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<document ID-CLB-Dataset="33918" ID-DOI="http://dx.doi.org/10.3897/phytokeys.13.3259" ID-PMC="PMC3391716" ID-Pensoft-Pub="1314-2003-13-39" ID-Pensoft-UUID="FFD7FF88FFAB9757DA70FF94254DB433" ID-PubMed="22787426" ID-Zenodo-Dep="576124" ModsDocID="1314-2003-13-39" checkinTime="1451252006930" checkinUser="pensoft" docAuthor="Wagstaff, Steven J. &amp; Clarkson, Beverley R." docDate="2012" docId="CB9F5650305B56AFB48FD96B0DFE19F6" docLanguage="en" docName="PhytoKeys 13: 39-79" docOrigin="PhytoKeys 13" docPubDate="2012-07-03" docSource="http://dx.doi.org/10.3897/phytokeys.13.3259" docTitle="Empodisma minus L. A. S. Johnson &amp; D. F. Cutler, Kew Bull. 28, 383 1973" docType="treatment" docVersion="5" id="FFD7FF88FFAB9757DA70FF94254DB433" lastPageNumber="59" masterDocId="FFD7FF88FFAB9757DA70FF94254DB433" masterDocTitle="Systematics and ecology of the Australasian genus Empodisma (Restionaceae) and description of a new species from peatlands in northern New Zealand" masterLastPageNumber="79" masterPageNumber="39" pageNumber="58" updateTime="1720490797670" updateUser="ExternalLinkService">
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<mods:title id="D19502B939607192039088A0D15D5007">Systematics and ecology of the Australasian genus Empodisma (Restionaceae) and description of a new species from peatlands in northern New Zealand</mods:title>
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<mods:namePart id="828A66A9CF28244ABCEA4806312A4AC3">Wagstaff, Steven J.</mods:namePart>
<mods:affiliation id="623C387039F76245800C9E8E7CF6B6E3">Allan Herbarium, PO Box 40 Landcare Research, Lincoln 7640, New Zealand</mods:affiliation>
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<mods:namePart id="588E0504993F1F56311F8E84AC140745">Clarkson, Beverley R.</mods:namePart>
<mods:affiliation id="3D875B3CD41BD4B860B1B2D0104DEA51">Landcare Research, Private Bag 3127, Hamilton 3240, New Zealand</mods:affiliation>
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<mods:date id="F2A58FC6B84D35FB31B5FD32E4DE983F">2012</mods:date>
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<mods:number id="1ADFA31E778EC3794B52C6FA46E99ADD">13</mods:number>
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<treatment id="CB9F5650305B56AFB48FD96B0DFE19F6" ID-GBIF-Taxon="182229350" LSID="urn:lsid:plazi:treatment:CB9F5650305B56AFB48FD96B0DFE19F6" httpUri="http://treatment.plazi.org/id/CB9F5650305B56AFB48FD96B0DFE19F6" lastPageId="20" lastPageNumber="59" pageId="19" pageNumber="58">
<subSubSection id="DC73B75D3D518D4378617EFCB3FDE8AD" pageId="19" pageNumber="58" type="nomenclature">
<paragraph id="4F37BAA2A6F8E4D81C59F1766CF3453F" pageId="19" pageNumber="58">
2.
<taxonomicName id="E020ADD1A9F3D16006EF321B364C4D41" ID-CoL="39LK3" LSID="CB9F5650-305B-56AF-B48F-D96B0DFE19F6" authority="(Hook. f.) L. A. S. Johnson &amp; D. F. Cutler, Kew Bull. 28, 383 (1973)" authorityName="L. A. S. Johnson &amp; D. F. Cutler, Kew Bull. 28, 383" authorityYear="1973" baseAuthorityName="Hook. f." baseAuthorityYear="1973" class="Liliopsida" family="Restionaceae" genus="Empodisma" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Empodisma minus" order="Poales" pageId="19" pageNumber="58" phylum="Tracheophyta" rank="species" species="minus">Empodisma minus (Hook.f.) L.A.S.Johnson &amp; D.F.Cutler, Kew Bull. 28, 383 (1973)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="24E5FD9ADE19066E35BA35874B36B012" pageId="19" pageNumber="58" type="reference_group">
<paragraph id="1F46E139C464D7F2566491F64C9B0202" pageId="19" pageNumber="58">
<taxonomicName id="A821B14F0D259506FF0D85E247FB7552" class="Liliopsida" family="Restionaceae" genus="Calorophus" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="19" pageNumber="58" phylum="Tracheophyta" rank="species" species="minor">Calorophus minor</taxonomicName>
Hook.f.,
<emphasis id="F89352BBD59135709824D402EBAFF690" italics="true" pageId="19" pageNumber="58">Fl. Nov. Zel. I</emphasis>
, 267 (1852-1853).
</paragraph>
<paragraph id="191731928E6F83E583C38CACB2BF1CDA" pageId="19" pageNumber="58">
<taxonomicName id="CB8CC86EC8C3EC11EA45808246C6D167" class="Liliopsida" family="Restionaceae" genus="Calorophus" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="19" pageNumber="58" phylum="Tracheophyta" rank="species" species="elongatus">Calorophus elongatus</taxonomicName>
var.
<emphasis id="7D972DA270F7FD13B590CF6F317DC6C0" italics="true" pageId="19" pageNumber="58">minor</emphasis>
(Hook.f.) Hook.f.,
<emphasis id="042D9DFC757F263094E7FE5E692A8922" italics="true" pageId="19" pageNumber="58">Fl. Tas. II</emphasis>
, 75 (1858-1859).
</paragraph>
<paragraph id="851A6C3BB44054EF8B5D2158C540E6BF" pageId="19" pageNumber="58">
<taxonomicName id="DFC9D373703824E5FB31F2F6F2E0CBDD" authorityName="Benth" authorityYear="1878" baseAuthorityName="R. Br." class="Liliopsida" family="Restionaceae" genus="Hypolaena" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="19" pageNumber="58" phylum="Tracheophyta" rank="species" species="lateriflora">Hypolaena lateriflora</taxonomicName>
var.
<emphasis id="1AB8E9F27F2D42D358A6254713157410" italics="true" pageId="19" pageNumber="58">minor</emphasis>
(Hook.f.) Cheeseman,
<emphasis id="D7545963D5617DF3AF32D39AC6F1FEFA" italics="true" pageId="19" pageNumber="58">Manual N.Z. Flora</emphasis>
ed. 1, 762 (1906)
</paragraph>
</subSubSection>
<subSubSection id="9218904E138DC387EC0EBFDC0FE60C61" pageId="19" pageNumber="58" type="lectotype">
<paragraph id="8140D82741A9EA05717AF167150F007E" pageId="19" pageNumber="58">Lectotype.</paragraph>
<paragraph id="38681A8CEF3FC288CE475E759030F6D7" pageId="19" pageNumber="58">
New Zealand, near Nelson, Bidwell no. 84, K000441989; (
<figureCitation id="D3D4276B88516041958084A15FA73702" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. High resolution photograph of the lectotype of Empodisma minus (Hook. f.) L. A. S. Johnson &amp; D. F. Cutler. Reproduced with the consent of the Royal Botanic Gardens, Kew, © The Board of Trustees of the Royal Botanic Gardens. Hooker (1853) described the new species Calorophus minor Hook. f. based upon Bidwell, Colenso and Lyall specimens. A specimen collected near Nelson by Bidwell, no. 84, K 000441989, was chosen as the lectotype by Moore and Edgar (1970)." httpUri="https://binary.pensoft.net/fig/10091" pageId="19" pageNumber="58">Fig. 10</figureCitation>
; designated by
<bibRefCitation id="5259FD21CC06E72A3A33381B7CA5342A" author="Moore, LB" journalOrPublisher="Palaeogeography, Palaeoclimatology, Palaeoecology" pageId="24" pageNumber="63" refId="B66" refString="Moore, LB, Edgar, E, 1970. Flora of New Zealand II. Government Printer, Wellington, New Zealand." title="Flora of New Zealand II. Government Printer, Wellington, New Zealand." year="1970">Moore and Edgar 1970</bibRefCitation>
, pg 89).
</paragraph>
<caption id="D02264E26EF62284A74CED6BACAC08D3" httpUri="https://binary.pensoft.net/fig/10091" pageId="19" pageNumber="58" start="Figure 10" startId="F10">
<paragraph id="F109EAE1F8B47C2F83C67C7D5F4DBD35" pageId="19" pageNumber="58">
<emphasis id="E4EC092B43E26AF252C61864E494B5F3" bold="true" pageId="19" pageNumber="58">Figure 10.</emphasis>
High resolution photograph of the lectotype of
<taxonomicName id="FF83F1FA5371DCDCAFA18FA0194615F6" authorityName="L. A. S. Johnson &amp; D. F. Cutler, Kew Bull. 28, 383" authorityYear="1973" baseAuthorityName="Hook. f." class="Liliopsida" family="Restionaceae" genus="Empodisma" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Poales" pageId="19" pageNumber="58" phylum="Tracheophyta" rank="species" species="minus">
<emphasis id="407CA2F2788DDB38FF532C71A7CFAE4B" italics="true" pageId="19" pageNumber="58">Empodisma minus</emphasis>
</taxonomicName>
(Hook.f.) L.A.S. Johnson &amp; D.F.Cutler. Reproduced with the consent of the Royal Botanic Gardens, Kew, © The Board of Trustees of the Royal Botanic Gardens.
<bibRefCitation id="CBAF9A53A66DD408D83E3D6ECEFD1864" author="Hooker, JD" journalOrPublisher="Systematic Biology" pageId="23" pageNumber="62" refId="B41" refString="Hooker, JD, 1852-1853. The Botany of the Antarctic Voyage of H.M. Discovery Ships Erebus and Terror in the Years 1839-1843.Vol. 2. Flora Novae-Zelandiae Part I. Flowering Plants. London, Lovell Reeve. 312 p." title="The Botany of the Antarctic Voyage of H. M. Discovery Ships Erebus and Terror in the Years 1839 - 1843. Vol. 2. Flora Novae-Zelandiae Part I. Flowering Plants. London, Lovell Reeve. 312 p." year="1852 - 1853">Hooker (1853)</bibRefCitation>
described the new species
<taxonomicName id="3F9D043437B72A484B719DA2AEBFFBB2" class="Liliopsida" family="Restionaceae" genus="Calorophus" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="19" pageNumber="58" phylum="Tracheophyta" rank="species" species="minor">
<emphasis id="B337E862A263B8498F8D4C8AC202700F" italics="true" pageId="19" pageNumber="58">Calorophus minor</emphasis>
</taxonomicName>
Hook.f. based upon Bidwell, Colenso and Lyall specimens. A specimen collected near Nelson by Bidwell, no. 84, K000441989, was chosen as the lectotype by
<bibRefCitation id="20C3C65CE9368ADF6310801F613D90FE" author="Moore, LB" journalOrPublisher="Palaeogeography, Palaeoclimatology, Palaeoecology" pageId="24" pageNumber="63" refId="B66" refString="Moore, LB, Edgar, E, 1970. Flora of New Zealand II. Government Printer, Wellington, New Zealand." title="Flora of New Zealand II. Government Printer, Wellington, New Zealand." year="1970">Moore and Edgar (1970)</bibRefCitation>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="75D29FF46513091B432F0CFB7B4B8206" pageId="19" pageNumber="58" type="etymology">
<paragraph id="B9A3EE832E1DF60FC3FB13155CA5F7D9" pageId="19" pageNumber="58">Etymology.</paragraph>
<paragraph id="9C94A3D31D4FEE87575FB3C1F7785A06" pageId="19" pageNumber="58">
minus describes the small stature of
<taxonomicName id="2BFF035C77AE9F6392854534479A0ECB" authorityName="L. A. S. Johnson &amp; D. F. Cutler, Kew Bull. 28, 383" authorityYear="1973" baseAuthorityName="Hook. f." class="Liliopsida" family="Restionaceae" genus="Empodisma" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Poales" pageId="19" pageNumber="58" phylum="Tracheophyta" rank="species" species="minus">
<emphasis id="A20A61038A5393FDD724395C85ED7807" italics="true" pageId="19" pageNumber="58">Empodisma minus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="BB5F28BA359A11ABDAED2D12A7F891D1" lastPageId="20" lastPageNumber="59" pageId="19" pageNumber="58" type="description">
<paragraph id="901E60838CD5270EB80D441B7144D4A1" pageId="19" pageNumber="58">Description.</paragraph>
<paragraph id="8036958FA8DA7B869B88D5F4F96C4E85" lastPageId="20" lastPageNumber="59" pageId="19" pageNumber="58">
Culms dark green, 12-81 cm in height, 0.7-1.3 mm in diameter, branching profusely. Leaf sheaths closely appressed, 3.5-10.2 mm in length, borne at
<pageBreakToken id="D10ABC70A75C8003E4D0CE4B88E737D2" pageId="20" pageNumber="59" start="start">short</pageBreakToken>
intervals 15.0-48.0 mm; light green to light brown early in the season maturing dark brown; mouth ciliate with a prominent tuft of woolly white hairs in New Zealand specimens, spare or lacking in Australian specimens. Lamina strongly reflexed from leaf sheath, 1.5-4.2 mm long, persistent light green when young maturing dark brown. Spikelets brown, male spikelet 3.9-8 mm long, anthers 1.2-2.0 mm long; female spikelet 3.5-7.0 mm long; nutlets dark brown approximately 2.6 mm long. 2n = 24. Flowering Aug.- Apr. [
<figureCitation id="1AD2F8FBE8E6F93DC604623A95A3AD2D" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Morphological characteristics of Empodisma minus. A Vegetative shoot with attached leaves and flowers (2.5 x actual size) B Rhizomes with emerging vegetative shoots (2.5 x actual size) C Vegetative shoot with attached pistillate flower D Pistillate flower with attached bracts E Gynoecium F Mature nut G Vegetative shoot with attached staminate spikelet H Staminate flower with attached bracts I Staminate flower. Scale bar = 1 mm." httpUri="https://binary.pensoft.net/fig/10092" pageId="20" pageNumber="59">Fig. 11</figureCitation>
; see also illustration in
<bibRefCitation id="C2103D09FD05C23DA2E09420CE5C5A89" author="Meney, KA" journalOrPublisher="Journal of Biogeography" pageId="24" pageNumber="63" refId="B61" refString="Meney, KA, Pate, JS, 1999. Australian Rushes-Biology, Identification and Conservation of Restionaceae and Allied Families. University of Western Australia Press, Perth." title="Australian Rushes-Biology, Identification and Conservation of Restionaceae and Allied Families. University of Western Australia Press, Perth." year="1999">Meney and Pate (1999)</bibRefCitation>
].
</paragraph>
<caption id="80D89C276196F471C99CE0EB394CAEC8" httpUri="https://binary.pensoft.net/fig/10092" pageId="20" pageNumber="59" start="Figure 11" startId="F11">
<paragraph id="5F30D9CA4A6175F2531715EB5D0BECDE" pageId="20" pageNumber="59">
<emphasis id="9189D40A3FCDF1BA82B8D5A5DDDA4194" bold="true" pageId="20" pageNumber="59">Figure 11.</emphasis>
Morphological characteristics of
<taxonomicName id="22659A3302BD7977216D31D62639C0A8" authorityName="L. A. S. Johnson &amp; D. F. Cutler, Kew Bull. 28, 383" authorityYear="1973" baseAuthorityName="Hook. f." class="Liliopsida" family="Restionaceae" genus="Empodisma" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Poales" pageId="20" pageNumber="59" phylum="Tracheophyta" rank="species" species="minus">
<emphasis id="4D8E1584A456DD17C2FFD94F3683596E" italics="true" pageId="20" pageNumber="59">Empodisma minus</emphasis>
</taxonomicName>
.
<emphasis id="0515E85A893170520F40549F9721AB5B" bold="true" pageId="20" pageNumber="59">A</emphasis>
Vegetative shoot with attached leaves and flowers (2.5
<normalizedToken id="A19DE11F7E7824710A7506F2B0DBFD39" originalValue="×">x</normalizedToken>
actual size)
<emphasis id="C92ACC8329230201106BF76FA0C191D7" bold="true" pageId="20" pageNumber="59">B</emphasis>
Rhizomes with emerging vegetative shoots (2.5
<normalizedToken id="F88AA2AF6A3B60DC3A662EBC62BEC899" originalValue="×">x</normalizedToken>
actual size)
<emphasis id="C638C1B78E7BB79B1FA7F61EF2AF5D9D" bold="true" pageId="20" pageNumber="59">C</emphasis>
Vegetative shoot with attached pistillate flower
<emphasis id="A6329B8977CDA3A649768B035C67B1DE" bold="true" pageId="20" pageNumber="59">D</emphasis>
Pistillate flower with attached bracts
<emphasis id="8AE4A489F0605053E1B46B42FD1F7DB8" bold="true" pageId="20" pageNumber="59">E</emphasis>
Gynoecium
<emphasis id="55666AE98300F2FCD77C936360CBD3AB" bold="true" pageId="20" pageNumber="59">F</emphasis>
Mature nut
<emphasis id="BE5F24559AE9DE1B937EF3E491A63700" bold="true" pageId="20" pageNumber="59">G</emphasis>
Vegetative shoot with attached staminate spikelet
<emphasis id="1A59E952EE3774057D05B8A5EFE62D1E" bold="true" pageId="20" pageNumber="59">H</emphasis>
Staminate flower with attached bracts
<emphasis id="198E07AAFBB70E05BA5F99647382E604" bold="true" pageId="20" pageNumber="59">I</emphasis>
Staminate flower. Scale bar = 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="8EEC52AA5C0764B101567CA6D6213C63" pageId="20" pageNumber="59" type="comments">
<paragraph id="4E34A746E453079B54E2B9C405A12314" pageId="20" pageNumber="59">Comments.</paragraph>
<paragraph id="7D72D1685F295BA65D6371FC29CDD2DF" pageId="20" pageNumber="59">
Morphologically similar to small forms of
<taxonomicName id="CF5A342DFDA578664182565E1AFFF2AC" authorityName="Wagstaff &amp; B. R. Clarkson" authorityYear="2012" class="Liliopsida" family="Restionaceae" genus="Empodisma" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Poales" pageId="20" pageNumber="59" phylum="Tracheophyta" rank="species" species="robustum">
<emphasis id="4A3A9EEEEE2C68B4A0EECBB57A2AB0D4" italics="true" pageId="20" pageNumber="59">Empodisma robustum</emphasis>
</taxonomicName>
distinguished by its smaller stature (though plants from 0.4 to 1 m tall are noted from eastern Australia), more delicate culms and smaller spikelets. Most collections are sterile or male, and the few females generally lack mature fruits. Chromosome counts of 2n=24 were reported from plants collected from the NW slope of Mt. Ruapehu and three counts from NSW (
<bibRefCitation id="1BFCA4F9113229270F3F80973AF07BE8" author="Briggs, BG" journalOrPublisher="The Botanical Review" pageId="22" pageNumber="61" refId="B5" refString="Briggs, BG, 1966. Chromosome numbers of some Australian monocotyledons. Contributions of the N.S.W. National Herbarium IV: 24-34." title="Chromosome numbers of some Australian monocotyledons. Contributions of the N. S. W. National Herbarium IV: 24 - 34." year="1966">Briggs 1966</bibRefCitation>
,
<bibRefCitation id="B95E6C83972021C5ACE1D581B4D64BB0" author="Johnson, LAS" journalOrPublisher="Kew Bulletin" pageId="23" pageNumber="62" pagination="381 - 385" publicationUrl="10.2307/4108881" refId="B49" refString="Johnson, LAS, Cutler, DF, 1973. Empodisma: a new genus of Australasian Restionaceae. Kew Bulletin 28: 381 - 385, 10.2307/4108881" title="Empodisma: a new genus of Australasian Restionaceae." url="10.2307/4108881" volume="28" year="1973">Johnson and Cutler 1973</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C0999D905E1A03453D20A9718BBF18AC" pageId="20" pageNumber="59" type="representative specimens">
<paragraph id="8E308BDC3F538FB48FBEF962B11A6544" pageId="20" pageNumber="59">Representative specimens.</paragraph>
<paragraph id="65443EA60D443E300D6083D7A7996C10" pageId="20" pageNumber="59">Australia, Tasmania, near Margate, ♂ flowers, D.A. &amp; A. Ratkowsky 1474, CHR303032, Australia, Queensland, Moreton Island, ♂ flowers, L. Durrington 1114 &amp; S. Levine, CHR272564, Australia, Tasmania, Newdegate Pass, not in flower, T. Dobson 77107, CHR313744; New Zealand, S. Westland, ♂ flowers, G.C. Kelly, Oct 1966, CHR177206; New Zealand, Rahu Saddle, with few ♀ flowers, E.J. Godley, 1 July 1958, CHR108315; New Zealand, Bell Hill Plains, ♂ flowers, J. Clarke 1 Feb 1969, CHR189013; New Zealand, Ngamatea, ♂ flowers, N.J. Moar, 12 Jan 1949, CHR70144; New Zealand, Waikareiti, ♂ flowers, A.P. Druce, Feb 1968, CHR180674; New Zealand, Silica Springs Track, ♂ flowers G. Rennison, A61/36,, CHR535708; New Zealand, Makerikeri tarns, ♂ flowers, A.P. Druce, Nov 1973, CHR260376; New Zealand, Tussock Creek, ♂ flowers, L.B. Moore, 28 July 1968, CHR188099; New Zealand, Mokoreta, with few ♀ flowers, W.R. Sykes 41/94, CHR497058; New Zealand, Bayswater Bog, not flowering, B.R. Clarkson19 Feb.2009, CHR605146; New Zealand, Awarua Bay, not flowering, P.N. Johnson 653, CHR437892; New Zealand, West Cape, A.F. Mark, 5 Feb 1972, CHR218694; New Zealand, Coal Creek, ♂ flowers; I. Payton, 13 Sept 1976, CHR520808; New Zealand, Mt. Rockport, not flowering, I.A. McNew, 31 July 1942, CHR35234; New Zealand, Lake Sylvester, ♂ flowers, R. Melville 5915, CHR142781; New Zealand, Lake Sylvester, ♀ flowers, R. Mason &amp; N. Moar 4658, CHR95709; New Zealand, Bealey spur, with immature ♀ flowers, P. Douglas 26 Nov 1979, CHR362302; New Zealand, Lake Tennyson, ♂ flowers, M.J.A. Simpson 6315, CHR22759; New Zealand, Patterson Inlet, ♂ flowers, L.J. Dumbleton &amp; E. Edgar, CHR182509; New Zealand, ♂ flowers, Fosberg, Feb 1949; CHR30378; New Zealand, Kaitangata, plants with ♀ and ♂ flowers, R. Mason &amp; N.T. Moar 953, CHR 75833; New Zealand, Kapuka, ♀ flowers, W.H. Harbond 20 Nov 1968, CHR183615.</paragraph>
</subSubSection>
<subSubSection id="E0B733BFDDA39D5FB16B8D10BF48C16E" pageId="20" pageNumber="59" type="distribution">
<paragraph id="E49ED99B5342F709399C53C9D9CF6C73" pageId="20" pageNumber="59">Distribution.</paragraph>
<paragraph id="DEC0AE2BA2DBA0A2530AF6DDDFFE86B5" pageId="20" pageNumber="59">Widely distributed in Tasmania and all mainland Australian States except Western Australia and the Northern Territory; in New Zealand extending north to approximately 38°S latitude.</paragraph>
</subSubSection>
<subSubSection id="890F1F9DAC0C45E5F4BC6F8553FE27A5" pageId="20" pageNumber="59" type="habitat">
<paragraph id="0D44232DA60D43CE48194F10B5D07D7F" pageId="20" pageNumber="59">Habitat.</paragraph>
<paragraph id="5D29F5B58DCDDBA177C2224BC1F67AF1" pageId="20" pageNumber="59">Locally abundant in seasonally or permanently inundated wetlands, heathlands, fens and peat bogs from sea level to alpine</paragraph>
</subSubSection>
<subSubSection id="CF22A0A7B9969EC27DBAB7FDEC756E0F" pageId="20" pageNumber="59" type="conservation status">
<paragraph id="9309AD8EB45AF58991D10B18DC7C2A3E" pageId="20" pageNumber="59">Conservation status.</paragraph>
<paragraph id="5F6BE955C9470C6029C23C9E17682C0D" pageId="20" pageNumber="59">Not threatened.</paragraph>
</subSubSection>
</treatment>
</document>