382 lines
81 KiB
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382 lines
81 KiB
XML
<document id="5C42D6A6561ED93C0FF5EB6DEB01A267" ID-CLB-Dataset="298715" ID-DOI="10.11646/zootaxa.5471.2.1" ID-GBIF-Dataset="3a469638-dc21-45a3-80f6-cb307fbf5941" ID-ISSN="1175-5326" ID-Zenodo-Dep="12189904" ID-ZooBank="BAA047F4-2D65-4E52-8798-D426E3B26DA1" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1718892848818" checkinUser="plazi" docAuthor="Kosterin, Oleg E., Onishko, Vladimir V., Ilyina, Elena V., Chepurnov, Grigory Yu. & Blinov, Alexander G." docDate="2024" docId="DB0F87E19105F108A7D3FBDE78C67F5B" docLanguage="en" docName="zootaxa.5471.2.1.pdf" docOrigin="Zootaxa 5471 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5471.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coenagrion ponticum" docType="treatment" docVersion="5" lastPageNumber="172" masterDocId="2736FF999111F11EA744F92E7A657E46" masterDocTitle="The genus Coenagrion Kirby, 1890 (Odonata: Coenagrionidae) in the Russian part of the Caucasus" masterLastPageNumber="190" masterPageNumber="151" pageNumber="171" updateTime="1718988639371" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E87200E0175D39CEC6CDB0B980275ACC">The genus Coenagrion Kirby, 1890 (Odonata: Coenagrionidae) in the Russian part of the Caucasus</mods:title>
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<mods:namePart id="1CB3A133F5A0A49AC6537AE364D69352">Kosterin, Oleg E.</mods:namePart>
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<mods:affiliation id="706D8CACB5F9FCEB4F23313480FBFBAD">Institute of Cytology & Genetics SB RAS, Acad. Lavrentyev ave. 10, Novosibirsk, 630090, Russia.</mods:affiliation>
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<mods:nameIdentifier id="ABFC241A4F667EBE6B698C2B453AAFAD" type="email">kosterin@bionet.nsc.ru</mods:nameIdentifier>
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<mods:namePart id="5A87C4756535A9E17B47A44C907749CA">Onishko, Vladimir V.</mods:namePart>
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<mods:affiliation id="BBED1710D7B4742B4A75BF40F65D6AD2">Moscow Zoo, Department of Herpetology, Bolshaya Gruzinskaya Str. 1, Moscow 123242 Russia.</mods:affiliation>
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<mods:nameIdentifier id="32783D275244398B1AD28101C2907F2E" type="email">wervolf999@yandex.ru</mods:nameIdentifier>
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<mods:namePart id="795D895A4AEF91314D594453B5BE035E">Ilyina, Elena V.</mods:namePart>
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<mods:affiliation id="5947E4460C596EC4E92C97BF1807C2F0">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, Dagestanskiy State Nature Reserve, Gagarina Street 120, Makhachkala, Republic of Dagestan, Russia.</mods:affiliation>
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<mods:nameIdentifier id="CF83BF1B07BC01FEA60F6767A5EC8693" type="email">carabus@list.ru</mods:nameIdentifier>
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<mods:namePart id="3A88FD0C45C7B1E3B81CFF6E247A0139">Chepurnov, Grigory Yu.</mods:namePart>
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<mods:affiliation id="64BB5AA4A2AF5B43E5331416BAE08C18">Institute of Cytology & Genetics SB RAS, Acad. Lavrentyev ave. 10, Novosibirsk, 630090, Russia. & Institute of Molecular and Cellular Biology SB RAS, Acad. Lavrentyev ave. 8 / 2, Novosibirsk, 630090, Russia.</mods:affiliation>
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<mods:nameIdentifier id="CA782F363478557E0B414EC0BEAC21D0" type="email">grigory.tchepurnov@yandex.ru</mods:nameIdentifier>
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<mods:namePart id="498144758AA3280D87752EA894290EAD">Blinov, Alexander G.</mods:namePart>
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<mods:affiliation id="8618E74D3CCACBDE4FC3445A11F9CB7A">Institute of Cytology & Genetics SB RAS, Acad. Lavrentyev ave. 10, Novosibirsk, 630090, Russia. & Institute of Molecular and Cellular Biology SB RAS, Acad. Lavrentyev ave. 8 / 2, Novosibirsk, 630090, Russia.</mods:affiliation>
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<mods:nameIdentifier id="9A9721E890CAFEA7152D16869BD88234" type="email">blinov@bionet.nsc.ru</mods:nameIdentifier>
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<mods:date id="8A050B8B8926037277B1D1F369A0BCAE">2024</mods:date>
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<treatment id="DB0F87E19105F108A7D3FBDE78C67F5B" ID-DOI="http://doi.org/10.5281/zenodo.12190723" ID-GBIF-Taxon="233309488" ID-Zenodo-Dep="12190723" LSID="urn:lsid:plazi:treatment:DB0F87E19105F108A7D3FBDE78C67F5B" httpUri="http://treatment.plazi.org/id/DB0F87E19105F108A7D3FBDE78C67F5B" lastPageId="22" lastPageNumber="172" pageId="20" pageNumber="171">
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<subSubSection id="1BBC657C9105F10AA7D3FBDE78EA7D4C" box="[151,655,752,778]" pageId="20" pageNumber="171" type="nomenclature">
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<paragraph id="531936F79105F10AA7D3FBDE78EA7D4C" blockId="20.[151,655,752,815]" box="[151,655,752,778]" pageId="20" pageNumber="171">
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4.
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<taxonomicName id="94A64D749105F10AA7F1FBDE78EA7D4C" ID-CoL="WTXX" authority="(Bartenef, 1929)" baseAuthorityName="Bartenef" baseAuthorityYear="1929" box="[181,655,752,778]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="ponticum">
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<emphasis id="61D2EAE59105F10AA7F1FBDE7BD87D4C" bold="true" box="[181,445,752,778]" italics="true" pageId="20" pageNumber="171">Coenagrion ponticum</emphasis>
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(
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<bibRefCitation id="37374B069105F10AA68AFBDE78E37D4C" author="Bartenef, A. N." box="[462,646,752,778]" pageId="20" pageNumber="171" pagination="54 - 68" refId="ref27207" refString="Bartenef, A. N. (1929) Neue Arten und Varietaten der Odonata des West Kaukasus. Zoologischer Anzeiger, 85, 54 - 68." type="journal article" year="1929">Bartenef, 1929</bibRefCitation>
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)
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<paragraph id="531936F79105F10AA7D3FA3A787B7D68" blockId="20.[151,655,752,815]" box="[151,542,788,815]" pageId="20" pageNumber="171">
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<figureCitation id="CB9D2A729105F10AA7D3FA3A7B6E7D68" box="[151,267,788,814]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="20" pageNumber="171">Figs 2l–m</figureCitation>
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,
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<figureCitation id="CB9D2A729105F10AA652FA3A7B377D68" box="[278,338,788,814]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="20" pageNumber="171">3l–m</figureCitation>
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,
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<figureCitation id="CB9D2A729105F10AA61AFA3A7BF27D68" box="[350,407,788,814]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="20" pageNumber="171">4c–d</figureCitation>
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,
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||
<figureCitation id="CB9D2A729105F10AA6E6FA3A7BBE7D69" box="[418,475,788,815]" captionStart="FIGURE 5" captionStartId="13.[151,250,1570,1594]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@13.[151,1436,181,1546]" captionTargetPageId="13" captionText="FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="20" pageNumber="171">5d–e</figureCitation>
|
||
,
|
||
<figureCitation id="CB9D2A729105F10AA6A2FA3A78607D68" box="[486,517,788,814]" captionStart="FIGURE 7" captionStartId="15.[151,250,1565,1589]" captionTargetBox="[447,1140,184,1541]" captionTargetId="figure-19@15.[447,1140,181,1541]" captionTargetPageId="15" captionText="FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="20" pageNumber="171">7h</figureCitation>
|
||
,
|
||
<figureCitation id="CB9D2A729105F10AA554FA3A787B7D68" box="[528,542,788,814]" captionStart="FIGURE 9" captionStartId="21.[151,250,1715,1739]" captionTargetBox="[151,1436,181,1691]" captionTargetId="figure-19@21.[151,1436,181,1691]" captionTargetPageId="21" captionText="FIGURE 9. Males (a–b) and female (c) of Coenagrion ponticum in nature by a pond in the Fanagoriyskoe village NW environs, Goryachiy Klyuch Municipality, Krasnodarskiy Kray, Russia, 27.04.2019 (108924871, 37680876, 37681071, respectively); a—photo by OEK, b, c—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189928" httpUri="https://zenodo.org/record/12189928/files/figure.png" pageId="20" pageNumber="171">9</figureCitation>
|
||
</paragraph>
|
||
<paragraph id="531936F79105F10AA7D3FA727B3C7DF8" blockId="20.[151,1437,860,2038]" pageId="20" pageNumber="171">
|
||
<emphasis id="61D2EAE59105F10AA7D3FA727B627D30" bold="true" box="[151,263,860,886]" pageId="20" pageNumber="171">Reported</emphasis>
|
||
(as
|
||
<taxonomicName id="94A64D749105F10AA670FA727B957D30" authority="Kosterin & Onishko & Ilyina & Chepurnov & Blinov, 2024" authorityName="Kosterin & Onishko & Ilyina & Chepurnov & Blinov" authorityYear="2024" box="[308,496,860,886]" class="Insecta" family="Coenagrionidae" genus="Agrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="ponticum" status="sp. nov.">
|
||
<emphasis id="61D2EAE59105F10AA670FA727B957D30" box="[308,496,860,886]" italics="true" pageId="20" pageNumber="171">Agrion ponticum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7AE1579E9105F10AA6B3FA7378577D30" box="[503,562,860,887]" pageId="20" pageNumber="171" rank="species">sp. n.</taxonomicNameLabel>
|
||
)
|
||
<emphasis id="61D2EAE59105F10AA505FA7278E07D30" bold="true" box="[577,645,860,886]" pageId="20" pageNumber="171">from:</emphasis>
|
||
just three locations in Krasnodarskiy Kray: the environs of Sochy (five localities), Tuapse and Adler (one locality in each), all by the species’ author, A.N. Bartenev (
|
||
<bibRefCitation id="37374B069105F10AA3E8FAAE7F347DDC" author="Bartenef, A. N." box="[1196,1361,896,922]" pageId="20" pageNumber="171" pagination="54 - 68" refId="ref27207" refString="Bartenef, A. N. (1929) Neue Arten und Varietaten der Odonata des West Kaukasus. Zoologischer Anzeiger, 85, 54 - 68." type="journal article" year="1929">Bartenef 1929</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="37374B069105F10AA218FAAE7FF27DDC" author="Bartenef, A. N." box="[1372,1431,896,922]" pageId="20" pageNumber="171" pagination="1 - 138" refId="ref27235" refString="Bartenef, A. N. (1930) Materialy k poznaniyu Zapadnogo Kavkaza v odonatologicheskom otnoshenii [Materials for the knowledge on West Caucasus in odonatological respect]. Trudy Zapadnokavkazskoi assotsiatsii nauchno-issledovatel'skikh institutov [Proceedings of the West Caucasian Association of scientific Research Institutes], 72 (14), 1 - 138. [in Russian, with German summary but without German title]" type="journal article" year="1930">1930</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="37374B069105F10AA7D3FA8A7B287DF8" author="Skvortsov, V. E." box="[151,333,932,958]" pageId="20" pageNumber="171" refId="ref29513" refString="Skvortsov, V. E. (2010) The dragonflies of Eastern Europe and Caucasus: An illustrated guide. KMK Scientific Press Ltd. Moscow [bilingual Russian and English], 623 pp." type="book" year="2010">Skvortsov 2010</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="531936F79105F10AA783FAE6795A7A34" blockId="20.[151,1437,860,2038]" pageId="20" pageNumber="171">
|
||
<emphasis id="61D2EAE59105F10AA783FAE6782C7DA4" bold="true" box="[199,585,968,994]" pageId="20" pageNumber="171">
|
||
Specimens.
|
||
<materialsCitation id="E3CE3CAA9105F10AA616FAE67EF97A6C" ID-GBIF-Occurrence="4897992363" collectingDate="2019-06-27" elevation="101" location="Goryachiy Klyuch Municipality" municipality="Fanagoriyskoe village" pageId="20" pageNumber="171" specimenCount="3" specimenCount-female="3">
|
||
Krasnodarskiy Kray:
|
||
<specimenCount id="45A0FD7E9105F10AA513FAE678C67DA5" box="[599,675,968,995]" count="22" pageId="20" pageNumber="171" type="male">22 ♂♂</specimenCount>
|
||
(
|
||
<figureCitation id="CB9D2A729105F10AA5F5FAE679407DA4" box="[689,805,968,994]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="20" pageNumber="171">Figs 2l–m</figureCitation>
|
||
,
|
||
<figureCitation id="CB9D2A729105F10AA46BFAE6790E7DA4" box="[815,875,968,994]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="20" pageNumber="171">3l–m</figureCitation>
|
||
,
|
||
<figureCitation id="CB9D2A729105F10AA432FAE679CA7DA4" box="[886,943,968,994]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="20" pageNumber="171">4c–d</figureCitation>
|
||
,
|
||
<figureCitation id="CB9D2A729105F10AA4FDFAE679947DA5" box="[953,1009,968,995]" captionStart="FIGURE 5" captionStartId="13.[151,250,1570,1594]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@13.[151,1436,181,1546]" captionTargetPageId="13" captionText="FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="20" pageNumber="171">5d–e</figureCitation>
|
||
) (108924869, 108924870, 108924871 (
|
||
<figureCitation id="CB9D2A729105F10AA7DBFAC27A9D7A40" box="[159,248,1004,1030]" captionStart="FIGURE 9" captionStartId="21.[151,250,1715,1739]" captionTargetBox="[151,1436,181,1691]" captionTargetId="figure-19@21.[151,1436,181,1691]" captionTargetPageId="21" captionText="FIGURE 9. Males (a–b) and female (c) of Coenagrion ponticum in nature by a pond in the Fanagoriyskoe village NW environs, Goryachiy Klyuch Municipality, Krasnodarskiy Kray, Russia, 27.04.2019 (108924871, 37680876, 37681071, respectively); a—photo by OEK, b, c—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189928" httpUri="https://zenodo.org/record/12189928/files/figure.png" pageId="20" pageNumber="171">Fig. 9a</figureCitation>
|
||
), 37680876 (
|
||
<figureCitation id="CB9D2A729105F10AA6DEFAC27B917A40" box="[410,500,1004,1030]" captionStart="FIGURE 9" captionStartId="21.[151,250,1715,1739]" captionTargetBox="[151,1436,181,1691]" captionTargetId="figure-19@21.[151,1436,181,1691]" captionTargetPageId="21" captionText="FIGURE 9. Males (a–b) and female (c) of Coenagrion ponticum in nature by a pond in the Fanagoriyskoe village NW environs, Goryachiy Klyuch Municipality, Krasnodarskiy Kray, Russia, 27.04.2019 (108924871, 37680876, 37681071, respectively); a—photo by OEK, b, c—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189928" httpUri="https://zenodo.org/record/12189928/files/figure.png" pageId="20" pageNumber="171">Fig. 9b</figureCitation>
|
||
), 37680891),
|
||
<specimenCount id="45A0FD7E9105F10AA5D9FAC278867A41" box="[669,739,1004,1031]" count="3" pageId="20" pageNumber="171" type="female">3 ♀♀</specimenCount>
|
||
(
|
||
<figureCitation id="CB9D2A729105F10AA5BEFAC279367A40" box="[762,851,1004,1030]" captionStart="FIGURE 7" captionStartId="15.[151,250,1565,1589]" captionTargetBox="[447,1140,184,1541]" captionTargetId="figure-19@15.[447,1140,181,1541]" captionTargetPageId="15" captionText="FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="20" pageNumber="171">Fig. 7h</figureCitation>
|
||
),
|
||
<location id="5679602C9105F10AA42BFAC27E837A40" LSID="urn:lsid:plazi:treatment:DB0F87E19105F108A7D3FBDE78C67F5B:5679602C9105F10AA42BFAC27E837A40" box="[879,1254,1004,1030]" municipality="Fanagoriyskoe village" name="Goryachiy Klyuch Municipality" pageId="20" pageNumber="171">Goryachiy Klyuch Municipality</location>
|
||
,
|
||
<collectingMunicipality id="B37DAC8D9105F10AA3B3FAC27A867A6C" pageId="20" pageNumber="171">Fanagoriyskoe village</collectingMunicipality>
|
||
NW environs, ponds, 44.515–516 N, 39.108–109 E,
|
||
<elevation id="D88BD1C49105F10AA40BFD3E7E697A6C" box="[847,1036,1040,1067]" metricMagnitude="2" metricUnit="m" metricValue="1.015" metricValueMax="1.02" metricValueMin="1.01" pageId="20" pageNumber="171" unit="m" value="101.5" valueMax="102.0" valueMin="101.0">
|
||
<quantity id="945E9B129105F10AA40BFD3E79AA7A6D" box="[847,975,1040,1067]" metricMagnitude="2" metricUnit="m" metricValue="1.015" metricValueMax="1.02" metricValueMin="1.01" pageId="20" pageNumber="171" unit="m" value="101.5" valueMax="102.0" valueMin="101.0">101–102 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="271810379105F10AA35EFD3E7EF97A6C" box="[1050,1180,1040,1066]" pageId="20" pageNumber="171" value="2019-06-27">
|
||
<collectingDate id="375CE9DF9105F10AA35EFD3E7EF97A6C" box="[1050,1180,1040,1066]" pageId="20" pageNumber="171" value="2019-06-27">27.06.2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
</emphasis>
|
||
;
|
||
<materialsCitation id="E3CE3CAA9105F10AA3ECFD3F795A7A34" ID-GBIF-Occurrence="4897992327" collectingDate="2019-06-27" collectionCode="V" collectorName="Onishko & O. Kosterin" elevation="155" latitude="39.055" location="Goryachiy Klyuch Municipality" longLatPrecision="70" longitude="-39.058" municipality="Chepsi River" pageId="20" pageNumber="171" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="45A0FD7E9105F10AA3ECFD3F7EDA7A6D" box="[1192,1215,1041,1067]" count="1" pageId="20" pageNumber="171" type="male">♂</specimenCount>
|
||
,
|
||
<location id="5679602C9105F10AA388FD3E7B4C7A08" LSID="urn:lsid:plazi:treatment:DB0F87E19105F108A7D3FBDE78C67F5B:5679602C9105F10AA388FD3E7B4C7A08" latitude="39.055" longLatPrecision="70" longitude="-39.058" municipality="Chepsi River" name="Goryachiy Klyuch Municipality" pageId="20" pageNumber="171">Goryachiy Klyuch Municipality</location>
|
||
, the
|
||
<collectingMunicipality id="B37DAC8D9105F10AA61FFD1A7B887A08" box="[347,493,1076,1102]" pageId="20" pageNumber="171">Chepsi River</collectingMunicipality>
|
||
banks
|
||
<locationDeviation id="42AB2FD59105F10AA57EFD1A798A7A08" box="[570,1007,1076,1103]" location="Fanagoriyskoe village" pageId="20" pageNumber="171">
|
||
<quantity id="945E9B129105F10AA57EFD1A78C67A08" box="[570,675,1076,1103]" metricMagnitude="3" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="20" pageNumber="171" unit="km" value="6.75" valueMax="7.0" valueMin="6.5">6.5–7 km</quantity>
|
||
SW of Fanagoriyskoe village
|
||
</locationDeviation>
|
||
,
|
||
<geoCoordinate id="369250309105F10AA4BDFD1A7E237A08" box="[1017,1094,1076,1102]" degrees="44.464" direction="north" orientation="latitude" pageId="20" pageNumber="171" precision="55" value="44.464">44.464</geoCoordinate>
|
||
–44.469 N, 39.055–
|
||
<geoCoordinate id="369250309105F10AA26EFD1A7FF27A09" box="[1322,1431,1076,1103]" degrees="39.058" direction="west" orientation="longitude" pageId="20" pageNumber="171" precision="55" value="-39.058">39.058 E</geoCoordinate>
|
||
,
|
||
<elevation id="D88BD1C49105F10AA7D3FD767B367A34" box="[151,339,1112,1139]" metricMagnitude="2" metricUnit="m" metricValue="1.55" metricValueMax="1.6199999999999999" metricValueMin="1.48" pageId="20" pageNumber="171" unit="m" value="155.0" valueMax="162.0" valueMin="148.0">
|
||
<quantity id="945E9B129105F10AA7D3FD767B737A35" box="[151,278,1112,1139]" metricMagnitude="2" metricUnit="m" metricValue="1.55" metricValueMax="1.6199999999999999" metricValueMin="1.48" pageId="20" pageNumber="171" unit="m" value="155.0" valueMax="162.0" valueMin="148.0">148–162 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="271810379105F10AA61BFD767B857A34" box="[351,480,1112,1138]" pageId="20" pageNumber="171" value="2019-06-27">
|
||
<collectingDate id="375CE9DF9105F10AA61BFD767B857A34" box="[351,480,1112,1138]" pageId="20" pageNumber="171" value="2019-06-27">27.06.2019</collectingDate>
|
||
</date>
|
||
,
|
||
<collectionCode id="35B7AE329105F10AA6AFFD777B9A7A35" box="[491,511,1113,1139]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="20" pageNumber="171" type="Museum">V</collectionCode>
|
||
.
|
||
<collectorName id="FE5353219105F10AA54DFD76780D7A34" box="[521,616,1112,1138]" pageId="20" pageNumber="171">Onishko</collectorName>
|
||
&
|
||
<collectorName id="FE5353219105F10AA5C8FD76796B7A34" box="[652,782,1112,1138]" pageId="20" pageNumber="171">O. Kosterin</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="531936F79105F10AA783FD5279497B28" blockId="20.[151,1437,860,2038]" pageId="20" pageNumber="171">
|
||
<emphasis id="61D2EAE59105F10AA783FD5279F37AD0" bold="true" box="[199,918,1148,1174]" pageId="20" pageNumber="171">Additional photographic registrations: Krasnodarskiy Kray</emphasis>
|
||
:
|
||
<materialsCitation id="E3CE3CAA9105F10AA4E2FD527B1D7A98" ID-GBIF-Occurrence="4897992346" collectingDate="2019-04-27" elevation="101" latitude="39.108" location="Goryachiy Klyuch Municipality" longLatPrecision="70" longitude="39.109" municipality="Fanagoriyskoe village" pageId="20" pageNumber="171" specimenCount="1" specimenCount-female="1">
|
||
1 teneral
|
||
<specimenCount id="45A0FD7E9105F10AA34BFD537E417AD1" box="[1039,1060,1149,1175]" count="1" pageId="20" pageNumber="171" type="female">♀</specimenCount>
|
||
(37681071 (
|
||
<figureCitation id="CB9D2A729105F10AA3F1FD527F637AD0" box="[1205,1286,1148,1174]" captionStart="FIGURE 9" captionStartId="21.[151,250,1715,1739]" captionTargetBox="[151,1436,181,1691]" captionTargetId="figure-19@21.[151,1436,181,1691]" captionTargetPageId="21" captionText="FIGURE 9. Males (a–b) and female (c) of Coenagrion ponticum in nature by a pond in the Fanagoriyskoe village NW environs, Goryachiy Klyuch Municipality, Krasnodarskiy Kray, Russia, 27.04.2019 (108924871, 37680876, 37681071, respectively); a—photo by OEK, b, c—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189928" httpUri="https://zenodo.org/record/12189928/files/figure.png" pageId="20" pageNumber="171">Fig. 9c</figureCitation>
|
||
)),
|
||
<location id="5679602C9105F10AA262FD527BE17AFC" LSID="urn:lsid:plazi:treatment:DB0F87E19105F108A7D3FBDE78C67F5B:5679602C9105F10AA262FD527BE17AFC" latitude="39.108" longLatPrecision="70" longitude="39.109" municipality="Fanagoriyskoe village" name="Goryachiy Klyuch Municipality" pageId="20" pageNumber="171">Goryachiy Klyuch Municipality</location>
|
||
,
|
||
<collectingMunicipality id="B37DAC8D9105F10AA6D4FD8E78EC7AFC" box="[400,649,1184,1210]" pageId="20" pageNumber="171">Fanagoriyskoe village</collectingMunicipality>
|
||
NW environs, ponds,
|
||
<geoCoordinate id="369250309105F10AA4CDFD8E79B07AFD" box="[905,981,1184,1211]" degrees="44.515" direction="north" orientation="latitude" pageId="20" pageNumber="171" precision="55" value="44.515">44.515</geoCoordinate>
|
||
–44.516 N, 39.108–
|
||
<geoCoordinate id="369250309105F10AA3FAFD8E7F487AFD" box="[1214,1325,1184,1211]" degrees="39.109" direction="east" orientation="longitude" pageId="20" pageNumber="171" precision="55" value="39.109">39.109 E</geoCoordinate>
|
||
,
|
||
<elevation id="D88BD1C49105F10AA27EFD8E7A8F7A98" metricMagnitude="2" metricUnit="m" metricValue="1.015" metricValueMax="1.02" metricValueMin="1.01" pageId="20" pageNumber="171" unit="m" value="101.5" valueMax="102.0" valueMin="101.0">
|
||
<quantity id="945E9B129105F10AA27EFD8E7AC87A99" metricMagnitude="2" metricUnit="m" metricValue="1.015" metricValueMax="1.02" metricValueMin="1.01" pageId="20" pageNumber="171" unit="m" value="101.5" valueMax="102.0" valueMin="101.0">101–102 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="271810379105F10AA7B2FDEA7B1D7A98" box="[246,376,1220,1246]" pageId="20" pageNumber="171" value="2019-04-27">
|
||
<collectingDate id="375CE9DF9105F10AA7B2FDEA7B1D7A98" box="[246,376,1220,1246]" pageId="20" pageNumber="171" value="2019-04-27">27.04.2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="E3CE3CAA9105F10AA6C7FDEA79227B60" ID-GBIF-Occurrence="4897992325" collectingDate="2024-04-08" elevation="55" latitude="39.125" location="Goryachiy Klyuch Town" longLatPrecision="70" longitude="39.129" pageId="20" pageNumber="171" specimenCount="1" specimenCount-female="1">
|
||
1 teneral
|
||
<specimenCount id="45A0FD7E9105F10AA6A8FDEB78647A99" box="[492,513,1221,1247]" count="1" pageId="20" pageNumber="171" type="male">♂</specimenCount>
|
||
(37681035), the same place,
|
||
<date id="271810379105F10AA408FDEA79AB7A98" box="[844,974,1220,1246]" pageId="20" pageNumber="171" value="2019-04-28">28.04.2019</date>
|
||
;
|
||
<specimenCount id="45A0FD7E9105F10AA49DFDEA7E7D7A99" box="[985,1048,1220,1247]" count="4" pageId="20" pageNumber="171" type="male">4 ♂♂</specimenCount>
|
||
(37680916, 37680932, 37680994, 37681023), the same place,
|
||
<date id="271810379105F10AA693FDC6783C7B44" box="[471,601,1256,1282]" pageId="20" pageNumber="171" value="2019-04-28">28.04.2019</date>
|
||
; 2 teneral
|
||
<specimenCount id="45A0FD7E9105F10AA594FDC7789F7B45" box="[720,762,1257,1283]" count="1" pageId="20" pageNumber="171" type="female">♀♀</specimenCount>
|
||
(207535828, 207535790),
|
||
<location id="5679602C9105F10AA374FDC67F2D7B45" LSID="urn:lsid:plazi:treatment:DB0F87E19105F108A7D3FBDE78C67F5B:5679602C9105F10AA374FDC67F2D7B45" box="[1072,1352,1256,1283]" latitude="39.125" longLatPrecision="70" longitude="39.129" name="Goryachiy Klyuch Town" pageId="20" pageNumber="171">Goryachiy Klyuch Town</location>
|
||
centre, a pond,
|
||
<geoCoordinate id="369250309105F10AA7B1FC227B0C7B61" box="[245,361,1292,1319]" degrees="44.638" direction="north" orientation="latitude" pageId="20" pageNumber="171" precision="55" value="44.638">44.638 N</geoCoordinate>
|
||
, 39.125-
|
||
<geoCoordinate id="369250309105F10AA68FFC2278587B61" box="[459,573,1292,1319]" degrees="39.129" direction="east" orientation="longitude" pageId="20" pageNumber="171" precision="55" value="39.129">39.129 E</geoCoordinate>
|
||
,
|
||
<elevation id="D88BD1C49105F10AA50FFC2278A17B60" box="[587,708,1292,1319]" metricMagnitude="1" metricUnit="m" metricValue="5.5" pageId="20" pageNumber="171" unit="m" value="55.0">
|
||
<quantity id="945E9B129105F10AA50FFC2278E37B61" box="[587,646,1292,1319]" metricMagnitude="1" metricUnit="m" metricValue="5.5" pageId="20" pageNumber="171" unit="m" value="55.0">55 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="271810379105F10AA597FC2279227B60" box="[723,839,1292,1318]" pageId="20" pageNumber="171" value="2024-04-08">
|
||
<collectingDate id="375CE9DF9105F10AA597FC2279227B60" box="[723,839,1292,1318]" pageId="20" pageNumber="171" value="2024-04-08">8.04.2024</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
; 2 teneral
|
||
<specimenCount id="45A0FD7E9105F10AA484FC23798F7B61" box="[960,1002,1293,1319]" count="1" pageId="20" pageNumber="171" type="male">♂♂</specimenCount>
|
||
(75178528, 75178636), 2 teneral
|
||
<specimenCount id="45A0FD7E9105F10AA236FC237FF97B61" box="[1394,1436,1293,1319]" count="1" pageId="20" pageNumber="171" type="female">♀♀</specimenCount>
|
||
(75178417, 75179228), the same place,
|
||
<date id="271810379105F10AA52AFC1E78957B0C" box="[622,752,1328,1354]" pageId="20" pageNumber="171" value="2021-04-23">23.04.2021</date>
|
||
;
|
||
<specimenCount id="45A0FD7E9105F10AA5BBFC1E79267B0D" box="[767,835,1328,1355]" count="3" pageId="20" pageNumber="171" type="female">3 ♀♀</specimenCount>
|
||
(75178494, 75178577, 75178395), the same place,
|
||
<date id="271810379105F10AA7D3FC7A7B7C7B28" box="[151,281,1364,1390]" pageId="20" pageNumber="171" value="2021-04-24">24.04.2021</date>
|
||
;
|
||
<specimenCount id="45A0FD7E9105F10AA660FC7A7B2B7B29" box="[292,334,1364,1391]" count="1" pageId="20" pageNumber="171" type="male">1 ♂</specimenCount>
|
||
(108928403), the same place,
|
||
<date id="271810379105F10AA5E2FC7A794D7B28" box="[678,808,1364,1390]" pageId="20" pageNumber="171" value="2019-06-27">27.06.2019</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="531936F79105F10AA783FC567F437BD4" blockId="20.[151,1437,860,2038]" box="[199,1318,1400,1427]" pageId="20" pageNumber="171">
|
||
<emphasis id="61D2EAE59105F10AA783FC567B317BD4" bold="true" box="[199,340,1400,1426]" pageId="20" pageNumber="171">Characters.</emphasis>
|
||
The male appendages resemble those of
|
||
<taxonomicName id="94A64D749105F10AA465FC5679DB7BD4" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[801,958,1400,1426]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="pulchellum">
|
||
<emphasis id="61D2EAE59105F10AA465FC5679DB7BD4" box="[801,958,1400,1426]" italics="true" pageId="20" pageNumber="171">C. pulchellum</emphasis>
|
||
</taxonomicName>
|
||
, with the following differences:
|
||
</paragraph>
|
||
<paragraph id="531936F79105F10AA783FCB27E6B782C" blockId="20.[151,1437,860,2038]" pageId="20" pageNumber="171">
|
||
Male cercus trunk much shorter and less processed, in lateral view (
|
||
<figureCitation id="CB9D2A729105F10AA482FCB27E537BF0" box="[966,1078,1436,1462]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="20" pageNumber="171">Fig. 3l–m</figureCitation>
|
||
) not raised, with slanting upper surface and blunter apex, so that tubercle is entirely seen in dorsal view (and is remarkably narrow); trunk in dorsal view (
|
||
<figureCitation id="CB9D2A729105F10AA7A4FCCA7B377BB8" box="[224,338,1508,1534]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="20" pageNumber="171">Fig. 2l–m</figureCitation>
|
||
) somewhat expanding posteriorly while of almost an even width in
|
||
<taxonomicName id="94A64D749105F10AA32EFCCA7F6C7BB8" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[1130,1289,1508,1534]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="pulchellum">
|
||
<emphasis id="61D2EAE59105F10AA32EFCCA7F6C7BB8" box="[1130,1289,1508,1534]" italics="true" pageId="20" pageNumber="171">C. pulchellum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="CB9D2A729105F10AA258FCCA7FEA7BB8" box="[1308,1423,1508,1534]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="20" pageNumber="171">Fig. 2o–q</figureCitation>
|
||
); prominence well expressed but broadly rounded, those of the two cerci also contact each other (see in dorsal view,
|
||
<figureCitation id="CB9D2A729105F10AA7D3FF027B637800" box="[151,262,1580,1606]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="20" pageNumber="171">Fig. 2l–m</figureCitation>
|
||
) as in
|
||
<taxonomicName id="94A64D749105F10AA60BFF027B897800" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[335,492,1580,1606]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="pulchellum">
|
||
<emphasis id="61D2EAE59105F10AA60BFF027B897800" box="[335,492,1580,1606]" italics="true" pageId="20" pageNumber="171">C. pulchellum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="CB9D2A729105F10AA6B8FF0278097800" box="[508,620,1580,1606]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="20" pageNumber="171">Fig. 2o–q</figureCitation>
|
||
). Process nearly straight as in
|
||
<taxonomicName id="94A64D749105F10AA4FBFF027E397800" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[959,1116,1580,1606]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="pulchellum">
|
||
<emphasis id="61D2EAE59105F10AA4FBFF027E397800" box="[959,1116,1580,1606]" italics="true" pageId="20" pageNumber="171">C. pulchellum</emphasis>
|
||
</taxonomicName>
|
||
but slenderer and somewhat shorter (
|
||
<figureCitation id="CB9D2A729105F10AA7B0FF7E7B06782C" box="[244,355,1616,1642]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="20" pageNumber="171">Fig. 4c–d</figureCitation>
|
||
), slightly not reaching the level of the paraproct prominence.
|
||
</paragraph>
|
||
<paragraph id="531936F79105F10AA783FF5A7A847890" blockId="20.[151,1437,860,2038]" pageId="20" pageNumber="171">
|
||
Male paraproct spine directed more behind than up and protrudes caudad much farer than tubercle; incision between spine and prominence as deep but not as broad, and prominence as attenuated as in
|
||
<taxonomicName id="94A64D749105F10AA3FDFFB67F3278F4" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[1209,1367,1688,1714]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="pulchellum">
|
||
<emphasis id="61D2EAE59105F10AA3FDFFB67F3278F4" box="[1209,1367,1688,1714]" italics="true" pageId="20" pageNumber="171">C. pulchellum</emphasis>
|
||
</taxonomicName>
|
||
. (
|
||
<figureCitation id="CB9D2A729105F10AA22BFFB67AB07890" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="20" pageNumber="171">Fig. 3l–m</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="531936F79105F10AA783FFCE7BB07904" blockId="20.[151,1437,860,2038]" pageId="20" pageNumber="171">
|
||
Male pronotum hind margin is generally as in
|
||
<taxonomicName id="94A64D749105F10AA596FFCE795C78BC" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[722,825,1760,1786]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="61D2EAE59105F10AA596FFCE795C78BC" box="[722,825,1760,1786]" italics="true" pageId="20" pageNumber="171">C. puella</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="CB9D2A729105F10AA40DFFCE79D278BD" box="[841,951,1760,1787]" captionStart="FIGURE 5" captionStartId="13.[151,250,1570,1594]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@13.[151,1436,181,1546]" captionTargetPageId="13" captionText="FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="20" pageNumber="171">Fig. 5d–e</figureCitation>
|
||
), female pronotum as well but with a more prominent, roundish triangular central lobe and very slightly deeper and somewhat asymmetrical incisions between it and lateral lobes (
|
||
<figureCitation id="CB9D2A729105F10AA631FE067BAC7904" box="[373,457,1832,1858]" captionStart="FIGURE 7" captionStartId="15.[151,250,1565,1589]" captionTargetBox="[447,1140,184,1541]" captionTargetId="figure-19@15.[447,1140,181,1541]" captionTargetPageId="15" captionText="FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="20" pageNumber="171">Fig. 7h</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="531936F79105F10AA783FE62785B79B0" blockId="20.[151,1437,860,2038]" pageId="20" pageNumber="171">
|
||
The black maculation development in males varies greatly, including in the same population in Fanagoriyskoe village. It is best reflected by the relative middorsal length of the posterior black spot in S3, which varies, quite continuously, ca from 20 to 70% of the S3 length, usually ca 50% (
|
||
<figureCitation id="CB9D2A729105F10AA4EDFEBA799879E8" box="[937,1021,1940,1966]" captionStart="FIGURE 9" captionStartId="21.[151,250,1715,1739]" captionTargetBox="[151,1436,181,1691]" captionTargetId="figure-19@21.[151,1436,181,1691]" captionTargetPageId="21" captionText="FIGURE 9. Males (a–b) and female (c) of Coenagrion ponticum in nature by a pond in the Fanagoriyskoe village NW environs, Goryachiy Klyuch Municipality, Krasnodarskiy Kray, Russia, 27.04.2019 (108924871, 37680876, 37681071, respectively); a—photo by OEK, b, c—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189928" httpUri="https://zenodo.org/record/12189928/files/figure.png" pageId="20" pageNumber="171">Fig. 9a</figureCitation>
|
||
). Along with its regular long lateral projections, as a rule broad in this species, it forms a cenral tooth (
|
||
<figureCitation id="CB9D2A729105F10AA43FFE9679AB7994" box="[891,974,1976,2002]" captionStart="FIGURE 9" captionStartId="21.[151,250,1715,1739]" captionTargetBox="[151,1436,181,1691]" captionTargetId="figure-19@21.[151,1436,181,1691]" captionTargetPageId="21" captionText="FIGURE 9. Males (a–b) and female (c) of Coenagrion ponticum in nature by a pond in the Fanagoriyskoe village NW environs, Goryachiy Klyuch Municipality, Krasnodarskiy Kray, Russia, 27.04.2019 (108924871, 37680876, 37681071, respectively); a—photo by OEK, b, c—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189928" httpUri="https://zenodo.org/record/12189928/files/figure.png" pageId="20" pageNumber="171">Fig. 9b</figureCitation>
|
||
), in some specimens produced into short, but of variable length, middorsal line.
|
||
</paragraph>
|
||
<caption id="07D9667F9104F10BA7D3FF9D78187955" ID-DOI="http://doi.org/10.5281/zenodo.12189928" ID-Zenodo-Dep="12189928" httpUri="https://zenodo.org/record/12189928/files/figure.png" pageId="21" pageNumber="172" startId="21.[151,250,1715,1739]" targetBox="[151,1436,181,1691]" targetPageId="21" targetType="figure">
|
||
<paragraph id="531936F79104F10BA7D3FF9D78187955" blockId="21.[151,1436,1715,1811]" pageId="21" pageNumber="172">
|
||
<emphasis id="61D2EAE59104F10BA7D3FF9D7B77788D" bold="true" box="[151,274,1715,1739]" pageId="21" pageNumber="172">FIGURE 9.</emphasis>
|
||
Males (a–b) and female (c) of
|
||
<taxonomicName id="94A64D749104F10BA502FF9D7947788D" baseAuthorityName="Bartenef" baseAuthorityYear="1929" box="[582,802,1715,1739]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="172" phylum="Arthropoda" rank="species" species="ponticum">
|
||
<emphasis id="61D2EAE59104F10BA502FF9D7947788D" box="[582,802,1715,1739]" italics="true" pageId="21" pageNumber="172">Coenagrion ponticum</emphasis>
|
||
</taxonomicName>
|
||
in nature by a pond in the Fanagoriyskoe village NW environs, Goryachiy Klyuch Municipality, Krasnodarskiy Kray, Russia, 27.04.2019 (108924871, 37680876, 37681071, respectively); a—photo by OEK, b, c—by VVO. Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="531936F79104F10BA783FE4B7F3779E5" blockId="21.[151,1437,1893,2028]" pageId="21" pageNumber="172">
|
||
Male S9 with a posterior black bar of variable shape, which usually has two (at sides, most frequent version) (
|
||
<figureCitation id="CB9D2A729104F10BA7DBFEA77B6979E5" box="[159,268,1929,1955]" captionStart="FIGURE 9" captionStartId="21.[151,250,1715,1739]" captionTargetBox="[151,1436,181,1691]" captionTargetId="figure-19@21.[151,1436,181,1691]" captionTargetPageId="21" captionText="FIGURE 9. Males (a–b) and female (c) of Coenagrion ponticum in nature by a pond in the Fanagoriyskoe village NW environs, Goryachiy Klyuch Municipality, Krasnodarskiy Kray, Russia, 27.04.2019 (108924871, 37680876, 37681071, respectively); a—photo by OEK, b, c—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189928" httpUri="https://zenodo.org/record/12189928/files/figure.png" pageId="21" pageNumber="172">Fig. 9a, b</figureCitation>
|
||
), one (central) or three (all above mentioned) anterior projections. S10 black above, blue at sides.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1BBC657C9104F108A783FE8378C67F5B" lastPageId="22" lastPageNumber="173" pageId="21" pageNumber="172" type="distribution">
|
||
<paragraph id="531936F79104F108A783FE8378C67F5B" blockId="21.[151,1437,1893,2028]" lastBlockId="22.[151,1436,151,285]" lastPageId="22" lastPageNumber="173" pageId="21" pageNumber="172">
|
||
<emphasis id="61D2EAE59104F10BA783FE837B917981" bold="true" box="[199,500,1965,1991]" pageId="21" pageNumber="172">Distribution and habitat.</emphasis>
|
||
In
|
||
<collectingCountry id="2BB176679104F10BA558FE8378027981" box="[540,615,1965,1991]" name="Russia" pageId="21" pageNumber="172">Russia</collectingCountry>
|
||
found only in Krasnodarskiy Kray, both at the SE part of the Black Sea coast as well as inland, behind the Main Caucasus Range, in Goryachiy Klyuch District. All records were at ponds situated below
|
||
<quantity id="945E9B129107F108A606F9B97BEF7EF7" box="[322,394,151,177]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="22" pageNumber="173" unit="m" value="200.0">200 m</quantity>
|
||
a.s.l. and surrounded by forest vegetation. The earliest registered date of imaginal emerging was
|
||
<date id="271810379107F108A78EF9957B587E93" box="[202,317,187,213]" pageId="22" pageNumber="173" value="2024-04-08">8.04.2024</date>
|
||
. No other
|
||
<taxonomicName id="94A64D749107F108A6F5F99578097E93" box="[433,620,187,213]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="61D2EAE59107F108A6F5F99578527E93" box="[433,567,187,213]" italics="true" pageId="22" pageNumber="173">Coenagrion</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
were found at these localities. Outside the region considered, the species ranges along the Black Sea coast of
|
||
<collectingCountry id="2BB176679107F108A569F9F178E27EBF" box="[557,647,223,249]" name="Georgia" pageId="22" pageNumber="173">Georgia</collectingCountry>
|
||
and has been reported from
|
||
<collectingCountry id="2BB176679107F108A482F9F17E217EBF" box="[966,1092,223,249]" name="Azerbaijan" pageId="22" pageNumber="173">Azerbaijan</collectingCountry>
|
||
, Northern
|
||
<collectingCountry id="2BB176679107F108A3FFF9F17E8D7EBF" box="[1211,1256,223,249]" name="Iran" pageId="22" pageNumber="173">Iran</collectingCountry>
|
||
(
|
||
<collectingRegion id="9162F8159107F108A3B3F9F17F5D7EBF" box="[1271,1336,223,249]" country="Iran" name="Gilan" pageId="22" pageNumber="173">Gilan</collectingRegion>
|
||
) and NE
|
||
<collectingCountry id="2BB176679107F108A7D3F82D7A927F5B" box="[151,247,259,285]" name="Turkey" pageId="22" pageNumber="173">Anatolia</collectingCountry>
|
||
(east of
|
||
<collectingRegion id="9162F8159107F108A61DF82D7BD67F5B" box="[345,435,259,285]" country="Turkey" name="Giresun" pageId="22" pageNumber="173">Giresun</collectingRegion>
|
||
) (
|
||
<bibRefCitation id="37374B069107F108A68DF82D78FE7F5B" author="Boudot, J. - P. & Borisov, S. & De Knijf, G. & Van Grunsven, R. & Schroter, A. & Kalkman, V. J." box="[457,667,259,285]" pageId="22" pageNumber="173" pagination="3 - 248" refId="ref27573" refString="Boudot, J. - P., Borisov, S., De Knijf, G., Van Grunsven, R., Schroter, A. & Kalkman, V. J. (2021) Atlas of the dragonflies and damselflies of West and Central Asia. Brachytron, 22, Supplement, 3 - 248." type="journal article" year="2021">Boudot et al. 2021</bibRefCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |