageNumber="64" pageId="61" pageNumber="62" type="description">
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<paragraph id="8B9336ADE33AFFACFF25FE7FBA37FD92" blockId="61.[151,1437,258,895]" pageId="61" pageNumber="62">
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<emphasis id="B958EABFE33AFFACFF25FE7FB86DFE76" bold="true" box="[199,337,474,499]" pageId="61" pageNumber="62">Description</emphasis>
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. Size:
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<typeStatus id="5497880FE33AFFACFE41FE7FBB39FE76" box="[419,517,474,499]" pageId="61" pageNumber="62" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.55/0.70; head width 0.37; pronotum 0.35/0.52, PA 0.41, PB 0.44; elytra 0.93/0.70.
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</paragraph>
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<paragraph id="8B9336ADE33AFFACFF75FD87B8C9FD06" blockId="61.[151,1437,258,895]" pageId="61" pageNumber="62">
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<typeStatus id="5497880FE33AFFACFF75FD87B9C3FDBE" box="[151,255,546,571]" pageId="61" pageNumber="62" type="holotype">Holotype</typeStatus>
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teneral, mature color most likely with dorsum reddish brown, head darker than pronotum (piceous in darkest specimens); pronotum lighter than elytra; legs light brown to testaceous; maxillary palpi testaceous, distal ½ of last palpomere not darker.
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</paragraph>
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<paragraph id="8B9336ADE33AFFACFF25FD2BBB42FCB2" blockId="61.[151,1437,258,895]" pageId="61" pageNumber="62">Head with clypeus finely sparsely punctate medially, laterally very weakly microreticulate; frons more coarsely punctate laterally than medially where interstices ca. 1–4xpd. Pronotum in anterior and posterior 1/3 more densely punctate than on disc, interstices ca. 0.5–1xpd; punctures on disc distinctly smaller, interstices ca. 1–3xpd, shining. Mentum finely microreticulate, dull; postmentum finely microreticulate medially. Genae raised, shining, with posterior ridge, interrupted in midline.</paragraph>
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<paragraph id="8B9336ADE33AFFACFF25FCE6BAE5FCFA" blockId="61.[151,1437,258,895]" pageId="61" pageNumber="62">Pronotum bisinuate laterally; anterior margin straight behind eyes, emarginate behind frons; PF1 and PF2 very shallow, almost obsolete; PF3 and PF4 deep and very shallowly confluent.</paragraph>
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<caption id="DF536625E33AFFACFF75F865BADFF853" box="[151,995,1984,2006]" pageId="61" pageNumber="62">
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<paragraph id="8B9336ADE33AFFACFF75F865BADFF853" blockId="61.[151,995,1984,2006]" box="[151,995,1984,2006]" pageId="61" pageNumber="62">
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<emphasis id="B958EABFE33AFFACFF75F865B81CF850" bold="true" box="[151,288,1984,2006]" pageId="61" pageNumber="62">FIGURE 50.</emphasis>
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<taxonomicName id="4C2C4D2EE33AFFACFEC7F865BB2FF853" authorityName="Perkins" authorityYear="2017" box="[293,531,1984,2006]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="multiarcuata">
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<emphasis id="B958EABFE33AFFACFEC7F865BB2FF853" box="[293,531,1984,2006]" italics="true" pageId="61" pageNumber="62">Hydraena multiarcuata</emphasis>
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</taxonomicName>
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, holotype, habitus dorsal, lateral, and ventral.
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</paragraph>
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</caption>
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<caption id="DF536625E339FFAFFF75F8F1BB98F8EE" box="[151,676,1876,1899]" pageId="62" pageNumber="63">
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<paragraph id="8B9336ADE339FFAFFF75F8F1BB98F8EE" blockId="62.[151,676,1876,1899]" box="[151,676,1876,1899]" pageId="62" pageNumber="63">
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<emphasis id="B958EABFE339FFAFFF75F8F1B86FF8EF" bold="true" box="[151,339,1876,1899]" pageId="62" pageNumber="63">FIGURES 51–52.</emphasis>
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<taxonomicName id="4C2C4D2EE339FFAFFEB8F8F0B883F8EE" box="[346,447,1877,1899]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EABFE339FFAFFEB8F8F0B883F8EE" box="[346,447,1877,1899]" italics="true" pageId="62" pageNumber="63">Hydraena</emphasis>
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</taxonomicName>
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, aedeagi of holotypes.
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</paragraph>
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</caption>
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<paragraph id="8B9336ADE338FFAEFF25FF32BDA6FE98" blockId="63.[151,1436,151,680]" pageId="63" pageNumber="64">Elytra weakly arcuate laterally; summit of steep posterior declivity slightly before midlength; lateral explanate margins wide; serial punctures round, large and deeply impressed, especially on basal 0.5, separated by ca. 1xpd, larger than largest pronotal punctures. Intervals not raised, shining, ca. 0.5–1xpd. Apices in dorsal aspect rather sharply conjointly rounded, in posterior aspect margins not forming angle with one another.</paragraph>
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<paragraph id="8B9336ADE338FFAEFF25FE82BAD0FDE5" blockId="63.[151,1436,151,680]" pageId="63" pageNumber="64">
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Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/2/2. P1 as wide as P2; median carina very weakly sinuate in profile. P2 l/w ca. 2/1, sides parallel, apex blunt and on same level as anteromedian carina of metaventrite. Plaques narrow, subcarinate, arcuate and slightly converging anteriorly, separated basally by ca. 2–
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<date id="FF92106DE338FFAEFA60FED5BCA0FE0D" box="[1410,1436,368,392]" pageId="63" pageNumber="64">3x</date>
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plaque width, located at sides of median depression. Metaventrite anteromedian carina slightly longer than plaques, attaining anterior level of plaques; long longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at arcuate posterior margin ca.
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<date id="FF92106DE338FFAEFCA9FE79BA5AFE71" box="[843,870,476,500]" pageId="63" pageNumber="64">2x</date>
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P2. Protibia slightly arcuate. Mesotibia distinctly arcuate, medial margin with 4 or 5 short sharp spines, spines longer and stronger from most proximal to most distal, surface of tibia notched between spines. Metatibia arcuate, slender. Metafemur slender, arcuate on lateral margin, sinuate on medial margin. Abdominal apex with narrow apicomedian notch.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E338FFAEFF25FDC9B843FD2D" pageId="63" pageNumber="64" type="etymology">
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<paragraph id="8B9336ADE338FFAEFF25FDC9B843FD2D" blockId="63.[151,1436,151,680]" pageId="63" pageNumber="64">
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<emphasis id="B958EABFE338FFAEFF25FDC9B872FD00" bold="true" box="[199,334,620,645]" pageId="63" pageNumber="64">Etymology.</emphasis>
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Named in reference to the arcuate meso- and metatibiae, the arcuate elytra, and the arcuate metaventral plaques.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |