192 lines
22 KiB
XML
192 lines
22 KiB
XML
<document id="0DCC48BEC1FED7FE363EBFC0E15AB817" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE317FF81FF75FD87BBB1F8A5" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena Micromadraena" docType="treatment" docVersion="9" lastPageNumber="17" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="17" updateTime="1698480684898" updateUser="ExternalLinkService">
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<mods:titleInfo id="4C14642FBD0C90B1645FFBDCF4C24121">
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<mods:title id="5A96400C94945BAFF3C0443AF46323D8">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
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<mods:name id="622D4595AD7AA1ADF8EFDB05E7F70D9B" type="personal">
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<mods:namePart id="F6C0F180EC56DB1BA9BE3CD7BF05043C">Perkins, Philip D.</mods:namePart>
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<mods:title id="AB1931C563C15E501527CE87A5BD4471">Zootaxa</mods:title>
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<mods:part id="8CE50F8C78959F9516648B6785A1377F">
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<mods:date id="2CD8E42315627573574F4C2F543911DA">2017</mods:date>
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<mods:number id="178BBEBBE1AC16E0C49A5F93DD1AA36F">2017-11-03</mods:number>
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<mods:number id="232E5564ABDA78ABF042ACBE8BF4D8FC">4342</mods:number>
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<mods:identifier id="4B1859B56088AC5A4FB906600788508C" type="DOI">10.11646/zootaxa.4342.1.1</mods:identifier>
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<treatment id="038587BBE317FF81FF75FD87BBB1F8A5" ID-DOI="http://doi.org/10.5281/zenodo.5323377" ID-GBIF-Taxon="137233415" ID-Zenodo-Dep="5323377" LSID="urn:lsid:plazi:treatment:038587BBE317FF81FF75FD87BBB1F8A5" httpUri="http://treatment.plazi.org/id/038587BBE317FF81FF75FD87BBB1F8A5" lastPageNumber="17" pageId="16" pageNumber="17">
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<subSubSection id="C3366526E317FF81FF75FD87BB96FDB9" box="[151,682,546,572]" pageId="16" pageNumber="17" type="nomenclature">
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<paragraph id="8B9336ADE317FF81FF75FD87BB96FDB9" blockId="16.[151,682,546,572]" box="[151,682,546,572]" pageId="16" pageNumber="17">
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<heading id="D0DB81C1E317FF81FF75FD87BB96FDB9" bold="true" box="[151,682,546,572]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
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<emphasis id="B958EABFE317FF81FF75FD87BB96FDB9" bold="true" box="[151,682,546,572]" pageId="16" pageNumber="17">
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<taxonomicName id="4C2C4D2EE317FF81FF75FD87B8D2FDB9" ID-CoL="8VRR8" authority="(Micromadraena)" authorityName="Micromadraena" box="[151,494,546,572]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EABFE317FF81FF75FD87B82CFDB9" bold="true" box="[151,272,546,572]" italics="true" pageId="16" pageNumber="17">Hydraena</emphasis>
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||
(
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<emphasis id="B958EABFE317FF81FEC2FD87B8DFFDB9" bold="true" box="[288,483,546,572]" italics="true" pageId="16" pageNumber="17">Micromadraena</emphasis>
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)
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</taxonomicName>
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, new subgenus
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E317FF81FF75FDCFBBFDF95D" pageId="16" pageNumber="17" type="materials_examined">
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<paragraph id="8B9336ADE317FF81FF75FDCFBA78FDFA" blockId="16.[151,836,617,639]" box="[151,836,617,639]" pageId="16" pageNumber="17">
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Type Species:
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<taxonomicName id="4C2C4D2EE317FF81FEC8FDCCBBF8FDFA" authority="Perkins, 2017" authorityName="Perkins" authorityYear="2017" box="[298,708,617,639]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="parvipalpis" status="sp. nov." subGenus="Micromadraena">
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<emphasis id="B958EABFE317FF81FEC8FDCCB8B3FDFA" box="[298,399,617,639]" italics="true" pageId="16" pageNumber="17">Hydraena</emphasis>
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(
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<emphasis id="B958EABFE317FF81FE7CFDCCBB7EFDFA" box="[414,578,617,639]" italics="true" pageId="16" pageNumber="17">Micromadraena</emphasis>
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)
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<emphasis id="B958EABFE317FF81FDB2FDCCBBF8FDFA" box="[592,708,617,639]" italics="true" pageId="16" pageNumber="17">parvipalpis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26B57C4E317FF81FD28FDCFBA78FDFA" box="[714,836,617,639]" pageId="16" pageNumber="17" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8B9336ADE317FF81FF75FD09B867FD62" blockId="16.[151,1436,684,1824]" pageId="16" pageNumber="17">
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This unusual group comprises six species:
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<emphasis id="B958EABFE317FF81FD90FD08B868FD6D" italics="true" pageId="16" pageNumber="17">
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<taxonomicName id="4C2C4D2EE317FF81FD90FD08BA2AFD40" authorityName="Perkins" authorityYear="2017" box="[626,790,684,709]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="parvipalpis">H. parvipalpis</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE317FF81FCC2FD08BAA6FD41" authorityName="Perkins" authorityYear="2017" box="[800,922,684,708]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="fortipes">H. fortipes</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE317FF81FC47FD08BD13FD40" authorityName="Perkins" authorityYear="2017" box="[933,1071,684,709]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="genuvela">H. genuvela</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE317FF81FBDBFD08BDE0FD41" authorityName="Perkins" authorityYear="2017" box="[1081,1244,685,708]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="serripennis">H. serripennis</taxonomicName>
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,
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<taxonomicName id="4C2C4D2EE317FF81FB04FD08BCA4FD40" authorityName="Perkins" authorityYear="2017" box="[1254,1432,684,709]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rubridentata">H. rubridentata</taxonomicName>
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, and
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<taxonomicName id="4C2C4D2EE317FF81FF2AFD74B868FD6D" authorityName="Perkins" authorityYear="2017" box="[200,340,719,744]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="breviceps">H. breviceps</taxonomicName>
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</emphasis>
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.
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</paragraph>
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<paragraph id="8B9336ADE317FF81FF25FD51BB03FBA9" blockId="16.[151,1436,684,1824]" pageId="16" pageNumber="17">
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All of the specimens in this group have been collected from "sifted litter (leaf mold, rotten wood)", are nonaquatic, and have a distinctive habitus related to the microhabitat. The maxillary palpi and legs are short, and can consequently be hidden beneath the laterally widened head and the very wide elytral explanate margin. The labrum is small, directed ventrad, and hidden in dorsal view. Despite these modifications, the derived characters of
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<taxonomicName id="4C2C4D2EE317FF81FF75FC21B83BFC18" box="[151,263,900,925]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EABFE317FF81FF75FC21B83BFC18" box="[151,263,900,925]" italics="true" pageId="16" pageNumber="17">Hydraena</emphasis>
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</taxonomicName>
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are still present (see
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<bibRefCitation id="EFBD4B5CE317FF81FDF2FC21BB90FC19" author="Perkins" box="[528,684,900,925]" pageId="16" pageNumber="17" refString="Perkins, P. D. (1989) Adelphydraena, new genus, and two new species from Venezuela, and remarks on phylogenetic relationships within the subtribe Hydraenina (Coleoptera: Hydraenidae). Proceedings of the Biological Society of Washington, 102, 447 - 457." type="journal article" year="1989">Perkins 1989</bibRefCitation>
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,
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<bibRefCitation id="EFBD4B5CE317FF81FD5EFC21BBC5FC19" author="Perkins" box="[700,761,900,924]" pageId="16" pageNumber="17" refString="Perkins, P. D. (1997) Life on the effective bubble: exocrine secretion delivery systems (ESDS) and the evolution and classification of beetles in the family Hydraenidae (Insecta: Coleoptera). Annals of Carnegie Museum, 66 (2), 89 - 207." type="journal article" year="1997">1997</bibRefCitation>
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): 1) the labrum, although reduced in size, still forms an interlocking device with the mandibles; 2) the procoxal cavities are closed behind by the widened apex of the intercoxal process; 3) the mentum is produced in the midline; 4) the hypomeron has the characteristic
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<taxonomicName id="4C2C4D2EE317FF81FACEFC69BCA0FC60" box="[1324,1436,972,997]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EABFE317FF81FACEFC69BCA0FC60" box="[1324,1436,972,997]" italics="true" pageId="16" pageNumber="17">Hydraena</emphasis>
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</taxonomicName>
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morphology; 5) females have a pair of sensory clusters on the gonocoxite; and 6) the spermatheca does not have unusual morphology (
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<figureCitation id="13172A28E317FF81FE6FFBB1BB0FFBA9" box="[397,563,1044,1069]" captionStart="FIGURES 228 – 232" captionStartId="227.[151,264,1701,1724]" captionTargetBox="[297,1284,202,1671]" captionTargetId="figure@227.[283,1303,193,1681]" captionTargetPageId="227" captionText="FIGURES 228 – 232. Hydraena species, female gonocoxites, spermathecae, antenna and maxillary palpus." httpUri="https://zenodo.org/record/1041150/files/figure.png" pageId="16" pageNumber="17">Figs. 231, 232</figureCitation>
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).
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</paragraph>
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<paragraph id="8B9336ADE317FF81FF25FB92BC6FFA3D" blockId="16.[151,1436,684,1824]" pageId="16" pageNumber="17">
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The unusual reduced appendage morphology is matched by the unusual aedeagal morphology: the aedeagus has very short parameres, which originate from the base; each paramere has two short setae. Very short parameres with reduced number of setae and having the origination basally appear in a few other groups (
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<figureCitation id="13172A28E317FF81FB54FBDABC25FB1D" box="[1206,1305,1151,1176]" captionStart="FIGURES 344 – 345. 344" captionStartId="261.[151,264,1715,1738]" captionTargetBox="[266,1317,241,1662]" captionTargetId="figure@261.[226,1360,193,1694]" captionTargetPageId="261" captionText="FIGURES 344 – 345. 344. Modified basal portion of DNA sequence data phylogeny of Hydraena, showing identity of Madagascar species, and new subgenera. Used by permission of Ignacio Ribera. 345. Three examples of aedeagi having very short parameres that originate near the basal orifice, and have only one or two setae." httpUri="https://zenodo.org/record/1041202/files/figure.png" pageId="16" pageNumber="17">Fig. 345</figureCitation>
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):
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<taxonomicName id="4C2C4D2EE317FF81FACEFBDAB9CAFB38" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="monikae">
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<emphasis id="B958EABFE317FF81FACEFBDAB9CAFB38" italics="true" pageId="16" pageNumber="17">Hydraena monikae</emphasis>
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</taxonomicName>
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group (
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<bibRefCitation id="EFBD4B5CE317FF81FEADFB01BA07FB38" author="Perkins" box="[335,827,1188,1213]" pageId="16" pageNumber="17" refString="Perkins, P. D. (2014) A revision of the water beetle genus Hydraena Kugelann for southern Africa (Coleoptera: Hydraenidae). Zootaxa, 3758 (1), 1 - 92. https: // doi. org / 10.11646 / zootaxa. 3758.1.1" type="journal article" year="2014">
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Perkins 2014; four species in
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<collectingCountry id="F33B763DE317FF81FD44FB01BA07FB38" box="[678,827,1188,1213]" name="South Africa" pageId="16" pageNumber="17">
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<collectingRegion id="49E8F84FE317FF81FD44FB01BBDBFB38" box="[678,743,1188,1213]" country="Tanzania" name="Kusini Unguja" pageId="16" pageNumber="17">South</collectingRegion>
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Africa
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</collectingCountry>
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</bibRefCitation>
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), and some
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<taxonomicName id="4C2C4D2EE317FF81FC27FB01BD5EFB38" authorityName="Perkins" authorityYear="1997" box="[965,1122,1188,1213]" class="Insecta" family="Hydraenidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="tribe" tribe="Parhydraenini">Parhydraenini</taxonomicName>
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including
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<taxonomicName id="4C2C4D2EE317FF81FB3FFB01BCA0FB38" authorityName="Perkins" authorityYear="2009" box="[1245,1436,1188,1213]" class="Insecta" family="Hydraenidae" genus="Parhydraenopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EABFE317FF81FB3FFB01BCA0FB38" box="[1245,1436,1188,1213]" italics="true" pageId="16" pageNumber="17">Parhydraenopsis</emphasis>
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</taxonomicName>
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species in northeast Africa, and
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<taxonomicName id="4C2C4D2EE317FF81FDEFFB6CBB8DFB65" box="[525,689,1225,1248]" class="Insecta" family="Hydraenidae" genus="Protozantaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EABFE317FF81FDEFFB6CBB8DFB65" box="[525,689,1225,1248]" italics="true" pageId="16" pageNumber="17">Protozantaena</emphasis>
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</taxonomicName>
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species in
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<collectingCountry id="F33B763DE317FF81FCD4FB62BA81FB65" box="[822,957,1223,1248]" name="Madagascar" pageId="16" pageNumber="17">Madagascar</collectingCountry>
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and southern Africa (
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<bibRefCitation id="EFBD4B5CE317FF81FB5CFB62BC67FB65" author="Perkins" box="[1214,1371,1223,1248]" pageId="16" pageNumber="17" refString="Perkins, P. D. (2009) Revisions of the genera Parhydraena Orchymont, Protozantaena Perkins, Decarthrocerus Orchymont, and Parhydraenopsis nomen novum, aquatic and humicolous beetles from Africa and Madagascar, and comparative morphology of the tribe Parhydraenini (Coleoptera: Hydraenidae). Zootaxa, 2038, 1 - 119." type="journal article" year="2009">Perkins 2009</bibRefCitation>
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). The shape of the basal orifice in
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<taxonomicName id="4C2C4D2EE317FF81FDF3FB49BB92FA80" authorityName="Perkins" authorityYear="1997" box="[529,686,1260,1285]" class="Insecta" family="Hydraenidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="tribe" tribe="Parhydraenini">Parhydraenini</taxonomicName>
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is characteristic and differs distinctively from the shape in
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<emphasis id="B958EABFE317FF81FF75FAAAB870FAAD" box="[151,332,1295,1320]" italics="true" pageId="16" pageNumber="17">Micromadraena</emphasis>
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, which is more like that present in other
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<taxonomicName id="4C2C4D2EE317FF81FCA6FAAABA88FAAD" box="[836,948,1295,1320]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EABFE317FF81FCA6FAAABA88FAAD" box="[836,948,1295,1320]" italics="true" pageId="16" pageNumber="17">Hydraena</emphasis>
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</taxonomicName>
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. The similarity in the short parameres is discussed in the
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<taxonomicName id="4C2C4D2EE317FF81FEADFA91B883FAC8" box="[335,447,1332,1357]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EABFE317FF81FEADFA91B883FAC8" box="[335,447,1332,1357]" italics="true" pageId="16" pageNumber="17">Hydraena</emphasis>
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</taxonomicName>
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subgenera section. In short, it is possible that
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<emphasis id="B958EABFE317FF81FC24FA91BD47FAC8" box="[966,1147,1332,1357]" italics="true" pageId="16" pageNumber="17">Micromadraena</emphasis>
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is more closely related to the
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<taxonomicName id="4C2C4D2EE317FF81FF21FAF2B8A0FAF5" box="[195,412,1367,1392]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="monikae">
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<emphasis id="B958EABFE317FF81FF21FAF2B8A0FAF5" box="[195,412,1367,1392]" italics="true" pageId="16" pageNumber="17">Hydraena monikae</emphasis>
|
||
</taxonomicName>
|
||
group than to the other
|
||
<collectingCountry id="F33B763DE317FF81FD59FAF2BA7EFAF5" box="[699,834,1367,1392]" name="Madagascar" pageId="16" pageNumber="17">Madagascar</collectingCountry>
|
||
<taxonomicName id="4C2C4D2EE317FF81FCAEFAF2BA80FAF5" box="[844,956,1367,1392]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B958EABFE317FF81FCAEFAF2BA80FAF5" box="[844,956,1367,1392]" italics="true" pageId="16" pageNumber="17">Hydraena</emphasis>
|
||
</taxonomicName>
|
||
. The aedeagi in both of these groups are similar in having a main piece that is tubular in cross section, a distal piece that is a rather simple, a rigid flagellum, a similarly shaped basal orifice, and the very short parameres. In habitus the two groups are quite dissimilar.
|
||
</paragraph>
|
||
<paragraph id="8B9336ADE317FF81FF25FA61BCB1F985" blockId="16.[151,1436,684,1824]" pageId="16" pageNumber="17">
|
||
Both sexes of
|
||
<emphasis id="B958EABFE317FF81FE8CFA60B8B6FA59" box="[366,394,1477,1500]" italics="true" pageId="16" pageNumber="17">H.</emphasis>
|
||
(
|
||
<emphasis id="B958EABFE317FF81FE7EFA61BB6DFA58" box="[412,593,1476,1501]" italics="true" pageId="16" pageNumber="17">Micromadraena</emphasis>
|
||
) have very wide explanate elytral margins, males having the edges more strongly serrate than those of females. The femoral-tibial articulations are shielded by the wide explanate margin.
|
||
</paragraph>
|
||
<paragraph id="8B9336ADE317FF81FF25F9A9BBFDF95D" blockId="16.[151,1436,684,1824]" pageId="16" pageNumber="17">
|
||
The morphology of concealed appendages and thick cuticle of species in this group could possibly be termed an anti-ant defense system; the collector of most of these hydraenids, Brian L. Fisher, was targeting litter dwelling ants. There are other groups of terrestrial
|
||
<taxonomicName id="4C2C4D2EE317FF81FD61F9F1BBCFF9E8" box="[643,755,1620,1645]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B958EABFE317FF81FD61F9F1BBCFF9E8" box="[643,755,1620,1645]" italics="true" pageId="16" pageNumber="17">Hydraena</emphasis>
|
||
</taxonomicName>
|
||
that have shortened appendages, for example undescribed species in New Caledonia (Perkins, in prep.), but these species have a very different aedeagus, indicating relationship with
|
||
<emphasis id="B958EABFE317FF81FE80F938B841F931" box="[354,381,1693,1716]" italics="true" pageId="16" pageNumber="17">H.</emphasis>
|
||
(
|
||
<taxonomicName id="4C2C4D2EE317FF81FE73F939BB10F930" box="[401,556,1692,1717]" class="Insecta" family="Hydraenidae" genus="Hydraenopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B958EABFE317FF81FE73F939BB10F930" box="[401,556,1692,1717]" italics="true" pageId="16" pageNumber="17">Hydraenopsis</emphasis>
|
||
</taxonomicName>
|
||
). Unfortunately, no specimens collected in absolute alcohol were available; DNA sequence data must await more collections.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3366526E317FF81FF25F941BBB1F8A5" pageId="16" pageNumber="17" type="etymology">
|
||
<paragraph id="8B9336ADE317FF81FF25F941BBB1F8A5" blockId="16.[151,1436,684,1824]" pageId="16" pageNumber="17">
|
||
<emphasis id="B958EABFE317FF81FF25F941B872F978" bold="true" box="[199,334,1764,1789]" pageId="16" pageNumber="17">Etymology.</emphasis>
|
||
Named in reference to the short appendages, the short parameres, the geographical distribution (
|
||
<collectingCountry id="F33B763DE317FF81FF7CF8A2B817F8A5" box="[158,299,1799,1824]" name="Madagascar" pageId="16" pageNumber="17">Madagascar</collectingCountry>
|
||
), and relationship to
|
||
<taxonomicName id="4C2C4D2EE317FF81FDF4F8A2BBBAF8A5" box="[534,646,1799,1824]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B958EABFE317FF81FDF4F8A2BBBAF8A5" box="[534,646,1799,1824]" italics="true" pageId="16" pageNumber="17">Hydraena</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |