159 lines
20 KiB
XML
159 lines
20 KiB
XML
<document id="AE521CA3A7CA739B61CAE0CC9B31B8B8" ID-CLB-Dataset="45537" ID-DOI="10.5281/zenodo.279941" ID-GBIF-Dataset="9841826a-d893-45f3-9bc6-f4396bf43661" ID-ISSN="1175-5326" ID-Zenodo-Dep="279941" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460434601243" checkinUser="plazi" docAuthor="Høisaeter, Tore" docDate="2012" docId="038587B47E5065198487941933CEFEC8" docLanguage="en" docName="zt03178p067.pdf" docOrigin="Zootaxa 3178" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cimaria vargasi Høisaeter, 2012, sp. nov." docType="treatment" docVersion="9" lastPageNumber="66" masterDocId="FFBCFFCC7E52651A841096063778FFAE" masterDocTitle="Cimaria vargasi n. gen, n. sp. (Gastropoda: Pyramidellidae: Odostomiinae) from the Pacific Coast of Costa Rica, Central America" masterLastPageNumber="67" masterPageNumber="63" pageNumber="65" updateTime="1698331887839" updateUser="ExternalLinkService">
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<mods:mods id="76C45649032CBB9F73A09E4C774A3BF7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5EE474F34B95C3AD30CDAA2C825EF75F">
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<mods:title id="0934B10FB9AC9695E9EFECA7C514ACE4">Cimaria vargasi n. gen, n. sp. (Gastropoda: Pyramidellidae: Odostomiinae) from the Pacific Coast of Costa Rica, Central America</mods:title>
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<mods:name id="79ABEE0D87D7A58EA0DA21198397F2A4" type="personal">
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<mods:role id="5B48720786D44402DCDE3E567AF8D821">
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<mods:roleTerm id="08E2CE3E66F48322ED894E3D2870C591">Author</mods:roleTerm>
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<mods:namePart id="4333FA013627FECEF4C0D77F709C1CBB">Høisaeter, Tore</mods:namePart>
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<mods:typeOfResource id="25B4FA93CEFE75BD89740B3ABB83F154">text</mods:typeOfResource>
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<mods:title id="D0EDC8E1A1590312D2D5041BACFB5E08">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1A0232E457140A4E6BD1283C37E27D70">
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<mods:date id="2D45682DE1D4B72B30FEE780C1DB334A">2012</mods:date>
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<mods:number id="8D0178D0FFD502628E79A73E206A1D46">3178</mods:number>
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<mods:classification id="8FAE173248CCBBD062E2A75232ED11A3">journal article</mods:classification>
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<mods:identifier id="A2F9109CBE0CA57DA2B14F090BD41E7B" type="CLB-Dataset">45537</mods:identifier>
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<mods:identifier id="CA75CD57288FA223E6214895C8736DE5" type="DOI">10.5281/zenodo.279941</mods:identifier>
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<mods:identifier id="498CE54B3B68A6ACEB94A9833D7E0D09" type="GBIF-Dataset">9841826a-d893-45f3-9bc6-f4396bf43661</mods:identifier>
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<mods:identifier id="DB693D4535373F6758693D23661EDA02" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="8AC008605113E81767DF82B793B571BA" type="Zenodo-Dep">279941</mods:identifier>
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<treatment id="038587B47E5065198487941933CEFEC8" ID-DOI="http://doi.org/10.5281/zenodo.5689065" ID-GBIF-Taxon="119431406" ID-Zenodo-Dep="5689065" LSID="urn:lsid:plazi:treatment:038587B47E5065198487941933CEFEC8" httpUri="http://treatment.plazi.org/id/038587B47E5065198487941933CEFEC8" lastPageId="3" lastPageNumber="66" pageId="2" pageNumber="65">
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<subSubSection id="C33665297E506518848794193624FDF4" pageId="2" pageNumber="65" type="nomenclature">
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<paragraph id="8B9336A27E5065188487941936C6FD97" blockId="2.[151,446,543,602]" box="[151,446,543,569]" pageId="2" pageNumber="65">
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<heading id="D0DB81CE7E5065188487941936C6FD97" bold="true" box="[151,446,543,569]" fontSize="11" level="1" pageId="2" pageNumber="65" reason="1">
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<emphasis id="B958EAB07E5065188487941936C6FD97" bold="true" box="[151,446,543,569]" pageId="2" pageNumber="65">
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<taxonomicName id="4C2C4D217E506518848794193621FD97" ID-CoL="VBZR" box="[151,345,543,569]" class="Gastropoda" family="Pyramidellidae" genus="Cimaria" kingdom="Animalia" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="vargasi" status="sp. nov.">
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<emphasis id="B958EAB07E506518848794193621FD97" bold="true" box="[151,345,543,569]" italics="true" pageId="2" pageNumber="65">Cimaria vargasi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26B57CB7E5065188571941936C6FD97" box="[353,446,543,569]" pageId="2" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9336A27E506518848794443624FDF4" blockId="2.[151,446,543,602]" box="[151,348,578,602]" pageId="2" pageNumber="65">
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<figureCitation id="13172A277E5065188487944437A4FDF4" box="[151,220,578,602]" captionStart="FIGURE 2" captionStartId="2.[151,244,1920,1941]" captionTargetBox="[171,1419,1268,1896]" captionTargetId="figure@2.[158,1429,1253,1903]" captionTargetPageId="2" captionText="FIGURE 2. Cimaria vargasi, scanning electron microscopy. A, D - Holotype (MZUCR 8955); 2.2 × 1.3 mm. A. Whole shell in apertural view. D. Detail of sculpture. B, C - Paratype, juvenile (MZUCR 8956); 0.8 × 0.6 mm. B. Semiapertural view, showing shell scar. C. Apical view. Scale bars = 100 µm." httpUri="https://zenodo.org/record/279943/files/figure.png" pageId="2" pageNumber="65">Figs 2</figureCitation>
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and
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<figureCitation id="13172A277E506518850394443658FDF4" box="[275,288,578,602]" captionStart="FIGURE 3" captionStartId="3.[151,244,1561,1582]" captionTargetBox="[167,1408,753,1507]" captionTargetId="figure@3.[151,1436,740,1537]" captionTargetPageId="3" captionText="FIGURE 3. Light microscopy. A & B - Cimaria vargasi. Two views of largest shell found (ZMBN 87909); 2.7 mm long. Showing the thickened outer lip in fully grown shells. C - Cimaria sp. Shell from Macau, Brazil (ZMBN 87911); 1.4 mm long." httpUri="https://zenodo.org/record/279944/files/figure.png" pageId="2" pageNumber="65">3</figureCitation>
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A, B.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33665297E5065188487948D3542FC94" pageId="2" pageNumber="65" type="materials_examined">
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<paragraph id="8B9336A27E5065188487948D3580FD11" blockId="2.[151,1436,650,1224]" pageId="2" pageNumber="65">
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<emphasis id="B958EAB07E5065188487948D364FFD0F" bold="true" box="[151,311,650,673]" pageId="2" pageNumber="65">
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<typeStatus id="549788007E5065188487948D37B4FD0E" box="[151,204,651,672]" pageId="2" pageNumber="65">Type</typeStatus>
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material:
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</emphasis>
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<typeStatus id="549788007E506518852E948C36E4FD0E" box="[318,412,650,672]" pageId="2" pageNumber="65" type="holotype">Holotype</typeStatus>
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(adult, empty shell)
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<collectionCode id="ED3DAE677E5065188662948D35B4FD0E" box="[626,716,651,672]" pageId="2" pageNumber="65">MZUCR</collectionCode>
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8955 and one
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<typeStatus id="549788007E5065188775948D34C4FD0E" box="[869,956,651,672]" pageId="2" pageNumber="65" type="paratype">paratype</typeStatus>
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(living juvenile)
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<collectionCode id="ED3DAE677E506518807F948D33B1FD0E" box="[1135,1225,651,672]" pageId="2" pageNumber="65">MZUCR</collectionCode>
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8956 (both on same SEM-stub), and 32
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<typeStatus id="549788007E506518854D94AC36B9FD11" box="[349,449,682,703]" pageId="2" pageNumber="65" type="paratype">paratypes</typeStatus>
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,
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<collectionCode id="ED3DAE677E50651885DA94AC356BFD11" box="[458,531,682,703]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="2" pageNumber="65">ZMBN</collectionCode>
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87909, 87912–87916.
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</paragraph>
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<paragraph id="8B9336A27E50651884D794CE33E9FD70" blockId="2.[151,1436,650,1224]" box="[199,1169,711,734]" pageId="2" pageNumber="65">
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<emphasis id="B958EAB07E50651884D794CE3622FD70" bold="true" box="[199,346,711,734]" pageId="2" pageNumber="65">
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<typeStatus id="549788007E50651884D794CE3784FD73" box="[199,252,712,733]" pageId="2" pageNumber="65">Type</typeStatus>
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locality:
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</emphasis>
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Punta Morales,
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<geoCoordinate id="EE1850657E50651885EF94CE352DFD73" box="[511,597,712,733]" direction="north" orientation="latitude" pageId="2" pageNumber="65" precision="925" value="10.066667">10°04'N</geoCoordinate>
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,
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<geoCoordinate id="EE1850657E506518864F94CE35C0FD70" box="[607,696,712,734]" direction="west" orientation="longitude" pageId="2" pageNumber="65" precision="925" value="-84.96667">84°58'W</geoCoordinate>
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, Golfo de Nicoya, Pacific coast of
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<collectingCountry id="F33B76327E506518800B94CE33F5FD70" box="[1051,1165,712,734]" name="Costa Rica" pageId="2" pageNumber="65">Costa Rica</collectingCountry>
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.
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</paragraph>
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<paragraph id="8B9336A27E50651884D794E03542FC94" blockId="2.[151,1436,650,1224]" pageId="2" pageNumber="65">
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<emphasis id="B958EAB07E50651884D794E036EFFD53" bold="true" box="[199,407,742,765]" pageId="2" pageNumber="65">Material examined</emphasis>
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(
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<tableCitation id="C6AE03197E50651885B894E0368EFD53" box="[424,502,742,765]" captionStart="TABLE 1" captionStartId="1.[151,231,953,974]" captionTargetBox="[174,1412,1166,1612]" captionTargetPageId="1" captionText="TABLE 1. Samples from the Pacific coast of Costa Rica and from northeastern Brazil, containing shells of Cimaria spp. Localities: A - Punta Morales, Golfo de Nicoya, high water, 10 ° 04 ' N, 84 ° 58 ' W; B - Cocorocas, river inlet, Golfo de Nicoya, low water, 10 ° 04 ' N, 84 ° 57 ' W; C - Puerto Jiménez, Golfo Dulce, 8 ° 32 ' N, 83 ° 18 ' W; D - Across opening of mangrove creek. Max. depth 1.5 m Salinas Cristao, Macau, Brazil, 5 ° 05 ' S, 36 ° 30 ' W." httpUri="http://table.plazi.org/id/DF53662A7E53651B848795BF3680FB88" pageId="2" pageNumber="65" tableUuid="DF53662A7E53651B848795BF3680FB88">Table 1</tableCitation>
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): The
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<typeStatus id="549788007E506518865494E035E5FD52" box="[580,669,742,764]" pageId="2" pageNumber="65" type="holotype">holotype</typeStatus>
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and 33
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<typeStatus id="549788007E50651886E894E13421FD52" box="[760,857,743,764]" pageId="2" pageNumber="65" type="paratype">paratypes</typeStatus>
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from Punta Morales; two worn shells from the Estero Cocorocas, about
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<quantity id="4CD49B477E5065188543950336EEFCB5" box="[339,406,773,795]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="65" unit="m" value="500.0">500 m</quantity>
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east of Punta Morales (
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<collectionCode id="ED3DAE677E5065188684950335A7FCB4" box="[660,735,773,794]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="2" pageNumber="65">ZMBN</collectionCode>
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87917), and two juveniles found alive at Puerto Jiménez in Golfo Dulce,
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<geoCoordinate id="EE1850657E50651884F095223651FC97" box="[224,297,804,825]" direction="north" orientation="latitude" pageId="2" pageNumber="65" precision="925" value="8.533334">8°32'N</geoCoordinate>
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,
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<geoCoordinate id="EE1850657E5065188524952236F5FC97" box="[308,397,804,825]" direction="west" orientation="longitude" pageId="2" pageNumber="65" precision="925" value="-83.3">83°18'W</geoCoordinate>
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(
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<collectionCode id="ED3DAE677E506518858A9522369DFC97" box="[410,485,804,825]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="2" pageNumber="65">ZMBN</collectionCode>
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87910).
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</paragraph>
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</subSubSection>
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<subSubSection id="C33665297E50651884D795453493FCD9" pageId="2" pageNumber="65" type="etymology">
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<paragraph id="8B9336A27E50651884D795453493FCD9" blockId="2.[151,1436,650,1224]" pageId="2" pageNumber="65">
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<emphasis id="B958EAB07E50651884D79545363CFCF6" bold="true" box="[199,324,835,856]" pageId="2" pageNumber="65">Etymology:</emphasis>
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The species is named for José Antonio Vargas, former director of CIMAR, and dedicated marine biologist, who has studied the fauna of the mud flats at Punta Morales for more than a decade.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33665297E50651984D7957933CEFEC8" lastPageId="3" lastPageNumber="66" pageId="2" pageNumber="65" type="description">
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<paragraph id="8B9336A27E50651884D7957932F9FB66" blockId="2.[151,1436,650,1224]" pageId="2" pageNumber="65">
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<emphasis id="B958EAB07E50651884D795793634FC38" bold="true" box="[199,332,895,918]" pageId="2" pageNumber="65">Description:</emphasis>
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Shell (
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<figureCitation id="13172A277E5065188589958636A0FC38" box="[409,472,896,918]" captionStart="FIGURE 2" captionStartId="2.[151,244,1920,1941]" captionTargetBox="[171,1419,1268,1896]" captionTargetId="figure@2.[158,1429,1253,1903]" captionTargetPageId="2" captionText="FIGURE 2. Cimaria vargasi, scanning electron microscopy. A, D - Holotype (MZUCR 8955); 2.2 × 1.3 mm. A. Whole shell in apertural view. D. Detail of sculpture. B, C - Paratype, juvenile (MZUCR 8956); 0.8 × 0.6 mm. B. Semiapertural view, showing shell scar. C. Apical view. Scale bars = 100 µm." httpUri="https://zenodo.org/record/279943/files/figure.png" pageId="2" pageNumber="65">Fig. 2</figureCitation>
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) is globose-conical, with inflated body whorl. Whorls slightly shouldered, with marked and deep suture. Just below the suture a distinct constriction. Whorls regular and round in young specimens, but tends to get slightly irregular in fully grown shells. Sculpture of teleoconch whorls characteristic, consisting of opisthocline, slightly irregular rows of circular, close-set pits, covering entire surface of the teleoconch except for the subsutural ridge (
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<figureCitation id="13172A277E506518808995DD33A0FC5C" box="[1177,1240,987,1010]" captionStart="FIGURE 2" captionStartId="2.[151,244,1920,1941]" captionTargetBox="[171,1419,1268,1896]" captionTargetId="figure@2.[158,1429,1253,1903]" captionTargetPageId="2" captionText="FIGURE 2. Cimaria vargasi, scanning electron microscopy. A, D - Holotype (MZUCR 8955); 2.2 × 1.3 mm. A. Whole shell in apertural view. D. Detail of sculpture. B, C - Paratype, juvenile (MZUCR 8956); 0.8 × 0.6 mm. B. Semiapertural view, showing shell scar. C. Apical view. Scale bars = 100 µm." httpUri="https://zenodo.org/record/279943/files/figure.png" pageId="2" pageNumber="65">Fig. 2</figureCitation>
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D). Pits increase in diameter with growth of shell, to approximately 40 µm in fully grown specimens. Except for occasional growth lines at about 30° angle to rows of pits, no other sculpture visible. Shell unusually thick, when alive translucent bluish white. Empty, worn shells white and opaque. Protoconch (
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<figureCitation id="13172A277E506518860F923E3524FBE0" box="[543,604,1080,1102]" captionStart="FIGURE 2" captionStartId="2.[151,244,1920,1941]" captionTargetBox="[171,1419,1268,1896]" captionTargetId="figure@2.[158,1429,1253,1903]" captionTargetPageId="2" captionText="FIGURE 2. Cimaria vargasi, scanning electron microscopy. A, D - Holotype (MZUCR 8955); 2.2 × 1.3 mm. A. Whole shell in apertural view. D. Detail of sculpture. B, C - Paratype, juvenile (MZUCR 8956); 0.8 × 0.6 mm. B. Semiapertural view, showing shell scar. C. Apical view. Scale bars = 100 µm." httpUri="https://zenodo.org/record/279943/files/figure.png" pageId="2" pageNumber="65">Fig. 2</figureCitation>
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C) small, smooth and glossy, with a distinct demarcation line to first postnuclear whorl. Obliquely immersed (almost 180°) in the following whorl, with only its base visible. Protoconch probably of less than one full whorl, and measures roughly 300 µm across. Aperture broadly ovate, rather spatulate in young specimens. Inner lip fairly straight, with a distinct columellar fold near upper edge, sometimes somewhat retracted into the mouth. No real umbilicus, but a distinct chink where inner lip meets the shell. In fully grown shells, outer lip thickened and varix-like (
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<figureCitation id="13172A277E506518811E92B43231FB66" box="[1294,1353,1202,1224]" captionStart="FIGURE 3" captionStartId="3.[151,244,1561,1582]" captionTargetBox="[167,1408,753,1507]" captionTargetId="figure@3.[151,1436,740,1537]" captionTargetPageId="3" captionText="FIGURE 3. Light microscopy. A & B - Cimaria vargasi. Two views of largest shell found (ZMBN 87909); 2.7 mm long. Showing the thickened outer lip in fully grown shells. C - Cimaria sp. Shell from Macau, Brazil (ZMBN 87911); 1.4 mm long." httpUri="https://zenodo.org/record/279944/files/figure.png" pageId="2" pageNumber="65">Fig. 3</figureCitation>
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A,B).
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</paragraph>
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<caption id="DF53662A7E506518848791863505F876" httpUri="https://zenodo.org/record/279943/files/figure.png" pageId="2" pageNumber="65" targetBox="[171,1419,1268,1896]" targetPageId="2">
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||
<paragraph id="8B9336A27E506518848791863505F876" blockId="2.[151,1436,1919,2008]" pageId="2" pageNumber="65">
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||
<emphasis id="B958EAB07E506518848791863674F83B" bold="true" box="[151,268,1920,1941]" pageId="2" pageNumber="65">FIGURE 2.</emphasis>
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<taxonomicName id="4C2C4D217E5065188502918636CFF838" box="[274,439,1920,1942]" class="Gastropoda" family="Pyramidellidae" genus="Cimaria" kingdom="Animalia" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="vargasi">
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<emphasis id="B958EAB07E5065188502918636CFF838" box="[274,439,1920,1942]" italics="true" pageId="2" pageNumber="65">Cimaria vargasi</emphasis>
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</taxonomicName>
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, scanning electron microscopy. A, D - Holotype (MZUCR 8955); 2.2 × 1.3 mm. A. Whole shell in apertural view. D. Detail of sculpture. B, C - Paratype, juvenile (MZUCR 8956); 0.8 × 0.6 mm. B. Semiapertural view, showing shell scar. C. Apical view. Scale bars = 100 µm.
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</paragraph>
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</caption>
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<paragraph id="8B9336A27E51651984D7969F32E5FF62" blockId="3.[151,1437,152,358]" pageId="3" pageNumber="66">
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<emphasis id="B958EAB07E51651984D7969F3647FF00" box="[199,319,153,174]" italics="true" pageId="3" pageNumber="66">Dimensions</emphasis>
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: Of 40 shells found, the largest was c. 2.7 ×
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<quantity id="4CD49B477E516519870A969E3414FF00" box="[794,876,152,174]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="66" unit="mm" value="1.5">1.5 mm</quantity>
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, with a little more than four teleoconch whorls. Body whorl very large,
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<quantity id="4CD49B477E516519855996B036DAFF62" box="[329,418,182,204]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="3" pageNumber="66" unit="mm" value="1.95">1.95 mm</quantity>
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long, aperture
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<quantity id="4CD49B477E516519862A96B135F1FF62" box="[570,649,183,204]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="66" unit="mm" value="1.3">1.3 mm</quantity>
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.
|
||
<typeStatus id="549788007E516519868296B03588FF62" box="[658,752,182,204]" pageId="3" pageNumber="66" type="holotype">Holotype</typeStatus>
|
||
2.2 ×
|
||
<quantity id="4CD49B477E516519872096B034F4FF62" box="[816,908,182,204]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="3" pageNumber="66" unit="mm" value="1.25">1.25 mm</quantity>
|
||
, body whorl
|
||
<quantity id="4CD49B477E516519801F96B03313FF62" box="[1039,1131,182,204]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="3" pageNumber="66" unit="mm" value="1.75">1.75 mm</quantity>
|
||
, aperture almost
|
||
<quantity id="4CD49B477E516519810696B1321BFF62" box="[1302,1379,183,204]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="66" unit="mm" value="1.2">1.2 mm</quantity>
|
||
long.
|
||
</paragraph>
|
||
<paragraph id="8B9336A27E51651984D796D333CEFEC8" blockId="3.[151,1437,152,358]" pageId="3" pageNumber="66">
|
||
<emphasis id="B958EAB07E51651984D796D3364AFF45" box="[199,306,213,235]" italics="true" pageId="3" pageNumber="66">Soft parts:</emphasis>
|
||
The snails were not observed alive, but through the partly transparent shell in alcohol preserved specimens the eyes were seen to be very close together. Distance between the small eyes less than their diameter. Digestive gland tends to be dark brown, filling almost 1 ½ teleoconch whorls. Rest of body dirty yellowish white after preservation. A fuzzy darkish spot on the base, somewhat to the left (in ventral view) of the columella, could be interpreted as the remnants of the pigmented mantle organ (which usually fades quickly in alcohol). Operculum thin and shiny, with no visible apophysis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |