271 lines
37 KiB
XML
271 lines
37 KiB
XML
<document id="9807199BCCF6334DCE187543B9AA8145" ID-DOI="10.5281/zenodo.188890" ID-GBIF-Dataset="179e0d3c-544e-4194-baf6-8e49f27e708e" ID-ISSN="1175-5326" ID-Zenodo-Dep="188890" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461098343359" checkinUser="plazi" docAuthor="Karaman, Ivo M., Bedek, Jana & Horvatović, Mladen" docDate="2009" docId="F8478790FF80FFC4FF6CF7A1D79E7C1E" docLanguage="en" docName="zt02158p064.pdf" docOrigin="Zootaxa 2158" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Thaumatoniscellus speluncae Karaman, Bedek & Horvatović, 2009, n. sp." docType="treatment" docUuid="A6CBECDF-EFBA-4073-A30B-542A45217397" docUuidSource="ZooBank" docVersion="10" lastPageNumber="61" masterDocId="047EFFE8FF82FFC0FFFBF736D5097D6D" masterDocTitle="Thaumatoniscellus speluncae n. sp. (Isopoda: Oniscidea: Trichoniscidae), a new troglobitic oniscid species from Croatia" masterLastPageNumber="64" masterPageNumber="57" pageNumber="59" updateTime="1698590142322" updateUser="plazi">
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<mods:mods id="E51DF6931FBAE0DE45952DB10F390564" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A5A5CAEC07D3A833D3CD8FCCAC6BD176">
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<mods:title id="809B49F56AF62BAE5A3A798D411A6FBC">Thaumatoniscellus speluncae n. sp. (Isopoda: Oniscidea: Trichoniscidae), a new troglobitic oniscid species from Croatia</mods:title>
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</mods:titleInfo>
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<mods:name id="48342D1395A1D6F79485B51960ED21C8" type="personal">
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<mods:role id="94BA3D7BA85BAF4829661DCFCBAF6DFF">
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<mods:roleTerm id="96E1FA10CCB3EB67AF8E2501A25A4F08">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="98A2522DB629BBE926719B78D6A4070D">Karaman, Ivo M.</mods:namePart>
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</mods:name>
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<mods:name id="9F156886808450CED1FD82F1442640D4" type="personal">
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<mods:role id="C2F4D38AF24315907F801126EB799907">
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<mods:roleTerm id="20AC5AD887325121F94CFC0FB8EEE238">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B422C3B76C4FD38ED5E19D7319A47675">Bedek, Jana</mods:namePart>
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</mods:name>
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<mods:name id="0FA7D8AFC3831F2FDEAC79FD632FB187" type="personal">
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<mods:role id="A0B37A9B1AE5879D01993E62BB173F1C">
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<mods:roleTerm id="205333E7008014DE20125348BCD0407F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E08578D300D895791525BF05960E6DA6">Horvatović, Mladen</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="E2D779D82BA72E63FBED2A6FB27A3DAA">text</mods:typeOfResource>
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<mods:titleInfo id="33491208BAAE0B84B49DEC9DCFCDE009">
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<mods:title id="F02CBE84D36E50C322044980AF36F89C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0328AB4049DAFFD8F09220FA1965CDFF">
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<mods:date id="52E7E828B7628F51ED5B1F21665FA238">2009</mods:date>
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<mods:detail id="C151D806BFEBFB13AC28F9A4B1AED883" type="volume">
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<mods:number id="DFD8BEAFEC6C5CB6DD7E3A3E69EEE8B5">2158</mods:number>
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<mods:end id="63A3679496B7B356D03530967E1494EF">64</mods:end>
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<mods:classification id="3916A3437E59D5BBAAD435E94B9B2EBF">journal article</mods:classification>
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<mods:identifier id="0E3FCDCF8EBECB08BB6A0DB97364A213" type="DOI">10.5281/zenodo.188890</mods:identifier>
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<mods:identifier id="30A156C17D188F7D89C61CB049598871" type="GBIF-Dataset">179e0d3c-544e-4194-baf6-8e49f27e708e</mods:identifier>
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<mods:identifier id="291B632CDB24679BDB588A525F4ED678" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="F1A6F255AC60BB4E9B9444B8CE51A60E" type="Zenodo-Dep">188890</mods:identifier>
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<mods:identifier id="B50558A4C32020072F370D6ECB10AB1A" type="ZooBank">DA70B147-780E-4814-9D2D-856FF7850482</mods:identifier>
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</mods:mods>
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<treatment id="F8478790FF80FFC4FF6CF7A1D79E7C1E" ID-DOI="http://doi.org/10.5281/zenodo.6224088" ID-GBIF-Taxon="119620989" ID-Zenodo-Dep="6224088" LSID="urn:lsid:zoobank.org:act:A6CBECDF-EFBA-4073-A30B-542A45217397" httpUri="http://treatment.plazi.org/id/F8478790FF80FFC4FF6CF7A1D79E7C1E" lastPageId="4" lastPageNumber="61" pageId="2" pageNumber="59">
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<subSubSection id="38F4650DFF80FFC2FF6CF7A1D4177DB9" pageId="2" pageNumber="59" type="nomenclature">
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<paragraph id="70513686FF80FFC2FF6CF7A1D74E7DDC" blockId="2.[151,583,151,212]" box="[151,583,151,177]" pageId="2" pageNumber="59">
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<heading id="2B1981EAFF80FFC2FF6CF7A1D74E7DDC" bold="true" box="[151,583,151,177]" fontSize="11" level="1" pageId="2" pageNumber="59" reason="1">
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<emphasis id="429AEA94FF80FFC2FF6CF7A1D74E7DDC" bold="true" box="[151,583,151,177]" pageId="2" pageNumber="59">
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<taxonomicName id="B7EE4D05FF80FFC2FF6CF7A1D4F47DDC" ID-CoL="7C72R" box="[151,509,151,177]" class="Malacostraca" family="Trichoniscidae" genus="Thaumatoniscellus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="speluncae" status="sp. nov.">
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<emphasis id="429AEA94FF80FFC2FF6CF7A1D4F47DDC" bold="true" box="[151,509,151,177]" italics="true" pageId="2" pageNumber="59">Thaumatoniscellus speluncae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="59A957EFFF80FFC2FDFEF7A1D74E7DDC" box="[517,583,151,177]" pageId="2" pageNumber="59" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="70513686FF80FFC2FF6CF78CD4177DB9" blockId="2.[151,583,151,212]" box="[151,286,186,212]" pageId="2" pageNumber="59">
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(
|
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<figureCitation id="E8D52A03FF80FFC2FF64F78CD41C7DB9" box="[159,277,186,212]" captionStart-0="FIGURES 1 – 6" captionStart-1="FIGURES 7 – 11" captionStart-2="FIGURES 12 – 15" captionStart-3="FIGURES 16 – 21" captionStartId-0="1.[151,269,1866,1890]" captionStartId-1="3.[151,273,1665,1689]" captionStartId-2="4.[151,269,1924,1948]" captionStartId-3="5.[151,271,1882,1906]" captionTargetBox-0="[151,1436,390,1839]" captionTargetBox-1="[166,1406,222,1605]" captionTargetBox-2="[192,1403,459,1870]" captionTargetBox-3="[215,1380,230,1811]" captionTargetId-0="figure@1.[151,1436,390,1840]" captionTargetId-1="figure@3.[151,1436,197,1640]" captionTargetId-2="figure@4.[151,1436,429,1898]" captionTargetId-3="figure@5.[151,1436,194,1858]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURES 1 – 6. Thaumatoniscellus speluncae n. sp., paratype female (1.31 mm). 1, lateral view of the body. 2, cephalon and pereonites 1 – 3, lateral view. 3, anterior part of the animal, frontal view. 4, pleon and pereonites 5 – 7, lateral view. 5, pleon and pereonite 7, dorsal view. 6, detail of the integument ornamentic with characteristic scales." captionText-1="FIGURES 7 – 11. Thaumatoniscellus speluncae n. sp., mouth parts of paratype female (1.38 mm). 7, endopodite of maxilla 1. 8, exopodite of maxilla 1. 9, maxilla 2, dorsal view. 10, maxilla 2, ventral view. 11, maxilliped. Scale bar = 0.1 mm." captionText-2="FIGURES 12 – 15. Thaumatoniscellus speluncae n. sp., 12, antennula, paratype female (1.38 mm). 13 – 15. Holotype. 13, antenna. 14, pereopod 1. 15, pereopod 7. Scale bar = 0.2 mm." captionText-3="FIGURES 16 – 21. Thaumatoniscellus speluncae n. sp., holotype. 16, genital papilla, ventral view. 17, genital papilla, lateral view. 18, exopodite of pleopod 1, ventral view. 19, pleopod 1, dorsal view. 20, pleopod 2. 21, endopodite of pleopod 1, lateral view. Scale bar = 0.1 mm." httpUri-0="https://zenodo.org/record/188891/files/figure.png" httpUri-1="https://zenodo.org/record/188892/files/figure.png" httpUri-2="https://zenodo.org/record/188893/files/figure.png" httpUri-3="https://zenodo.org/record/188894/files/figure.png" pageId="2" pageNumber="59">Figs 1–21</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="38F4650DFF80FFC2FF6CF631D6B57C8E" pageId="2" pageNumber="59" type="materials_examined">
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<paragraph id="70513686FF80FFC2FF6CF631D6B57C8E" blockId="2.[151,1438,263,1991]" pageId="2" pageNumber="59">
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<emphasis id="429AEA94FF80FFC2FF6CF631D7127C4C" bold="true" box="[151,539,263,289]" pageId="2" pageNumber="59">
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Material examined.
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<typeStatus id="AF558824FF80FFC2FE66F631D71C7C4C" box="[413,533,263,289]" pageId="2" pageNumber="59" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Male, Pit 3 near Banina Stream, Veprinac, Učka, Istria,
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<collectingCountry id="08F97616FF80FFC2FB28F63ED0397C4F" box="[1235,1328,264,290]" name="Croatia" pageId="2" pageNumber="59">Croatia</collectingCountry>
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, (Gauss- Krieger coordinates x=5019084, y=5440933),
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<date id="04501046FF80FFC2FD46F618D64B7C25" box="[701,834,302,328]" pageId="2" pageNumber="59" value="2008-04-05">05.IV.2008</date>
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, leg. J. Bedek (Inv. No IT 1287,
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<collectionCode id="16FFAE43FF80FFC2FB33F618D02F7C25" box="[1224,1318,302,328]" pageId="2" pageNumber="59">CBSSC</collectionCode>
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,
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<collectionCode id="16FFAE43FF80FFC2FAC8F618D0997C25" LSID="urn:lsid:biocol.org:col:33139" box="[1331,1424,302,328]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33139" name="Cincinnati Museum of Natural History" pageId="2" pageNumber="59">CNHM</collectionCode>
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).
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<emphasis id="429AEA94FF80FFC2FF6CF663D4127C02" bold="true" box="[151,283,341,367]" pageId="2" pageNumber="59">
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<typeStatus id="AF558824FF80FFC2FF6CF663D41E7C02" box="[151,279,341,367]" pageId="2" pageNumber="59" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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<specimenCount id="66E8FD0FFF80FFC2FED8F663D4837C02" box="[291,394,341,367]" pageId="2" pageNumber="59" type="female">1 female</specimenCount>
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and
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<specimenCount id="66E8FD0FFF80FFC2FE3FF663D7327C02" box="[452,571,341,367]" pageId="2" pageNumber="59" type="juvenile">1 juvenile</specimenCount>
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, leg. J. Bedek (Inv. No IT 996;
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<collectionCode id="16FFAE43FF80FFC2FC4AF663D1067C02" box="[945,1039,341,367]" pageId="2" pageNumber="59">CBSSC</collectionCode>
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,
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<collectionCode id="16FFAE43FF80FFC2FBE7F663D1707C02" LSID="urn:lsid:biocol.org:col:33139" box="[1052,1145,341,367]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33139" name="Cincinnati Museum of Natural History" pageId="2" pageNumber="59">CNHM</collectionCode>
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);
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<specimenCount id="66E8FD0FFF80FFC2FB75F663D1FE7C02" box="[1166,1271,341,367]" pageId="2" pageNumber="59" type="female">1 female</specimenCount>
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, leg. J. Bedek (Inv. No 1092, DBE).
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<emphasis id="429AEA94FF80FFC2FE57F64DD73E7CF8" bold="true" box="[428,567,379,405]" pageId="2" pageNumber="59">Topotypes.</emphasis>
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<specimenCount id="66E8FD0FFF80FFC2FDB9F64AD7B87CFB" box="[578,689,380,406]" pageId="2" pageNumber="59" type="female">1 female</specimenCount>
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, ibid.,
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<date id="04501046FF80FFC2FCFCF64AD69F7CFB" box="[775,918,380,406]" pageId="2" pageNumber="59" value="2006-11-04">04.XI.2006</date>
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, leg. M. Pavlek (Inv. No IT 772,
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<collectionCode id="16FFAE43FF80FFC2FACDF64AD0917CFB" box="[1334,1432,380,406]" pageId="2" pageNumber="59">CBSSC</collectionCode>
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,
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<collectionCode id="16FFAE43FF80FFC2FF6CF694D5FD7CD1" LSID="urn:lsid:biocol.org:col:33139" box="[151,244,418,444]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33139" name="Cincinnati Museum of Natural History" pageId="2" pageNumber="59">CNHM</collectionCode>
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);
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<specimenCount id="66E8FD0FFF80FFC2FEF3F694D46B7CD1" box="[264,354,418,444]" pageId="2" pageNumber="59" type="male">2 males</specimenCount>
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and
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<specimenCount id="66E8FD0FFF80FFC2FE60F694D7067CD1" box="[411,527,418,444]" pageId="2" pageNumber="59" type="female">2 females</specimenCount>
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, ibid.,
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<date id="04501046FF80FFC2FDA4F694D7EF7CD1" box="[607,742,418,444]" pageId="2" pageNumber="59" value="2007-09-17">17.IX.2007</date>
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, leg. J. Bedek (Inv. No IT 1056,
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<collectionCode id="16FFAE43FF80FFC2FB9DF694D1CD7CD1" box="[1126,1220,418,444]" pageId="2" pageNumber="59">CBSSC</collectionCode>
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,
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<collectionCode id="16FFAE43FF80FFC2FB2BF694D0247CD1" LSID="urn:lsid:biocol.org:col:33139" box="[1232,1325,418,444]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33139" name="Cincinnati Museum of Natural History" pageId="2" pageNumber="59">CNHM</collectionCode>
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);
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<specimenCount id="66E8FD0FFF80FFC2FABAF694D0927CD1" box="[1345,1435,418,444]" pageId="2" pageNumber="59" type="male">2 males</specimenCount>
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and
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<specimenCount id="66E8FD0FFF80FFC2FF33F6FFD4357C8E" box="[200,316,457,483]" pageId="2" pageNumber="59" type="female">3 females</specimenCount>
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, ibid.,
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<date id="04501046FF80FFC2FE77F6FFD71D7C8E" box="[396,532,457,483]" pageId="2" pageNumber="59" value="2007-09-17">17.IX.2007</date>
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, leg. J. Bedek (Inv. No 1093, DBE).
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</paragraph>
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</subSubSection>
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<subSubSection id="38F4650DFF80FFC2FF3DF6D9D73F7FA6" pageId="2" pageNumber="59" type="diagnosis">
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<paragraph id="70513686FF80FFC2FF3DF6D9D73F7FA6" blockId="2.[151,1438,263,1991]" pageId="2" pageNumber="59">
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<emphasis id="429AEA94FF80FFC2FF3DF6D9D44F7F64" bold="true" box="[198,326,495,521]" pageId="2" pageNumber="59">Diagnosis.</emphasis>
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Small species (up to 1.45 mm long). Median pair of tubercles on posterior part of cephalon small. Pereon tergites with 2 pairs of prominent tubercles per side. Male pleopod 1 endopodite constricted in median part; exopodite stout, terminally widened, without setae. Maxilla 1 exopodite with elongated teeth, compared with
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<taxonomicName id="B7EE4D05FF80FFC2FEB4F552D4D67F10" box="[335,479,612,637]" class="Malacostraca" family="Trichoniscidae" genus="Thaumatoniscellus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="orghidani">
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<emphasis id="429AEA94FF80FFC2FEB4F552D4D67F10" box="[335,479,612,637]" italics="true" pageId="2" pageNumber="59">T. orghidani</emphasis>
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</taxonomicName>
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. Maxilla 2 outer lobe remarkably shorter than inner lobe, with 1 rod-like seta on tip. Anchor-like lateral protrusions weakly developed in comparison with
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<taxonomicName id="B7EE4D05FF80FFC2FBDDF5BCD1B57FCE" box="[1062,1212,650,675]" class="Malacostraca" family="Trichoniscidae" genus="Thaumatoniscellus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="orghidani">
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||
<emphasis id="429AEA94FF80FFC2FBDDF5BCD1B57FCE" box="[1062,1212,650,675]" italics="true" pageId="2" pageNumber="59">T. orghidani</emphasis>
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</taxonomicName>
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. Male pereopod 7 propodus twice as long as dactylus.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="38F4650DFF80FFC4FF3DF5E1D41B7C4B" lastPageId="4" lastPageNumber="61" pageId="2" pageNumber="59" type="description">
|
||
<paragraph id="70513686FF80FFC2FF3DF5E1D5F67E0B" blockId="2.[151,1438,263,1991]" pageId="2" pageNumber="59">
|
||
<emphasis id="429AEA94FF80FFC2FF3DF5E1D7497F9C" bold="true" box="[198,576,727,753]" pageId="2" pageNumber="59">
|
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Description of male
|
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<typeStatus id="AF558824FF80FFC2FE35F5E1D7357F9C" box="[462,572,727,753]" pageId="2" pageNumber="59" type="holotype">holotype</typeStatus>
|
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.
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</emphasis>
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Body length 1.45 mm.
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<emphasis id="429AEA94FF80FFC2FC91F5EED6D57F9C" box="[874,988,728,753]" italics="true" pageId="2" pageNumber="59">Cephalon</emphasis>
|
||
(
|
||
<figureCitation id="E8D52A03FF80FFC2FC14F5EED1527F9F" box="[1007,1115,728,754]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1866,1890]" captionTargetBox="[151,1436,390,1839]" captionTargetId="figure@1.[151,1436,390,1840]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Thaumatoniscellus speluncae n. sp., paratype female (1.31 mm). 1, lateral view of the body. 2, cephalon and pereonites 1 – 3, lateral view. 3, anterior part of the animal, frontal view. 4, pleon and pereonites 5 – 7, lateral view. 5, pleon and pereonite 7, dorsal view. 6, detail of the integument ornamentic with characteristic scales." httpUri="https://zenodo.org/record/188891/files/figure.png" pageId="2" pageNumber="59">Figs 2–3</figureCitation>
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) short and wide, frontally compressed, bearing 2 rows of rounded tubercles: frontal row with large median tubercle and 2 smaller tubercles; posterior row of 6 tubercles, median pair very small. Lateral lobes short and wide, ventrally oriented.
|
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</paragraph>
|
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<paragraph id="70513686FF80FFC2FF3EF444D1B17EE1" blockId="2.[151,1438,263,1991]" box="[197,1208,882,908]" pageId="2" pageNumber="59">
|
||
<emphasis id="429AEA94FF80FFC2FF3EF444D4107EE6" box="[197,281,882,907]" italics="true" pageId="2" pageNumber="59">Pereon</emphasis>
|
||
(
|
||
<figureCitation id="E8D52A03FF80FFC2FED3F444D4867EE1" box="[296,399,882,908]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1866,1890]" captionTargetBox="[151,1436,390,1839]" captionTargetId="figure@1.[151,1436,390,1840]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Thaumatoniscellus speluncae n. sp., paratype female (1.31 mm). 1, lateral view of the body. 2, cephalon and pereonites 1 – 3, lateral view. 3, anterior part of the animal, frontal view. 4, pleon and pereonites 5 – 7, lateral view. 5, pleon and pereonite 7, dorsal view. 6, detail of the integument ornamentic with characteristic scales." httpUri="https://zenodo.org/record/188891/files/figure.png" pageId="2" pageNumber="59">Figs 1–4</figureCitation>
|
||
) prominently convex with a row of 4 pronounced rounded tubercles.
|
||
</paragraph>
|
||
<paragraph id="70513686FF80FFC2FF3EF4AFD6B97EB7" blockId="2.[151,1438,263,1991]" pageId="2" pageNumber="59">
|
||
<emphasis id="429AEA94FF80FFC2FF3EF4AFD4007EDF" box="[197,265,921,946]" italics="true" pageId="2" pageNumber="59">Pleon</emphasis>
|
||
(
|
||
<figureCitation id="E8D52A03FF80FFC2FEE1F4AFD48A7EDE" box="[282,387,921,947]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1866,1890]" captionTargetBox="[151,1436,390,1839]" captionTargetId="figure@1.[151,1436,390,1840]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Thaumatoniscellus speluncae n. sp., paratype female (1.31 mm). 1, lateral view of the body. 2, cephalon and pereonites 1 – 3, lateral view. 3, anterior part of the animal, frontal view. 4, pleon and pereonites 5 – 7, lateral view. 5, pleon and pereonite 7, dorsal view. 6, detail of the integument ornamentic with characteristic scales." httpUri="https://zenodo.org/record/188891/files/figure.png" pageId="2" pageNumber="59">Figs 4–5</figureCitation>
|
||
) prominently convex; pleonal segments 3–5 with developed epimera. Pleonal segment 3 with medial pair of rounded tubercles and traces of paramedial pair.
|
||
</paragraph>
|
||
<paragraph id="70513686FF80FFC2FF3EF4D1D6E4796D" blockId="2.[151,1438,263,1991]" box="[197,1005,998,1024]" pageId="2" pageNumber="59">
|
||
<emphasis id="429AEA94FF80FFC2FF3EF4D1D448796D" box="[197,321,999,1024]" italics="true" pageId="2" pageNumber="59">Pleotelson</emphasis>
|
||
(
|
||
<figureCitation id="E8D52A03FF80FFC2FEABF4D0D49F796D" box="[336,406,998,1024]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1866,1890]" captionTargetBox="[151,1436,390,1839]" captionTargetId="figure@1.[151,1436,390,1840]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Thaumatoniscellus speluncae n. sp., paratype female (1.31 mm). 1, lateral view of the body. 2, cephalon and pereonites 1 – 3, lateral view. 3, anterior part of the animal, frontal view. 4, pleon and pereonites 5 – 7, lateral view. 5, pleon and pereonite 7, dorsal view. 6, detail of the integument ornamentic with characteristic scales." httpUri="https://zenodo.org/record/188891/files/figure.png" pageId="2" pageNumber="59">Fig. 5</figureCitation>
|
||
) post dorsal surface and posterior margin concave.
|
||
</paragraph>
|
||
<paragraph id="70513686FF80FFC2FF3EF33BD72079F6" blockId="2.[151,1438,263,1991]" pageId="2" pageNumber="59">
|
||
<emphasis id="429AEA94FF80FFC2FF3EF33BD45E794B" box="[197,343,1037,1062]" italics="true" pageId="2" pageNumber="59">Integument</emphasis>
|
||
ornamented with net of polygonal cells bordered with line of granules (
|
||
<figureCitation id="E8D52A03FF80FFC2FAEFF33BD086794A" box="[1300,1423,1037,1063]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1866,1890]" captionTargetBox="[151,1436,390,1839]" captionTargetId="figure@1.[151,1436,390,1840]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Thaumatoniscellus speluncae n. sp., paratype female (1.31 mm). 1, lateral view of the body. 2, cephalon and pereonites 1 – 3, lateral view. 3, anterior part of the animal, frontal view. 4, pleon and pereonites 5 – 7, lateral view. 5, pleon and pereonite 7, dorsal view. 6, detail of the integument ornamentic with characteristic scales." httpUri="https://zenodo.org/record/188891/files/figure.png" pageId="2" pageNumber="59">Figs 3–6</figureCitation>
|
||
). Characteristic scale-setae with a spindle-shaped basal part and fan-shaped apical part (
|
||
<figureCitation id="E8D52A03FF80FFC2FB78F302D1C17923" box="[1155,1224,1076,1102]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1866,1890]" captionTargetBox="[151,1436,390,1839]" captionTargetId="figure@1.[151,1436,390,1840]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Thaumatoniscellus speluncae n. sp., paratype female (1.31 mm). 1, lateral view of the body. 2, cephalon and pereonites 1 – 3, lateral view. 3, anterior part of the animal, frontal view. 4, pleon and pereonites 5 – 7, lateral view. 5, pleon and pereonite 7, dorsal view. 6, detail of the integument ornamentic with characteristic scales." httpUri="https://zenodo.org/record/188891/files/figure.png" pageId="2" pageNumber="59">Fig. 6</figureCitation>
|
||
) on exposed parts of dorsal body surface (tubercles, cephalon, pleotelson and margins of the body segments), frontal sides of antenna and uropodal protopodite.
|
||
</paragraph>
|
||
<paragraph id="70513686FF80FFC2FF3EF39ED09579AF" blockId="2.[151,1438,263,1991]" box="[197,1436,1192,1218]" pageId="2" pageNumber="59">
|
||
<emphasis id="429AEA94FF80FFC2FF3EF39ED43479AC" box="[197,317,1192,1217]" italics="true" pageId="2" pageNumber="59">Antennula</emphasis>
|
||
(
|
||
<figureCitation id="E8D52A03FF80FFC2FEB6F39ED4A879AF" box="[333,417,1192,1218]" captionStart="FIGURES 12 – 15" captionStartId="4.[151,269,1924,1948]" captionTargetBox="[192,1403,459,1870]" captionTargetId="figure@4.[151,1436,429,1898]" captionTargetPageId="4" captionText="FIGURES 12 – 15. Thaumatoniscellus speluncae n. sp., 12, antennula, paratype female (1.38 mm). 13 – 15. Holotype. 13, antenna. 14, pereopod 1. 15, pereopod 7. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/188893/files/figure.png" pageId="2" pageNumber="59">Fig. 12</figureCitation>
|
||
) of 3 articles, similar in length; apex bearing 3 aesthetascs with dorsal surface grooved.
|
||
</paragraph>
|
||
<paragraph id="70513686FF80FFC2FF3EF3F9D0147985" blockId="2.[151,1438,263,1991]" box="[197,1309,1230,1256]" pageId="2" pageNumber="59">
|
||
<emphasis id="429AEA94FF80FFC2FF3EF3F9D42E7985" box="[197,295,1231,1256]" italics="true" pageId="2" pageNumber="59">Antenna</emphasis>
|
||
(
|
||
<figureCitation id="E8D52A03FF80FFC2FECDF3F8D4827985" box="[310,395,1230,1256]" captionStart="FIGURES 12 – 15" captionStartId="4.[151,269,1924,1948]" captionTargetBox="[192,1403,459,1870]" captionTargetId="figure@4.[151,1436,429,1898]" captionTargetPageId="4" captionText="FIGURES 12 – 15. Thaumatoniscellus speluncae n. sp., 12, antennula, paratype female (1.38 mm). 13 – 15. Holotype. 13, antenna. 14, pereopod 1. 15, pereopod 7. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/188893/files/figure.png" pageId="2" pageNumber="59">Fig. 13</figureCitation>
|
||
) with stout pedunclar articles; flagellum three segmented, as long as article 5.
|
||
</paragraph>
|
||
<paragraph id="70513686FF80FFC2FF3EF3C3D1657862" blockId="2.[151,1438,263,1991]" box="[197,1132,1269,1295]" pageId="2" pageNumber="59">
|
||
<emphasis id="429AEA94FF80FFC2FF3EF3C3D4707863" box="[197,377,1269,1294]" italics="true" pageId="2" pageNumber="59">Right mandible</emphasis>
|
||
with 1 penicil; pars incisiva with 3 teeth; lacinia mobilis absent.
|
||
</paragraph>
|
||
<paragraph id="70513686FF80FFC2FF3EF22AD4D97831" blockId="2.[151,1438,263,1991]" pageId="2" pageNumber="59">
|
||
<emphasis id="429AEA94FF80FFC2FF3EF22AD4607858" box="[197,361,1308,1333]" italics="true" pageId="2" pageNumber="59">Left mandible</emphasis>
|
||
without penicils; pars incisiva lateralis with 3 teeth; pars incisiva medialis of molar form; pars molaris oval in shape.
|
||
</paragraph>
|
||
<paragraph id="70513686FF80FFC2FF3EF25FD71A78BD" blockId="2.[151,1438,263,1991]" pageId="2" pageNumber="59">
|
||
<emphasis id="429AEA94FF80FFC2FF3EF25FD43378EF" box="[197,314,1385,1410]" italics="true" pageId="2" pageNumber="59">Maxilla 1</emphasis>
|
||
exopodite (
|
||
<figureCitation id="E8D52A03FF80FFC2FE31F25FD71A78EE" box="[458,531,1385,1411]" captionStart="FIGURES 7 – 11" captionStartId="3.[151,273,1665,1689]" captionTargetBox="[166,1406,222,1605]" captionTargetId="figure@3.[151,1436,197,1640]" captionTargetPageId="3" captionText="FIGURES 7 – 11. Thaumatoniscellus speluncae n. sp., mouth parts of paratype female (1.38 mm). 7, endopodite of maxilla 1. 8, exopodite of maxilla 1. 9, maxilla 2, dorsal view. 10, maxilla 2, ventral view. 11, maxilliped. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/188892/files/figure.png" pageId="2" pageNumber="59">Fig. 8</figureCitation>
|
||
) with 3 strong elongated outer teeth, 5 inner teeth, 3 of them very short; 2 stick-like setae paramedially, 1 very elongated and 1 short; endopodite (
|
||
<figureCitation id="E8D52A03FF80FFC2FC24F2A6D12C78C7" box="[991,1061,1424,1450]" captionStart="FIGURES 7 – 11" captionStartId="3.[151,273,1665,1689]" captionTargetBox="[166,1406,222,1605]" captionTargetId="figure@3.[151,1436,197,1640]" captionTargetPageId="3" captionText="FIGURES 7 – 11. Thaumatoniscellus speluncae n. sp., mouth parts of paratype female (1.38 mm). 7, endopodite of maxilla 1. 8, exopodite of maxilla 1. 9, maxilla 2, dorsal view. 10, maxilla 2, ventral view. 11, maxilliped. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/188892/files/figure.png" pageId="2" pageNumber="59">Fig. 7</figureCitation>
|
||
) with 3 short and stout penicils, terminal one with rounded apex.
|
||
</paragraph>
|
||
<paragraph id="70513686FF80FFC2FF3EF2EBD4927B29" blockId="2.[151,1438,263,1991]" pageId="2" pageNumber="59">
|
||
<emphasis id="429AEA94FF80FFC2FF3EF2EBD43F789B" box="[197,310,1501,1526]" italics="true" pageId="2" pageNumber="59">Maxilla 2</emphasis>
|
||
(
|
||
<figureCitation id="E8D52A03FF80FFC2FEBFF2EBD4B2789A" box="[324,443,1501,1527]" captionStart="FIGURES 7 – 11" captionStartId="3.[151,273,1665,1689]" captionTargetBox="[166,1406,222,1605]" captionTargetId="figure@3.[151,1436,197,1640]" captionTargetPageId="3" captionText="FIGURES 7 – 11. Thaumatoniscellus speluncae n. sp., mouth parts of paratype female (1.38 mm). 7, endopodite of maxilla 1. 8, exopodite of maxilla 1. 9, maxilla 2, dorsal view. 10, maxilla 2, ventral view. 11, maxilliped. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/188892/files/figure.png" pageId="2" pageNumber="59">Figs 9–10</figureCitation>
|
||
) outer lobe narrow, remarkably shorter than inner lobe, with 1 strong rod-like seta on tip; inner lobe with row of elongated rod-like setae on dorsal side, ventral side with brush-like surface of dense, very tiny setae.
|
||
</paragraph>
|
||
<paragraph id="70513686FF80FFC2FF3EF167D4F27BB2" blockId="2.[151,1438,263,1991]" pageId="2" pageNumber="59">
|
||
<emphasis id="429AEA94FF80FFC2FF3EF167D44D7B07" box="[197,324,1617,1642]" italics="true" pageId="2" pageNumber="59">Maxilliped</emphasis>
|
||
(
|
||
<figureCitation id="E8D52A03FF80FFC2FEA8F167D4AF7B06" box="[339,422,1617,1643]" captionStart="FIGURES 7 – 11" captionStartId="3.[151,273,1665,1689]" captionTargetBox="[166,1406,222,1605]" captionTargetId="figure@3.[151,1436,197,1640]" captionTargetPageId="3" captionText="FIGURES 7 – 11. Thaumatoniscellus speluncae n. sp., mouth parts of paratype female (1.38 mm). 7, endopodite of maxilla 1. 8, exopodite of maxilla 1. 9, maxilla 2, dorsal view. 10, maxilla 2, ventral view. 11, maxilliped. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/188892/files/figure.png" pageId="2" pageNumber="59">Fig. 11</figureCitation>
|
||
) basipodite with rounded latero-distal margin with rows of sparse elongated thin setae; palp with three lobed apex, bearing strong setae terminally, row on medial edge and 1 on outer edge; endite conical, with elongate penicil terminally and spine-like protrusion on medio-distal margin, large seta subapically near inner margin.
|
||
</paragraph>
|
||
<paragraph id="70513686FF80FFC2FF3EF1DAD4647A17" blockId="2.[151,1438,263,1991]" pageId="2" pageNumber="59">
|
||
<emphasis id="429AEA94FF80FFC2FF3EF1DAD47F7A68" box="[197,374,1772,1797]" italics="true" pageId="2" pageNumber="59">Pereopods 1–7</emphasis>
|
||
similar in shape, with stout articles; without noticeable secondary sexual characteristics; merus as long as wide, carpus longer than wide (
|
||
<figureCitation id="E8D52A03FF80FFC2FD29F025D6527A40" box="[722,859,1811,1837]" captionStart="FIGURES 12 – 15" captionStartId="4.[151,269,1924,1948]" captionTargetBox="[192,1403,459,1870]" captionTargetId="figure@4.[151,1436,429,1898]" captionTargetPageId="4" captionText="FIGURES 12 – 15. Thaumatoniscellus speluncae n. sp., 12, antennula, paratype female (1.38 mm). 13 – 15. Holotype. 13, antenna. 14, pereopod 1. 15, pereopod 7. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/188893/files/figure.png" pageId="2" pageNumber="59">Figs 14–15</figureCitation>
|
||
). Pereopod 1 with 6 comb-like rows of fine setae on frontal surface of propodus; one row distally on carpus (
|
||
<figureCitation id="E8D52A03FF80FFC2FC9FF00FD6B77A3E" box="[868,958,1849,1875]" captionStart="FIGURES 12 – 15" captionStartId="4.[151,269,1924,1948]" captionTargetBox="[192,1403,459,1870]" captionTargetId="figure@4.[151,1436,429,1898]" captionTargetPageId="4" captionText="FIGURES 12 – 15. Thaumatoniscellus speluncae n. sp., 12, antennula, paratype female (1.38 mm). 13 – 15. Holotype. 13, antenna. 14, pereopod 1. 15, pereopod 7. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/188893/files/figure.png" pageId="2" pageNumber="59">Fig. 14</figureCitation>
|
||
). Pereopod 7 propodus twice as long as dactylus (
|
||
<figureCitation id="E8D52A03FF80FFC2FEF3F056D4567A17" box="[264,351,1888,1914]" captionStart="FIGURES 12 – 15" captionStartId="4.[151,269,1924,1948]" captionTargetBox="[192,1403,459,1870]" captionTargetId="figure@4.[151,1436,429,1898]" captionTargetPageId="4" captionText="FIGURES 12 – 15. Thaumatoniscellus speluncae n. sp., 12, antennula, paratype female (1.38 mm). 13 – 15. Holotype. 13, antenna. 14, pereopod 1. 15, pereopod 7. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/188893/files/figure.png" pageId="2" pageNumber="59">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="70513686FF80FFC2FF3EF0B1D4CB7AAA" blockId="2.[151,1438,263,1991]" pageId="2" pageNumber="59">
|
||
<emphasis id="429AEA94FF80FFC2FF3EF0B1D4757ACD" box="[197,380,1927,1952]" italics="true" pageId="2" pageNumber="59">Genital papilla</emphasis>
|
||
(
|
||
<figureCitation id="E8D52A03FF80FFC2FE77F0B0D71C7ACD" box="[396,533,1926,1952]" captionStart="FIGURES 16 – 21" captionStartId="5.[151,271,1882,1906]" captionTargetBox="[215,1380,230,1811]" captionTargetId="figure@5.[151,1436,194,1858]" captionTargetPageId="5" captionText="FIGURES 16 – 21. Thaumatoniscellus speluncae n. sp., holotype. 16, genital papilla, ventral view. 17, genital papilla, lateral view. 18, exopodite of pleopod 1, ventral view. 19, pleopod 1, dorsal view. 20, pleopod 2. 21, endopodite of pleopod 1, lateral view. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/188894/files/figure.png" pageId="2" pageNumber="59">Figs 16–17</figureCitation>
|
||
) with narrowed terminal part elbows to dorsal side, anchor-like terminally, with tip directed dorsally.
|
||
</paragraph>
|
||
<caption id="2491660EFF81FFC3FF6CF1B7D5C17BB3" httpUri="https://zenodo.org/record/188892/files/figure.png" pageId="3" pageNumber="60" targetBox="[166,1406,222,1605]" targetPageId="3">
|
||
<paragraph id="70513686FF81FFC3FF6CF1B7D5C17BB3" blockId="3.[151,1436,1665,1758]" pageId="3" pageNumber="60">
|
||
<emphasis id="429AEA94FF81FFC3FF6CF1B7D45C7BF4" bold="true" box="[151,341,1665,1689]" pageId="3" pageNumber="60">FIGURES 7–11.</emphasis>
|
||
<taxonomicName id="B7EE4D05FF81FFC3FEA6F1B4D7AC7BF4" box="[349,677,1666,1689]" class="Malacostraca" family="Trichoniscidae" genus="Thaumatoniscellus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="speluncae" status="sp. nov.">
|
||
<emphasis id="429AEA94FF81FFC3FEA6F1B4D7AC7BF4" box="[349,677,1666,1689]" italics="true" pageId="3" pageNumber="60">Thaumatoniscellus speluncae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="59A957EFFF81FFC3FD56F1B7D7E57BF4" box="[685,748,1665,1689]" pageId="3" pageNumber="60" rank="species">
|
||
<emphasis id="429AEA94FF81FFC3FD56F1B7D7E57BF4" bold="true" box="[685,748,1665,1689]" pageId="3" pageNumber="60">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, mouth parts of paratype female (1.38 mm). 7, endopodite of maxilla 1. 8, exopodite of maxilla 1. 9, maxilla 2, dorsal view. 10, maxilla 2, ventral view. 11, maxilliped. Scale bar = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="70513686FF81FFC3FF3EF027D6447AA8" blockId="3.[151,1436,1808,2028]" pageId="3" pageNumber="60">
|
||
<emphasis id="429AEA94FF81FFC3FF3EF027D4377A47" box="[197,318,1809,1834]" italics="true" pageId="3" pageNumber="60">Pleopod 1</emphasis>
|
||
(
|
||
<figureCitation id="E8D52A03FF81FFC3FEB4F026D4DF7A47" box="[335,470,1808,1834]" captionStart="FIGURES 16 – 21" captionStartId="5.[151,271,1882,1906]" captionTargetBox="[215,1380,230,1811]" captionTargetId="figure@5.[151,1436,194,1858]" captionTargetPageId="5" captionText="FIGURES 16 – 21. Thaumatoniscellus speluncae n. sp., holotype. 16, genital papilla, ventral view. 17, genital papilla, lateral view. 18, exopodite of pleopod 1, ventral view. 19, pleopod 1, dorsal view. 20, pleopod 2. 21, endopodite of pleopod 1, lateral view. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/188894/files/figure.png" pageId="3" pageNumber="60">Figs 18–19</figureCitation>
|
||
) exopodite voluminous, laterally constricted in the medial part; lateral margin of basal part widely convex; terminal part with complex three-dimensional structure; with a wide and short conical apophysis distally, laterally-oriented elongated terminal lobe bottle-shaped, mediodistally folded and without apical denticles; wide tetragonal lobe dorsally oriented. Endopodite (
|
||
<figureCitation id="E8D52A03FF81FFC3FBDDF0B2D1757AF3" box="[1062,1148,1924,1950]" captionStart="FIGURES 16 – 21" captionStartId="5.[151,271,1882,1906]" captionTargetBox="[215,1380,230,1811]" captionTargetId="figure@5.[151,1436,194,1858]" captionTargetPageId="5" captionText="FIGURES 16 – 21. Thaumatoniscellus speluncae n. sp., holotype. 16, genital papilla, ventral view. 17, genital papilla, lateral view. 18, exopodite of pleopod 1, ventral view. 19, pleopod 1, dorsal view. 20, pleopod 2. 21, endopodite of pleopod 1, lateral view. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/188894/files/figure.png" pageId="3" pageNumber="60">Fig. 19</figureCitation>
|
||
) shorter than exopodite, stout, terminally slightly widened, without spine-like setae.
|
||
</paragraph>
|
||
<paragraph id="70513686FF81FFC4FF3EF0E4D1787DB5" blockId="3.[151,1436,1808,2028]" lastBlockId="4.[151,1437,152,371]" lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60">
|
||
<emphasis id="429AEA94FF81FFC3FF3EF0E4D4377A86" box="[197,318,2002,2027]" italics="true" pageId="3" pageNumber="60">Pleopod 2</emphasis>
|
||
(
|
||
<figureCitation id="E8D52A03FF81FFC3FEB5F0E4D4DC7A81" box="[334,469,2002,2028]" captionStart="FIGURES 16 – 21" captionStartId="5.[151,271,1882,1906]" captionTargetBox="[215,1380,230,1811]" captionTargetId="figure@5.[151,1436,194,1858]" captionTargetPageId="5" captionText="FIGURES 16 – 21. Thaumatoniscellus speluncae n. sp., holotype. 16, genital papilla, ventral view. 17, genital papilla, lateral view. 18, exopodite of pleopod 1, ventral view. 19, pleopod 1, dorsal view. 20, pleopod 2. 21, endopodite of pleopod 1, lateral view. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/188894/files/figure.png" pageId="3" pageNumber="60">Figs 20–21</figureCitation>
|
||
) exopodite short with protruded mediodistal margin and spine-like setae on distal margin. Endopodite, triarticulated, slightly sigmoidal (lateral view) (
|
||
<figureCitation id="E8D52A03FF86FFC4FC06F7AED1557DDF" box="[1021,1116,152,178]" captionStart="FIGURES 16 – 21" captionStartId="5.[151,271,1882,1906]" captionTargetBox="[215,1380,230,1811]" captionTargetId="figure@5.[151,1436,194,1858]" captionTargetPageId="5" captionText="FIGURES 16 – 21. Thaumatoniscellus speluncae n. sp., holotype. 16, genital papilla, ventral view. 17, genital papilla, lateral view. 18, exopodite of pleopod 1, ventral view. 19, pleopod 1, dorsal view. 20, pleopod 2. 21, endopodite of pleopod 1, lateral view. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/188894/files/figure.png" pageId="4" pageNumber="61">Fig. 21</figureCitation>
|
||
); distal article elongated, spiniform; medial article twice as long as basal article; basal article wider than long.
|
||
</paragraph>
|
||
<paragraph id="70513686FF86FFC4FF3DF7D3D41B7C4B" blockId="4.[151,1437,152,371]" pageId="4" pageNumber="61">
|
||
<emphasis id="429AEA94FF86FFC4FF3DF7D3D4297D93" box="[198,288,229,254]" italics="true" pageId="4" pageNumber="61">Uropod</emphasis>
|
||
exopodite and endopodite conical, subequal in size. Endopodite articulated more proximal than exopodite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="38F4650DFF86FFC4FF3DF604D79E7C1E" pageId="4" pageNumber="61" type="etymology">
|
||
<paragraph id="70513686FF86FFC4FF3DF604D79E7C1E" blockId="4.[151,1437,152,371]" pageId="4" pageNumber="61">
|
||
<emphasis id="429AEA94FF86FFC4FF3DF604D45A7C21" bold="true" box="[198,339,306,332]" pageId="4" pageNumber="61">Etymology.</emphasis>
|
||
The species is named after the Speleological Organisation “Spelunka“ from Veprinac, Istria. The name is a noun in the genitive singular.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |