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<document id="8B81116A6A16262A9D0D1B4BAC09779E" ID-CLB-Dataset="298715" ID-DOI="10.11646/zootaxa.5471.2.1" ID-GBIF-Dataset="3a469638-dc21-45a3-80f6-cb307fbf5941" ID-ISSN="1175-5326" ID-Zenodo-Dep="12189904" ID-ZooBank="BAA047F4-2D65-4E52-8798-D426E3B26DA1" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1718892848818" checkinUser="plazi" docAuthor="Kosterin, Oleg E., Onishko, Vladimir V., Ilyina, Elena V., Chepurnov, Grigory Yu. &amp; Blinov, Alexander G." docDate="2024" docId="DB0F87E19107F108A7D3F84A7F557999" docLanguage="en" docName="zootaxa.5471.2.1.pdf" docOrigin="Zootaxa 5471 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5471.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coenagrion ornatum" docType="treatment" docVersion="5" lastPageNumber="173" masterDocId="2736FF999111F11EA744F92E7A657E46" masterDocTitle="The genus Coenagrion Kirby, 1890 (Odonata: Coenagrionidae) in the Russian part of the Caucasus" masterLastPageNumber="190" masterPageNumber="151" pageNumber="173" updateTime="1718988639371" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="48A930681C981FCF470F02C77213F7F5">The genus Coenagrion Kirby, 1890 (Odonata: Coenagrionidae) in the Russian part of the Caucasus</mods:title>
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<mods:namePart id="1D471A74BA09FF76100C1B831A3AA45C">Kosterin, Oleg E.</mods:namePart>
<mods:nameIdentifier id="CF0252B587B45645409C0A60641B3F1B" type="ORCID">0000-0001-5955-4057</mods:nameIdentifier>
<mods:affiliation id="700D6A2DA04602819E93A54E56FCA88B">Institute of Cytology &amp; Genetics SB RAS, Acad. Lavrentyev ave. 10, Novosibirsk, 630090, Russia.</mods:affiliation>
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<mods:namePart id="FB858DA737726AF696D047516EA9B0AC">Onishko, Vladimir V.</mods:namePart>
<mods:nameIdentifier id="259770E15784937EDCE27A3B01A1A24D" type="ORCID">0000-0002-6469-6778</mods:nameIdentifier>
<mods:affiliation id="3820AF309206E553224527210F64E826">Moscow Zoo, Department of Herpetology, Bolshaya Gruzinskaya Str. 1, Moscow 123242 Russia.</mods:affiliation>
<mods:nameIdentifier id="0ACC8C2D3DE7B9805CD0BA0910360973" type="email">wervolf999@yandex.ru</mods:nameIdentifier>
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<mods:namePart id="0A2C18CD7F2E59B4C21A345085ADE1DD">Ilyina, Elena V.</mods:namePart>
<mods:nameIdentifier id="5D4E9BAAF0A5786CE89C9A74D11CC6C3" type="ORCID">0000-0002-5426-4015</mods:nameIdentifier>
<mods:affiliation id="5AB8AB220006BFC6FAB2BA912AEBD353">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, Dagestanskiy State Nature Reserve, Gagarina Street 120, Makhachkala, Republic of Dagestan, Russia.</mods:affiliation>
<mods:nameIdentifier id="463B7960655DC91570BA7983067BA9D7" type="email">carabus@list.ru</mods:nameIdentifier>
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<mods:namePart id="0896BC46A03B7CD47651F91A7DFC9987">Chepurnov, Grigory Yu.</mods:namePart>
<mods:nameIdentifier id="EAD54EBA207626CC92D0A2E81E9669B6" type="ORCID">0009-0000-8268-0289</mods:nameIdentifier>
<mods:affiliation id="FDA70887CA9D2FA796973F115C178553">Institute of Cytology &amp; Genetics SB RAS, Acad. Lavrentyev ave. 10, Novosibirsk, 630090, Russia. &amp; Institute of Molecular and Cellular Biology SB RAS, Acad. Lavrentyev ave. 8 / 2, Novosibirsk, 630090, Russia.</mods:affiliation>
<mods:nameIdentifier id="36F37FA46C158D00F12C0F17772B18E6" type="email">grigory.tchepurnov@yandex.ru</mods:nameIdentifier>
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<mods:namePart id="7AFD9E42503D8FE2218585B2C5163651">Blinov, Alexander G.</mods:namePart>
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<mods:affiliation id="F460C40FFE1A96B5184CF010B7402C2F">Institute of Cytology &amp; Genetics SB RAS, Acad. Lavrentyev ave. 10, Novosibirsk, 630090, Russia. &amp; Institute of Molecular and Cellular Biology SB RAS, Acad. Lavrentyev ave. 8 / 2, Novosibirsk, 630090, Russia.</mods:affiliation>
<mods:nameIdentifier id="2F248A4744259B862C6A145E5348FE43" type="email">blinov@bionet.nsc.ru</mods:nameIdentifier>
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<treatment id="DB0F87E19107F108A7D3F84A7F557999" ID-DOI="http://doi.org/10.5281/zenodo.12190725" ID-GBIF-Taxon="233309485" ID-Zenodo-Dep="12190725" LSID="urn:lsid:plazi:treatment:DB0F87E19107F108A7D3F84A7F557999" httpUri="http://treatment.plazi.org/id/DB0F87E19107F108A7D3F84A7F557999" lastPageNumber="173" pageId="22" pageNumber="173">
<subSubSection id="1BBC657C9107F108A7D3F84A78327F38" box="[151,599,356,382]" pageId="22" pageNumber="173" type="nomenclature">
<paragraph id="531936F79107F108A7D3F84A78327F38" blockId="22.[151,599,356,418]" box="[151,599,356,382]" pageId="22" pageNumber="173">
<heading id="0851819B9107F108A7D3F84A78327F38" bold="true" box="[151,599,356,382]" fontSize="11" level="1" pageId="22" pageNumber="173" reason="1">
<emphasis id="61D2EAE59107F108A7D3F84A78327F38" bold="true" box="[151,599,356,382]" pageId="22" pageNumber="173">
5.
<taxonomicName id="94A64D749107F108A7F1F84A78327F38" ID-CoL="5ZJR2" authority="(Selys, 1850)" baseAuthorityName="Selys" baseAuthorityYear="1850" box="[181,599,356,382]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="ornatum">
<emphasis id="61D2EAE59107F108A7F1F84A7BD67F38" bold="true" box="[181,435,356,382]" italics="true" pageId="22" pageNumber="173">Coenagrion ornatum</emphasis>
(
<bibRefCitation id="37374B069107F108A687F84A782A7F38" author="Selys Lonchamps, E." box="[451,591,356,382]" pageId="22" pageNumber="173" refId="ref29427" refString="Selys Lonchamps, E. (de) (1850) Revue des Odonates ou Libellules d'Europe. C. Murquardt, Bruxelles et Leipzig &amp; Roret, Paris, 408 pp." type="book" year="1850">Selys, 1850</bibRefCitation>
)
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</paragraph>
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<paragraph id="531936F79107F108A7D3F8A67B597FE4" blockId="22.[151,599,356,418]" box="[151,316,392,418]" pageId="22" pageNumber="173">
<figureCitation id="CB9D2A729107F108A7D3F8A67A897FE4" box="[151,236,392,418]" captionStart="FIGURE 10" captionStartId="22.[151,250,1641,1665]" captionTargetBox="[163,1425,1352,1618]" captionTargetId="figure-399@22.[151,1436,1342,1618]" captionTargetPageId="22" captionText="FIGURE 10. Male appendages of Coenagrion ornatum specimens from Dagestan in dorsal view: a–b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; c–d—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, a swampy Dzharag-Khyan Spring, 28.05.2023. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189930" httpUri="https://zenodo.org/record/12189930/files/figure.png" pageId="22" pageNumber="173">Figs 10</figureCitation>
,
<figureCitation id="CB9D2A729107F108A7BCF8A67B597FE4" box="[248,316,392,418]" captionStart="FIGURE 11" captionStartId="24.[151,250,1356,1380]" captionTargetBox="[213,1375,181,1333]" captionTargetId="figure-24@24.[213,1375,181,1333]" captionTargetPageId="24" captionText="FIGURE 11. Coenagrion ornatum (a–d) and C. scitulum (e–k) in different populations in the Russian part of the Caucasus in nature; a–c, e–f—males; d—copula; g–j—females; k—tandems; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (180159191); b—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag-Khyan Spring, Sirtych, 28.05.2023 (178695635); c—Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021 (178693662); d—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (98025103); e, j—Krasnodarskiy Kray, Anapa Municipality, E suburbs of Supsekh settlement, Lake Zmeinoe env., 24.06.2019 (108802758, 108804470, respectively); f—Karachay-Cherkes Rpeublic, Malokarachaevsk District, Uchkeken village E environs, 9.07.2019 (109974039, Fig. 11f); g, k—Dagestan, Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (176987626, 182772933, respectively); h—Derbent District, 3.5 km NE of Belidzhi village, thickets at a swamp, 25.05.2023 (186590343); i—Dagestan, Buynaksk District, Nizhnee Kazanishche village environs, a water reservoir, 29.05.2023 (176987387); a, g–i, k—photo by VVO, b–f, j—by OEK. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189932" httpUri="https://zenodo.org/record/12189932/files/figure.png" pageId="22" pageNumber="173">11a–d</figureCitation>
</paragraph>
<paragraph id="531936F79107F108A7D3F8FE79197C74" blockId="22.[151,1437,464,1319]" pageId="22" pageNumber="173">
<emphasis id="61D2EAE59107F108A7D3F8FE7B357FAC" bold="true" box="[151,336,464,490]" pageId="22" pageNumber="173">Reported from:</emphasis>
Krasnodarskiy Kray, environs of Gryaznukhinskaya Stanitsa (sub.
<taxonomicName id="94A64D749107F108A375F8FE7F437FAC" authority="Selys" authorityName="Selys" authorityYear="1850" box="[1073,1318,464,490]" class="Insecta" family="Coenagrionidae" genus="Agrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="ornatum">
<emphasis id="61D2EAE59107F108A375F8FE7E867FAC" box="[1073,1251,464,490]" italics="true" pageId="22" pageNumber="173">Agrion ornatum</emphasis>
Selys
</taxonomicName>
) (
<bibRefCitation id="37374B069107F108A27EF8FE7AB67C48" author="Bartenef, A. N." pageId="22" pageNumber="173" pagination="1 - 138" refId="ref27235" refString="Bartenef, A. N. (1930) Materialy k poznaniyu Zapadnogo Kavkaza v odonatologicheskom otnoshenii [Materials for the knowledge on West Caucasus in odonatological respect]. Trudy Zapadnokavkazskoi assotsiatsii nauchno-issledovatel'skikh institutov [Proceedings of the West Caucasian Association of scientific Research Institutes], 72 (14), 1 - 138. [in Russian, with German summary but without German title]" type="journal article" year="1930">Bartenef 1930</bibRefCitation>
), the Pslukh River in Sochi Municipality (
<bibRefCitation id="37374B069107F108A58EF8DA79E37C48" author="Skvortsov, V. E." box="[714,902,500,526]" pageId="22" pageNumber="173" refId="ref29513" refString="Skvortsov, V. E. (2010) The dragonflies of Eastern Europe and Caucasus: An illustrated guide. KMK Scientific Press Ltd. Moscow [bilingual Russian and English], 623 pp." type="book" year="2010">Skvortsov 2010</bibRefCitation>
); Kellermesskaya and Khanskaya Stanitsas in
<collectingRegion id="9162F8159107F108A7D3FB367A8B7C74" box="[151,238,536,562]" country="Russia" name="Adygeya" pageId="22" pageNumber="173">Adygea</collectingRegion>
Republic (
<bibRefCitation id="37374B069107F108A62DFB3678687C74" author="Bartenef, A. N." box="[361,525,536,562]" pageId="22" pageNumber="173" pagination="1 - 138" refId="ref27235" refString="Bartenef, A. N. (1930) Materialy k poznaniyu Zapadnogo Kavkaza v odonatologicheskom otnoshenii [Materials for the knowledge on West Caucasus in odonatological respect]. Trudy Zapadnokavkazskoi assotsiatsii nauchno-issledovatel'skikh institutov [Proceedings of the West Caucasian Association of scientific Research Institutes], 72 (14), 1 - 138. [in Russian, with German summary but without German title]" type="journal article" year="1930">Bartenef 1930</bibRefCitation>
;
<bibRefCitation id="37374B069107F108A55DFB3679157C74" author="Shapovalov, M. I. &amp; Korotkov, E. A." box="[537,880,536,562]" pageId="22" pageNumber="173" pagination="341 - 349" refId="ref29460" refString="Shapovalov, M. I. &amp; Korotkov, E. A. (2019) Update of the Odonata fauna of Georgia, southern Caucasus ecoregion. Russian Entomological Journal, 28 (4), 341 - 349. https: // doi. org / 10.15298 / rusentj. 28.4.01" type="journal article" year="2019">Shapovalov &amp; Korotkov 2019</bibRefCitation>
).
</paragraph>
<paragraph id="531936F79107F108A783FB1278B37B60" blockId="22.[151,1437,464,1319]" pageId="22" pageNumber="173">
<emphasis id="61D2EAE59107F108A783FB127B297C10" bold="true" box="[199,332,572,598]" pageId="22" pageNumber="173">Specimens.</emphasis>
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<emphasis id="61D2EAE59107F108A61DFB1278D67C10" bold="true" box="[345,691,572,598]" pageId="22" pageNumber="173">
<collectingRegion id="9162F8159107F108A61DFB1278D67C10" box="[345,691,572,598]" name="Karachay-Cherkes Republic" pageId="22" pageNumber="173">Karachay-Cherkes Republic</collectingRegion>
</emphasis>
:
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(37680843, 37680856),
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,
<collectingCountry id="2BB176679107F108A36EFB127E1D7C10" box="[1066,1144,572,598]" name="Russia" pageId="22" pageNumber="173">Russia</collectingCountry>
, N
<collectingCounty id="BA784E7B9107F108A3EDFB127F737C10" box="[1193,1302,572,598]" pageId="22" pageNumber="173">Caucasus</collectingCounty>
,
<collectingMunicipality id="B37DAC8D9107F108A262FB127A8B7C3C" pageId="22" pageNumber="173">Zelenchuk District</collectingMunicipality>
,
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W environs, S banks of a large water reserve at the
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, 43.793–796 N, 41.536–540 E,
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a.s.l.
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,
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<collectingDate id="375CE9DF9107F108A532FBAA788C7CD8" box="[630,745,644,670]" pageId="22" pageNumber="173" value="2019-07-03">3.07.2019</collectingDate>
</date>
</materialsCitation>
.
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<collectingRegion id="9162F8159107F108A5BCFBAA7E617CD8" box="[760,1028,644,670]" country="Russia" name="Dagestan" pageId="22" pageNumber="173">
<emphasis id="61D2EAE59107F108A5BCFBAA79E07CD8" bold="true" box="[760,901,644,670]" pageId="22" pageNumber="173">Republic of</emphasis>
<emphasis id="61D2EAE59107F108A4D5FBAA7E6C7CD8" bold="true" box="[913,1033,644,670]" pageId="22" pageNumber="173">Dagestan:</emphasis>
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<specimenCount id="45A0FD7E9107F108A350FBAB7E4E7CD9" box="[1044,1067,645,671]" count="1" pageId="22" pageNumber="173" type="male">♂</specimenCount>
,
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(178696749, 180155503, 180155777, 1180155989, 180156091, 80156210, 180156435, 180157387, 180159191 (
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), 180160258, 180160572, 180160723, 180160831, 180160934),
<collectingCounty id="BA784E7B9107F108A589FBE279FF7CA0" box="[717,922,716,742]" pageId="22" pageNumber="173">Buynaksk District</collectingCounty>
,
<collectingMunicipality id="B37DAC8D9107F108A4E0FBE27E717CA0" box="[932,1044,716,742]" pageId="22" pageNumber="173">Buynaksk</collectingMunicipality>
S environs,
<location id="5679602C9107F108A3D9FBE27A867D4C" LSID="urn:lsid:plazi:treatment:DB0F87E19107F108A7D3F84A7F557999:5679602C9107F108A3D9FBE27A867D4C" country="Russia" county="Buynaksk District" latitude="47.1157" longLatPrecision="6" longitude="47.1169" municipality="Buynaksk" name="Verkhnee Kazanishche village" pageId="22" pageNumber="173" stateProvince="Dagestan">Verkhnee Kazanishche village</location>
environs, a swampy, reedy brook on a gentle left slope of the
<location id="5679602C9107F108A4FFFBDE7EAF7D4C" LSID="urn:lsid:plazi:treatment:DB0F87E19107F108A7D3F84A7F557999:5679602C9107F108A4FFFBDE7EAF7D4C" box="[955,1226,752,778]" country="Russia" county="Buynaksk District" latitude="47.1157" longLatPrecision="6" longitude="47.1169" municipality="Buynaksk" name="Akpeozen' River valley" pageId="22" pageNumber="173" stateProvince="Dagestan">Akpeozen’ River valley</location>
,
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–42.7992 N, 47.1157–
<geoCoordinate id="369250309107F108A661FA3A7BC17D69" box="[293,420,788,815]" degrees="47.1169" direction="east" orientation="longitude" pageId="22" pageNumber="173" precision="5" value="47.1169">47.1169 E</geoCoordinate>
,
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a.s.l.
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,
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<collectingDate id="375CE9DF9107F108A5CCFA3A796F7D69" box="[648,778,788,815]" pageId="22" pageNumber="173" value="2023-05-29">29.05.2023</collectingDate>
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;
<materialsCitation id="E3CE3CAA9107F108A45DFA3A79657DA5" ID-GBIF-Occurrence="4897992333" collectingDate="2023-05-28" country="Russia" county="Tabasaran District" elevation="104" latitude="48.2396" location="Dzharag-Khyan Spring" longLatPrecision="6" longitude="48.2411" municipality="Rubas River valley" pageId="22" pageNumber="173" specimenCount="6" specimenCount-male="6" stateProvince="Dagestan">
<specimenCount id="45A0FD7E9107F108A45DFA3A79397D69" box="[793,860,788,815]" count="6" pageId="22" pageNumber="173" type="male">6 ♂♂</specimenCount>
(
<figureCitation id="CB9D2A729107F108A42BFA3A79887D68" box="[879,1005,788,814]" captionStart="FIGURE 10" captionStartId="22.[151,250,1641,1665]" captionTargetBox="[163,1425,1352,1618]" captionTargetId="figure-399@22.[151,1436,1342,1618]" captionTargetPageId="22" captionText="FIGURE 10. Male appendages of Coenagrion ornatum specimens from Dagestan in dorsal view: a–b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; c–d—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, a swampy Dzharag-Khyan Spring, 28.05.2023. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189930" httpUri="https://zenodo.org/record/12189930/files/figure.png" pageId="22" pageNumber="173">Fig. 10d–e</figureCitation>
) (178695634, 178695635 (
<figureCitation id="CB9D2A729107F108A26FFA3A7FEB7D68" box="[1323,1422,788,814]" captionStart="FIGURE 11" captionStartId="24.[151,250,1356,1380]" captionTargetBox="[213,1375,181,1333]" captionTargetId="figure-24@24.[213,1375,181,1333]" captionTargetPageId="24" captionText="FIGURE 11. Coenagrion ornatum (a–d) and C. scitulum (e–k) in different populations in the Russian part of the Caucasus in nature; a–c, e–f—males; d—copula; g–j—females; k—tandems; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (180159191); b—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag-Khyan Spring, Sirtych, 28.05.2023 (178695635); c—Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021 (178693662); d—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (98025103); e, j—Krasnodarskiy Kray, Anapa Municipality, E suburbs of Supsekh settlement, Lake Zmeinoe env., 24.06.2019 (108802758, 108804470, respectively); f—Karachay-Cherkes Rpeublic, Malokarachaevsk District, Uchkeken village E environs, 9.07.2019 (109974039, Fig. 11f); g, k—Dagestan, Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (176987626, 182772933, respectively); h—Derbent District, 3.5 km NE of Belidzhi village, thickets at a swamp, 25.05.2023 (186590343); i—Dagestan, Buynaksk District, Nizhnee Kazanishche village environs, a water reservoir, 29.05.2023 (176987387); a, g–i, k—photo by VVO, b–f, j—by OEK. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189932" httpUri="https://zenodo.org/record/12189932/files/figure.png" pageId="22" pageNumber="173">Fig. 11b</figureCitation>
), 178695636, 180231099, 180231185, 180643335, 180643481, 180643723, 180643834, 180643926, 180644391, 180644477, 180644677, 180644769, 180644838, 180644922, 180645019, 180645088, 180950128, 180951966, 180952452, 180952577, 180952701, 180952786, 180952885, 180953036, 180953178, 180953270),
<collectingCounty id="BA784E7B9107F108A26FFAAE7A8B7DF8" pageId="22" pageNumber="173">Tabasaran District</collectingCounty>
, the
<collectingMunicipality id="B37DAC8D9107F108A666FA8A7B9D7DF8" box="[290,504,932,958]" pageId="22" pageNumber="173">Rubas River valley</collectingMunicipality>
<locationDeviation id="42AB2FD59107F108A6BBFA8A79217DF8" box="[511,836,932,959]" location="Sirtych village" pageId="22" pageNumber="173">
<quantity id="945E9B129107F108A6BBFA8A78287DF8" box="[511,589,932,959]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="22" pageNumber="173" unit="km" value="2.5">2.5 km</quantity>
NE of Sirtych village
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, a swampy
<location id="5679602C9107F108A482FA8A7EA87DF8" LSID="urn:lsid:plazi:treatment:DB0F87E19107F108A7D3F84A7F557999:5679602C9107F108A482FA8A7EA87DF8" box="[966,1229,932,958]" country="Russia" county="Tabasaran District" latitude="48.2396" longLatPrecision="6" longitude="48.2411" municipality="Rubas River valley" name="Dzharag-Khyan Spring" pageId="22" pageNumber="173" stateProvince="Dagestan">Dzharag-Khyan Spring</location>
,
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–41.8527 N, 48.2396–
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,
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<quantity id="945E9B129107F108A6F4FAE678547DA5" box="[432,561,968,995]" metricMagnitude="2" metricUnit="m" metricValue="1.04" metricValueMax="1.05" metricValueMin="1.03" pageId="22" pageNumber="173" unit="m" value="104.0" valueMax="105.0" valueMin="103.0">103–105 m</quantity>
a.s.l.
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,
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<collectingDate id="375CE9DF9107F108A53AFAE679657DA5" box="[638,768,968,995]" pageId="22" pageNumber="173" value="2023-05-28">28.05.2023</collectingDate>
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;
<materialsCitation id="E3CE3CAA9107F108A448FAE67B4C7A6C" ID-GBIF-Occurrence="4897992380" collectingDate="2022-06-04" collectorName="E. Ilyina" country="Russia" county="Magaramkent District" elevation="22" latitude="48.547" location="Bil' - Bil' Kazmalyar village" longLatPrecision="66" longitude="48.548" municipality="Samur Delta Forest" pageId="22" pageNumber="173" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Dagestan">
<specimenCount id="45A0FD7E9107F108A448FAE679357DA5" box="[780,848,968,995]" count="4" pageId="22" pageNumber="173" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="45A0FD7E9107F108A41AFAE679C57DA5" box="[862,928,968,995]" count="3" pageId="22" pageNumber="173" type="female">3 ♀♀</specimenCount>
,
<collectingCounty id="BA784E7B9107F108A4EBFAE67ECF7DA4" box="[943,1194,968,994]" pageId="22" pageNumber="173">Magaramkent District</collectingCounty>
,
<collectingMunicipality id="B37DAC8D9107F108A3F3FAE67FFD7DA5" box="[1207,1432,968,995]" pageId="22" pageNumber="173">Samur Delta Forest</collectingMunicipality>
,
<location id="5679602C9107F108A7D3FAC27BAA7A40" LSID="urn:lsid:plazi:treatment:DB0F87E19107F108A7D3F84A7F557999:5679602C9107F108A7D3FAC27BAA7A40" box="[151,463,1004,1030]" country="Russia" county="Magaramkent District" latitude="48.547" longLatPrecision="66" longitude="48.548" municipality="Samur Delta Forest" name="Bil' - Bil' Kazmalyar village" pageId="22" pageNumber="173" stateProvince="Dagestan">Bil’-Bil’ Kazmalyar village</location>
, a brook and a pond,
<geoCoordinate id="369250309107F108A581FAC279777A40" box="[709,786,1004,1030]" degrees="41.818" direction="north" orientation="latitude" pageId="22" pageNumber="173" precision="55" value="41.818">41.818</geoCoordinate>
–41.819 N, 48.547–
<geoCoordinate id="369250309107F108A4BEFAC27E0C7A41" box="[1018,1129,1004,1031]" degrees="48.548" direction="east" orientation="longitude" pageId="22" pageNumber="173" precision="55" value="48.548">48.548 E</geoCoordinate>
,
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<quantity id="945E9B129107F108A332FAC27EBF7A41" box="[1142,1242,1004,1031]" metricMagnitude="1" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="22" pageNumber="173" unit="m" value="21.5" valueMax="22.0" valueMin="21.0">21–22 m</quantity>
a.s.l.
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,
<date id="271810379107F108A261FAC27FFD7A40" box="[1317,1432,1004,1030]" pageId="22" pageNumber="173" value="2022-06-04">
<collectingDate id="375CE9DF9107F108A261FAC27FFD7A40" box="[1317,1432,1004,1030]" pageId="22" pageNumber="173" value="2022-06-04">4.06.2022</collectingDate>
</date>
,
<collectorName id="FE5353219107F108A7D3FD3F7A9F7A6C" box="[151,250,1040,1067]" pageId="22" pageNumber="173">E. Ilyina</collectorName>
leg
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;
<specimenCount id="45A0FD7E9107F108A67CFD3E7B1E7A6D" box="[312,379,1040,1067]" count="2" pageId="22" pageNumber="173" type="male">2 ♂♂</specimenCount>
, the same place,
<date id="271810379107F108A501FD3E78DD7A6D" box="[581,696,1040,1067]" pageId="22" pageNumber="173" value="2023-06-05">5.06.2023</date>
, E. Ilyina leg.;
<specimenCount id="45A0FD7E9107F108A429FD3E79F97A6D" box="[877,924,1040,1067]" count="2" pageId="22" pageNumber="173" type="male">2 ♂</specimenCount>
, the same place,
<date id="271810379107F108A322FD3E7EBD7A6D" box="[1126,1240,1040,1067]" pageId="22" pageNumber="173" value="2023-05-05">5.05.2023</date>
, E. Ilyina leg.;
<specimenCount id="45A0FD7E9107F108A2CAFD3E7AA47A09" count="3" pageId="22" pageNumber="173" type="male">3 ♂♂</specimenCount>
(178693662 (
<figureCitation id="CB9D2A729107F108A61BFD1A7BD87A08" box="[351,445,1076,1102]" captionStart="FIGURE 11" captionStartId="24.[151,250,1356,1380]" captionTargetBox="[213,1375,181,1333]" captionTargetId="figure-24@24.[213,1375,181,1333]" captionTargetPageId="24" captionText="FIGURE 11. Coenagrion ornatum (a–d) and C. scitulum (e–k) in different populations in the Russian part of the Caucasus in nature; a–c, e–f—males; d—copula; g–j—females; k—tandems; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (180159191); b—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag-Khyan Spring, Sirtych, 28.05.2023 (178695635); c—Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021 (178693662); d—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (98025103); e, j—Krasnodarskiy Kray, Anapa Municipality, E suburbs of Supsekh settlement, Lake Zmeinoe env., 24.06.2019 (108802758, 108804470, respectively); f—Karachay-Cherkes Rpeublic, Malokarachaevsk District, Uchkeken village E environs, 9.07.2019 (109974039, Fig. 11f); g, k—Dagestan, Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (176987626, 182772933, respectively); h—Derbent District, 3.5 km NE of Belidzhi village, thickets at a swamp, 25.05.2023 (186590343); i—Dagestan, Buynaksk District, Nizhnee Kazanishche village environs, a water reservoir, 29.05.2023 (176987387); a, g–i, k—photo by VVO, b–f, j—by OEK. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189932" httpUri="https://zenodo.org/record/12189932/files/figure.png" pageId="22" pageNumber="173">Fig. 11c</figureCitation>
), 178693663, 181380130, 181380245, 181380516, 181380589, 181380955, 181381118, 182680720, 182680856, 182681001, 182681109, 182681639, 182681821, 182682133, 182682287, 182683094, 182683374, 182683704, 182683883, 182683987, 182684102, 182771447, 182771534, 182771582), the same place,
<date id="271810379107F108A7A6FD8E7B017AFD" box="[226,356,1184,1211]" pageId="22" pageNumber="173" value="2023-05-27">27.05.2023</date>
;
<specimenCount id="45A0FD7E9107F108A635FD8E7BD77AFD" box="[369,434,1184,1211]" count="3" pageId="22" pageNumber="173" type="male">3 ♂♂</specimenCount>
(
<figureCitation id="CB9D2A729107F108A680FD8E78277AFC" box="[452,578,1184,1210]" captionStart="FIGURE 10" captionStartId="22.[151,250,1641,1665]" captionTargetBox="[163,1425,1352,1618]" captionTargetId="figure-399@22.[151,1436,1342,1618]" captionTargetPageId="22" captionText="FIGURE 10. Male appendages of Coenagrion ornatum specimens from Dagestan in dorsal view: a–b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; c–d—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, a swampy Dzharag-Khyan Spring, 28.05.2023. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189930" httpUri="https://zenodo.org/record/12189930/files/figure.png" pageId="22" pageNumber="173">Fig. 10a–b</figureCitation>
),
<materialsCitation id="E3CE3CAA9107F108A51CFD8E78B77B60" ID-GBIF-Occurrence="4897992381" collectingDate="2021-06-26" country="Russia" county="Rutul District" elevation="1370" latitude="47.358" location="Kufa" longLatPrecision="66" longitude="47.362" municipality="Samur River" pageId="22" pageNumber="173" specimenCount="4" specimenCount-female="4" stateProvince="Dagestan">
<specimenCount id="45A0FD7E9107F108A51CFD8E78FC7AFD" box="[600,665,1184,1211]" count="4" pageId="22" pageNumber="173" type="female">4 ♀♀</specimenCount>
(98025103 (
<figureCitation id="CB9D2A729107F108A469FD8E79EA7AFC" box="[813,911,1184,1210]" captionStart="FIGURE 11" captionStartId="24.[151,250,1356,1380]" captionTargetBox="[213,1375,181,1333]" captionTargetId="figure-24@24.[213,1375,181,1333]" captionTargetPageId="24" captionText="FIGURE 11. Coenagrion ornatum (a–d) and C. scitulum (e–k) in different populations in the Russian part of the Caucasus in nature; a–c, e–f—males; d—copula; g–j—females; k—tandems; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (180159191); b—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag-Khyan Spring, Sirtych, 28.05.2023 (178695635); c—Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021 (178693662); d—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (98025103); e, j—Krasnodarskiy Kray, Anapa Municipality, E suburbs of Supsekh settlement, Lake Zmeinoe env., 24.06.2019 (108802758, 108804470, respectively); f—Karachay-Cherkes Rpeublic, Malokarachaevsk District, Uchkeken village E environs, 9.07.2019 (109974039, Fig. 11f); g, k—Dagestan, Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (176987626, 182772933, respectively); h—Derbent District, 3.5 km NE of Belidzhi village, thickets at a swamp, 25.05.2023 (186590343); i—Dagestan, Buynaksk District, Nizhnee Kazanishche village environs, a water reservoir, 29.05.2023 (176987387); a, g–i, k—photo by VVO, b–f, j—by OEK. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189932" httpUri="https://zenodo.org/record/12189932/files/figure.png" pageId="22" pageNumber="173">Fig. 11d</figureCitation>
), 98116240, 98963192, 98963230, 98963408, 98963476, 98963562, 98963695, 98963732, 98964853, 98964859, 98964915, 98964939, 98964993, 98965076, 98968166, 98968201),
<collectingCounty id="BA784E7B9107F108A6DFFDC678527B44" box="[411,567,1256,1282]" pageId="22" pageNumber="173">Rutul District</collectingCounty>
, the
<collectingMunicipality id="B37DAC8D9107F108A52EFDC678927B44" box="[618,759,1256,1282]" pageId="22" pageNumber="173">Samur River</collectingMunicipality>
left arm between
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and
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villages,
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–41.570 N, 47.358–
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,
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a.s.l.
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,
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<collectingDate id="375CE9DF9107F108A514FC2278B77B60" box="[592,722,1292,1318]" pageId="22" pageNumber="173" value="2021-06-26">26.06.2021</collectingDate>
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.
</paragraph>
</subSubSection>
<caption id="07D9667F9107F108A7D3FF477995788F" ID-DOI="http://doi.org/10.5281/zenodo.12189930" ID-Zenodo-Dep="12189930" httpUri="https://zenodo.org/record/12189930/files/figure.png" pageId="22" pageNumber="173" startId="22.[151,250,1641,1665]" targetBox="[163,1425,1352,1618]" targetPageId="22" targetType="figure">
<paragraph id="531936F79107F108A7D3FF477995788F" blockId="22.[151,1436,1641,1738]" pageId="22" pageNumber="173">
<emphasis id="61D2EAE59107F108A7D3FF477B4678C4" bold="true" box="[151,291,1641,1666]" pageId="22" pageNumber="173">FIGURE 10.</emphasis>
Male appendages of
<taxonomicName id="94A64D749107F108A543FF4478BA78C7" baseAuthorityName="Selys" baseAuthorityYear="1850" box="[519,735,1642,1665]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="ornatum">
<emphasis id="61D2EAE59107F108A543FF4478BA78C7" box="[519,735,1642,1665]" italics="true" pageId="22" pageNumber="173">Coenagrion ornatum</emphasis>
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specimens from Dagestan in dorsal view: a–b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; c–d—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, a swampy Dzharag-Khyan Spring, 28.05.2023. Scale bar 0.5 mm.
</paragraph>
</caption>
<subSubSection id="1BBC657C9107F108A783FE3F7F557999" pageId="22" pageNumber="173" type="distribution">
<paragraph id="531936F79107F108A783FE3F79427916" blockId="22.[151,1437,1809,2016]" pageId="22" pageNumber="173">
<emphasis id="61D2EAE59107F108A783FE3F7B96796D" bold="true" box="[199,499,1809,1835]" pageId="22" pageNumber="173">Distribution and habitat.</emphasis>
Widespread but local in the Caucasus, although the populations found were rather large. The species invariably occurs at small slow streams.
</paragraph>
<paragraph id="531936F79107F108A783FE777F557999" blockId="22.[151,1437,1809,2016]" pageId="22" pageNumber="173">
<emphasis id="61D2EAE59107F108A783FE777B317935" bold="true" box="[199,340,1881,1907]" pageId="22" pageNumber="173">Characters.</emphasis>
The characters are unmistakable for this species (
<bibRefCitation id="37374B069107F108A4CCFE777E507935" author="Kalkman, V. J." box="[904,1077,1881,1907]" pageId="22" pageNumber="173" pagination="3 - 82" refId="ref28232" refString="Kalkman, V. J. (2006) Key to the dragonflies of Turkey including species known from Greece, Bulgaria, Lebanon, Syria, the Trans-Caucasus and Iran. Brachytron, 10 (1), 3 - 82." type="journal article" year="2006">Kalkman 2006</bibRefCitation>
;
<bibRefCitation id="37374B069107F108A304FE777E937935" author="Skvortsov, V. E." box="[1088,1270,1881,1907]" pageId="22" pageNumber="173" refId="ref29513" refString="Skvortsov, V. E. (2010) The dragonflies of Eastern Europe and Caucasus: An illustrated guide. KMK Scientific Press Ltd. Moscow [bilingual Russian and English], 623 pp." type="book" year="2010">Skvortsov 2010</bibRefCitation>
;
<bibRefCitation id="37374B069107F108A245FE777AB479D1" author="Dijkstra, K. - D. &amp; Schroter, A. &amp; Lewington, R." pageId="22" pageNumber="173" refId="ref27712" refString="Dijkstra, K. - D., Schroter, A. &amp; Lewington, R. (2020) Field Guide to the Dragonflies of Britain and Europe, including western Turkey and north-western Africa. 2 nd Edition. Bloomsbury Publishing, London, Oxford etc., 336 pp." type="book" year="2020">
Dijkstra
<emphasis id="61D2EAE59107F108A227FE747FF87935" box="[1379,1437,1881,1907]" italics="true" pageId="22" pageNumber="173">et al.</emphasis>
2020
</bibRefCitation>
) and deserve no attention beyond a particular problem with the position of the male cerci (see ‘Discussion’). In our male specimens, the cerci are more or less divergent to a variable extent (
<figureCitation id="CB9D2A729107F108A4B6FE8F7E2179FD" box="[1010,1092,1953,1979]" captionStart="FIGURE 10" captionStartId="22.[151,250,1641,1665]" captionTargetBox="[163,1425,1352,1618]" captionTargetId="figure-399@22.[151,1436,1342,1618]" captionTargetPageId="22" captionText="FIGURE 10. Male appendages of Coenagrion ornatum specimens from Dagestan in dorsal view: a–b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; c–d—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, a swampy Dzharag-Khyan Spring, 28.05.2023. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189930" httpUri="https://zenodo.org/record/12189930/files/figure.png" pageId="22" pageNumber="173">Fig. 10</figureCitation>
); in specimens from the Rubas River valley the divergence is on average less expressed, with the least divergent case shown in
<figureCitation id="CB9D2A729107F108A38FFEEB7F497999" box="[1227,1324,1989,2015]" captionStart="FIGURE 10" captionStartId="22.[151,250,1641,1665]" captionTargetBox="[163,1425,1352,1618]" captionTargetId="figure-399@22.[151,1436,1342,1618]" captionTargetPageId="22" captionText="FIGURE 10. Male appendages of Coenagrion ornatum specimens from Dagestan in dorsal view: a–b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; c–d—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, a swampy Dzharag-Khyan Spring, 28.05.2023. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189930" httpUri="https://zenodo.org/record/12189930/files/figure.png" pageId="22" pageNumber="173">Fig. 10d</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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