293 lines
60 KiB
XML
293 lines
60 KiB
XML
<document id="A025ABFC05CC743038B9799486ADAD00" ID-CLB-Dataset="298715" ID-DOI="10.11646/zootaxa.5471.2.1" ID-GBIF-Dataset="3a469638-dc21-45a3-80f6-cb307fbf5941" ID-ISSN="1175-5326" ID-Zenodo-Dep="12189904" ID-ZooBank="BAA047F4-2D65-4E52-8798-D426E3B26DA1" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1718892848818" checkinUser="plazi" docAuthor="Kosterin, Oleg E., Onishko, Vladimir V., Ilyina, Elena V., Chepurnov, Grigory Yu. & Blinov, Alexander G." docDate="2024" docId="DB0F87E1910EF101A7D3F80979D87817" docLanguage="en" docName="zootaxa.5471.2.1.pdf" docOrigin="Zootaxa 5471 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5471.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coenagrion australocaspicum Dumont & Heidari 1996" docType="treatment" docVersion="4" lastPageNumber="182" masterDocId="2736FF999111F11EA744F92E7A657E46" masterDocTitle="The genus Coenagrion Kirby, 1890 (Odonata: Coenagrionidae) in the Russian part of the Caucasus" masterLastPageNumber="190" masterPageNumber="151" pageNumber="182" updateTime="1718988639371" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A71310D64637C6C3C34196DC97D3675B">The genus Coenagrion Kirby, 1890 (Odonata: Coenagrionidae) in the Russian part of the Caucasus</mods:title>
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<mods:namePart id="4BAEA84DD8BCA61AF52A466B0833D99B">Kosterin, Oleg E.</mods:namePart>
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<mods:affiliation id="5DBACDEE68D01F82C7696A6724D50E8A">Institute of Cytology & Genetics SB RAS, Acad. Lavrentyev ave. 10, Novosibirsk, 630090, Russia.</mods:affiliation>
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<mods:nameIdentifier id="F1D085245B318874957E079BE073B2D4" type="email">kosterin@bionet.nsc.ru</mods:nameIdentifier>
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<mods:namePart id="89A4D08E616B14310833F28E7D06831A">Onishko, Vladimir V.</mods:namePart>
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<mods:nameIdentifier id="3C90EDD2915C9FC1EEB96998979D5CAC" type="ORCID">0000-0002-6469-6778</mods:nameIdentifier>
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<mods:affiliation id="0F913D74B663C8C28F72CF414E955683">Moscow Zoo, Department of Herpetology, Bolshaya Gruzinskaya Str. 1, Moscow 123242 Russia.</mods:affiliation>
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<mods:nameIdentifier id="2B1A0B2BC935FC7212470646F18BFD99" type="email">wervolf999@yandex.ru</mods:nameIdentifier>
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<mods:namePart id="F6C0B2DE462D4374E882A8C86D1401B9">Ilyina, Elena V.</mods:namePart>
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<mods:affiliation id="3BCA2B58704E168D95000DDA4329AC44">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, Dagestanskiy State Nature Reserve, Gagarina Street 120, Makhachkala, Republic of Dagestan, Russia.</mods:affiliation>
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<mods:nameIdentifier id="44587593FCE4F4C0B8292D5C9ACF5838" type="email">carabus@list.ru</mods:nameIdentifier>
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<mods:namePart id="779A8BD2466CBF63C216CB899D4F0FA8">Chepurnov, Grigory Yu.</mods:namePart>
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<mods:affiliation id="82C912B1B8DDAEB8C95562BAF0237EEA">Institute of Cytology & Genetics SB RAS, Acad. Lavrentyev ave. 10, Novosibirsk, 630090, Russia. & Institute of Molecular and Cellular Biology SB RAS, Acad. Lavrentyev ave. 8 / 2, Novosibirsk, 630090, Russia.</mods:affiliation>
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<mods:nameIdentifier id="797758341B4AC7DEAC355FAC115444C2" type="email">grigory.tchepurnov@yandex.ru</mods:nameIdentifier>
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<mods:namePart id="511CE7C9BB5AD45A2BA52FFE94250EFE">Blinov, Alexander G.</mods:namePart>
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<mods:affiliation id="B149099A39F1F3084C823C31AA0CB829">Institute of Cytology & Genetics SB RAS, Acad. Lavrentyev ave. 10, Novosibirsk, 630090, Russia. & Institute of Molecular and Cellular Biology SB RAS, Acad. Lavrentyev ave. 8 / 2, Novosibirsk, 630090, Russia.</mods:affiliation>
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<mods:nameIdentifier id="AA74322BE70002273EB935278F820B70" type="email">blinov@bionet.nsc.ru</mods:nameIdentifier>
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<mods:title id="F7E4D01BD7F39262D846ACE0C6CA9699">Zootaxa</mods:title>
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<mods:part id="EAB95432325EAD8AFE63B6325BD550D4">
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<mods:date id="9948AA9262442208BB771B129973EADA">2024</mods:date>
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<mods:number id="EEB3CCB7B00D36A02B80A01B2315060C">2024-06-18</mods:number>
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<mods:number id="5000F30755A899699F8FA057185F8EAB">5471</mods:number>
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<mods:url id="282B964F2F403495BE49538110EE9E0C">http://dx.doi.org/10.11646/zootaxa.5471.2.1</mods:url>
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<treatment id="DB0F87E1910EF101A7D3F80979D87817" ID-DOI="http://doi.org/10.5281/zenodo.12206851" ID-GBIF-Taxon="233309484" ID-Zenodo-Dep="12206851" LSID="urn:lsid:plazi:treatment:DB0F87E1910EF101A7D3F80979D87817" httpUri="http://treatment.plazi.org/id/DB0F87E1910EF101A7D3F80979D87817" lastPageNumber="182" pageId="31" pageNumber="182">
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<subSubSection id="1BBC657C910EF101A7D3F80978617F07" box="[151,516,294,321]" pageId="31" pageNumber="182" type="nomenclature">
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<paragraph id="531936F7910EF101A7D3F80978617F07" blockId="31.[151,516,294,321]" box="[151,516,294,321]" pageId="31" pageNumber="182">
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<heading id="0851819B910EF101A7D3F80978617F07" bold="true" box="[151,516,294,321]" fontSize="11" level="1" pageId="31" pageNumber="182" reason="1">
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<taxonomicName id="94A64D74910EF101A7D3F80978617F07" ID-CoL="WTWH" authority="Dumont & Heidari, 1996" authorityName="Dumont & Heidari" authorityYear="1996" box="[151,516,294,321]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="australocaspicum">
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<emphasis id="61D2EAE5910EF101A7D3F80978617F07" bold="true" box="[151,516,294,321]" italics="true" pageId="31" pageNumber="182">Coenagrion australocaspicum</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="1BBC657C910EF101A7D3F8417E307A1F" pageId="31" pageNumber="182" type="reference_group">
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<paragraph id="531936F7910EF101A7D3F8417F2F7D8F" blockId="31.[151,1437,367,1617]" pageId="31" pageNumber="182">
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<taxonomicName id="94A64D74910EF101A7D3F8417BE57FCF" ID-CoL="WTWH" authorityName="Dumont & Heidari" authorityYear="1996" box="[151,384,367,393]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="australocaspicum">
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<emphasis id="61D2EAE5910EF101A7D3F8417BE57FCF" box="[151,384,367,393]" italics="true" pageId="31" pageNumber="182">C. australocaspicum</emphasis>
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</taxonomicName>
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stands aside from other
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<taxonomicName id="94A64D74910EF101A5DDF84179307FCF" box="[665,853,367,393]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="61D2EAE5910EF101A5DDF841797A7FCF" box="[665,799,367,393]" italics="true" pageId="31" pageNumber="182">Coenagrion</emphasis>
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spp.
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</taxonomicName>
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in the area considered with respect to its very long male paraprocts, a character seen even by a naked eye and shared only by
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<taxonomicName id="94A64D74910EF101A488F8BD7E367FEA" ID-CoL="WTYC" baseAuthorityName="Morton" baseAuthorityYear="1924" box="[972,1107,403,429]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="syriacum">
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<emphasis id="61D2EAE5910EF101A488F8BD7E367FEA" box="[972,1107,403,429]" italics="true" pageId="31" pageNumber="182">C. syriacum</emphasis>
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</taxonomicName>
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ranging in the Mediterranean coastal areas of
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<collectingCountry id="2BB17667910EF101A609F8997BFB7F97" box="[333,414,439,465]" name="Turkey" pageId="31" pageNumber="182">Turkey</collectingCountry>
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,
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<collectingCountry id="2BB17667910EF101A6EEF8997B8D7F97" box="[426,488,439,465]" name="Syria" pageId="31" pageNumber="182">Syria</collectingCountry>
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,
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<collectingCountry id="2BB17667910EF101A6B0F89978337F97" box="[500,598,439,465]" name="Lebanon" pageId="31" pageNumber="182">Lebanon</collectingCountry>
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and
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<collectingCountry id="2BB17667910EF101A5CAF89978A97F97" box="[654,716,439,465]" name="Israel" pageId="31" pageNumber="182">Israel</collectingCountry>
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(
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<bibRefCitation id="37374B06910EF101A59FF89979EC7F97" author="Kalkman, V. J." box="[731,905,439,465]" pageId="31" pageNumber="182" pagination="3 - 82" refId="ref28232" refString="Kalkman, V. J. (2006) Key to the dragonflies of Turkey including species known from Greece, Bulgaria, Lebanon, Syria, the Trans-Caucasus and Iran. Brachytron, 10 (1), 3 - 82." type="journal article" year="2006">Kalkman 2006</bibRefCitation>
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). It is remarkable that the only difference of
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<taxonomicName id="94A64D74910EF101A2C7F8997B3B7FB3" ID-CoL="WTWH" authorityName="Dumont & Heidari" authorityYear="1996" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="australocaspicum">
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<emphasis id="61D2EAE5910EF101A2C7F8997B3B7FB3" italics="true" pageId="31" pageNumber="182">C. australocaspicum</emphasis>
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</taxonomicName>
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from
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<taxonomicName id="94A64D74910EF101A6E5F8F5784D7FB2" ID-CoL="WTYC" baseAuthorityName="Morton" baseAuthorityYear="1924" box="[417,552,475,501]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="syriacum">
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<emphasis id="61D2EAE5910EF101A6E5F8F5784D7FB2" box="[417,552,475,501]" italics="true" pageId="31" pageNumber="182">C. syriacum</emphasis>
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</taxonomicName>
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stated in the original description is as follows: “It differs, from
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<taxonomicName id="94A64D74910EF101A3A2F8F57F0B7FB2" ID-CoL="WTXX" baseAuthorityName="Bartenef" baseAuthorityYear="1929" box="[1254,1390,475,501]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="ponticum">
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<emphasis id="61D2EAE5910EF101A3A2F8F57F0B7FB2" box="[1254,1390,475,501]" italics="true" pageId="31" pageNumber="182">C. ponticum</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="94A64D74910EF101A7D3F8D17B7B7C5E" ID-CoL="WTYC" baseAuthorityName="Morton" baseAuthorityYear="1924" box="[151,286,511,537]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="syriacum">
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<emphasis id="61D2EAE5910EF101A7D3F8D17B7B7C5E" box="[151,286,511,537]" italics="true" pageId="31" pageNumber="182">C. syriacum</emphasis>
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</taxonomicName>
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by the widely separated app. sup. of the males, a character it only shares with
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<taxonomicName id="94A64D74910EF101A33EF8D17E857C5F" ID-CoL="WTXY" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1146,1248,511,537]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="puella">
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<emphasis id="61D2EAE5910EF101A33EF8D17E857C5F" box="[1146,1248,511,537]" italics="true" pageId="31" pageNumber="182">C. puella</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="7AE1579E910EF101A3A1F8D17F6D7C5F" box="[1253,1288,511,537]" pageId="31" pageNumber="182" sensu="stricto">s.s.</taxonomicNameLabel>
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” (
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<bibRefCitation id="37374B06910EF101A267F8D17B487C7B" author="Dumont, H. J. & Heidari, H." pageId="31" pageNumber="182" pagination="63 - 78" refId="ref27800" refString="Dumont, H. J. & Heidari, H. (1996) On a collection of spring Odonata from Iran, with the description of Coenagrion australocaspicum n. sp. Bulletin et Annales de la Societe Royale Belge d'Entomologie, 132, 63 - 78." type="journal article" year="1996">Dumont & Heidari 1996</bibRefCitation>
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). Unfortunarely, this character was not well illustrated in the original descripion, where the illustrations contradict each other. The drawings in the dorsal and caudal views show the bases of the cerci set apart from each other and laterally of the central prominence of the S10 inner side (
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<bibRefCitation id="37374B06910EF101A4C0FB457EF97CC3" author="Dumont, H. J. & Heidari, H." box="[900,1180,619,645]" pageId="31" pageNumber="182" pagination="63 - 78" refId="ref27800" refString="Dumont, H. J. & Heidari, H. (1996) On a collection of spring Odonata from Iran, with the description of Coenagrion australocaspicum n. sp. Bulletin et Annales de la Societe Royale Belge d'Entomologie, 132, 63 - 78." type="journal article" year="1996">Dumont & Heidari 1996</bibRefCitation>
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: figs 7, 8). At the same time, the scanned electronic microphotograph shows, in caudal view, their bases disposed close to each other under the central prominence (Ibid.: fig. 4). Exactly this is seen also in our caudal view photos of
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<taxonomicName id="94A64D74910EF101A3F7FB9D7FF97C8B" ID-CoL="WTWH" authorityName="Dumont & Heidari" authorityYear="1996" box="[1203,1436,691,717]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="australocaspicum">
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<emphasis id="61D2EAE5910EF101A3F7FB9D7FF97C8B" box="[1203,1436,691,717]" italics="true" pageId="31" pageNumber="182">C. australocaspicum</emphasis>
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</taxonomicName>
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from Dagestan (
|
||
<figureCitation id="CB9D2A72910EF101A614FBF97BDB7CB7" box="[336,446,727,753]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="31" pageNumber="182">Fig. 4a–b</figureCitation>
|
||
) and in the photos of male specimens of
|
||
<taxonomicName id="94A64D74910EF101A4DEFBF97E467CB6" ID-CoL="WTYC" baseAuthorityName="Morton" baseAuthorityYear="1924" box="[922,1059,727,753]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="syriacum">
|
||
<emphasis id="61D2EAE5910EF101A4DEFBF97E467CB6" box="[922,1059,727,753]" italics="true" pageId="31" pageNumber="182">C. syriacum</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="9162F815910EF101A32DFBF97ED77CB7" box="[1129,1202,727,753]" country="Turkey" name="Adana" pageId="31" pageNumber="182">Adana</collectingRegion>
|
||
Il of
|
||
<collectingCountry id="2BB17667910EF101A3B1FBF97F207CB7" box="[1269,1349,727,753]" name="Turkey" pageId="31" pageNumber="182">Turkey</collectingCountry>
|
||
kept in RMNH, kindly provided by the curator of Odonata collection Charlotte Hartong (
|
||
<figureCitation id="CB9D2A72910EF101A36BFBD57ECC7D53" box="[1071,1193,763,789]" captionStart="FIGURE 16" captionStartId="32.[151,250,1220,1244]" captionTargetBox="[164,1423,181,1196]" captionTargetId="figure-24@32.[164,1423,181,1196]" captionTargetPageId="32" captionText="FIGURE 16. Morphological details of male specimens of Coenagrion syriacum kept in the RMNH collection, courtesy by Charlotte Hartong: a–c—“Turkey, Adana / River Ceyhan, Ca. 20 km NW of / Osmaniye (37SBB42) River and / pools. / 11 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH.INS.1468579); d–f—“Turkey, Adana / Aydinlar, 15 km W of Kadirli / (37SBB33) Well vegetated, slowly flowing / channel. / 10 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH. INS.1468580); a, d—end of abbdomen in lateral view; b, e—ditto, dorsal view; a, d—ditto, caudal view." figureDoi="http://doi.org/10.5281/zenodo.12189942" httpUri="https://zenodo.org/record/12189942/files/figure.png" pageId="31" pageNumber="182">Fig. 16d,f</figureCitation>
|
||
). Taking into account the subjectivity of drawings, we conclude that the very bases of the cerci are set tightly close to each other in both species. The actual difference between
|
||
<taxonomicName id="94A64D74910EF101A511FA6D795B7D1B" ID-CoL="WTWH" authorityName="Dumont & Heidari" authorityYear="1996" box="[597,830,835,861]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="australocaspicum">
|
||
<emphasis id="61D2EAE5910EF101A511FA6D795B7D1B" box="[597,830,835,861]" italics="true" pageId="31" pageNumber="182">C. australocaspicum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="94A64D74910EF101A43CFA6D7E647D1A" ID-CoL="WTYC" baseAuthorityName="Morton" baseAuthorityYear="1924" box="[888,1025,835,861]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="syriacum">
|
||
<emphasis id="61D2EAE5910EF101A43CFA6D7E647D1A" box="[888,1025,835,861]" italics="true" pageId="31" pageNumber="182">C. syriacum</emphasis>
|
||
</taxonomicName>
|
||
is seen in dorsal view (
|
||
<figureCitation id="CB9D2A72910EF101A254FA6D7F1E7D1B" box="[1296,1403,835,861]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="31" pageNumber="182">Fig. 2a–c</figureCitation>
|
||
vs
|
||
<figureCitation id="CB9D2A72910EF101A7D3FA497B6B7DC7" box="[151,270,871,897]" captionStart="FIGURE 16" captionStartId="32.[151,250,1220,1244]" captionTargetBox="[164,1423,181,1196]" captionTargetId="figure-24@32.[164,1423,181,1196]" captionTargetPageId="32" captionText="FIGURE 16. Morphological details of male specimens of Coenagrion syriacum kept in the RMNH collection, courtesy by Charlotte Hartong: a–c—“Turkey, Adana / River Ceyhan, Ca. 20 km NW of / Osmaniye (37SBB42) River and / pools. / 11 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH.INS.1468579); d–f—“Turkey, Adana / Aydinlar, 15 km W of Kadirli / (37SBB33) Well vegetated, slowly flowing / channel. / 10 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH. INS.1468580); a, d—end of abbdomen in lateral view; b, e—ditto, dorsal view; a, d—ditto, caudal view." figureDoi="http://doi.org/10.5281/zenodo.12189942" httpUri="https://zenodo.org/record/12189942/files/figure.png" pageId="31" pageNumber="182">Fig. 16b, e</figureCitation>
|
||
) and consist of the presence of inner bulges of the basal parts of the cerci in
|
||
<taxonomicName id="94A64D74910EF101A31BFA497E827DC6" ID-CoL="WTYC" baseAuthorityName="Morton" baseAuthorityYear="1924" box="[1119,1255,871,897]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="syriacum">
|
||
<emphasis id="61D2EAE5910EF101A31BFA497E827DC6" box="[1119,1255,871,897]" italics="true" pageId="31" pageNumber="182">C. syriacum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="CB9D2A72910EF101A3B1FA497F0B7DC7" box="[1269,1390,871,897]" captionStart="FIGURE 16" captionStartId="32.[151,250,1220,1244]" captionTargetBox="[164,1423,181,1196]" captionTargetId="figure-24@32.[164,1423,181,1196]" captionTargetPageId="32" captionText="FIGURE 16. Morphological details of male specimens of Coenagrion syriacum kept in the RMNH collection, courtesy by Charlotte Hartong: a–c—“Turkey, Adana / River Ceyhan, Ca. 20 km NW of / Osmaniye (37SBB42) River and / pools. / 11 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH.INS.1468579); d–f—“Turkey, Adana / Aydinlar, 15 km W of Kadirli / (37SBB33) Well vegetated, slowly flowing / channel. / 10 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH. INS.1468580); a, d—end of abbdomen in lateral view; b, e—ditto, dorsal view; a, d—ditto, caudal view." figureDoi="http://doi.org/10.5281/zenodo.12189942" httpUri="https://zenodo.org/record/12189942/files/figure.png" pageId="31" pageNumber="182">Fig. 16b, e</figureCitation>
|
||
), so that they contact to each other.
|
||
<taxonomicName id="94A64D74910EF101A6B2FAA578BB7DE3" ID-CoL="WTWH" authorityName="Dumont & Heidari" authorityYear="1996" box="[502,734,907,933]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="australocaspicum">
|
||
<emphasis id="61D2EAE5910EF101A6B2FAA578BB7DE3" box="[502,734,907,933]" italics="true" pageId="31" pageNumber="182">C. australocaspicum</emphasis>
|
||
</taxonomicName>
|
||
misses these bulges (
|
||
<figureCitation id="CB9D2A72910EF101A495FAA57E5B7DE3" box="[977,1086,907,933]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="31" pageNumber="182">Fig. 2a–c</figureCitation>
|
||
), so its cerci are well separated from each other indeed in dorsal view, but this is a matter of their shape rather than positions of their bases.
|
||
</paragraph>
|
||
<paragraph id="531936F7910EF101A783FAFD7E307A1F" blockId="31.[151,1437,367,1617]" pageId="31" pageNumber="182">
|
||
The original description of
|
||
<taxonomicName id="94A64D74910EF101A6B4FAFD78B07DAB" ID-CoL="WTWH" authorityName="Dumont & Heidari" authorityYear="1996" box="[496,725,979,1005]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="australocaspicum">
|
||
<emphasis id="61D2EAE5910EF101A6B4FAFD78B07DAB" box="[496,725,979,1005]" italics="true" pageId="31" pageNumber="182">C. australocaspicum</emphasis>
|
||
</taxonomicName>
|
||
also mentions (but not as diagnostic) and illustrates the paraproct basoventral part with a rounded prominence, rather than an angular one, as in
|
||
<taxonomicName id="94A64D74910EF101A4B9FAD97EE07A56" ID-CoL="WTYC" baseAuthorityName="Morton" baseAuthorityYear="1924" box="[1021,1157,1015,1041]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="syriacum">
|
||
<emphasis id="61D2EAE5910EF101A4B9FAD97EE07A56" box="[1021,1157,1015,1041]" italics="true" pageId="31" pageNumber="182">C. syriacum</emphasis>
|
||
</taxonomicName>
|
||
, but our specimens of
|
||
<taxonomicName id="94A64D74910EF101A2C7FAD97B3B7A73" ID-CoL="WTWH" authorityName="Dumont & Heidari" authorityYear="1996" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="australocaspicum">
|
||
<emphasis id="61D2EAE5910EF101A2C7FAD97B3B7A73" italics="true" pageId="31" pageNumber="182">C. australocaspicum</emphasis>
|
||
</taxonomicName>
|
||
from Dagestan (
|
||
<figureCitation id="CB9D2A72910EF101A55BFD3578EE7A73" box="[543,651,1051,1077]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="31" pageNumber="182">Fig. 3a–c</figureCitation>
|
||
) and those of Skvortzov & Snegovaya (2015) from
|
||
<collectingCountry id="2BB17667910EF101A3A4FD357F3B7A73" box="[1248,1374,1051,1077]" name="Azerbaijan" pageId="31" pageNumber="182">Azerbaijan</collectingCountry>
|
||
, both series being from the Caspian coast, exhibit the latter condition of angular projection.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1BBC657C910EF101A783FD4D79D87817" pageId="31" pageNumber="182" type="description">
|
||
<paragraph id="531936F7910EF101A783FD4D78D07A83" blockId="31.[151,1437,367,1617]" pageId="31" pageNumber="182">
|
||
The comparison of the photographs of male specimens of
|
||
<taxonomicName id="94A64D74910EF101A433FD4D7E667A3A" baseAuthorityName="Morton" baseAuthorityYear="1924" box="[887,1027,1123,1149]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="syriacum">
|
||
<emphasis id="61D2EAE5910EF101A433FD4D7E667A3A" box="[887,1027,1123,1149]" italics="true" pageId="31" pageNumber="182">C. syriacum</emphasis>
|
||
</taxonomicName>
|
||
kept in RMNH (
|
||
<figureCitation id="CB9D2A72910EF101A395FD4D7F427A3B" box="[1233,1319,1123,1149]" captionStart="FIGURE 16" captionStartId="32.[151,250,1220,1244]" captionTargetBox="[164,1423,181,1196]" captionTargetId="figure-24@32.[164,1423,181,1196]" captionTargetPageId="32" captionText="FIGURE 16. Morphological details of male specimens of Coenagrion syriacum kept in the RMNH collection, courtesy by Charlotte Hartong: a–c—“Turkey, Adana / River Ceyhan, Ca. 20 km NW of / Osmaniye (37SBB42) River and / pools. / 11 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH.INS.1468579); d–f—“Turkey, Adana / Aydinlar, 15 km W of Kadirli / (37SBB33) Well vegetated, slowly flowing / channel. / 10 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH. INS.1468580); a, d—end of abbdomen in lateral view; b, e—ditto, dorsal view; a, d—ditto, caudal view." figureDoi="http://doi.org/10.5281/zenodo.12189942" httpUri="https://zenodo.org/record/12189942/files/figure.png" pageId="31" pageNumber="182">Fig. 16</figureCitation>
|
||
) with our specimens of
|
||
<taxonomicName id="94A64D74910EF101A675FDA9787C7AE7" authorityName="Dumont & Heidari" authorityYear="1996" box="[305,537,1159,1185]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="australocaspicum">
|
||
<emphasis id="61D2EAE5910EF101A675FDA9787C7AE7" box="[305,537,1159,1185]" italics="true" pageId="31" pageNumber="182">C. australocaspicum</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="9162F815910EF101A519FDA978A07AE7" box="[605,709,1159,1185]" country="Russia" name="Dagestan" pageId="31" pageNumber="182">Dagestan</collectingRegion>
|
||
(
|
||
<figureCitation id="CB9D2A72910EF101A591FDA979217AE7" box="[725,836,1159,1185]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="31" pageNumber="182">Figs 2a–c</figureCitation>
|
||
,
|
||
<figureCitation id="CB9D2A72910EF101A414FDA979ED7AE7" box="[848,904,1159,1185]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="31" pageNumber="182">3a–c</figureCitation>
|
||
,
|
||
<figureCitation id="CB9D2A72910EF101A4D7FDA979A97AE7" box="[915,972,1159,1185]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="31" pageNumber="182">4a–b</figureCitation>
|
||
,
|
||
<figureCitation id="CB9D2A72910EF101A49CFDA879957AE7" box="[984,1008,1158,1185]" captionStart="FIGURE 5" captionStartId="13.[151,250,1570,1594]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@13.[151,1436,181,1546]" captionTargetPageId="13" captionText="FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="31" pageNumber="182">5f</figureCitation>
|
||
) showed the following morphological differences of the former species from the latter:
|
||
</paragraph>
|
||
<paragraph id="531936F7910EF101A783FDE1797A7AAF" blockId="31.[151,1437,367,1617]" box="[199,799,1231,1257]" pageId="31" pageNumber="182">
|
||
—the cercus has a well expressed bulge (
|
||
<figureCitation id="CB9D2A72910EF101A5D1FDE179757AAF" box="[661,784,1231,1257]" captionStart="FIGURE 16" captionStartId="32.[151,250,1220,1244]" captionTargetBox="[164,1423,181,1196]" captionTargetId="figure-24@32.[164,1423,181,1196]" captionTargetPageId="32" captionText="FIGURE 16. Morphological details of male specimens of Coenagrion syriacum kept in the RMNH collection, courtesy by Charlotte Hartong: a–c—“Turkey, Adana / River Ceyhan, Ca. 20 km NW of / Osmaniye (37SBB42) River and / pools. / 11 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH.INS.1468579); d–f—“Turkey, Adana / Aydinlar, 15 km W of Kadirli / (37SBB33) Well vegetated, slowly flowing / channel. / 10 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH. INS.1468580); a, d—end of abbdomen in lateral view; b, e—ditto, dorsal view; a, d—ditto, caudal view." figureDoi="http://doi.org/10.5281/zenodo.12189942" httpUri="https://zenodo.org/record/12189942/files/figure.png" pageId="31" pageNumber="182">Fig. 16b, e</figureCitation>
|
||
);
|
||
</paragraph>
|
||
<paragraph id="531936F7910EF101A783FDDD7BB17B77" blockId="31.[151,1437,367,1617]" pageId="31" pageNumber="182">
|
||
—the paraproct spine is thicker, blunter, a bit relatively shorter and very slightly skewed up rather than directed strictly caudad (
|
||
<figureCitation id="CB9D2A72910EF101A60FFC397BA37B77" box="[331,454,1303,1329]" captionStart="FIGURE 16" captionStartId="32.[151,250,1220,1244]" captionTargetBox="[164,1423,181,1196]" captionTargetId="figure-24@32.[164,1423,181,1196]" captionTargetPageId="32" captionText="FIGURE 16. Morphological details of male specimens of Coenagrion syriacum kept in the RMNH collection, courtesy by Charlotte Hartong: a–c—“Turkey, Adana / River Ceyhan, Ca. 20 km NW of / Osmaniye (37SBB42) River and / pools. / 11 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH.INS.1468579); d–f—“Turkey, Adana / Aydinlar, 15 km W of Kadirli / (37SBB33) Well vegetated, slowly flowing / channel. / 10 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH. INS.1468580); a, d—end of abbdomen in lateral view; b, e—ditto, dorsal view; a, d—ditto, caudal view." figureDoi="http://doi.org/10.5281/zenodo.12189942" httpUri="https://zenodo.org/record/12189942/files/figure.png" pageId="31" pageNumber="182">Fig. 16a, d</figureCitation>
|
||
);
|
||
</paragraph>
|
||
<paragraph id="531936F7910EF101A783FC157AD37B3F" blockId="31.[151,1437,367,1617]" pageId="31" pageNumber="182">
|
||
—the tubercle of the cerci is slanting down in
|
||
<taxonomicName id="94A64D74910EF101A583FC15792B7B12" baseAuthorityName="Morton" baseAuthorityYear="1924" box="[711,846,1339,1365]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="syriacum">
|
||
<emphasis id="61D2EAE5910EF101A583FC15792B7B12" box="[711,846,1339,1365]" italics="true" pageId="31" pageNumber="182">C. syriacum</emphasis>
|
||
</taxonomicName>
|
||
but slightly raised in
|
||
<taxonomicName id="94A64D74910EF101A379FC157F417B13" authorityName="Dumont & Heidari" authorityYear="1996" box="[1085,1316,1339,1365]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="australocaspicum">
|
||
<emphasis id="61D2EAE5910EF101A379FC157F417B13" box="[1085,1316,1339,1365]" italics="true" pageId="31" pageNumber="182">C. australocaspicum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="CB9D2A72910EF101A27DFC157ACD7B3F" captionStart="FIGURE 16" captionStartId="32.[151,250,1220,1244]" captionTargetBox="[164,1423,181,1196]" captionTargetId="figure-24@32.[164,1423,181,1196]" captionTargetPageId="32" captionText="FIGURE 16. Morphological details of male specimens of Coenagrion syriacum kept in the RMNH collection, courtesy by Charlotte Hartong: a–c—“Turkey, Adana / River Ceyhan, Ca. 20 km NW of / Osmaniye (37SBB42) River and / pools. / 11 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH.INS.1468579); d–f—“Turkey, Adana / Aydinlar, 15 km W of Kadirli / (37SBB33) Well vegetated, slowly flowing / channel. / 10 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH. INS.1468580); a, d—end of abbdomen in lateral view; b, e—ditto, dorsal view; a, d—ditto, caudal view." figureDoi="http://doi.org/10.5281/zenodo.12189942" httpUri="https://zenodo.org/record/12189942/files/figure.png" pageId="31" pageNumber="182">Fig. 16a, d</figureCitation>
|
||
);
|
||
</paragraph>
|
||
<paragraph id="531936F7910EF101A783FCAD7E1E7B87" blockId="31.[151,1437,367,1617]" pageId="31" pageNumber="182">
|
||
—the processes of the cerci in
|
||
<taxonomicName id="94A64D74910EF101A55DFCAD78C57BDA" baseAuthorityName="Morton" baseAuthorityYear="1924" box="[537,672,1411,1437]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="syriacum">
|
||
<emphasis id="61D2EAE5910EF101A55DFCAD78C57BDA" box="[537,672,1411,1437]" italics="true" pageId="31" pageNumber="182">C. syriacum</emphasis>
|
||
</taxonomicName>
|
||
are much longer than in
|
||
<taxonomicName id="94A64D74910EF101A4F4FCAD7EF27BDB" authorityName="Dumont & Heidari" authorityYear="1996" box="[944,1175,1411,1437]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="australocaspicum">
|
||
<emphasis id="61D2EAE5910EF101A4F4FCAD7EF27BDB" box="[944,1175,1411,1437]" italics="true" pageId="31" pageNumber="182">C. australocaspicum</emphasis>
|
||
</taxonomicName>
|
||
and protrude below the middle of the paraproct bases (
|
||
<figureCitation id="CB9D2A72910EF101A6B4FC8978027B87" box="[496,615,1447,1473]" captionStart="FIGURE 16" captionStartId="32.[151,250,1220,1244]" captionTargetBox="[164,1423,181,1196]" captionTargetId="figure-24@32.[164,1423,181,1196]" captionTargetPageId="32" captionText="FIGURE 16. Morphological details of male specimens of Coenagrion syriacum kept in the RMNH collection, courtesy by Charlotte Hartong: a–c—“Turkey, Adana / River Ceyhan, Ca. 20 km NW of / Osmaniye (37SBB42) River and / pools. / 11 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH.INS.1468579); d–f—“Turkey, Adana / Aydinlar, 15 km W of Kadirli / (37SBB33) Well vegetated, slowly flowing / channel. / 10 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH. INS.1468580); a, d—end of abbdomen in lateral view; b, e—ditto, dorsal view; a, d—ditto, caudal view." figureDoi="http://doi.org/10.5281/zenodo.12189942" httpUri="https://zenodo.org/record/12189942/files/figure.png" pageId="31" pageNumber="182">Fig. 16c, f</figureCitation>
|
||
), they are straight almost like in
|
||
<taxonomicName id="94A64D74910EF101A492FC897E167B87" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[982,1139,1447,1473]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="pulchellum">
|
||
<emphasis id="61D2EAE5910EF101A492FC897E167B87" box="[982,1139,1447,1473]" italics="true" pageId="31" pageNumber="182">C. pulchellum</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
</paragraph>
|
||
<paragraph id="531936F7910EF101A783FCE579C67BA3" blockId="31.[151,1437,367,1617]" box="[199,931,1483,1509]" pageId="31" pageNumber="182">
|
||
—the S10 dorsal margin more raised in lateral view (
|
||
<figureCitation id="CB9D2A72910EF101A45EFCE579F37BA3" box="[794,918,1483,1509]" captionStart="FIGURE 16" captionStartId="32.[151,250,1220,1244]" captionTargetBox="[164,1423,181,1196]" captionTargetId="figure-24@32.[164,1423,181,1196]" captionTargetPageId="32" captionText="FIGURE 16. Morphological details of male specimens of Coenagrion syriacum kept in the RMNH collection, courtesy by Charlotte Hartong: a–c—“Turkey, Adana / River Ceyhan, Ca. 20 km NW of / Osmaniye (37SBB42) River and / pools. / 11 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH.INS.1468579); d–f—“Turkey, Adana / Aydinlar, 15 km W of Kadirli / (37SBB33) Well vegetated, slowly flowing / channel. / 10 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH. INS.1468580); a, d—end of abbdomen in lateral view; b, e—ditto, dorsal view; a, d—ditto, caudal view." figureDoi="http://doi.org/10.5281/zenodo.12189942" httpUri="https://zenodo.org/record/12189942/files/figure.png" pageId="31" pageNumber="182">Fig. 16a, d</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="531936F7910EF101A783FCC179D87817" blockId="31.[151,1437,367,1617]" pageId="31" pageNumber="182">
|
||
Also there exists a steady difference in the abdominal black pattern: the males of
|
||
<taxonomicName id="94A64D74910EF101A32FFCC17E91784E" baseAuthorityName="Morton" baseAuthorityYear="1924" box="[1131,1268,1519,1545]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="syriacum">
|
||
<emphasis id="61D2EAE5910EF101A32FFCC17E91784E" box="[1131,1268,1519,1545]" italics="true" pageId="31" pageNumber="182">C. syriacum</emphasis>
|
||
</taxonomicName>
|
||
always have a broad dorsal black bar on the hind part of S9 (
|
||
<figureCitation id="CB9D2A72910EF101A5C9FF3D78BA786B" box="[653,735,1555,1581]" captionStart="FIGURE 16" captionStartId="32.[151,250,1220,1244]" captionTargetBox="[164,1423,181,1196]" captionTargetId="figure-24@32.[164,1423,181,1196]" captionTargetPageId="32" captionText="FIGURE 16. Morphological details of male specimens of Coenagrion syriacum kept in the RMNH collection, courtesy by Charlotte Hartong: a–c—“Turkey, Adana / River Ceyhan, Ca. 20 km NW of / Osmaniye (37SBB42) River and / pools. / 11 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH.INS.1468579); d–f—“Turkey, Adana / Aydinlar, 15 km W of Kadirli / (37SBB33) Well vegetated, slowly flowing / channel. / 10 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH. INS.1468580); a, d—end of abbdomen in lateral view; b, e—ditto, dorsal view; a, d—ditto, caudal view." figureDoi="http://doi.org/10.5281/zenodo.12189942" httpUri="https://zenodo.org/record/12189942/files/figure.png" pageId="31" pageNumber="182">Fig. 16</figureCitation>
|
||
; iNaturalist 2023) while the males of
|
||
<taxonomicName id="94A64D74910EF101A33FFF3D7F07786B" authorityName="Dumont & Heidari" authorityYear="1996" box="[1147,1378,1555,1581]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="australocaspicum">
|
||
<emphasis id="61D2EAE5910EF101A33FFF3D7F07786B" box="[1147,1378,1555,1581]" italics="true" pageId="31" pageNumber="182">C. australocaspicum</emphasis>
|
||
</taxonomicName>
|
||
have S9 entirely blue, with at most a pair of black dots (
|
||
<figureCitation id="CB9D2A72910EF101A58AFF19795C7817" box="[718,825,1591,1617]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="31" pageNumber="182">Fig. 2a–c</figureCitation>
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,
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<figureCitation id="CB9D2A72910EF101A401FF1979D57817" box="[837,944,1591,1617]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="31" pageNumber="182">6a–c, j–k</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |