treatments-xml/data/03/85/87/03858791FFEAFFF1FF67F908FF14F0D9.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.6213074" ID-GBIF-Dataset="e04117e5-9306-489d-a0a3-9dbd97774f74" ID-GBIF-Taxon="119613330" ID-ISSN="1175-5326" ID-Zenodo-Dep="6213074" checkinTime="1461057042134" checkinUser="plazi" docAuthor="Hong, Yong &amp; James, Samuel W." docDate="2009" docId="03858791FFEAFFF1FF67F908FF14F0D9" docLanguage="en" docName="zt02059p045.pdf" docOrigin="Zootaxa 2059" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@46f0bf0d" docStyleName="zootaxa.2007.monograph" docTitle="Pheretima bicolensis Hong &amp; James, 2009, sp. nov." docType="treatment" docVersion="8" lastPageId="4" lastPageNumber="37" masterDocId="FFBCFFE9FFEBFFF5FFF0FF8AFFEBF41B" masterDocTitle="New earthworms of the Pheretima urceolata species group (Oligochaeta: Megascolecidae) from southern Luzon, Philippines" masterLastPageNumber="45" masterPageNumber="33" pageId="1" pageNumber="34" updateTime="1645486389336" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>New earthworms of the Pheretima urceolata species group (Oligochaeta: Megascolecidae) from southern Luzon, Philippines</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hong, Yong</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>James, Samuel W.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="volume">
<mods:number>2059</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>33</mods:start>
<mods:end>45</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.186796</mods:identifier>
<mods:identifier type="GBIF-Dataset">e04117e5-9306-489d-a0a3-9dbd97774f74</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">186796</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6213074" ID-GBIF-Taxon="119613330" ID-Zenodo-Dep="6213074" LSID="urn:lsid:plazi:treatment:03858791FFEAFFF1FF67F908FF14F0D9" httpUri="http://treatment.plazi.org/id/03858791FFEAFFF1FF67F908FF14F0D9" lastPageId="4" lastPageNumber="37" pageId="1" pageNumber="34">
<subSubSection pageId="1" pageNumber="34" type="nomenclature">
<paragraph blockId="1.[151,503,1666,1727]" box="[151,503,1666,1692]" pageId="1" pageNumber="34">
<heading bold="true" box="[151,503,1666,1692]" fontSize="11" level="1" pageId="1" pageNumber="34" reason="1">
<emphasis bold="true" box="[151,503,1666,1692]" pageId="1" pageNumber="34">
<taxonomicName box="[151,403,1666,1692]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="34" phylum="Annelida" rank="species" species="bicolensis" status="sp. nov.">
<emphasis bold="true" box="[151,403,1666,1692]" italics="true" pageId="1" pageNumber="34">Pheretima bicolensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[410,503,1666,1692]" pageId="1" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="1.[151,503,1666,1727]" box="[151,294,1701,1727]" pageId="1" pageNumber="34">
(
<figureCitation box="[159,224,1701,1727]" captionStart="FIGURE 1" captionStartId="2.[151,255,1523,1547]" captionTargetBox="[218,1357,245,1468]" captionTargetId="figure@2.[208,1365,237,1489]" captionTargetPageId="2" captionText="FIGURE 1. Pheretima bicolensis sp. nov. A: ventral view; B: spermatheca. Pheretima viracensis sp. nov. C: ventral view; D: spermatheca. Pheretima camarinensis sp. nov. E: ventral view; F: spermatheca. Scale bars = 2.5 mm (ventral views), 1 mm (spermathecae)." httpUri="https://zenodo.org/record/186797/files/figure.png" pageId="1" pageNumber="34">
Fig. 
<number box="[214,224,1701,1727]" pageId="1" pageNumber="34" value="1.0">1</number>
</figureCitation>
A–B)
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="34" type="materials_examined">
<paragraph blockId="1.[151,1436,1781,2007]" pageId="1" pageNumber="34">
<emphasis bold="true" box="[151,267,1781,1807]" pageId="1" pageNumber="34">Material.</emphasis>
<typeStatus box="[276,390,1781,1807]" pageId="1" pageNumber="34">Holotype</typeStatus>
: Clitellate (NMA 004315): 
<collectingCountry box="[717,852,1781,1807]" name="Philippines" pageId="1" pageNumber="34">Philippines</collectingCountry>
, Catanduanes province, Catanduanes Island, near village of Summit, 
<quantity box="[377,454,1821,1847]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="1" pageNumber="34" unit="m" value="275.0">275 m</quantity>
, lowland forest, 
<date box="[649,807,1821,1847]" pageId="1" pageNumber="34" value="2001-05-22">
<number box="[649,678,1821,1847]" pageId="1" pageNumber="34" value="22.0">22</number>
May 
<number box="[746,807,1821,1847]" pageId="1" pageNumber="34" value="2001.0">2001</number>
</date>
, S. W. James coll. Other material: 
<number box="[1213,1227,1821,1847]" pageId="1" pageNumber="34" value="2.0">2</number>
clitellates, 
<number box="[1362,1377,1821,1847]" pageId="1" pageNumber="34" value="1.0">1</number>
aclitellate, same data as for 
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.
</paragraph>
</subSubSection>
<subSubSection box="[197,996,1901,1927]" pageId="1" pageNumber="34" type="etymology">
<paragraph blockId="1.[151,1436,1781,2007]" box="[197,996,1901,1927]" pageId="1" pageNumber="34">
<emphasis bold="true" box="[197,338,1901,1927]" pageId="1" pageNumber="34">Etymology.</emphasis>
The species is named for the region of the 
<typeStatus box="[844,895,1901,1927]" pageId="1" pageNumber="34">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="34" type="diagnosis">
<paragraph blockId="1.[151,1436,1781,2007]" pageId="1" pageNumber="34">
<emphasis bold="true" box="[197,325,1941,1967]" pageId="1" pageNumber="34">Diagnosis.</emphasis>
One pair of narrowly separated spermathecal pores in 
<number box="[978,992,1941,1967]" pageId="1" pageNumber="34" value="5.0">5</number>
/ 
<number box="[1003,1018,1941,1967]" pageId="1" pageNumber="34" value="6.0">6</number>
, no typhlosole, interior bursal pads two on the floor and one on the roof.
</paragraph>
</subSubSection>
<caption httpUri="https://zenodo.org/record/186797/files/figure.png" pageId="2" pageNumber="35" targetBox="[218,1357,245,1468]" targetPageId="2">
<paragraph blockId="2.[151,1436,1523,1616]" pageId="2" pageNumber="35">
<emphasis bold="true" box="[151,283,1523,1547]" pageId="2" pageNumber="35">FIGURE 1.</emphasis>
<taxonomicName box="[292,518,1523,1546]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="2" pageNumber="35" phylum="Annelida" rank="species" species="bicolensis" status="sp. nov.">
<emphasis box="[292,518,1523,1546]" italics="true" pageId="2" pageNumber="35">Pheretima bicolensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[527,615,1523,1547]" pageId="2" pageNumber="35">
<taxonomicNameLabel box="[527,615,1523,1547]" pageId="2" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A: ventral view; B: spermatheca. 
<taxonomicName box="[995,1222,1523,1546]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="2" pageNumber="35" phylum="Annelida" rank="species" species="viracensis" status="sp. nov.">
<emphasis box="[995,1222,1523,1546]" italics="true" pageId="2" pageNumber="35">Pheretima viracensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1231,1318,1523,1547]" pageId="2" pageNumber="35">
<taxonomicNameLabel box="[1231,1318,1523,1547]" pageId="2" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
C: ventral view; D: spermatheca. 
<taxonomicName box="[400,660,1558,1581]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="2" pageNumber="35" phylum="Annelida" rank="species" species="camarinensis" status="sp. nov.">
<emphasis box="[400,660,1558,1581]" italics="true" pageId="2" pageNumber="35">Pheretima camarinensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[668,755,1557,1581]" pageId="2" pageNumber="35">
<taxonomicNameLabel box="[668,755,1557,1581]" pageId="2" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
E: ventral view; F: spermatheca. Scale bars = 2.5 mm (ventral views), 1 mm (spermathecae).
</paragraph>
</caption>
<subSubSection lastPageId="4" lastPageNumber="37" pageId="2" pageNumber="35" type="description">
<paragraph blockId="2.[151,1436,1662,2014]" pageId="2" pageNumber="35">
<emphasis bold="true" box="[197,349,1662,1688]" pageId="2" pageNumber="35">Description.</emphasis>
Medium brown to dark brown dorsal pigment, unpigmented ventrally except on I–III. Dimensions 
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by 
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at segment X, 4.0– 
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or 4.0 mm at XXX, 
<quantity box="[1085,1228,1702,1728]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" metricValueMax="4.2" metricValueMin="3.5" pageId="2" pageNumber="35" unit="mm" value="3.85" valueMax="4.2" valueMin="3.5">3.5–4.2 mm</quantity>
at clitellum, segments 
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; body circular in cross-section. Setae numbering 
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at VII, 
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at XX; 
<number box="[1224,1267,1742,1768]" pageId="2" pageNumber="35" value="2.5" valueMax="3.0" valueMin="2.0">2–3</number>
between male pores, size larger in anterior dorsal segments, dorsal spacing greater, setal formula AA:AB:YZ:ZZ= 
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: 
<number box="[169,212,1822,1848]" pageId="2" pageNumber="35" value="2.5">2.5</number>
: 
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: 
<number box="[263,278,1822,1848]" pageId="2" pageNumber="35" value="5.0">5</number>
at XIII. Clitellum annular XIV–XVI; setae invisible externally.
</paragraph>
<paragraph blockId="2.[151,1436,1662,2014]" pageId="2" pageNumber="35">
First dorsal pores 
<number box="[414,445,1865,1891]" pageId="2" pageNumber="35" value="12.0">12</number>
/ 
<number box="[455,484,1865,1891]" pageId="2" pageNumber="35" value="13.0">13</number>
. One pair of spermathecal pores in 
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/ 
<number box="[937,951,1865,1891]" pageId="2" pageNumber="35" value="6.0">6</number>
at 
<number box="[991,1006,1865,1891]" pageId="2" pageNumber="35" value="3.0">3</number>
rd setal lines, near mid-ventral; spermathecal pores 
<number box="[344,461,1905,1931]" pageId="2" pageNumber="35" value="0.055" valueMax="0.06" valueMin="0.05">0.05–0.06</number>
circumference apart, distance between spermathecal pores around 
<quantity box="[1282,1432,1905,1931]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="2" pageNumber="35" unit="mm" value="0.75" valueMax="0.9" valueMin="0.6">0.6–0.9 mm</quantity>
; 
<quantity box="[151,293,1948,1974]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="35" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.3–0.4 mm</quantity>
openings of copulatory bursae paired in XVIII at 
<number box="[883,897,1948,1974]" pageId="2" pageNumber="35" value="7.0">7</number>
th setal lines, ventrally placed, 
<number box="[1254,1305,1948,1974]" pageId="2" pageNumber="35" value="0.14">0.14</number>
circumference apart, distance between openings 
<quantity box="[606,700,1988,2014]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="35" unit="mm" value="1.7">1.7 mm</quantity>
. Genital markings lacking.
</paragraph>
<paragraph blockId="3.[151,1170,191,218]" box="[151,1170,191,218]" pageId="3" pageNumber="36">
<emphasis bold="true" box="[151,277,191,217]" pageId="3" pageNumber="36">
<collectionCode box="[151,248,191,217]" pageId="3" pageNumber="36">TABLE</collectionCode>
<number box="[255,270,191,217]" pageId="3" pageNumber="36" value="1.0">1</number>
.
</emphasis>
Comparison of species of the 
<taxonomicName box="[634,745,192,217]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="36" phylum="Annelida" rank="species" species="urceolata">
<emphasis box="[634,745,192,217]" italics="true" pageId="3" pageNumber="36">urceolata</emphasis>
</taxonomicName>
group (modified from James 
<number box="[1094,1158,192,218]" pageId="3" pageNumber="36" value="2004.0">2004</number>
).
</paragraph>
<paragraph pageId="3" pageNumber="36">
<table box="[151,1418,250,1327]" gridcols="7" gridrows="22" pageId="3" pageNumber="36">
<tr box="[151,1418,250,301]" gridrow="0" pageId="3" pageNumber="36">
<th box="[151,452,250,301]" gridcol="0" gridrow="0" pageId="3" pageNumber="36">
<taxonomicName box="[353,452,250,301]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="36" phylum="Annelida" rank="species" species="abiadai">
<emphasis box="[353,452,250,301]" italics="true" pageId="3" pageNumber="36">Pheretima abiadai</emphasis>
</taxonomicName>
</th>
<th box="[472,588,250,301]" gridcol="1" gridrow="0" pageId="3" pageNumber="36">
<taxonomicName box="[492,588,250,271]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="36" phylum="Annelida" rank="species" species="nagaensis">
<emphasis box="[492,588,250,271]" italics="true" pageId="3" pageNumber="36">nagaensis</emphasis>
</taxonomicName>
</th>
<th box="[628,725,250,301]" gridcol="2" gridrow="0" pageId="3" pageNumber="36">
<taxonomicName box="[632,681,250,271]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="36" phylum="Annelida" rank="species" species="simsi">
<emphasis box="[632,681,250,271]" italics="true" pageId="3" pageNumber="36">simsi</emphasis>
</taxonomicName>
</th>
<th box="[748,1003,250,301]" gridcol="3" gridrow="0" pageId="3" pageNumber="36">
<emphasis box="[748,1003,250,301]" italics="true" pageId="3" pageNumber="36">
<taxonomicName box="[748,813,250,271]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="36" phylum="Annelida" rank="species" species="baletei">baletei</taxonomicName>
bukidnonen- sis
</emphasis>
</th>
<th box="[1023,1146,250,301]" gridcol="4" gridrow="0" pageId="3" pageNumber="36">
<emphasis box="[1023,1098,250,271]" italics="true" pageId="3" pageNumber="36">heaneyi</emphasis>
</th>
<th box="[1163,1284,250,301]" gridcol="5" gridrow="0" pageId="3" pageNumber="36">
<emphasis box="[1163,1284,250,301]" italics="true" pageId="3" pageNumber="36">kitangladen- sis</emphasis>
</th>
<th box="[1303,1418,250,301]" gridcol="6" gridrow="0" pageId="3" pageNumber="36">
<taxonomicName box="[1303,1408,250,271]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="36" phylum="Annelida" rank="species" species="monotheca">
<emphasis box="[1303,1408,250,271]" italics="true" pageId="3" pageNumber="36">monotheca</emphasis>
</taxonomicName>
</th>
</tr>
<tr box="[151,1418,319,370]" gridrow="1" pageId="3" pageNumber="36">
<th box="[151,452,319,370]" gridcol="0" gridrow="1" pageId="3" pageNumber="36">Setae vi, xx 32–42, 42–47</th>
<td box="[472,588,319,370]" gridcol="1" gridrow="1" pageId="3" pageNumber="36">31–38, 41–48</td>
<td box="[628,725,319,370]" gridcol="2" gridrow="1" pageId="3" pageNumber="36">31–33, 39–47</td>
<td box="[748,1003,319,370]" gridcol="3" gridrow="1" pageId="3" pageNumber="36">24, 40 38, 52</td>
<td box="[1023,1146,319,370]" gridcol="4" gridrow="1" pageId="3" pageNumber="36">38, 43</td>
<td box="[1163,1284,319,370]" gridcol="5" gridrow="1" pageId="3" pageNumber="36">44, 48</td>
<td box="[1303,1418,319,370]" gridcol="6" gridrow="1" pageId="3" pageNumber="36">36, 32</td>
</tr>
<tr box="[151,1418,389,410]" gridrow="2" pageId="3" pageNumber="36">
<th box="[151,452,389,410]" gridcol="0" gridrow="2" pageId="3" pageNumber="36">Setal gaps D, V D, V</th>
<td box="[472,588,389,410]" gridcol="1" gridrow="2" pageId="3" pageNumber="36">D, V</td>
<td box="[628,725,389,410]" gridcol="2" gridrow="2" pageId="3" pageNumber="36">D, V</td>
<td box="[748,1003,389,410]" gridcol="3" gridrow="2" pageId="3" pageNumber="36">none D</td>
<td box="[1023,1146,389,410]" gridcol="4" gridrow="2" pageId="3" pageNumber="36">D</td>
<td box="[1163,1284,389,410]" gridcol="5" gridrow="2" pageId="3" pageNumber="36">none</td>
<td box="[1303,1418,389,410]" gridcol="6" gridrow="2" pageId="3" pageNumber="36">D</td>
</tr>
<tr box="[151,1418,429,450]" gridrow="3" pageId="3" pageNumber="36">
<th box="[151,452,429,450]" gridcol="0" gridrow="3" pageId="3" pageNumber="36">Male pore setae 6–9</th>
<td box="[472,588,429,450]" gridcol="1" gridrow="3" pageId="3" pageNumber="36">6–9</td>
<td box="[628,725,429,450]" gridcol="2" gridrow="3" pageId="3" pageNumber="36">6–8</td>
<td box="[748,1003,429,450]" gridcol="3" gridrow="3" pageId="3" pageNumber="36">6 10</td>
<td box="[1023,1146,429,450]" gridcol="4" gridrow="3" pageId="3" pageNumber="36">6–8</td>
<td box="[1163,1284,429,450]" gridcol="5" gridrow="3" pageId="3" pageNumber="36">8</td>
<td box="[1303,1418,429,450]" gridcol="6" gridrow="3" pageId="3" pageNumber="36">3</td>
</tr>
<tr box="[151,1418,469,490]" gridrow="4" pageId="3" pageNumber="36">
<th box="[151,452,469,490]" gridcol="0" gridrow="4" pageId="3" pageNumber="36">Male pore spacing 0.19</th>
<td box="[472,588,469,490]" gridcol="1" gridrow="4" pageId="3" pageNumber="36">0.25</td>
<td box="[628,725,469,490]" gridcol="2" gridrow="4" pageId="3" pageNumber="36">0.28</td>
<td box="[748,1003,469,490]" gridcol="3" gridrow="4" pageId="3" pageNumber="36">0.24 0.17</td>
<td box="[1023,1146,469,490]" gridcol="4" gridrow="4" pageId="3" pageNumber="36">0.19–.23</td>
<td box="[1163,1284,469,490]" gridcol="5" gridrow="4" pageId="3" pageNumber="36">0.20</td>
<td box="[1303,1418,469,490]" gridcol="6" gridrow="4" pageId="3" pageNumber="36">0.11</td>
</tr>
<tr box="[151,1418,509,530]" gridrow="5" pageId="3" pageNumber="36">
<th box="[151,452,509,530]" gridcol="0" gridrow="5" pageId="3" pageNumber="36">Pigment Violet-red</th>
<td box="[472,588,509,530]" gridcol="1" gridrow="5" pageId="3" pageNumber="36">brown</td>
<td box="[628,725,509,530]" gridcol="2" gridrow="5" pageId="3" pageNumber="36">brown</td>
<td box="[748,1003,509,530]" gridcol="3" gridrow="5" pageId="3" pageNumber="36">Pale pink brown</td>
<td box="[1023,1146,509,530]" gridcol="4" gridrow="5" pageId="3" pageNumber="36">brown</td>
<td box="[1163,1284,509,530]" gridcol="5" gridrow="5" pageId="3" pageNumber="36">brown</td>
<td box="[1303,1418,509,530]" gridcol="6" gridrow="5" pageId="3" pageNumber="36">brown</td>
</tr>
<tr box="[151,1418,549,570]" gridrow="6" pageId="3" pageNumber="36">
<th box="[151,452,549,570]" gridcol="0" gridrow="6" pageId="3" pageNumber="36">Length (mm) 33–60</th>
<td box="[472,588,549,570]" gridcol="1" gridrow="6" pageId="3" pageNumber="36">36–53</td>
<td box="[628,725,549,570]" gridcol="2" gridrow="6" pageId="3" pageNumber="36">68–91</td>
<td box="[748,1003,549,570]" gridcol="3" gridrow="6" pageId="3" pageNumber="36">78 76</td>
<td box="[1023,1146,549,570]" gridcol="4" gridrow="6" pageId="3" pageNumber="36">83–116</td>
<td box="[1163,1284,549,570]" gridcol="5" gridrow="6" pageId="3" pageNumber="36">108</td>
<td box="[1303,1418,549,570]" gridcol="6" gridrow="6" pageId="3" pageNumber="36">62</td>
</tr>
<tr box="[151,1418,589,610]" gridrow="7" pageId="3" pageNumber="36">
<th box="[151,452,589,610]" gridcol="0" gridrow="7" pageId="3" pageNumber="36">Intestinal origin XVI</th>
<td box="[472,588,589,610]" gridcol="1" gridrow="7" pageId="3" pageNumber="36">XVI</td>
<td box="[628,725,589,610]" gridcol="2" gridrow="7" pageId="3" pageNumber="36">XV</td>
<td box="[748,1003,589,610]" gridcol="3" gridrow="7" pageId="3" pageNumber="36">XVII XVIII</td>
<td box="[1023,1146,589,610]" gridcol="4" gridrow="7" pageId="3" pageNumber="36">XVIII</td>
<td box="[1163,1284,589,610]" gridcol="5" gridrow="7" pageId="3" pageNumber="36">XVI</td>
<td box="[1303,1418,589,610]" gridcol="6" gridrow="7" pageId="3" pageNumber="36">XVIII</td>
</tr>
<tr box="[151,1418,629,650]" gridrow="8" pageId="3" pageNumber="36">
<th box="[151,452,629,650]" gridcol="0" gridrow="8" pageId="3" pageNumber="36">Typhlosole vestigial</th>
<td box="[472,588,629,650]" gridcol="1" gridrow="8" pageId="3" pageNumber="36">+</td>
<td box="[628,725,629,650]" gridcol="2" gridrow="8" pageId="3" pageNumber="36">+</td>
<td box="[748,1003,629,650]" gridcol="3" gridrow="8" pageId="3" pageNumber="36">- +</td>
<td box="[1023,1146,629,650]" gridcol="4" gridrow="8" pageId="3" pageNumber="36">+</td>
<td box="[1163,1284,629,650]" gridcol="5" gridrow="8" pageId="3" pageNumber="36">+</td>
<td box="[1303,1418,629,650]" gridcol="6" gridrow="8" pageId="3" pageNumber="36">+</td>
</tr>
<tr box="[151,1418,669,690]" gridrow="9" pageId="3" pageNumber="36" rowspan-1="1">
<th box="[151,452,669,690]" gridcol="0" gridrow="9" pageId="3" pageNumber="36">Intestinal vessels</th>
<td box="[628,725,669,690]" gridcol="2" gridrow="9" pageId="3" pageNumber="36">?</td>
<td box="[748,1003,669,690]" gridcol="3" gridrow="9" pageId="3" pageNumber="36">? 20</td>
<td box="[1023,1146,669,690]" gridcol="4" gridrow="9" pageId="3" pageNumber="36">28</td>
<td box="[1163,1284,669,690]" gridcol="5" gridrow="9" pageId="3" pageNumber="36">32</td>
<td box="[1303,1418,669,690]" gridcol="6" gridrow="9" pageId="3" pageNumber="36">36</td>
</tr>
<tr box="[151,1418,709,730]" gridrow="10" pageId="3" pageNumber="36">
<th box="[151,452,709,730]" gridcol="0" gridrow="10" pageId="3" pageNumber="36">Hearts XI – XIII</th>
<td box="[472,588,709,730]" gridcol="1" gridrow="10" pageId="3" pageNumber="36">XI–XIII</td>
<td box="[628,725,709,730]" gridcol="2" gridrow="10" pageId="3" pageNumber="36">X–XIII</td>
<td box="[748,1003,709,730]" gridcol="3" gridrow="10" pageId="3" pageNumber="36">X – XIII XII, XIII</td>
<td box="[1023,1146,709,730]" gridcol="4" gridrow="10" pageId="3" pageNumber="36">XI – XIII</td>
<td box="[1163,1284,709,730]" gridcol="5" gridrow="10" pageId="3" pageNumber="36">X – XIII</td>
<td box="[1303,1418,709,730]" gridcol="6" gridrow="10" pageId="3" pageNumber="36">X – XIII</td>
</tr>
<tr box="[151,1418,749,770]" gridrow="11" pageId="3" pageNumber="36">
<th box="[151,452,749,770]" gridcol="0" gridrow="11" pageId="3" pageNumber="36">Septa 8/9/10 -</th>
<td box="[472,588,749,770]" gridcol="1" gridrow="11" pageId="3" pageNumber="36">-</td>
<td box="[628,725,749,770]" gridcol="2" gridrow="11" pageId="3" pageNumber="36">+, -</td>
<td box="[748,1003,749,770]" gridcol="3" gridrow="11" pageId="3" pageNumber="36">-, + +, +</td>
<td box="[1023,1146,749,770]" gridcol="4" gridrow="11" pageId="3" pageNumber="36">+, +/-</td>
<td box="[1163,1284,749,770]" gridcol="5" gridrow="11" pageId="3" pageNumber="36">+, -</td>
<td box="[1303,1418,749,770]" gridcol="6" gridrow="11" pageId="3" pageNumber="36">-, -</td>
</tr>
<tr box="[151,1418,789,839]" gridrow="12" pageId="3" pageNumber="36">
<th box="[151,452,789,839]" gridcol="0" gridrow="12" pageId="3" pageNumber="36">Spermathecal 0.25 pore spacing</th>
<td box="[472,588,789,839]" gridcol="1" gridrow="12" pageId="3" pageNumber="36">0.29</td>
<td box="[628,725,789,839]" gridcol="2" gridrow="12" pageId="3" pageNumber="36">?</td>
<td box="[748,1003,789,839]" gridcol="3" gridrow="12" pageId="3" pageNumber="36">0.25 0.38</td>
<td box="[1023,1146,789,839]" gridcol="4" gridrow="12" pageId="3" pageNumber="36">0.19–.22</td>
<td box="[1163,1284,789,839]" gridcol="5" gridrow="12" pageId="3" pageNumber="36">0.28</td>
<td box="[1303,1418,789,839]" gridcol="6" gridrow="12" pageId="3" pageNumber="36">monothecal</td>
</tr>
<tr box="[151,1418,858,879]" gridrow="13" pageId="3" pageNumber="36">
<th box="[151,452,858,879]" gridcol="0" gridrow="13" pageId="3" pageNumber="36">Penes +</th>
<td box="[472,588,858,879]" gridcol="1" gridrow="13" pageId="3" pageNumber="36">+</td>
<td box="[628,725,858,879]" gridcol="2" gridrow="13" pageId="3" pageNumber="36">+</td>
<td box="[748,1003,858,879]" gridcol="3" gridrow="13" pageId="3" pageNumber="36">+ -</td>
<td box="[1023,1146,858,879]" gridcol="4" gridrow="13" pageId="3" pageNumber="36">+</td>
<td box="[1163,1284,858,879]" gridcol="5" gridrow="13" pageId="3" pageNumber="36">-</td>
<td box="[1303,1418,858,879]" gridcol="6" gridrow="13" pageId="3" pageNumber="36">+</td>
</tr>
<tr box="[151,1418,898,919]" gridrow="14" pageId="3" pageNumber="36">
<th box="[151,452,898,919]" gridcol="0" gridrow="14" pageId="3" pageNumber="36">Bursal pads 4R</th>
<td box="[472,588,898,919]" gridcol="1" gridrow="14" pageId="3" pageNumber="36">2S, 2F</td>
<td box="[628,725,898,919]" gridcol="2" gridrow="14" pageId="3" pageNumber="36">2F</td>
<td box="[748,1003,898,919]" gridcol="3" gridrow="14" pageId="3" pageNumber="36">0 0</td>
<td box="[1023,1146,898,919]" gridcol="4" gridrow="14" pageId="3" pageNumber="36">0</td>
<td box="[1163,1284,898,919]" gridcol="5" gridrow="14" pageId="3" pageNumber="36">2</td>
<td box="[1303,1418,898,919]" gridcol="6" gridrow="14" pageId="3" pageNumber="36">0</td>
</tr>
<tr box="[151,1418,938,959]" gridrow="15" pageId="3" pageNumber="36">
<th box="[151,452,938,959]" gridcol="0" gridrow="15" pageId="3" pageNumber="36">Bursal glands +</th>
<td box="[472,588,938,959]" gridcol="1" gridrow="15" pageId="3" pageNumber="36">+</td>
<td box="[628,725,938,959]" gridcol="2" gridrow="15" pageId="3" pageNumber="36">?</td>
<td box="[748,1003,938,959]" gridcol="3" gridrow="15" pageId="3" pageNumber="36">- +</td>
<td box="[1023,1146,938,959]" gridcol="4" gridrow="15" pageId="3" pageNumber="36">+</td>
<td box="[1163,1284,938,959]" gridcol="5" gridrow="15" pageId="3" pageNumber="36">-</td>
<td box="[1303,1418,938,959]" gridcol="6" gridrow="15" pageId="3" pageNumber="36">+</td>
</tr>
<tr box="[151,1418,1002,1025]" gridrow="16" pageId="3" pageNumber="36" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th box="[151,452,1002,1025]" gridcol="0" gridrow="16" pageId="3" pageNumber="36">continued</th>
</tr>
<tr box="[151,1418,1058,1109]" gridrow="17" pageId="3" pageNumber="36">
<th box="[151,452,1058,1109]" gridcol="0" gridrow="17" pageId="3" pageNumber="36">
<taxonomicName box="[349,440,1058,1079]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="36" phylum="Annelida" rank="species" species="urceolata">
<emphasis box="[349,440,1058,1079]" italics="true" pageId="3" pageNumber="36">urceolata</emphasis>
</taxonomicName>
</th>
<td box="[472,588,1058,1109]" gridcol="1" gridrow="17" pageId="3" pageNumber="36">
<taxonomicName box="[472,571,1058,1109]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="36" phylum="Annelida" rank="species" species="bicolensis">
<emphasis box="[472,571,1058,1109]" italics="true" pageId="3" pageNumber="36">Pheretima bicolensis</emphasis>
</taxonomicName>
</td>
<td box="[628,725,1058,1109]" gridcol="2" gridrow="17" pageId="3" pageNumber="36">
<taxonomicName box="[628,725,1058,1079]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="36" phylum="Annelida" rank="species" species="viracensis">
<emphasis box="[628,725,1058,1079]" italics="true" pageId="3" pageNumber="36">viracensis</emphasis>
</taxonomicName>
</td>
<td box="[748,1003,1058,1109]" gridcol="3" gridrow="17" pageId="3" pageNumber="36">
<emphasis box="[765,989,1058,1079]" italics="true" pageId="3" pageNumber="36">
<taxonomicName box="[765,893,1058,1079]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="36" phylum="Annelida" rank="species" species="camarinensis">camarinensis</taxonomicName>
<taxonomicName box="[926,989,1058,1079]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="36" phylum="Annelida" rank="species" species="doriae">doriae</taxonomicName>
</emphasis>
</td>
<td box="[1023,1146,1058,1109]" gridcol="4" gridrow="17" pageId="3" pageNumber="36">
<taxonomicName box="[1056,1146,1058,1079]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="36" phylum="Annelida" rank="species" species="batoensis">
<emphasis box="[1056,1146,1058,1079]" italics="true" pageId="3" pageNumber="36">batoensis</emphasis>
</taxonomicName>
</td>
<td box="[1163,1284,1058,1109]" gridcol="5" gridrow="17" pageId="3" pageNumber="36">
<taxonomicName box="[1189,1279,1058,1079]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="36" phylum="Annelida" rank="species" species="buhiensis">
<emphasis box="[1189,1279,1058,1079]" italics="true" pageId="3" pageNumber="36">buhiensis</emphasis>
</taxonomicName>
</td>
<td box="[1303,1418,1058,1109]" gridcol="6" gridrow="17" pageId="3" pageNumber="36">
<taxonomicName box="[1329,1390,1058,1079]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="36" phylum="Annelida" rank="species" species="gorasi">
<emphasis box="[1329,1390,1058,1079]" italics="true" pageId="3" pageNumber="36">gorasi</emphasis>
</taxonomicName>
</td>
</tr>
<tr box="[151,1418,1127,1178]" gridrow="18" pageId="3" pageNumber="36">
<th box="[151,452,1127,1178]" gridcol="0" gridrow="18" pageId="3" pageNumber="36">Setae vi, xx 20, 42</th>
<td box="[472,588,1127,1178]" gridcol="1" gridrow="18" pageId="3" pageNumber="36">29–34, 49–55</td>
<td box="[628,725,1127,1178]" gridcol="2" gridrow="18" pageId="3" pageNumber="36">37–38, 40–45</td>
<td box="[748,1003,1127,1178]" gridcol="3" gridrow="18" pageId="3" pageNumber="36">44–49, 35–40, 48–52 40–43</td>
<td box="[1023,1146,1127,1178]" gridcol="4" gridrow="18" pageId="3" pageNumber="36">36–43, 43–48</td>
<td box="[1163,1284,1127,1178]" gridcol="5" gridrow="18" pageId="3" pageNumber="36">31–50, 66–70</td>
<td box="[1303,1418,1127,1178]" gridcol="6" gridrow="18" pageId="3" pageNumber="36">24–32, 25–41</td>
</tr>
<tr box="[151,1418,1197,1218]" gridrow="19" pageId="3" pageNumber="36">
<th box="[151,452,1197,1218]" gridcol="0" gridrow="19" pageId="3" pageNumber="36">Setal gaps* D,?V</th>
<td box="[472,588,1197,1218]" gridcol="1" gridrow="19" pageId="3" pageNumber="36">V</td>
<td box="[628,725,1197,1218]" gridcol="2" gridrow="19" pageId="3" pageNumber="36">D</td>
<td box="[748,1003,1197,1218]" gridcol="3" gridrow="19" pageId="3" pageNumber="36">D D, V</td>
<td box="[1023,1146,1197,1218]" gridcol="4" gridrow="19" pageId="3" pageNumber="36">D, V</td>
<td box="[1163,1284,1197,1218]" gridcol="5" gridrow="19" pageId="3" pageNumber="36">D</td>
<td box="[1303,1418,1197,1218]" gridcol="6" gridrow="19" pageId="3" pageNumber="36">-</td>
</tr>
<tr box="[151,1418,1237,1258]" gridrow="20" pageId="3" pageNumber="36">
<th box="[151,452,1237,1258]" gridcol="0" gridrow="20" pageId="3" pageNumber="36">Male pore setae 6</th>
<td box="[472,588,1237,1258]" gridcol="1" gridrow="20" pageId="3" pageNumber="36">2–3</td>
<td box="[628,725,1237,1258]" gridcol="2" gridrow="20" pageId="3" pageNumber="36">7–10</td>
<td box="[748,1003,1237,1258]" gridcol="3" gridrow="20" pageId="3" pageNumber="36">6–9 5–8</td>
<td box="[1023,1146,1237,1258]" gridcol="4" gridrow="20" pageId="3" pageNumber="36">6–8</td>
<td box="[1163,1284,1237,1258]" gridcol="5" gridrow="20" pageId="3" pageNumber="36">0</td>
<td box="[1303,1418,1237,1258]" gridcol="6" gridrow="20" pageId="3" pageNumber="36">0</td>
</tr>
<tr box="[151,1418,1277,1327]" gridrow="21" pageId="3" pageNumber="36">
<th box="[151,452,1277,1327]" gridcol="0" gridrow="21" pageId="3" pageNumber="36">Male pore spac-? ing</th>
<td box="[472,588,1277,1327]" gridcol="1" gridrow="21" pageId="3" pageNumber="36">0.14</td>
<td box="[628,725,1277,1327]" gridcol="2" gridrow="21" pageId="3" pageNumber="36">0.20</td>
<td box="[748,1003,1277,1327]" gridcol="3" gridrow="21" pageId="3" pageNumber="36">0.19-.20 0.32</td>
<td box="[1023,1146,1277,1327]" gridcol="4" gridrow="21" pageId="3" pageNumber="36">0.15–.18</td>
<td box="[1163,1284,1277,1327]" gridcol="5" gridrow="21" pageId="3" pageNumber="36">0.07–.12</td>
<td box="[1303,1418,1277,1327]" gridcol="6" gridrow="21" pageId="3" pageNumber="36">0.10</td>
</tr>
</table>
</paragraph>
<paragraph blockId="3.[159,1390,1346,1396]" box="[159,1390,1346,1367]" pageId="3" pageNumber="36">Pigment red brown violet- light brown red-brown red-brown light brown brown</paragraph>
<paragraph blockId="3.[159,1390,1346,1396]" box="[628,689,1375,1396]" pageId="3" pageNumber="36">brown</paragraph>
<paragraph pageId="3" pageNumber="36">
<table box="[159,1418,1415,1556]" gridcols="8" gridrows="4" pageId="3" pageNumber="36">
<tr box="[159,1418,1415,1436]" gridrow="0" pageId="3" pageNumber="36">
<th box="[159,409,1415,1436]" gridcol="0" gridrow="0" pageId="3" pageNumber="36">Length (mm) 70–80</th>
<th box="[472,553,1415,1436]" gridcol="1" gridrow="0" pageId="3" pageNumber="36">71–98</th>
<th box="[628,688,1415,1436]" gridcol="2" gridrow="0" pageId="3" pageNumber="36">38–61</th>
<th box="[765,825,1415,1436]" gridcol="3" gridrow="0" pageId="3" pageNumber="36">54–61</th>
<th box="[926,987,1415,1436]" gridcol="4" gridrow="0" pageId="3" pageNumber="36">34–45</th>
<th box="[1056,1116,1415,1436]" gridcol="5" gridrow="0" pageId="3" pageNumber="36">45–65</th>
<th box="[1188,1273,1415,1436]" gridcol="6" gridrow="0" pageId="3" pageNumber="36">120–221</th>
<th box="[1329,1418,1415,1436]" gridcol="7" gridrow="0" pageId="3" pageNumber="36">155–177</th>
</tr>
<tr box="[159,1418,1455,1476]" gridrow="1" pageId="3" pageNumber="36">
<th box="[159,409,1455,1476]" gridcol="0" gridrow="1" pageId="3" pageNumber="36">Intestinal origin XVI</th>
<td box="[472,553,1455,1476]" gridcol="1" gridrow="1" pageId="3" pageNumber="36">XVI</td>
<td box="[628,688,1455,1476]" gridcol="2" gridrow="1" pageId="3" pageNumber="36">XVI</td>
<td box="[765,825,1455,1476]" gridcol="3" gridrow="1" pageId="3" pageNumber="36">XV</td>
<td box="[926,987,1455,1476]" gridcol="4" gridrow="1" pageId="3" pageNumber="36">XVI</td>
<td box="[1056,1116,1455,1476]" gridcol="5" gridrow="1" pageId="3" pageNumber="36">XVI</td>
<td box="[1188,1273,1455,1476]" gridcol="6" gridrow="1" pageId="3" pageNumber="36">XVI</td>
<td box="[1329,1418,1455,1476]" gridcol="7" gridrow="1" pageId="3" pageNumber="36">XVI</td>
</tr>
<tr box="[159,1418,1495,1516]" gridrow="2" pageId="3" pageNumber="36">
<th box="[159,409,1495,1516]" gridcol="0" gridrow="2" pageId="3" pageNumber="36">Typhlosole +</th>
<td box="[472,553,1495,1516]" gridcol="1" gridrow="2" pageId="3" pageNumber="36">vestigial</td>
<td box="[628,688,1495,1516]" gridcol="2" gridrow="2" pageId="3" pageNumber="36">+</td>
<td box="[765,825,1495,1516]" gridcol="3" gridrow="2" pageId="3" pageNumber="36">+</td>
<td box="[926,987,1495,1516]" gridcol="4" gridrow="2" pageId="3" pageNumber="36">+</td>
<td box="[1056,1116,1495,1516]" gridcol="5" gridrow="2" pageId="3" pageNumber="36">+</td>
<td box="[1188,1273,1495,1516]" gridcol="6" gridrow="2" pageId="3" pageNumber="36">+</td>
<td box="[1329,1418,1495,1516]" gridcol="7" gridrow="2" pageId="3" pageNumber="36">+</td>
</tr>
<tr box="[159,1418,1535,1556]" gridrow="3" pageId="3" pageNumber="36">
<th box="[159,409,1535,1556]" gridcol="0" gridrow="3" pageId="3" pageNumber="36">Intestinal vessels?</th>
<td box="[472,553,1535,1556]" gridcol="1" gridrow="3" pageId="3" pageNumber="36">?</td>
<td box="[628,688,1535,1556]" gridcol="2" gridrow="3" pageId="3" pageNumber="36">?</td>
<td box="[765,825,1535,1556]" gridcol="3" gridrow="3" pageId="3" pageNumber="36">?</td>
<td box="[926,987,1535,1556]" gridcol="4" gridrow="3" pageId="3" pageNumber="36">?</td>
<td box="[1056,1116,1535,1556]" gridcol="5" gridrow="3" pageId="3" pageNumber="36">?</td>
<td box="[1188,1273,1535,1556]" gridcol="6" gridrow="3" pageId="3" pageNumber="36">36</td>
<td box="[1329,1418,1535,1556]" gridcol="7" gridrow="3" pageId="3" pageNumber="36">38</td>
</tr>
</table>
</paragraph>
<paragraph pageId="3" pageNumber="36">
<table box="[159,1418,1575,1826]" gridcols="9" gridrows="6" pageId="3" pageNumber="36">
<tr box="[159,1418,1575,1596]" gridrow="0" pageId="3" pageNumber="36">
<th box="[159,307,1575,1596]" gridcol="0" gridrow="0" pageId="3" pageNumber="36">Hearts</th>
<th box="[349,431,1575,1596]" gridcol="1" gridrow="0" pageId="3" pageNumber="36">X – XIII</th>
<th box="[472,563,1575,1596]" gridcol="2" gridrow="0" pageId="3" pageNumber="36">XI – XIII</th>
<th box="[628,712,1575,1596]" gridcol="3" gridrow="0" pageId="3" pageNumber="36">XI–XIII</th>
<th box="[765,849,1575,1596]" gridcol="4" gridrow="0" pageId="3" pageNumber="36">X–XIII</th>
<th box="[926,1005,1575,1596]" gridcol="5" gridrow="0" pageId="3" pageNumber="36">XI–XIII</th>
<th box="[1056,1140,1575,1596]" gridcol="6" gridrow="0" pageId="3" pageNumber="36">XI–XIII</th>
<th box="[1188,1272,1575,1596]" gridcol="7" gridrow="0" pageId="3" pageNumber="36">XI–XIII</th>
<th box="[1329,1418,1575,1596]" gridcol="8" gridrow="0" pageId="3" pageNumber="36">XI–XIII</th>
</tr>
<tr box="[159,1418,1615,1636]" gridrow="1" pageId="3" pageNumber="36">
<th box="[159,307,1615,1636]" gridcol="0" gridrow="1" pageId="3" pageNumber="36">Septa 8/9/10**</th>
<td box="[349,431,1615,1636]" gridcol="1" gridrow="1" pageId="3" pageNumber="36">+, -</td>
<td box="[472,563,1615,1636]" gridcol="2" gridrow="1" pageId="3" pageNumber="36">-</td>
<td box="[628,712,1615,1636]" gridcol="3" gridrow="1" pageId="3" pageNumber="36">-</td>
<td box="[765,849,1615,1636]" gridcol="4" gridrow="1" pageId="3" pageNumber="36">+8/9, -</td>
<td box="[926,1005,1615,1636]" gridcol="5" gridrow="1" pageId="3" pageNumber="36">-, +9/10</td>
<td box="[1056,1140,1615,1636]" gridcol="6" gridrow="1" pageId="3" pageNumber="36">-</td>
<td box="[1188,1272,1615,1636]" gridcol="7" gridrow="1" pageId="3" pageNumber="36">+8/9, -</td>
<td box="[1329,1418,1615,1636]" gridcol="8" gridrow="1" pageId="3" pageNumber="36">+8/9, -</td>
</tr>
<tr box="[159,1418,1655,1706]" gridrow="2" pageId="3" pageNumber="36">
<th box="[159,307,1655,1706]" gridcol="0" gridrow="2" pageId="3" pageNumber="36">Spermathecal pore spacing</th>
<td box="[349,431,1655,1706]" gridcol="1" gridrow="2" pageId="3" pageNumber="36">?</td>
<td box="[472,563,1655,1706]" gridcol="2" gridrow="2" pageId="3" pageNumber="36">0.05–.06</td>
<td box="[628,712,1655,1706]" gridcol="3" gridrow="2" pageId="3" pageNumber="36">0.32–.36</td>
<td box="[765,849,1655,1706]" gridcol="4" gridrow="2" pageId="3" pageNumber="36">0.18–.20</td>
<td box="[926,1005,1655,1706]" gridcol="5" gridrow="2" pageId="3" pageNumber="36">0.32</td>
<td box="[1056,1140,1655,1706]" gridcol="6" gridrow="2" pageId="3" pageNumber="36">0.21–.23</td>
<td box="[1188,1272,1655,1706]" gridcol="7" gridrow="2" pageId="3" pageNumber="36">0.04–.06</td>
<td box="[1329,1418,1655,1706]" gridcol="8" gridrow="2" pageId="3" pageNumber="36">0.06–.07</td>
</tr>
<tr box="[159,1418,1725,1746]" gridrow="3" pageId="3" pageNumber="36">
<th box="[159,307,1725,1746]" gridcol="0" gridrow="3" pageId="3" pageNumber="36">Penes</th>
<td box="[349,431,1725,1746]" gridcol="1" gridrow="3" pageId="3" pageNumber="36">+</td>
<td box="[472,563,1725,1746]" gridcol="2" gridrow="3" pageId="3" pageNumber="36">+</td>
<td box="[628,712,1725,1746]" gridcol="3" gridrow="3" pageId="3" pageNumber="36">+</td>
<td box="[765,849,1725,1746]" gridcol="4" gridrow="3" pageId="3" pageNumber="36">+</td>
<td box="[926,1005,1725,1746]" gridcol="5" gridrow="3" pageId="3" pageNumber="36">+</td>
<td box="[1056,1140,1725,1746]" gridcol="6" gridrow="3" pageId="3" pageNumber="36">+</td>
<td box="[1188,1272,1725,1746]" gridcol="7" gridrow="3" pageId="3" pageNumber="36">+</td>
<td box="[1329,1418,1725,1746]" gridcol="8" gridrow="3" pageId="3" pageNumber="36">+</td>
</tr>
<tr box="[159,1418,1765,1786]" gridrow="4" pageId="3" pageNumber="36">
<th box="[159,307,1765,1786]" gridcol="0" gridrow="4" pageId="3" pageNumber="36">Bursal pads***</th>
<td box="[349,431,1765,1786]" gridcol="1" gridrow="4" pageId="3" pageNumber="36">2</td>
<td box="[472,563,1765,1786]" gridcol="2" gridrow="4" pageId="3" pageNumber="36">2F, 1R</td>
<td box="[628,712,1765,1786]" gridcol="3" gridrow="4" pageId="3" pageNumber="36">1F</td>
<td box="[765,849,1765,1786]" gridcol="4" gridrow="4" pageId="3" pageNumber="36">2S</td>
<td box="[926,1005,1765,1786]" gridcol="5" gridrow="4" pageId="3" pageNumber="36">3S</td>
<td box="[1056,1140,1765,1786]" gridcol="6" gridrow="4" pageId="3" pageNumber="36">2F</td>
<td box="[1188,1272,1765,1786]" gridcol="7" gridrow="4" pageId="3" pageNumber="36">1F, 2R</td>
<td box="[1329,1418,1765,1786]" gridcol="8" gridrow="4" pageId="3" pageNumber="36">1F-R, 1R</td>
</tr>
<tr box="[159,1418,1805,1826]" gridrow="5" pageId="3" pageNumber="36">
<th box="[159,307,1805,1826]" gridcol="0" gridrow="5" pageId="3" pageNumber="36">Bursal glands</th>
<td box="[349,431,1805,1826]" gridcol="1" gridrow="5" pageId="3" pageNumber="36">-</td>
<td box="[472,563,1805,1826]" gridcol="2" gridrow="5" pageId="3" pageNumber="36">+</td>
<td box="[628,712,1805,1826]" gridcol="3" gridrow="5" pageId="3" pageNumber="36">+</td>
<td box="[765,849,1805,1826]" gridcol="4" gridrow="5" pageId="3" pageNumber="36">+</td>
<td box="[926,1005,1805,1826]" gridcol="5" gridrow="5" pageId="3" pageNumber="36">+</td>
<td box="[1056,1140,1805,1826]" gridcol="6" gridrow="5" pageId="3" pageNumber="36">+</td>
<td box="[1188,1272,1805,1826]" gridcol="7" gridrow="5" pageId="3" pageNumber="36">+</td>
<td box="[1329,1418,1805,1826]" gridcol="8" gridrow="5" pageId="3" pageNumber="36">+</td>
</tr>
</table>
</paragraph>
<paragraph blockId="3.[151,768,1866,1959]" box="[151,640,1866,1889]" pageId="3" pageNumber="36">* Setal gaps presence: D (dorsal), V (ventral)</paragraph>
<paragraph blockId="3.[151,768,1866,1959]" box="[151,584,1901,1924]" pageId="3" pageNumber="36">** Septum present: +, Septum absent: -</paragraph>
<paragraph blockId="3.[151,768,1866,1959]" box="[151,768,1936,1959]" pageId="3" pageNumber="36">***F= floor R= roof, S= side of copulatory bursa interior.</paragraph>
<paragraph blockId="4.[151,1436,152,1218]" pageId="4" pageNumber="37">
<collectingCountry box="[197,262,152,178]" name="Spain" pageId="4" pageNumber="37">Septa</collectingCountry>
<number box="[269,282,152,178]" pageId="4" pageNumber="37" value="5.0">5</number>
/ 
<number box="[292,337,152,178]" pageId="4" pageNumber="37" value="6.5" valueMax="7.0" valueMin="6.0">6–7</number>
/ 
<number box="[347,360,152,178]" pageId="4" pageNumber="37" value="8.0">8</number>
thinly muscular; 
<number box="[565,579,152,178]" pageId="4" pageNumber="37" value="8.0">8</number>
/9, 9/ 
<number box="[640,668,152,178]" pageId="4" pageNumber="37" value="10.0">10</number>
absent, 
<date box="[765,913,152,178]" pageId="4" pageNumber="37" value="1914-10-11" valueMax="1914-10-13" valueMin="1914-10-11">
<number box="[765,794,152,178]" pageId="4" pageNumber="37" value="10.0">10</number>
/ 
<number box="[804,873,152,178]" pageId="4" pageNumber="37" value="12.0" valueMax="13.0" valueMin="11.0">11–13</number>
/ 
<number box="[884,913,152,178]" pageId="4" pageNumber="37" value="14.0">14</number>
</date>
thick. Gizzard in VIII–X, intestine begins in XVI, small paired lymph glands from XXVII along dorsal vessel; intestinal caeca simple originating in XXVII, and extending anteriorly about to XXIV, finger-shaped sac; typhlosole vestigial from XXVII. Hearts in XI–XIII esophageal; X absent.
</paragraph>
<paragraph blockId="4.[151,1436,152,1218]" pageId="4" pageNumber="37">
Ovaries and funnels in XIII, spermathecae in VI, with nephridia on ducts; spermatheca with large eggshaped ampulla, narrow non-muscular duct shorter than ampulla; diverticulum small oval chamber; iridescent, undulating slender stalk about half spermatheca length. Male sexual system holandric, testes and funnels in ventrally paired sacs in X–XI. Seminal vesicles two pairs in XI–XII, dorsal lobes large, pear-shaped or elongate, apically attached, prostates in XVII–XVIII, 
<number box="[756,771,472,498]" pageId="4" pageNumber="37" value="4.0">4</number>
main lobes lateral-anterior to copulatory bursae, thick slightly muscular short ducts, entering center of copulatory bursae without stalked glands; copulatory bursae openings flanked anterior, posterior by oval pads on floor; one smaller pad on roof medial to penis; glandular tissue in chamber, male pore on roughly ovate-conical penis from copulatory bursa roof.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="37" type="discussion">
<paragraph blockId="4.[151,1436,152,1218]" pageId="4" pageNumber="37">
<emphasis bold="true" box="[197,318,631,657]" pageId="4" pageNumber="37">Remarks.</emphasis>
<taxonomicName box="[328,574,632,657]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="37" phylum="Annelida" rank="species" species="bicolensis" status="sp. nov.">
<emphasis box="[328,574,632,657]" italics="true" pageId="4" pageNumber="37">Pheretima bicolensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[583,678,631,657]" pageId="4" pageNumber="37" rank="species">
<emphasis bold="true" box="[583,678,631,657]" pageId="4" pageNumber="37">sp. nov.</emphasis>
</taxonomicNameLabel>
keys to the 
<taxonomicName box="[826,937,632,657]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="37" phylum="Annelida" rank="species" species="urceolata">
<emphasis box="[826,937,632,657]" italics="true" pageId="4" pageNumber="37">urceolata</emphasis>
</taxonomicName>
group in 
<bibRefCitation author="Sims" box="[1054,1323,632,658]" pageId="4" pageNumber="45" refString="Sims, R. W. &amp; Easton, E. G. (1972) A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. The Biological Journal of the Linnean Society, London, 4, 169 - 268." type="journal article" year="1972">
Sims &amp; Easton (
<number box="[1251,1314,632,658]" pageId="4" pageNumber="37" value="1972.0">1972</number>
)
</bibRefCitation>
, which is composed of eight species, 
<taxonomicName authority="Horst, 1893" authorityName="Horst" authorityYear="1893" box="[479,791,672,698]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="37" phylum="Annelida" rank="species" species="urceolata">
<emphasis box="[479,620,672,697]" italics="true" pageId="4" pageNumber="37">P. urceolata</emphasis>
(
<bibRefCitation author="Horst" box="[637,782,672,698]" pageId="4" pageNumber="45" refString="Horst, R. (1893) Descriptions of earthworms. No. 7. On Malayan earthworms. Notes from the Leyden Museum, Leyden, 15, 316 - 329." type="journal article" year="1893">
Horst, 
<number box="[720,782,672,698]" pageId="4" pageNumber="37" value="1893.0">1893</number>
</bibRefCitation>
)
</taxonomicName>
, 
<emphasis box="[804,981,672,697]" italics="true" pageId="4" pageNumber="37">P. b a w e a n e n s i s</emphasis>
(
<bibRefCitation author="Michaelsen" box="[1001,1217,672,698]" pageId="4" pageNumber="45" refString="Michaelsen, W. (1924) Oligochaten von Niederlandisch-Indien. Treubia, Journal of the Zoological Museum and Laboratory, 5, 379 - 401." type="journal article" year="1924">
Michaelsen, 
<number box="[1154,1217,672,698]" pageId="4" pageNumber="37" value="1924.0">1924</number>
</bibRefCitation>
), 
<taxonomicName authority="James, 2004" authorityName="James" authorityYear="2004" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="37" phylum="Annelida" rank="species" species="baletei">
<emphasis box="[1238,1348,672,697]" italics="true" pageId="4" pageNumber="37">P. baletei</emphasis>
James, 
<number box="[151,212,712,738]" pageId="4" pageNumber="37" value="2004.0">2004</number>
</taxonomicName>
, 
<taxonomicName authority="James, 2004" authorityName="James" authorityYear="2004" box="[227,587,712,738]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="37" phylum="Annelida" rank="species" species="bukidnonensis">
<emphasis box="[227,427,712,737]" italics="true" pageId="4" pageNumber="37">P. bukidnonensis</emphasis>
James, 
<number box="[525,587,712,738]" pageId="4" pageNumber="37" value="2004.0">2004</number>
</taxonomicName>
, 
<emphasis box="[601,720,712,737]" italics="true" pageId="4" pageNumber="37">P. h e a n e y i</emphasis>
James, 
<number box="[823,885,712,738]" pageId="4" pageNumber="37" value="2004.0">2004</number>
, 
<taxonomicName authority="James, 2004" authorityName="James" authorityYear="2004" box="[900,1262,712,738]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="37" phylum="Annelida" rank="species" species="kitangladensis">
<emphasis box="[900,1100,712,737]" italics="true" pageId="4" pageNumber="37">P. kitangladensis</emphasis>
James, 
<number box="[1200,1262,712,738]" pageId="4" pageNumber="37" value="2004.0">2004</number>
</taxonomicName>
, 
<taxonomicName authority="James, 2004" authorityName="James" authorityYear="2004" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="37" phylum="Annelida" rank="species" species="monotheca">
<emphasis box="[1276,1436,712,737]" italics="true" pageId="4" pageNumber="37">P. monotheca</emphasis>
James, 
<number box="[237,299,752,778]" pageId="4" pageNumber="37" value="2004.0">2004</number>
</taxonomicName>
, and 
<emphasis box="[361,448,752,777]" italics="true" pageId="4" pageNumber="37">P. s i m s i</emphasis>
James &amp; Hong, 
<number box="[649,708,752,778]" pageId="4" pageNumber="37" value="2004.0">2004</number>
plus two more currently in press, 
<taxonomicName box="[1112,1231,752,778]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="37" phylum="Annelida" rank="species" species="abiadai">
<emphasis box="[1112,1126,752,777]" italics="true" pageId="4" pageNumber="37">P</emphasis>
. 
<emphasis box="[1141,1231,752,777]" italics="true" pageId="4" pageNumber="37">abiadai</emphasis>
</taxonomicName>
and 
<taxonomicName box="[1289,1436,752,778]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="37" phylum="Annelida" rank="species" species="nagaensis">
<emphasis box="[1289,1303,752,777]" italics="true" pageId="4" pageNumber="37">P</emphasis>
. 
<emphasis box="[1319,1436,752,777]" italics="true" pageId="4" pageNumber="37">nagaensis</emphasis>
</taxonomicName>
(
<bibRefCitation author="Hong" box="[159,453,792,818]" pageId="4" pageNumber="45" refString="Hong, Y. &amp; James, S. W. (2008 b) Two new earthworms of the genus Pheretima (Oligochaeta: Megascolecidae) from Mt. Isarog, Luzon Island, Philippines. Journal of Natural History, 42 (23 &amp; 24), 1565 - 1571." type="journal article" year="2008" yearSuffix="b">
Hong and James 
<number box="[375,438,792,818]" pageId="4" pageNumber="37" value="2008.0">2008</number>
b
</bibRefCitation>
). All comparisons to previously published species are confined to the 
<taxonomicName box="[1324,1435,792,817]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="37" phylum="Annelida" rank="species" species="urceolata">
<emphasis box="[1324,1435,792,817]" italics="true" pageId="4" pageNumber="37">urceolata</emphasis>
</taxonomicName>
group. Morphological data on this group is summarized in Table 
<number box="[911,927,832,858]" pageId="4" pageNumber="37" value="1.0">1</number>
. The reader should refer to this table for all the comparisons made below.
</paragraph>
<paragraph blockId="4.[151,1436,152,1218]" pageId="4" pageNumber="37">
<taxonomicName box="[197,450,912,937]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="37" phylum="Annelida" rank="species" species="bicolensis" status="sp. nov.">
<emphasis box="[197,450,912,937]" italics="true" pageId="4" pageNumber="37">Pheretima bicolensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[466,568,911,937]" pageId="4" pageNumber="37">
<taxonomicNameLabel box="[466,568,911,937]" pageId="4" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has narrowly spaced spermathecal pores, a condition it shares with 
<taxonomicName box="[151,393,952,977]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="37" phylum="Annelida" rank="species" species="buhiensis" status="sp. nov.">
<emphasis box="[151,393,952,977]" italics="true" pageId="4" pageNumber="37">Pheretima buhiensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[404,500,951,977]" pageId="4" pageNumber="37">
<taxonomicNameLabel box="[404,500,951,977]" pageId="4" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName box="[564,770,952,977]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="37" phylum="Annelida" rank="species" species="gorasi" status="sp. nov.">
<emphasis box="[564,770,952,977]" italics="true" pageId="4" pageNumber="37">Pheretima gorasi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[780,876,951,977]" pageId="4" pageNumber="37">
<taxonomicNameLabel box="[780,876,951,977]" pageId="4" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(See Table 
<number box="[1025,1039,952,978]" pageId="4" pageNumber="37" value="1.0">1</number>
). Compared to those species, its male pores are more widely spaced, it has only ventral setal gaps, has 
<number box="[966,980,992,1018]" pageId="4" pageNumber="37" value="2.0">2</number>
or 
<number box="[1019,1033,992,1018]" pageId="4" pageNumber="37" value="3.0">3</number>
rather than zero setae between the male openings, lacks a typhlosole, and lacks septa 
<date box="[767,845,1032,1058]" pageId="4" pageNumber="37" value="1910-09-08">
<number box="[767,780,1032,1058]" pageId="4" pageNumber="37" value="8.0">8</number>
/ 
<number box="[790,805,1032,1058]" pageId="4" pageNumber="37" value="9.0">9</number>
/ 
<number box="[815,845,1032,1058]" pageId="4" pageNumber="37" value="10.0">10</number>
</date>
. Pads in the copulatory bursae are also different, with those of 
<taxonomicName box="[313,559,1072,1097]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="37" phylum="Annelida" rank="species" species="bicolensis" status="sp. nov.">
<emphasis box="[313,559,1072,1097]" italics="true" pageId="4" pageNumber="37">Pheretima bicolensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[567,661,1071,1097]" pageId="4" pageNumber="37">
<taxonomicNameLabel box="[567,661,1071,1097]" pageId="4" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
being 
<number box="[742,756,1072,1098]" pageId="4" pageNumber="37" value="2.0">2</number>
on the bursal floor and one on the roof. The arrangement is reversed in 
<taxonomicName box="[315,555,1112,1137]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="37" phylum="Annelida" rank="species" species="buhiensis" status="sp. nov.">
<emphasis box="[315,555,1112,1137]" italics="true" pageId="4" pageNumber="37">Pheretima buhiensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[562,656,1111,1137]" pageId="4" pageNumber="37">
<taxonomicNameLabel box="[562,656,1111,1137]" pageId="4" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(one on the floor, two on the roof), and 
<taxonomicName box="[1131,1334,1112,1137]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="37" phylum="Annelida" rank="species" species="gorasi" status="sp. nov.">
<emphasis box="[1131,1334,1112,1137]" italics="true" pageId="4" pageNumber="37">Pheretima gorasi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1342,1436,1111,1137]" pageId="4" pageNumber="37">
<taxonomicNameLabel box="[1342,1436,1111,1137]" pageId="4" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has only two pads, one of which is on the side. Of these three species, 
<taxonomicName box="[1009,1256,1152,1177]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="37" phylum="Annelida" rank="species" species="bicolensis" status="sp. nov.">
<emphasis box="[1009,1256,1152,1177]" italics="true" pageId="4" pageNumber="37">Pheretima bicolensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1266,1361,1151,1177]" pageId="4" pageNumber="37">
<taxonomicNameLabel box="[1266,1361,1151,1177]" pageId="4" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the smallest.
</paragraph>
</subSubSection>
</treatment>
</document>