473 lines
63 KiB
XML
473 lines
63 KiB
XML
<document id="2D7150EC9940901F4A0F6964828738A7" ID-DOI="10.5281/zenodo.170178" ID-GBIF-Dataset="a04a166d-dde9-4a5c-a116-198b06eaafea" ID-ISSN="11755326" ID-Zenodo-Dep="170178" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459751443612" checkinUser="plazi" docAuthor="Kraus, Fred" docDate="2005" docId="038587AEFFF1FFAA303F022DFE5CD0A5" docLanguage="en" docName="zt01061p028.pdf" docOrigin="Zootaxa 1061" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Nactus sphaerodactylodes Kraus, 2005, sp. nov." docType="treatment" docVersion="10" lastPageNumber="17" masterDocId="FFBCFFD6FFF6FFBA3137070BFFD2D533" masterDocTitle="The genus Nactus (Lacertilia: Gekkonidae): a phylogenetic analysis and description of two new species from the Papuan Region" masterLastPageNumber="28" masterPageNumber="1" pageNumber="8" updateTime="1698211870861" updateUser="plazi">
|
||
<mods:mods id="B2CE6ED2136B0C7FAEA177B6A2C48B4E" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="AE167108C3428F19B87D94E286D89FFC">
|
||
<mods:title id="E8833B2FF937AE2372C88278250B18E1">The genus Nactus (Lacertilia: Gekkonidae): a phylogenetic analysis and description of two new species from the Papuan Region</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="9B1F77B2A5241D15EEE25A1DEE67D115" type="personal">
|
||
<mods:role id="3DF76BD293E7E38D9D364510A3F5F36C">
|
||
<mods:roleTerm id="4DF2AFB27595BB2E9BF5C60159DB8804">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="972A0C1A5E9EC7944D34E23CD75D7BD5">Kraus, Fred</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="B2E9682D99A45DBE7B3863449A158BC7">text</mods:typeOfResource>
|
||
<mods:relatedItem id="71932134EAD91A9AAED39E6A81E86B03" type="host">
|
||
<mods:titleInfo id="DA2F5F3CD9191A8C211AABF79281F298">
|
||
<mods:title id="424BF9372C27BE10DEF3BBF0465C269F">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="C27743C792BB09E74A22429AA6AD636A">
|
||
<mods:date id="F48DB6014E445B5F20E30B3CBAAA1386">2005</mods:date>
|
||
<mods:detail id="7A83A7A9D58EDBA29A3CDBB9E807E582" type="volume">
|
||
<mods:number id="45878CDC9AB4D733F28C38178DA796D0">1061</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="F685FB128406E0EEF90B010578D40F35" unit="page">
|
||
<mods:start id="FA8273E632D07233CC361CE3D47FB688">1</mods:start>
|
||
<mods:end id="364A6A2EE62A8C24F85F16FAD95AA4D2">28</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="2E5F3633A340B80D98E2A4D9D4D68AFD">journal article</mods:classification>
|
||
<mods:identifier id="9941B6174FAEA764F1B14E001441F8A5" type="DOI">10.5281/zenodo.170178</mods:identifier>
|
||
<mods:identifier id="336027C795F39DF534350658E0A5F1C0" type="GBIF-Dataset">a04a166d-dde9-4a5c-a116-198b06eaafea</mods:identifier>
|
||
<mods:identifier id="7D586201BF03C329C7BC681AFF209D26" type="ISSN">11755326</mods:identifier>
|
||
<mods:identifier id="C651F241278671817AF55E2C18C3333E" type="Zenodo-Dep">170178</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="038587AEFFF1FFAA303F022DFE5CD0A5" ID-DOI="http://doi.org/10.5281/zenodo.5689073" ID-GBIF-Taxon="119337798" ID-Zenodo-Dep="5689073" LSID="urn:lsid:plazi:treatment:038587AEFFF1FFAA303F022DFE5CD0A5" httpUri="http://treatment.plazi.org/id/038587AEFFF1FFAA303F022DFE5CD0A5" lastPageId="16" lastPageNumber="17" pageId="7" pageNumber="8">
|
||
<subSubSection id="C3366533FFF1FFBD303F022DFE7AD050" pageId="7" pageNumber="8" type="nomenclature">
|
||
<paragraph id="8B9336B8FFF1FFBD303F022DFD74D073" blockId="7.[264,678,1318,1379]" box="[264,678,1318,1344]" pageId="7" pageNumber="8">
|
||
<heading id="D0DB81D4FFF1FFBD303F022DFD74D073" bold="true" box="[264,678,1318,1344]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
|
||
<emphasis id="B958EAAAFFF1FFBD303F022DFD74D073" bold="true" box="[264,678,1318,1344]" pageId="7" pageNumber="8">
|
||
<taxonomicName id="4C2C4D3BFFF1FFBD303F022DFD93D073" box="[264,577,1318,1344]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="sphaerodactylodes" status="sp. nov.">
|
||
<emphasis id="B958EAAAFFF1FFBD303F022DFD93D073" bold="true" box="[264,577,1318,1344]" italics="true" pageId="7" pageNumber="8">Nactus sphaerodactylodes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B57D1FFF1FFBD337E022DFD74D073" box="[585,678,1318,1344]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9336B8FFF1FFBD303F0242FE7AD050" blockId="7.[264,678,1318,1379]" box="[264,424,1353,1379]" pageId="7" pageNumber="8">
|
||
<figureCitation id="13172A3DFFF1FFBD303F0242FEA5D050" box="[264,375,1353,1379]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3. A" captionStart-2="FIGURE 4" captionStartId-0="8.[264,367,1517,1541]" captionStartId-1="9.[264,367,1834,1858]" captionStartId-2="10.[264,368,1239,1263]" captionTargetBox-0="[304,1267,369,1439]" captionTargetBox-1="[313,1279,554,1757]" captionTargetBox-2="[392,1194,340,1219]" captionTargetId-0="figure@8.[281,1306,329,1491]" captionTargetId-1="figure@9.[304,1283,529,1773]" captionTargetId-2="figure@10.[392,1194,329,1225]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURE 2. Carination pattern of dorsal scales for holotype of Nactus sphaerodactylodes, BPBM 20764." captionText-1="FIGURE 3. A) Dorsal, B) lateral, and C) ventral views of head, and D) ventral view of precloacal region showing pores for holotype of Nactus sphaerodactylodes, BPBM 20764." captionText-2="FIGURE 4. Hyoid of Nactus sphaerodactylodes, BPBM 20774. Clear areas are ossified; lightly stippled areas are partly ossified; heavily stippled areas are cartilaginous. Scale bar is 5 mm." httpUri-0="https://zenodo.org/record/170180/files/figure.png" httpUri-1="https://zenodo.org/record/170181/files/figure.png" httpUri-2="https://zenodo.org/record/170182/files/figure.png" pageId="7" pageNumber="8">Figs. 2–4</figureCitation>
|
||
,
|
||
<figureCitation id="13172A3DFFF1FFBD30B30242FE7AD050" box="[388,424,1353,1379]" captionStart="FIGURE 5. A" captionStartId="11.[264,367,1767,1791]" captionTargetBox="[343,1243,329,1739]" captionTargetPageId="11" captionText="FIGURE 5. A) Nactus sphaerodactylodes, Sudest Island, photo by G. Shea, and B) Nactus acutus, Rossel Island, photo by F. Kraus, in life, showing distinctive appearance of each." httpUri="https://zenodo.org/record/170183/files/figure.png" pageId="7" pageNumber="8">5A</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3366533FFF1FFBD303F0292FB8CD258" pageId="7" pageNumber="8" type="materials_examined">
|
||
<paragraph id="8B9336B8FFF1FFBD303F0292FD77D330" blockId="7.[264,1323,1433,1899]" pageId="7" pageNumber="8">
|
||
<emphasis id="B958EAAAFFF1FFBD303F0292FEA1D081" box="[264,371,1433,1458]" italics="true" pageId="7" pageNumber="8">
|
||
<typeStatus id="5497881AFFF1FFBD303F0292FEA1D081" box="[264,371,1433,1458]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<collectionCode id="ED3DAE7DFFF1FFBD30B10292FE0AD080" LSID="urn:lsid:biocol.org:col:34697" box="[390,472,1433,1459]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="7" pageNumber="8">BPBM</collectionCode>
|
||
20764 (field tag FK 9887), adult male, collected by local villagers at Araeda, Sudest Island,
|
||
<geoCoordinate id="EE18507FFFF1FFBD332102CAFD7AD0E8" box="[534,680,1473,1499]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="1" value="-11.43624">11.43624 S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE18507FFFF1FFBD338302CAFC8AD0E8" box="[692,856,1473,1499]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="1" value="153.43008">153.43008 E</geoCoordinate>
|
||
,
|
||
<quantity id="4CD49B5DFFF1FFBD325302CAFC65D0E8" box="[868,951,1473,1499]" metricMagnitude="1" metricUnit="m" metricValue="1.05" metricValueMax="2.0" metricValueMin="0.1" pageId="7" pageNumber="8" unit="m" value="10.5" valueMax="20.0" valueMin="1.0">120 m</quantity>
|
||
elevation, Milne Bay Province,
|
||
<collectingCountry id="F33B7628FFF1FFBD303F02E2FE3FD330" box="[264,493,1513,1539]" name="Papua New Guinea" pageId="7" pageNumber="8">Papua New Guinea</collectingCountry>
|
||
,
|
||
<date id="FF921078FFF1FFBD30CE02E2FD73D330" box="[505,673,1513,1539]" pageId="7" pageNumber="8" value="2004-04-28">28 April 2004</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9336B8FFF1FFBD300F011AFB88D3F8" blockId="7.[264,1323,1433,1899]" pageId="7" pageNumber="8">
|
||
<emphasis id="B958EAAAFFF1FFBD300F011AFE7DD319" box="[312,431,1553,1578]" italics="true" pageId="7" pageNumber="8">
|
||
<typeStatus id="5497881AFFF1FFBD300F011AFE7DD319" box="[312,431,1553,1578]" pageId="7" pageNumber="8" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<collectionCode id="ED3DAE7DFFF1FFBD30F5011AFDC6D318" LSID="urn:lsid:biocol.org:col:34697" box="[450,532,1553,1579]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="7" pageNumber="8">BPBM</collectionCode>
|
||
2076063, 2076575, 2169296 (the last five to be repatriated to
|
||
<collectionCode id="ED3DAE7DFFF1FFBD303F0132FEA5D360" box="[264,375,1593,1619]" pageId="7" pageNumber="8">PNGNM</collectionCode>
|
||
), same data as
|
||
<typeStatus id="5497881AFFF1FFBD331A0132FD45D360" box="[557,663,1593,1619]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
;
|
||
<collectionCode id="ED3DAE7DFFF1FFBD33920132FD25D360" LSID="urn:lsid:biocol.org:col:34697" box="[677,759,1593,1619]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="7" pageNumber="8">BPBM</collectionCode>
|
||
20757, same data as
|
||
<typeStatus id="5497881AFFF1FFBD32CC0132FBB3D360" box="[1019,1121,1593,1619]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
except collected by F. Malesa
|
||
<date id="FF921078FFF1FFBD309D016AFD87D348" box="[426,597,1633,1659]" pageId="7" pageNumber="8" value="2004-04-10">10 April 2004</date>
|
||
;
|
||
<collectionCode id="ED3DAE7DFFF1FFBD3354016AFD67D348" LSID="urn:lsid:biocol.org:col:34697" box="[611,693,1633,1659]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="7" pageNumber="8">BPBM</collectionCode>
|
||
20758, same data as
|
||
<typeStatus id="5497881AFFF1FFBD3280016AFBCFD348" box="[951,1053,1633,1659]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
except collected by F. Kraus
|
||
<date id="FF921078FFF1FFBD30620182FE29D390" box="[341,507,1673,1699]" pageId="7" pageNumber="8" value="2004-04-12">12 April 2004</date>
|
||
;
|
||
<collectionCode id="ED3DAE7DFFF1FFBD33310182FD8AD390" LSID="urn:lsid:biocol.org:col:34697" box="[518,600,1673,1699]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="7" pageNumber="8">BPBM</collectionCode>
|
||
20759, same data as
|
||
<typeStatus id="5497881AFFF1FFBD32780182FC64D390" box="[847,950,1673,1699]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
except collected
|
||
<date id="FF921078FFF1FFBD35480182FAF7D390" box="[1151,1317,1673,1699]" pageId="7" pageNumber="8" value="2004-04-13">13 April 2004</date>
|
||
;
|
||
<collectionCode id="ED3DAE7DFFF1FFBD303F01BAFE88D3F8" LSID="urn:lsid:biocol.org:col:34697" box="[264,346,1713,1739]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="7" pageNumber="8">BPBM</collectionCode>
|
||
2077680, same data as
|
||
<typeStatus id="5497881AFFF1FFBD334C01BAFD30D3F8" box="[635,738,1713,1739]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
except collected
|
||
<date id="FF921078FFF1FFBD329901BAFB87D3F8" box="[942,1109,1713,1739]" pageId="7" pageNumber="8" value="2004-04-29">29 April 2004</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9336B8FFF1FFBD300F01D2FB8CD258" blockId="7.[264,1323,1433,1899]" pageId="7" pageNumber="8">
|
||
<emphasis id="B958EAAAFFF1FFBD300F01D2FDCCD3C1" box="[312,542,1753,1778]" italics="true" pageId="7" pageNumber="8">Referred specimens</emphasis>
|
||
:
|
||
<collectionCode id="ED3DAE7DFFF1FFBD330701D2FD50D3C0" LSID="urn:lsid:biocol.org:col:34697" box="[560,642,1753,1779]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="7" pageNumber="8">BPBM</collectionCode>
|
||
20756, collected by F. Malesa at Somusomuwa, Misima Island,
|
||
<geoCoordinate id="EE18507FFFF1FFBD3054000AFE2ED228" box="[355,508,1793,1819]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="1" value="-10.63791">10.63791 S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE18507FFFF1FFBD333A000AFD64D228" box="[525,694,1793,1819]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="1" value="152.82822">152.82821 E</geoCoordinate>
|
||
, Milne Bay Province,
|
||
<collectingCountry id="F33B7628FFF1FFBD32E5000AFB13D228" box="[978,1217,1793,1819]" name="Papua New Guinea" pageId="7" pageNumber="8">Papua New Guinea</collectingCountry>
|
||
,
|
||
<date id="FF921078FFF1FFBD35E5000AFE94D270" pageId="7" pageNumber="8" value="2004-04-08">8 April 2004</date>
|
||
.
|
||
<collectionCode id="ED3DAE7DFFF1FFBD30620022FE75D270" LSID="urn:lsid:biocol.org:col:34697" box="[341,423,1833,1859]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="7" pageNumber="8">BPBM</collectionCode>
|
||
17707, collected by A. Allison at Guasopa, Woodlark Island,
|
||
<geoCoordinate id="EE18507FFFF1FFBD359C0022FAF7D270" box="[1195,1317,1833,1859]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="5" value="-9.2233">9.2233 S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE18507FFFF1FFBD303F005AFE4FD258" box="[264,413,1873,1899]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="5" value="152.9433">152.9433 E</geoCoordinate>
|
||
, Milne Bay Province,
|
||
<collectingCountry id="F33B7628FFF1FFBD3393005AFC58D258" box="[676,906,1873,1899]" name="Papua New Guinea" pageId="7" pageNumber="8">Papua New Guinea</collectingCountry>
|
||
,
|
||
<date id="FF921078FFF1FFBD32A2005AFB88D258" box="[917,1114,1873,1899]" pageId="7" pageNumber="8" value="2003-01-24">24 January 2003</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF536630FFFEFFB2303F02E6FE80D314" httpUri="https://zenodo.org/record/170180/files/figure.png" pageId="8" pageNumber="9" targetBox="[304,1267,369,1439]" targetPageId="8">
|
||
<paragraph id="8B9336B8FFFEFFB2303F02E6FE80D314" blockId="8.[264,1322,1517,1575]" pageId="8" pageNumber="9">
|
||
<emphasis id="B958EAAAFFFEFFB2303F02E6FE59D336" bold="true" box="[264,395,1517,1541]" pageId="8" pageNumber="9">FIGURE 2.</emphasis>
|
||
Carination pattern of dorsal scales for holotype of
|
||
<taxonomicName id="4C2C4D3BFFFEFFB2328E02E5FB00D336" box="[953,1234,1518,1541]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="sphaerodactylodes">
|
||
<emphasis id="B958EAAAFFFEFFB2328E02E5FB00D336" box="[953,1234,1518,1541]" italics="true" pageId="8" pageNumber="9">Nactus sphaerodactylodes</emphasis>
|
||
</taxonomicName>
|
||
, BPBM 20764.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3366533FFFEFFB0300F0150FBD8D25F" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="diagnosis">
|
||
<paragraph id="8B9336B8FFFEFFB3300F0150FD82D4E5" blockId="8.[264,1324,1627,1896]" lastBlockId="9.[264,1323,284,470]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
|
||
<emphasis id="B958EAAAFFFEFFB2300F0150FE7DD347" box="[312,431,1627,1652]" italics="true" pageId="8" pageNumber="9">Diagnosis</emphasis>
|
||
: A small (SV = 20.4–26.0 mm, mean
|
||
<quantity id="4CD49B5DFFFEFFB232490150FC38D346" box="[894,1002,1627,1653]" metricMagnitude="-2" metricUnit="m" metricValue="2.42" pageId="8" pageNumber="9" unit="mm" value="24.2">24.2 mm</quantity>
|
||
) species of
|
||
<taxonomicName id="4C2C4D3BFFFEFFB2354D0150FB1ED347" box="[1146,1228,1627,1652]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958EAAAFFFEFFB2354D0150FB1ED347" box="[1146,1228,1627,1652]" italics="true" pageId="8" pageNumber="9">Nactus</emphasis>
|
||
</taxonomicName>
|
||
lacking dorsal tubercles; and having granular, largely tricarinate dorsal (
|
||
<figureCitation id="13172A3DFFFEFFB232CD0188FB92D3AE" box="[1018,1088,1667,1693]" captionStart="FIGURE 2" captionStartId="8.[264,367,1517,1541]" captionTargetBox="[304,1267,369,1439]" captionTargetId="figure@8.[281,1306,329,1491]" captionTargetPageId="8" captionText="FIGURE 2. Carination pattern of dorsal scales for holotype of Nactus sphaerodactylodes, BPBM 20764." httpUri="https://zenodo.org/record/170180/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
|
||
) and ventral scales; lamellate, unexpanded subdigital lamellae; 1013 lamellae under the 4th toe; a single row of scales between the orbit and supralabials; claws lying within four scales; granular postmentals; elongate first infralabials; 4–5 supralabials to center of eye and 6–7 to rictus; 5–7 infralabials; enlarged postnasals; smooth scales on the tail; oval ear opening; relatively broad head (HW/HL = 0.71–0.79); short, broad snout (EN/SV = 0.070–0.086, IN/SV = 0.038–0.049, EN/IN = 1.5–2.1); large eye (EE/SV = 0.085–0.103) and interorbital distance (ID/SV = 0.063–0.083); relatively broad tail (TW/TL = 0.12–0.16); four sternal/ xiphisternal ribs; and dark brown dorsal color, usually with scattered, obscure darker and lighter brown punctations and irregular markings, these sometimes coalescing to form obscure, irregular chevrons.
|
||
</paragraph>
|
||
<caption id="DF536630FFFFFFB3303F0021FB89D257" httpUri="https://zenodo.org/record/170181/files/figure.png" pageId="9" pageNumber="10" targetBox="[313,1279,554,1757]" targetPageId="9">
|
||
<paragraph id="8B9336B8FFFFFFB3303F0021FB89D257" blockId="9.[264,1322,1834,1892]" pageId="9" pageNumber="10">
|
||
<emphasis id="B958EAAAFFFFFFB3303F0021FE58D271" bold="true" box="[264,394,1834,1858]" pageId="9" pageNumber="10">FIGURE 3.</emphasis>
|
||
A) Dorsal, B) lateral, and C) ventral views of head, and D) ventral view of precloacal region showing pores for holotype of
|
||
<taxonomicName id="4C2C4D3BFFFFFFB333AC0046FC61D257" box="[667,947,1869,1892]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="sphaerodactylodes">
|
||
<emphasis id="B958EAAAFFFFFFB333AC0046FC61D257" box="[667,947,1869,1892]" italics="true" pageId="9" pageNumber="10">Nactus sphaerodactylodes</emphasis>
|
||
</taxonomicName>
|
||
, BPBM 20764.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF536630FFFCFFB0303F03DCFB32D022" httpUri="https://zenodo.org/record/170182/files/figure.png" pageId="10" pageNumber="11" targetBox="[392,1194,340,1219]" targetPageId="10">
|
||
<paragraph id="8B9336B8FFFCFFB0303F03DCFB32D022" blockId="10.[264,1323,1239,1297]" pageId="10" pageNumber="11">
|
||
<emphasis id="B958EAAAFFFCFFB0303F03DCFE5FD1DC" bold="true" box="[264,397,1239,1263]" pageId="10" pageNumber="11">FIGURE 4.</emphasis>
|
||
Hyoid of
|
||
<taxonomicName id="4C2C4D3BFFFCFFB0333403D3FCCFD1DC" box="[515,797,1240,1263]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="sphaerodactylodes">
|
||
<emphasis id="B958EAAAFFFCFFB0333403D3FCCFD1DC" box="[515,797,1240,1263]" italics="true" pageId="10" pageNumber="11">Nactus sphaerodactylodes</emphasis>
|
||
</taxonomicName>
|
||
, BPBM 20774. Clear areas are ossified; lightly stippled areas are partly ossified; heavily stippled areas are cartilaginous. Scale bar is 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9336B8FFFCFFB0300F024EFBD8D25F" blockId="10.[264,1324,1349,1900]" pageId="10" pageNumber="11">
|
||
<emphasis id="B958EAAAFFFCFFB0300F024EFD76D06D" box="[312,676,1349,1374]" italics="true" pageId="10" pageNumber="11">Comparison with other species</emphasis>
|
||
:
|
||
<taxonomicName id="4C2C4D3BFFFCFFB03383024EFC3BD06D" box="[692,1001,1349,1374]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="sphaerodactylodes">
|
||
<emphasis id="B958EAAAFFFCFFB03383024EFC3BD06D" box="[692,1001,1349,1374]" italics="true" pageId="10" pageNumber="11">Nactus sphaerodactylodes</emphasis>
|
||
</taxonomicName>
|
||
may be distinguished from all other members of the genus except
|
||
<taxonomicName id="4C2C4D3BFFFCFFB033E80266FC67D0B5" box="[735,949,1389,1414]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="coindemirensis">
|
||
<emphasis id="B958EAAAFFFCFFB033E80266FC67D0B5" box="[735,949,1389,1414]" italics="true" pageId="10" pageNumber="11">N. coindemirensis</emphasis>
|
||
</taxonomicName>
|
||
in 1) lacking rows of enlarged tubercles on the body, 2) having the claws lie within four (vs. two or three) scales, 3) having short digits, with 10–13 lamellae under 4th toe (vs. 15–25); and 4) having elongate infralabials such that the mental extends no further posteriorly than the center of the first infralabials (vs. to or past suture with 2nd infralabial) and the suture between the first and second infralabials lies behind the center of the second supralabial (vs. at beginning of second supralabial). From
|
||
<taxonomicName id="4C2C4D3BFFFCFFB033730168FCCAD34F" box="[580,792,1635,1660]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="coindemirensis">
|
||
<emphasis id="B958EAAAFFFCFFB033730168FCCAD34F" box="[580,792,1635,1660]" italics="true" pageId="10" pageNumber="11">N. coindemirensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2C4D3BFFFCFFB032100168FBF4D34F" box="[807,1062,1635,1660]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="sphaerodactylodes">
|
||
<emphasis id="B958EAAAFFFCFFB032100168FBF4D34F" box="[807,1062,1635,1660]" italics="true" pageId="10" pageNumber="11">N. sphaerodactylodes</emphasis>
|
||
</taxonomicName>
|
||
differs in the last two characters above as well as in having granular (vs. imbricate) ventral scales, carinate (vs. smooth) dorsal and ventral scales, lamellate (vs. granular) distal subdigital scales, unexpanded (vs. expanded) proximal subdigital lamellae, and 5–7 (vs. 10–11) infralabials. From the closely related
|
||
<taxonomicName id="4C2C4D3BFFFCFFB0330E0008FD39D22F" box="[569,747,1795,1820]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="vankampeni">
|
||
<emphasis id="B958EAAAFFFCFFB0330E0008FD39D22F" box="[569,747,1795,1820]" italics="true" pageId="10" pageNumber="11">N. vankampeni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2C4D3BFFFCFFB033C90008FC2CD22F" box="[766,1022,1795,1820]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="sphaerodactylodes">
|
||
<emphasis id="B958EAAAFFFCFFB033C90008FC2CD22F" box="[766,1022,1795,1820]" italics="true" pageId="10" pageNumber="11">N. sphaerodactylodes</emphasis>
|
||
</taxonomicName>
|
||
further differs in having tricarinate (vs. multicarinate) dorsal and ventral keeling, one (vs. two) scale row between orbit and supralabials, and four (vs. five) sternal/xiphisternal ribs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF536630FFFDFFB1303F01ECFBB5D212" httpUri="https://zenodo.org/record/170183/files/figure.png" pageId="11" pageNumber="12" targetBox="[343,1243,329,1739]" targetPageId="11">
|
||
<paragraph id="8B9336B8FFFDFFB1303F01ECFBB5D212" blockId="11.[264,1323,1767,1902]" pageId="11" pageNumber="12">
|
||
<emphasis id="B958EAAAFFFDFFB1303F01ECFE58D3CC" bold="true" box="[264,394,1767,1791]" pageId="11" pageNumber="12">FIGURE 5.</emphasis>
|
||
A)
|
||
<taxonomicName id="4C2C4D3BFFFDFFB1308201ECFCF0D3CD" authority="Sudest" authorityName="Sudest" box="[437,802,1767,1790]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="sphaerodactylodes">
|
||
<emphasis id="B958EAAAFFFDFFB1308201ECFD1CD3CD" box="[437,718,1767,1790]" italics="true" pageId="11" pageNumber="12">Nactus sphaerodactylodes</emphasis>
|
||
, Sudest
|
||
</taxonomicName>
|
||
Island, photo by G. Shea, and B)
|
||
<taxonomicName id="4C2C4D3BFFFDFFB135BA01ECFE9DD212" authority="Rossel" authorityName="Rossel" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B958EAAAFFFDFFB135BA01ECFAF1D3CD" box="[1165,1315,1767,1790]" italics="true" pageId="11" pageNumber="12">Nactus acutus</emphasis>
|
||
, Rossel
|
||
</taxonomicName>
|
||
Island, photo by F. Kraus, in life, showing distinctive appearance of each.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3366533FFFDFFAA300F0026FCE6D715" lastPageId="16" lastPageNumber="17" pageId="11" pageNumber="12" type="description">
|
||
<paragraph id="8B9336B8FFFDFFB6300F0026FE74D11D" blockId="11.[264,1323,1767,1902]" lastBlockId="12.[264,1325,284,1870]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||
<emphasis id="B958EAAAFFFDFFB1300F0026FD8ED275" box="[312,604,1837,1862]" italics="true" pageId="11" pageNumber="12">
|
||
Description of
|
||
<typeStatus id="5497881AFFFDFFB130C00026FD8ED275" box="[503,604,1837,1862]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
: An adult male with right lateral incision; part of liver removed. Animal small (SV = 22.0 mm, TrL = 9.3). Head relatively long (HL/SV = 0.27) and wide (HW/HL = 0.78), not depressed (HH/HL = 0.47), distinct from neck (
|
||
<figureCitation id="13172A3DFFFAFFB6358A0617FACFD405" box="[1213,1309,284,310]" captionStart="FIGURE 3. A" captionStartId="9.[264,367,1834,1858]" captionTargetBox="[313,1279,554,1757]" captionTargetId="figure@9.[304,1283,529,1773]" captionTargetPageId="9" captionText="FIGURE 3. A) Dorsal, B) lateral, and C) ventral views of head, and D) ventral view of precloacal region showing pores for holotype of Nactus sphaerodactylodes, BPBM 20764." httpUri="https://zenodo.org/record/170181/files/figure.png" pageId="12" pageNumber="13">Fig. 3A</figureCitation>
|
||
). Loreal and interorbital region slightly inflated; canthus rostralis absent. Snout relatively long (SN/HL = 0.41), much longer than eye diameter (SN/EY = 2.2) (
|
||
<figureCitation id="13172A3DFFFAFFB635B70667FB19D4B5" box="[1152,1227,364,390]" captionStart="FIGURE 3. A" captionStartId="9.[264,367,1834,1858]" captionTargetBox="[313,1279,554,1757]" captionTargetId="figure@9.[304,1283,529,1773]" captionTargetPageId="9" captionText="FIGURE 3. A) Dorsal, B) lateral, and C) ventral views of head, and D) ventral view of precloacal region showing pores for holotype of Nactus sphaerodactylodes, BPBM 20764." httpUri="https://zenodo.org/record/170181/files/figure.png" pageId="12" pageNumber="13">Fig. 3</figureCitation>
|
||
B). Eye relatively large (EY/HL = 0.19); pupil vertical, with crenellated margins; supraciliaries small, conical, not well differentiated from adjacent granules. Ear opening small (Ear/HL = 0.068), oval, with long axis running posterodorsal to anteroventral; distance between ear and eye twice eye diameter. Rostral half (
|
||
<quantity id="4CD49B5DFFFAFFB633D80507FC89D715" box="[751,859,524,550]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="12" pageNumber="13" unit="mm" value="0.55">0.55 mm</quantity>
|
||
) as high as wide (
|
||
<quantity id="4CD49B5DFFFAFFB635060507FB42D715" box="[1073,1168,524,550]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="13" unit="mm" value="1.1">1.1 mm</quantity>
|
||
), highest just medial to nares and depressed between these points; length (<
|
||
<quantity id="4CD49B5DFFFAFFB632C7053FFB82D77D" box="[1008,1104,564,590]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="13" unit="mm" value="0.5">0.5 mm</quantity>
|
||
) slightly less than height; partially divided dorsally by weak rostral groove. Two enlarged supranasals separated by three granular internasals. Rostral in contact with first supralabials, two supranasals, and three internasals. External nares oval, long dimension running anterodorsally to posteroventrally; each bordered by rostral, supranasal, first supralabial, and one nasal. First supralabial has an extension that reaches dorsally to behind the posteriormost limit of the naris. Nasal bordered posteriorly by two small granular scales. One row of small scales separates orbit from supralabials. Mental subtriangular, wider (
|
||
<quantity id="4CD49B5DFFFAFFB634300447FEE5D6BD" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="13" unit="mm" value="1.3">1.3 mm</quantity>
|
||
) than long (
|
||
<quantity id="4CD49B5DFFFAFFB630F0047FFDF5D6BD" box="[455,551,884,910]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="13" unit="mm" value="1.1">1.1 mm</quantity>
|
||
). No enlarged postmentals (
|
||
<figureCitation id="13172A3DFFFAFFB63247047FFC65D6BD" box="[880,951,884,910]" captionStart="FIGURE 3. A" captionStartId="9.[264,367,1834,1858]" captionTargetBox="[313,1279,554,1757]" captionTargetId="figure@9.[304,1283,529,1773]" captionTargetPageId="9" captionText="FIGURE 3. A) Dorsal, B) lateral, and C) ventral views of head, and D) ventral view of precloacal region showing pores for holotype of Nactus sphaerodactylodes, BPBM 20764." httpUri="https://zenodo.org/record/170181/files/figure.png" pageId="12" pageNumber="13">Fig. 3</figureCitation>
|
||
C); mental bordered posteriorly by nine tiny granules. Infralabials bordered by tiny granules, these somewhat larger posteriorly than anteriorly. Supralabials to midorbital position four on each side, to angle of jaw six on each side. Infralabials seven on right, six on left. Scale rows between center of orbits ~36.
|
||
</paragraph>
|
||
<paragraph id="8B9336B8FFFAFFB6300F0337FE83D02D" blockId="12.[264,1325,284,1870]" pageId="12" pageNumber="13">
|
||
Body rather slender (TrL/SV = 0.42), lacking ventrolateral folds (
|
||
<figureCitation id="13172A3DFFFAFFB635560337FB16D165" box="[1121,1220,1084,1110]" captionStart="FIGURE 5. A" captionStartId="11.[264,367,1767,1791]" captionTargetBox="[343,1243,329,1739]" captionTargetPageId="11" captionText="FIGURE 5. A) Nactus sphaerodactylodes, Sudest Island, photo by G. Shea, and B) Nactus acutus, Rossel Island, photo by F. Kraus, in life, showing distinctive appearance of each." httpUri="https://zenodo.org/record/170183/files/figure.png" pageId="12" pageNumber="13">Fig. 5A</figureCitation>
|
||
). Dorsal scales on head, body, and limbs tiny, conical, juxtaposed granules; each wider than long and with three sharp keels; granules on neck also usually higher than long; granules on venter longer than wide or roughly circular; otherwise, granules quite homogeneous. Organs on posterior end of each scale 3. Enlarged tubercles absent on head, body, and limbs.
|
||
</paragraph>
|
||
<paragraph id="8B9336B8FFFAFFB6300F0227FBA9D33D" blockId="12.[264,1325,284,1870]" pageId="12" pageNumber="13">
|
||
Precloacal pores in a shallow Vshaped series (
|
||
<figureCitation id="13172A3DFFFAFFB6326D0227FC4AD075" box="[858,920,1324,1350]" captionStart="FIGURE 3. A" captionStartId="9.[264,367,1834,1858]" captionTargetBox="[313,1279,554,1757]" captionTargetId="figure@9.[304,1283,529,1773]" captionTargetPageId="9" captionText="FIGURE 3. A) Dorsal, B) lateral, and C) ventral views of head, and D) ventral view of precloacal region showing pores for holotype of Nactus sphaerodactylodes, BPBM 20764." httpUri="https://zenodo.org/record/170181/files/figure.png" pageId="12" pageNumber="13">Fig 3</figureCitation>
|
||
D), 16, asymmetrically distributed with 12 on right side of apex of V and 4 on left, followed by a gap of 8 (left) or 10 (right) rows of enlarged femoral scales lacking pores, followed by 4 femoral pores on each side. Granules anterior to precloacal pore series and those in anterior femoral region enlarged relative to those on abdomen; granules immediately posterior to pore series conspicuously smaller. No precloacal groove. Scales on palms and soles rounded, smooth.
|
||
</paragraph>
|
||
<paragraph id="8B9336B8FFFAFFB6300F0117FB92D3CD" blockId="12.[264,1325,284,1870]" pageId="12" pageNumber="13">
|
||
Fore and hindlimbs relatively short, stout (FA/SV = 0.13, CS/SV = 0.17). Digits short, only modestly reflected at basal interphalangeal joints; all with robust, slightly recurved claws. Subdigital lamellae narrow, smooth, rounded, and slightly expanded distally. Lamellae on manus 47886 on right, 56997 on left; on pes
|
||
<date id="FF921078FFFAFFB6355B019FFAD7D39D" box="[1132,1285,1684,1710]" pageId="12" pageNumber="13" value="1911-05-11" valueMax="1911-08-11" valueMin="1911-05-11">58111111</date>
|
||
on right,
|
||
<date id="FF921078FFFAFFB6307901B7FE3BD3E5" box="[334,489,1724,1750]" pageId="12" pageNumber="13" value="1910-05-12" valueMax="1910-07-11" valueMin="1910-05-12">57101211</date>
|
||
on left. Relative lengths of digits on manus I <V <II <III <IV; on pes I <II <V <III <IV. No interdigital webbing present on manus or pes.
|
||
</paragraph>
|
||
<paragraph id="8B9336B8FFFAFFB7300F0007FC8ED4B5" blockId="12.[264,1325,284,1870]" lastBlockId="13.[264,1324,284,750]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">Tail relatively short (TL/SV = 0.96), with moderately wide base (TW/TL = 0.12), tapering to a point, slightly constricted at base. Cloacal sacs welldeveloped, external orifices large, approximately 3 scales in width, situated near lateral margins of vent. Scales of tail large, flat, smooth, imbricate; a pair of enlarged, rounded postcloacal spurs on each side of tailbase, the dorsal smaller than the ventral.</paragraph>
|
||
<paragraph id="8B9336B8FFFBFFB7300F069FFB19D7DD" blockId="13.[264,1324,284,750]" pageId="13" pageNumber="14">
|
||
<emphasis id="B958EAAAFFFBFFB7300F069FFDEFD49E" box="[312,573,404,429]" italics="true" pageId="13" pageNumber="14">Color in preservative</emphasis>
|
||
: Dorsal granules brown, each granule with 0–8 small black spots; keels of granules frequently margined with dark brown but often not. Interstitial skin darker brown. A few scattered granules light gray. Overall appearance is of a uniformly drab brown color. Rostral and labials chocolate brown with wide unpigmented strips along margins. Granules of venter much like those on dorsum except that many more are light gray, imparting to the venter an overall saltandpepper appearance. Light scales on chin and throat typically clustered together, giving the impression of scattered light gray spots on a dark ground. Scales on tail lighter brown, margined and speckled with black, or not, with dark color predominating proximally and disappearing distally.
|
||
</paragraph>
|
||
<paragraph id="8B9336B8FFFBFFB7300F042FFC6ED14D" blockId="13.[264,1322,804,1150]" pageId="13" pageNumber="14">
|
||
<emphasis id="B958EAAAFFFBFFB7300F042FFE14D60E" box="[312,454,804,829]" italics="true" pageId="13" pageNumber="14">Color in life</emphasis>
|
||
: Color pattern of BPBM 20756 was noted in life as: “Dorsum brown with middorsal row of dark brown chevrons bordered laterally by single row of yellowbrown scales. Venter dark gray with regular series of light gray blotches/flecks, these blotches being elongated on chin and fewer on tail. Under tail charcoal gray. Iris redorange.” Judging from a color image of this animal, the chevrons are relictual, not extending laterally very far beyond the middorsal region, and are frequently margined with yellow scales behind. An irregular array of tiny yellow spots can be seen over much of the head, body, and limbs. Such dorsal pattern as exists fades distally on the tail, which also gradually assumes a more orangebrown hue than the body
|
||
</paragraph>
|
||
<paragraph id="8B9336B8FFFBFFB7300F03BFFE52D3CD" blockId="13.[264,1323,1204,1870]" pageId="13" pageNumber="14">
|
||
<emphasis id="B958EAAAFFFBFFB7300F03BFFE7DD1FE" box="[312,431,1204,1229]" italics="true" pageId="13" pageNumber="14">Osteology</emphasis>
|
||
: The following is based primarily on BPBM 20774, a cleared and doublestained specimen, supplemented by additional notes on similarly cleared and doublestained BPBM 20773 and 20776 and Xrays of BPBM 17707 and 20764. Frontal with a narrow supraorbital waist that is onethird as wide as its posterior margin and narrower than the parietals; supraorbital ridges absent; anterior margin strongly tridentate. Parietals paired, longer than wide. Nasals fused. Premaxillae fused, with 13 teeth loci. Maxillae extend posterior to midorbit, each with approximately 25 teeth loci; mandibles each with approximately 30 teeth loci. Quadrate long, narrow, slightly curved. Cranial vault filled with endolymphatic space as far forward as to include the posterior 20% of parietals; solid subtriangular endolymphatic sacs extend from just behind quadrate posteriorly to cover anterior half of suprascapulae and extend dorsally to tops of suprascapulae; ducts to endolymphatic space of cranial vault not obvious. Phalangeal formula 23453 for manus, 23454 for pes. Presacral vertebrae 26, three of which are cervical vertebrae lacking ribs; two sacral vertebrae; anteriormost four or five caudal vertebrae with lateral processes.
|
||
</paragraph>
|
||
<paragraph id="8B9336B8FFFBFFB4300F0007FCA3D77D" blockId="13.[264,1323,1204,1870]" lastBlockId="14.[264,1323,284,1190]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||
Hyoid largely cartilage except for ossified first ceratobranchials (
|
||
<figureCitation id="13172A3DFFFBFFB735720007FB42D215" box="[1093,1168,1804,1830]" captionStart="FIGURE 4" captionStartId="10.[264,368,1239,1263]" captionTargetBox="[392,1194,340,1219]" captionTargetId="figure@10.[392,1194,329,1225]" captionTargetPageId="10" captionText="FIGURE 4. Hyoid of Nactus sphaerodactylodes, BPBM 20774. Clear areas are ossified; lightly stippled areas are partly ossified; heavily stippled areas are cartilaginous. Scale bar is 5 mm." httpUri="https://zenodo.org/record/170182/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
|
||
). Glossohyal long, extending forward to anterior end of orbits, continuous with basihyal and second ceratobranchials. Hypohyals and first ceratobranchials articulate at posterior end of basihyal. Ceratohyals followed by epihyals, extending posteriorly and dorsally to point just behind squamosal. First ceratobranchial long, tipped by cartilaginous first epibranchial, both extending posteriorly and dorsally to lie along the anterior face of the endolymphatic sacs. Second epibranchial separate from second ceratobranchial, recurved, with distal extremity lying anterior and deep to proximal extremity. Cricoid cartilage single, with lateral ala on each side, a large ventral foramen, a smaller anterodorsal foramen, and a pair of small posterodorsal foramina.
|
||
</paragraph>
|
||
<paragraph id="8B9336B8FFF8FFB4300F0557FD94D135" blockId="14.[264,1323,284,1190]" pageId="14" pageNumber="15">Scapula, coracoids, clavicles, and interclavicle ossified; sternum, epicoracoids, and suprascapulae cartilaginous. Scapula and coracoid separated from each other along approximately half their lengths, each articulating anteriorly with epicoracoid, which extends posteriorly to posterior extremity of coracoid. Ventral and separate from the epicoracoids lies the sternum, which is shieldshaped, with two anterior prongs, and four pairs of costal cartilages attaching along its posterior margin. Ventral to the anterior ends of the epicoracoids are two broad clavicles, each with a large fenestra near the point of articulation with interclavicle and each of which extends dorsolaterally in a long narrow arc to terminate at the base of the respective suprascapula. Interclavicle articulates deep to the clavicles anteriorly, is very narrow anteriorly, but broadens rapidly posteriorly to lie ventral to the epicoracoids.</paragraph>
|
||
<paragraph id="8B9336B8FFF8FFB4300F031FFE09D195" blockId="14.[264,1323,284,1190]" pageId="14" pageNumber="15">Ischium wider than long; ischial cartilage extends posterior to ischium for a length greater than length of ischium. BPBM 20776 is a male but I could find no cloacal bones in the specimen. However, those elements do appear as tiny, thin bones in Xrays of BPBM 17707 and 20764.</paragraph>
|
||
<paragraph id="8B9336B8FFF8FFB4300F03D7FC9BD305" blockId="14.[264,1323,1244,1870]" pageId="14" pageNumber="15">
|
||
<emphasis id="B958EAAAFFF8FFB4300F03D7FE73D1C6" box="[312,417,1244,1269]" italics="true" pageId="14" pageNumber="15">Var ia t io n</emphasis>
|
||
: Mensural variation is not great, nor is mensural sexual dimorphism apparent (
|
||
<tableCitation id="C6AE0303FFF8FFB43027020FFEB9D02D" box="[272,363,1284,1310]" captionStart="TABLE 2" captionStartId="15.[264,352,403,427]" captionTargetBox="[277,1287,512,1598]" captionTargetPageId="15" captionText="TABLE 2. Mensural variation in the type series of Nactus sphaerodactylodes from Sudest Island," httpUri="http://table.plazi.org/id/DF536630FFF9FFB5303F0698FAF9D498" pageId="14" pageNumber="15" tableUuid="DF536630FFF9FFB5303F0698FAF9D498">Table 2</tableCitation>
|
||
). Supralabials to mideye are either four or five, and total number of supralabials is either six or seven. No obvious trend in number is evident between the sexes. Number of infralabials varies from five to seven, with five being more common in my sample of females (n = 9/18 = 50% of individuals) than males (n = 2/11 = 18%). Number of lamellae under the fourth toe varies from 11–13 (mean = 11.6) in males and 10–13 (mean = 11.4) in females. In only two males is there a disparity in lamellae number between sides of the same animal as great as two. The large majority of animals have either the same number of lamellae on each fourth toe or only differ by one.
|
||
</paragraph>
|
||
<paragraph id="8B9336B8FFF8FFB4300F014FFC50D215" blockId="14.[264,1323,1244,1870]" pageId="14" pageNumber="15">The clearest sexual dimorphism lies in the preanal and femoral pores, which are present in males and lacking in females. Male preanal pore number varies from 14–18 (mean = 16.0) and femoral pore number varies from 4–18 (mean = 7.2), with all except one thigh having no more than nine pores. Femoral pore disparity between thighs of the same animal varies from 1.00 (equal numbers on each thigh) to 2.00 (twice as many on one thigh as on the other). The mean disparity is 1.28.</paragraph>
|
||
<paragraph id="8B9336B8FFF8FFB5300F003FFB94D405" blockId="14.[264,1323,1244,1870]" lastBlockId="15.[264,1094,284,310]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">Color and pattern are virtually identical among animals, with some animals in life showing only a bit more orangebrown wash on the tail than the mode.</paragraph>
|
||
<caption id="DF536630FFF9FFB5303F0698FAF9D498" ID-Table-UUID="DF536630FFF9FFB5303F0698FAF9D498" box="[264,1323,403,427]" httpUri="http://table.plazi.org/id/DF536630FFF9FFB5303F0698FAF9D498" pageId="15" pageNumber="16" targetBox="[277,1287,512,1598]" targetIsTable="true" targetPageId="15">
|
||
<paragraph id="8B9336B8FFF9FFB5303F0698FAF9D498" blockId="15.[264,1323,403,473]" box="[264,1323,403,427]" pageId="15" pageNumber="16">
|
||
<emphasis id="B958EAAAFFF9FFB5303F0698FEAED498" bold="true" box="[264,380,403,427]" pageId="15" pageNumber="16">TABLE 2.</emphasis>
|
||
Mensural variation in the type series of
|
||
<taxonomicName id="4C2C4D3BFFF9FFB53201069FFB9DD498" box="[822,1103,404,427]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="sphaerodactylodes">
|
||
<emphasis id="B958EAAAFFF9FFB53201069FFB9DD498" box="[822,1103,404,427]" italics="true" pageId="15" pageNumber="16">Nactus sphaerodactylodes</emphasis>
|
||
</taxonomicName>
|
||
from Sudest Island,
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9336B8FFF9FFB5303F06CAFD0CD4EA" blockId="15.[264,1323,403,473]" box="[264,734,447,473]" pageId="15" pageNumber="16">
|
||
Louisiade Archipelago,
|
||
<collectingCountry id="F33B7628FFF9FFB5333F06CAFD04D4EB" box="[520,726,449,472]" name="Papua New Guinea" pageId="15" pageNumber="16">Papua New Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9336B8FFF9FFB53022050BFB28D30E" pageId="15" pageNumber="16">
|
||
<table id="F92CC418FFF900453022050BFAD5D30D" box="[277,1287,512,1598]" gridcols="3" gridrows="23" pageId="15" pageNumber="16">
|
||
<tr id="351C34FAFFF900453022050BFAD5D724" box="[277,1287,512,535]" gridrow="0" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF900453022050BFEAFD724" box="[277,381,512,535]" gridcol="0" gridrow="0" pageId="15" pageNumber="16">Character</th>
|
||
<th id="76CD5D86FFF900453308050BFCBED724" box="[575,876,512,535]" gridcol="1" gridrow="0" pageId="15" pageNumber="16">Males (n = 11)</th>
|
||
<th id="76CD5D86FFF9004532EF050BFAD5D724" box="[984,1287,512,535]" gridcol="2" gridrow="0" pageId="15" pageNumber="16">Females (n = 18)</th>
|
||
</tr>
|
||
<tr id="351C34FAFFF9004530220526FAD5D777" box="[277,1287,557,580]" gridrow="1" pageId="15" pageNumber="16" rowspan-0="1">
|
||
<td id="76CD5D86FFF9004533080526FCBED777" box="[575,876,557,580]" gridcol="1" gridrow="1" pageId="15" pageNumber="16">mean range</td>
|
||
<td id="76CD5D86FFF9004532EF0526FAD5D777" box="[984,1287,557,580]" gridcol="2" gridrow="1" pageId="15" pageNumber="16">mean range</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF9004530220550FAD5D746" box="[277,1287,603,629]" gridrow="2" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF9004530220550FEAFD746" box="[277,381,603,629]" gridcol="0" gridrow="2" pageId="15" pageNumber="16">SV (mm)</th>
|
||
<td id="76CD5D86FFF9004533080550FCBED746" box="[575,876,603,629]" gridcol="1" gridrow="2" pageId="15" pageNumber="16">23.6 21.3–25.4</td>
|
||
<td id="76CD5D86FFF9004532EF0550FAD5D746" box="[984,1287,603,629]" gridcol="2" gridrow="2" pageId="15" pageNumber="16">24.6 20.4–26.0</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF9004530220580FAD5D795" box="[277,1287,651,678]" gridrow="3" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF9004530220580FEAFD795" box="[277,381,651,678]" gridcol="0" gridrow="3" pageId="15" pageNumber="16">TrL/SV</th>
|
||
<td id="76CD5D86FFF9004533080580FCBED795" box="[575,876,651,678]" gridcol="1" gridrow="3" pageId="15" pageNumber="16">0.43 0.42–0.46</td>
|
||
<td id="76CD5D86FFF9004532EF0580FAD5D795" box="[984,1287,651,678]" gridcol="2" gridrow="3" pageId="15" pageNumber="16">0.44 0.39–0.48</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF90045302205B7FAD5D7E5" box="[277,1287,700,726]" gridrow="4" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF90045302205B7FEAFD7E5" box="[277,381,700,726]" gridcol="0" gridrow="4" pageId="15" pageNumber="16">CS/SV</th>
|
||
<td id="76CD5D86FFF90045330805B7FCBED7E5" box="[575,876,700,726]" gridcol="1" gridrow="4" pageId="15" pageNumber="16">0.16 0.15–0.17</td>
|
||
<td id="76CD5D86FFF9004532EF05B7FAD5D7E5" box="[984,1287,700,726]" gridcol="2" gridrow="4" pageId="15" pageNumber="16">0.16 0.15–0.18</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF90045302205E7FAD5D634" box="[277,1287,748,775]" gridrow="5" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF90045302205E7FEAFD634" box="[277,381,748,775]" gridcol="0" gridrow="5" pageId="15" pageNumber="16">TL/SV</th>
|
||
<td id="76CD5D86FFF90045330805E7FCBED634" box="[575,876,748,775]" gridcol="1" gridrow="5" pageId="15" pageNumber="16">0.82 0.65–0.96</td>
|
||
<td id="76CD5D86FFF9004532EF05E7FAD5D634" box="[984,1287,748,775]" gridcol="2" gridrow="5" pageId="15" pageNumber="16">0.84 0.68–0.98</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF9004530220417FAD5D604" box="[277,1287,796,823]" gridrow="6" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF9004530220417FEAFD604" box="[277,381,796,823]" gridcol="0" gridrow="6" pageId="15" pageNumber="16">TW/SV</th>
|
||
<td id="76CD5D86FFF9004533080417FCBED604" box="[575,876,796,823]" gridcol="1" gridrow="6" pageId="15" pageNumber="16">0.11 0.10–0.12</td>
|
||
<td id="76CD5D86FFF9004532EF0417FAD5D604" box="[984,1287,796,823]" gridcol="2" gridrow="6" pageId="15" pageNumber="16">0.11 0.10–0.13</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF9004530220446FAD5D65B" box="[277,1287,845,872]" gridrow="7" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF9004530220446FEAFD65B" box="[277,381,845,872]" gridcol="0" gridrow="7" pageId="15" pageNumber="16">HL/SV</th>
|
||
<td id="76CD5D86FFF9004533080446FCBED65B" box="[575,876,845,872]" gridcol="1" gridrow="7" pageId="15" pageNumber="16">0.26 0.25–0.27</td>
|
||
<td id="76CD5D86FFF9004532EF0446FAD5D65B" box="[984,1287,845,872]" gridcol="2" gridrow="7" pageId="15" pageNumber="16">0.26 0.25–0.28</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF9004530220476FAD5D6AB" box="[277,1287,893,920]" gridrow="8" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF9004530220476FEAFD6AB" box="[277,381,893,920]" gridcol="0" gridrow="8" pageId="15" pageNumber="16">HW/SV</th>
|
||
<td id="76CD5D86FFF9004533080476FCBED6AB" box="[575,876,893,920]" gridcol="1" gridrow="8" pageId="15" pageNumber="16">0.20 0.19–0.21</td>
|
||
<td id="76CD5D86FFF9004532EF0476FAD5D6AB" box="[984,1287,893,920]" gridcol="2" gridrow="8" pageId="15" pageNumber="16">0.20 0.18–0.20</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF90045302204A5FAD5D6FA" box="[277,1287,942,969]" gridrow="9" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF90045302204A5FEAFD6FA" box="[277,381,942,969]" gridcol="0" gridrow="9" pageId="15" pageNumber="16">HH/SV</th>
|
||
<td id="76CD5D86FFF90045330804A5FCBED6FA" box="[575,876,942,969]" gridcol="1" gridrow="9" pageId="15" pageNumber="16">0.12 0.11–0.14</td>
|
||
<td id="76CD5D86FFF9004532EF04A5FAD5D6FA" box="[984,1287,942,969]" gridcol="2" gridrow="9" pageId="15" pageNumber="16">0.13 0.10–0.15</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF90045302204D5FAD5D6CA" box="[277,1287,990,1017]" gridrow="10" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF90045302204D5FEAFD6CA" box="[277,381,990,1017]" gridcol="0" gridrow="10" pageId="15" pageNumber="16">FA/ SV</th>
|
||
<td id="76CD5D86FFF90045330804D5FCBED6CA" box="[575,876,990,1017]" gridcol="1" gridrow="10" pageId="15" pageNumber="16">0.13 0.12–0.14</td>
|
||
<td id="76CD5D86FFF9004532EF04D5FAD5D6CA" box="[984,1287,990,1017]" gridcol="2" gridrow="10" pageId="15" pageNumber="16">0.13 0.11–0.14</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF9004530220304FAD5D11A" box="[277,1287,1039,1065]" gridrow="11" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF9004530220304FEAFD11A" box="[277,381,1039,1065]" gridcol="0" gridrow="11" pageId="15" pageNumber="16">EY/SV</th>
|
||
<td id="76CD5D86FFF9004533080304FCBED11A" box="[575,876,1039,1065]" gridcol="1" gridrow="11" pageId="15" pageNumber="16">0.051 0.047–0.054</td>
|
||
<td id="76CD5D86FFF9004532EF0304FAD5D11A" box="[984,1287,1039,1065]" gridcol="2" gridrow="11" pageId="15" pageNumber="16">0.050 0.046–0.054</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF9004530220334FAD5D169" box="[277,1287,1087,1114]" gridrow="12" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF9004530220334FEAFD169" box="[277,381,1087,1114]" gridcol="0" gridrow="12" pageId="15" pageNumber="16">EN/SV</th>
|
||
<td id="76CD5D86FFF9004533080334FCBED169" box="[575,876,1087,1114]" gridcol="1" gridrow="12" pageId="15" pageNumber="16">0.078 0.070–0.084</td>
|
||
<td id="76CD5D86FFF9004532EF0334FAD5D169" box="[984,1287,1087,1114]" gridcol="2" gridrow="12" pageId="15" pageNumber="16">0.077 0.073–0.086</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF900453022037BFAD5D1B9" box="[277,1287,1136,1162]" gridrow="13" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF900453022037BFEAFD1B9" box="[277,381,1136,1162]" gridcol="0" gridrow="13" pageId="15" pageNumber="16">IN/SV</th>
|
||
<td id="76CD5D86FFF900453308037BFCBED1B9" box="[575,876,1136,1162]" gridcol="1" gridrow="13" pageId="15" pageNumber="16">0.043 0.038–0.049</td>
|
||
<td id="76CD5D86FFF9004532EF037BFAD5D1B9" box="[984,1287,1136,1162]" gridcol="2" gridrow="13" pageId="15" pageNumber="16">0.043 0.038–0.049</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF90045302203ABFAD5D188" box="[277,1287,1184,1211]" gridrow="14" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF90045302203ABFEAFD188" box="[277,381,1184,1211]" gridcol="0" gridrow="14" pageId="15" pageNumber="16">SN/SV</th>
|
||
<td id="76CD5D86FFF90045330803ABFCBED188" box="[575,876,1184,1211]" gridcol="1" gridrow="14" pageId="15" pageNumber="16">0.11 0.09–0.11</td>
|
||
<td id="76CD5D86FFF9004532EF03ABFAD5D188" box="[984,1287,1184,1211]" gridcol="2" gridrow="14" pageId="15" pageNumber="16">0.11 0.10–0.12</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF90045302203DBFAD5D1D8" box="[277,1287,1232,1259]" gridrow="15" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF90045302203DBFEAFD1D8" box="[277,381,1232,1259]" gridcol="0" gridrow="15" pageId="15" pageNumber="16">EE/SV</th>
|
||
<td id="76CD5D86FFF90045330803DBFCBED1D8" box="[575,876,1232,1259]" gridcol="1" gridrow="15" pageId="15" pageNumber="16">0.091 0.085–0.100</td>
|
||
<td id="76CD5D86FFF9004532EF03DBFAD5D1D8" box="[984,1287,1232,1259]" gridcol="2" gridrow="15" pageId="15" pageNumber="16">0.093 0.074–0.103</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF900453022020AFAD5D02F" box="[277,1287,1281,1308]" gridrow="16" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF900453022020AFEAFD02F" box="[277,381,1281,1308]" gridcol="0" gridrow="16" pageId="15" pageNumber="16">IO/SV</th>
|
||
<td id="76CD5D86FFF900453308020AFCBED02F" box="[575,876,1281,1308]" gridcol="1" gridrow="16" pageId="15" pageNumber="16">0.072 0.068–0.077</td>
|
||
<td id="76CD5D86FFF9004532EF020AFAD5D02F" box="[984,1287,1281,1308]" gridcol="2" gridrow="16" pageId="15" pageNumber="16">0.074 0.063–0.083</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF900453022023AFAD5D07F" box="[277,1287,1329,1356]" gridrow="17" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF900453022023AFEAFD07F" box="[277,381,1329,1356]" gridcol="0" gridrow="17" pageId="15" pageNumber="16">TW/TL</th>
|
||
<td id="76CD5D86FFF900453308023AFCBED07F" box="[575,876,1329,1356]" gridcol="1" gridrow="17" pageId="15" pageNumber="16">0.14 0.12–0.16</td>
|
||
<td id="76CD5D86FFF9004532EF023AFAD5D07F" box="[984,1287,1329,1356]" gridcol="2" gridrow="17" pageId="15" pageNumber="16">0.13 0.12–0.17</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF9004530220269FAD5D04F" box="[277,1287,1378,1404]" gridrow="18" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF9004530220269FEAFD04F" box="[277,381,1378,1404]" gridcol="0" gridrow="18" pageId="15" pageNumber="16">HW/HL</th>
|
||
<td id="76CD5D86FFF9004533080269FCBED04F" box="[575,876,1378,1404]" gridcol="1" gridrow="18" pageId="15" pageNumber="16">0.75 0.71–0.79</td>
|
||
<td id="76CD5D86FFF9004532EF0269FAD5D04F" box="[984,1287,1378,1404]" gridcol="2" gridrow="18" pageId="15" pageNumber="16">0.75 0.71–0.82</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF9004530220299FAD5D09E" box="[277,1287,1426,1453]" gridrow="19" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF9004530220299FEAFD09E" box="[277,381,1426,1453]" gridcol="0" gridrow="19" pageId="15" pageNumber="16">EY/EN</th>
|
||
<td id="76CD5D86FFF9004533080299FCBED09E" box="[575,876,1426,1453]" gridcol="1" gridrow="19" pageId="15" pageNumber="16">0.65 0.60–0.76</td>
|
||
<td id="76CD5D86FFF9004532EF0299FAD5D09E" box="[984,1287,1426,1453]" gridcol="2" gridrow="19" pageId="15" pageNumber="16">0.65 0.58–0.74</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF90045302202C8FAD5D0EE" box="[277,1287,1475,1501]" gridrow="20" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF90045302202C8FEAFD0EE" box="[277,381,1475,1501]" gridcol="0" gridrow="20" pageId="15" pageNumber="16">EN/IN</th>
|
||
<td id="76CD5D86FFF90045330802C8FCBED0EE" box="[575,876,1475,1501]" gridcol="1" gridrow="20" pageId="15" pageNumber="16">1.8 1.5–2.1</td>
|
||
<td id="76CD5D86FFF9004532EF02C8FAD5D0EE" box="[984,1287,1475,1501]" gridcol="2" gridrow="20" pageId="15" pageNumber="16">1.8 1.6–2.0</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF90045302202F8FAD5D33D" box="[277,1287,1523,1550]" gridrow="21" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF90045302202F8FEAFD33D" box="[277,381,1523,1550]" gridcol="0" gridrow="21" pageId="15" pageNumber="16">SN/HL</th>
|
||
<td id="76CD5D86FFF90045330802F8FCBED33D" box="[575,876,1523,1550]" gridcol="1" gridrow="21" pageId="15" pageNumber="16">0.40 0.34–0.42</td>
|
||
<td id="76CD5D86FFF9004532EF02F8FAD5D33D" box="[984,1287,1523,1550]" gridcol="2" gridrow="21" pageId="15" pageNumber="16">0.41 0.40–0.43</td>
|
||
</tr>
|
||
<tr id="351C34FAFFF900453022012FFAD5D30D" box="[277,1287,1572,1598]" gridrow="22" pageId="15" pageNumber="16">
|
||
<th id="76CD5D86FFF900453022012FFEAFD30D" box="[277,381,1572,1598]" gridcol="0" gridrow="22" pageId="15" pageNumber="16">HH/HL</th>
|
||
<td id="76CD5D86FFF900453308012FFCBED30D" box="[575,876,1572,1598]" gridcol="1" gridrow="22" pageId="15" pageNumber="16">0.47 0.41–0.56</td>
|
||
<td id="76CD5D86FFF9004532EF012FFAD5D30D" box="[984,1287,1572,1598]" gridcol="2" gridrow="22" pageId="15" pageNumber="16">0.49 0.39–0.59</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8B9336B8FFF9FFAA300F0185FDCCD46D" blockId="15.[264,1322,1678,1904]" lastBlockId="16.[264,1323,284,550]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
|
||
<emphasis id="B958EAAAFFF9FFB5300F0185FE4AD394" box="[312,408,1678,1703]" italics="true" pageId="15" pageNumber="16">Ecology</emphasis>
|
||
: All individuals were found just inland from the coast at elevations no greater than
|
||
<quantity id="4CD49B5DFFF9FFB5307401BDFE53D3E3" box="[323,385,1718,1744]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="16" unit="m" value="40.0">40 m</quantity>
|
||
and inhabited disturbed forest or gardens in all cases. Animals were collected under rocks, logs, or rotted banana stumps on either moist or dry soil and would scamper into adjacent leaf litter when disturbed. One specimen was found adjacent (<
|
||
<quantity id="4CD49B5DFFF9FFB535B9000DFB65D213" box="[1166,1207,1798,1824]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="m" value="1.0">1 m</quantity>
|
||
distance) to a stream but most were found away from the immediate vicinity of water. Specimens were not found in closedcanopy rainforest despite more search effort being conducted in that habitat. These geckos often attempted escape by jumping (for admittedly short distances) or wriggling.
|
||
</paragraph>
|
||
<paragraph id="8B9336B8FFE6FFAA300F0667FCE6D715" blockId="16.[264,1323,284,550]" pageId="16" pageNumber="17">
|
||
To avoid damaging the specimens, no attempt was made to investigate reproductive status of the entire
|
||
<typeStatus id="5497881AFFE6FFAA30DE069FFDCED49D" box="[489,540,404,430]" pageId="16" pageNumber="17">type</typeStatus>
|
||
series. However, two examined males (BPBM 20762, 21692) had swollen testes and epididymes and three females (BPBM 20765, 20772, 21695) had single large eggs occupying most of their posterior abdominal cavities. Hence, reproduction occurs in at least some specimens during April.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3366533FFE6FFAA300F0557FDB8D655" pageId="16" pageNumber="17" type="etymology">
|
||
<paragraph id="8B9336B8FFE6FFAA300F0557FCFFD7F5" blockId="16.[264,1323,604,710]" pageId="16" pageNumber="17">
|
||
<emphasis id="B958EAAAFFE6FFAA300F0557FE67D746" box="[312,437,604,629]" italics="true" pageId="16" pageNumber="17">Etymology</emphasis>
|
||
: The name is a combination of the genus name “
|
||
<taxonomicName id="4C2C4D3BFFE6FFAA32CB0557FB6CD746" box="[1020,1214,604,629]" class="Reptilia" family="Sphaerodactylidae" genus="Sphaerodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
|
||
<emphasis id="B958EAAAFFE6FFAA32CB0557FB6CD746" box="[1020,1214,604,629]" italics="true" pageId="16" pageNumber="17">Sphaerodactylus</emphasis>
|
||
</taxonomicName>
|
||
” and the Greek suffix “odes”, meaning “like”, in reference to the superficial resemblance of the new species to geckos of that American genus.
|
||
</paragraph>
|
||
<paragraph id="8B9336B8FFE6FFAA300F05F7FDB8D655" blockId="16.[264,1325,764,870]" pageId="16" pageNumber="17">
|
||
<emphasis id="B958EAAAFFE6FFAA300F05F7FE51D626" box="[312,387,764,789]" italics="true" pageId="16" pageNumber="17">Range</emphasis>
|
||
: Known from Sudest, Misima, and Woodlark islands (
|
||
<figureCitation id="13172A3DFFE6FFAA352405F7FB8FD625" box="[1043,1117,764,790]" captionStart="FIGURE 7" captionStartId="20.[264,367,1608,1632]" captionTargetBox="[321,1266,969,1558]" captionTargetId="figure@20.[321,1267,969,1558]" captionTargetPageId="20" captionText="FIGURE 7. Map of the offshore islands of Milne Bay Province, Papua New Guinea, showing the known locations for Nactus sphaerodactylodes and N. acutus. Closed circle is type locality for N. sphaerodactylodes; open circles are other locations for N. sphaerodactylodes; closed square is type locality for N. acutus." httpUri="https://zenodo.org/record/170185/files/figure.png" pageId="16" pageNumber="17">Fig. 7</figureCitation>
|
||
). Likely to occur on the smaller Louisiades lying between Sudest and Misima, which apparently have yet to be explored by herpetologists.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3366533FFE6FFAA300F0497FE5CD0A5" pageId="16" pageNumber="17" type="discussion">
|
||
<paragraph id="8B9336B8FFE6FFAA300F0497FE5CD0A5" blockId="16.[264,1324,924,1430]" pageId="16" pageNumber="17">
|
||
<emphasis id="B958EAAAFFE6FFAA300F0497FE4CD686" box="[312,414,924,949]" italics="true" pageId="16" pageNumber="17">Remarks</emphasis>
|
||
: The single known specimen from Woodlark Island (BPBM 17707) differs from the Sudest and Misima specimens in the distribution of its femoral and preanal pores. Most specimens of
|
||
<taxonomicName id="4C2C4D3BFFE6FFAA30C404E7FCFBD136" box="[499,809,1004,1029]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="sphaerodactylodes">
|
||
<emphasis id="B958EAAAFFE6FFAA30C404E7FCFBD136" box="[499,809,1004,1029]" italics="true" pageId="16" pageNumber="17">Nactus sphaerodactylodes</emphasis>
|
||
</taxonomicName>
|
||
have 24–33 preanalfemoral pores arrayed in three clusters, with generally 4–9 pores in each femoral patch. In contrast, the Woodlark specimen has 46 pores, with 14 arrayed in each femoral patch. This is a striking difference and might suggest the presence of a sibling species on Woodlark. However, a single Sudest specimen also has a large number of femoral pores, with 41 pores in total and 9 of these in one femoral patch, with
|
||
<quantity id="4CD49B5DFFE6FFAA331B03BFFDBBD1FD" box="[556,617,1204,1230]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="16" pageNumber="17" unit="in" value="18.0">18 in</quantity>
|
||
the other. Hence, the increase in numbers of femoral pores may be an uncommon variant within
|
||
<taxonomicName id="4C2C4D3BFFE6FFAA33F203D7FC11D1C6" box="[709,963,1244,1269]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="sphaerodactylodes">
|
||
<emphasis id="B958EAAAFFE6FFAA33F203D7FC11D1C6" box="[709,963,1244,1269]" italics="true" pageId="16" pageNumber="17">N. sphaerodactylodes</emphasis>
|
||
</taxonomicName>
|
||
. Given the observed variation in this character within Sudest specimens and the fact that I can find no other feature to distinguish the sole Woodlark specimen from the Sudest and Misima lizards, I assign the Woodlark animal to
|
||
<taxonomicName id="4C2C4D3BFFE6FFAA3337025FFD2DD05E" box="[512,767,1364,1389]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="sphaerodactylodes">
|
||
<emphasis id="B958EAAAFFE6FFAA3337025FFD2DD05E" box="[512,767,1364,1389]" italics="true" pageId="16" pageNumber="17">N. sphaerodactylodes</emphasis>
|
||
</taxonomicName>
|
||
but note that further research might alter that conclusion.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |