129 lines
18 KiB
XML
129 lines
18 KiB
XML
<document id="7C995496FE57C13218884D18F3E8955B" ID-DOI="10.5281/zenodo.5391021" ID-ISSN="1638-9387" ID-Zenodo-Dep="5391021" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630314169453" checkinUser="marcus" docAuthor="Deuve, Thierry" docDate="2001" docId="CB2A87CE1C1DFFC4033FFE00FE9D80AD" docLanguage="fr" docName="Zoosystema.2001.23.3.547-578.pdf" docOrigin="Zoosystema 23 (3)" docStyle="DocumentStyle:64A3174DE33E974D4EB6573A0A793B12.4:Zoosystema.2000-2008.journal_article.type2" docStyleId="64A3174DE33E974D4EB6573A0A793B12" docStyleName="Zoosystema.2000-2008.journal_article.type2" docStyleVersion="4" docTitle="Eustra lebretoni Deuve 1987" docType="treatment" docVersion="4" lastPageNumber="568" masterDocId="3713FFB61C08FFD1023EFF9BFF908529" masterDocTitle="Le genre Eustra Schmidt-Goebel, 1846, insectes (Coleoptera, Paussidae, Ozaeninae) à genitalia femelles orthotopiques" masterLastPageNumber="578" masterPageNumber="547" pageNumber="568" updateTime="1699223122802" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A804937BE35E4F27BB2E88F5025FB6D0">Le genre Eustra Schmidt-Goebel, 1846, insectes (Coleoptera, Paussidae, Ozaeninae) à genitalia femelles orthotopiques</mods:title>
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<mods:namePart id="992BD6C2F28F4D07FB02980F05A18C7A">Deuve, Thierry</mods:namePart>
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<mods:affiliation id="9C74D89894C63EE42361DB63B915217B">Muséum national d’Histoire naturelle, Laboratoire d’Entomologie 45 rue Buffon, F- 75005 Paris deuve @ mnhn. fr.</mods:affiliation>
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<mods:nameIdentifier id="C1C9D5F774D76D111B6E206D310B54B3" type="email">deuve@mnhn.fr</mods:nameIdentifier>
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<mods:typeOfResource id="A5A848E6E9E99A7597C945E379859056">text</mods:typeOfResource>
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<mods:title id="C63D20D9E8B56D0210834490F7C2C024">Zoosystema</mods:title>
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</mods:titleInfo>
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<mods:part id="766687774C4556944D4B0E31AC7BC6A9">
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<mods:date id="4D0947D3BADBC861A1BA68E66EA53683">2001</mods:date>
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<mods:number id="5833CDBACC78D72C2B0673484601C4CA">23</mods:number>
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<mods:number id="A1E7CB4DB8EEF83488996B54DE209377">3</mods:number>
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<mods:start id="C6562187EF2589825AE284CEFEA55EFA">547</mods:start>
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<mods:end id="6919EE09A75DA9C8D8FA2F1C5054D9A0">578</mods:end>
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<mods:identifier id="7402E2917F98B26D1303CA2703A819E0" type="DOI">10.5281/zenodo.5391021</mods:identifier>
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<mods:identifier id="E83130F381DCCE88268A01B39D8EB548" type="ISSN">1638-9387</mods:identifier>
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<treatment id="CB2A87CE1C1DFFC4033FFE00FE9D80AD" ID-DOI="http://doi.org/10.5281/zenodo.5467675" ID-Zenodo-Dep="5467675" LSID="urn:lsid:plazi:treatment:CB2A87CE1C1DFFC4033FFE00FE9D80AD" httpUri="http://treatment.plazi.org/id/CB2A87CE1C1DFFC4033FFE00FE9D80AD" lastPageNumber="568" pageId="21" pageNumber="568">
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<paragraph id="433C36D81C1DFFC4033FFE00FDD2849F" blockId="21.[151,684,411,679]" box="[257,578,411,438]" pageId="21" pageNumber="568">
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<heading id="187481B41C1DFFC4033FFE00FDD2849F" box="[257,578,411,438]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="568" reason="2">
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<taxonomicName id="84834D5B1C1DFFC4033FFE00FDD2849F" ID-CoL="3DBDS" authority="Deuve, 1987" authorityName="Deuve" authorityYear="1987" box="[257,578,411,438]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="568" phylum="Arthropoda" rank="species" species="lebretoni">
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<emphasis id="71F7EACA1C1DFFC4033FFE00FE3F849F" bold="true" box="[257,431,411,438]" italics="true" pageId="21" pageNumber="568">Eustra lebretoni</emphasis>
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Deuve, 1987
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0B9965531C1DFFC402A9FE5DFEE78740" pageId="21" pageNumber="568" type="reference_group">
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<paragraph id="433C36D81C1DFFC402A9FE5DFDD184D1" blockId="21.[151,684,411,679]" pageId="21" pageNumber="568">
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<treatmentCitation id="C22210C91C1DFFC402A9FE5DFD8184F7" author="DEUVE TH" box="[151,529,454,478]" page="238" pageId="21" pageNumber="568" year="1987">
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<taxonomicName id="84834D5B1C1DFFC402A9FE5DFD8184F7" ID-CoL="3DBDS" authority="Deuve, 1987 a: 238" authorityName="Deuve" authorityPageNumber="238" authorityYear="1987" box="[151,529,454,478]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="568" phylum="Arthropoda" rank="species" species="lebretoni">
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<emphasis id="71F7EACA1C1DFFC402A9FE5DFEA784F7" box="[151,311,454,478]" italics="true" pageId="21" pageNumber="568">Eustra lebretoni</emphasis>
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<bibRefCitation id="27124B291C1DFFC4037EFE5DFD8184F7" author="DEUVE TH" box="[320,529,454,478]" pageId="21" pageNumber="568" pagination="237 - 242" refId="ref17629" refString="DEUVE TH. 1987 a. - Faune cavernicole de Thailande: coleopteres Paussidae Ozaeninae. Bulletin de la Societe entomologique de France 91 (1986): 237 - 242." type="journal article" year="1987">Deuve, 1987a: 238</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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;
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<treatmentCitation id="C22210C91C1DFFC40017FE5DFD0D84F7" author="DEUVE TH" box="[553,669,454,478]" page="18" pageId="21" pageNumber="568" year="1988">
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<bibRefCitation id="27124B291C1DFFC40017FE5DFD0D84F7" author="DEUVE TH" box="[553,669,454,478]" pageId="21" pageNumber="568" pagination="285 - 290" refId="ref17818" refString="DEUVE TH. 1988 d. - Trechiama siamensis n. sp., nouveau coleoptere cavernicole des karsts de Thailande (Trechidae). Bulletin de la Societe entomologique de France 92: 285 - 290." type="journal article" year="1988">1988d: 18</bibRefCitation>
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</treatmentCitation>
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;
|
||
<treatmentCitation id="C22210C91C1DFFC402A9FE7BFF6884D1" author="DEUVE TH" box="[151,248,479,504]" page="70" pageId="21" pageNumber="568" year="1993">
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<bibRefCitation id="27124B291C1DFFC402A9FE7BFF6884D1" author="DEUVE TH" box="[151,248,479,504]" pageId="21" pageNumber="568" refId="ref17905" refString="DEUVE TH. 1993. - L'abdomen et les genitalia des femelles de coleopteres Adephaga. Memoires du Museum national d'Histoire naturelle 155: 184 p." type="book" year="1993">1993: 70</bibRefCitation>
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</treatmentCitation>
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, 71. —
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<treatmentCitation id="C22210C91C1DFFC40376FE7BFDAD84D1" author="CASALE A. & VIGNA TAGLIANTI A. & JUBERTHIE CHR" box="[328,573,479,504]" page="1056" pageId="21" pageNumber="568" year="1998">
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<bibRefCitation id="27124B291C1DFFC40376FE7BFDAD84D1" author="CASALE A. & VIGNA TAGLIANTI A. & JUBERTHIE CHR" box="[328,573,479,504]" pageId="21" pageNumber="568" pagination="1047 - 1081" refId="ref17414" refString="CASALE A., VIGNA TAGLIANTI A. & JUBERTHIE CHR. 1998. - Coleoptera Carabidae, in JUBERTHIE CHR. & DECU V. (eds), Encyclopaedia Biospeologica 2: 1047 - 1081." type="journal article" year="1998">
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Casale
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<emphasis id="71F7EACA1C1DFFC403B3FE44FE2484D1" box="[397,436,479,504]" italics="true" pageId="21" pageNumber="568">et al</emphasis>
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. 1998: 1056
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</bibRefCitation>
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="433C36D81C1DFFC402A9FD9FFEE78740" blockId="21.[151,684,411,679]" pageId="21" pageNumber="568">
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<materialsCitation id="F3EB3C851C1DFFC402A9FD9FFEE78740" collectingDate="1985-07-11" collectionCode="MNHN" collectorName="de Chiang Dao & de Tham Tab Tao, L" country="Thailand" location="Thailande" pageId="21" pageNumber="568" specimenCount="2" typeStatus="holotype">
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TYPES. —
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<typeStatus id="9C38887A1C1DFFC402C0FD9EFECB8734" box="[254,347,517,541]" pageId="21" pageNumber="568" type="holotype">Holotype</typeStatus>
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1,
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<collectingCountry id="3B9476481C1DFFC403A9FD9EFE6D8734" box="[407,509,517,541]" name="Thailand" pageId="21" pageNumber="568">Thaïlande</collectingCountry>
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, environs
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<collectorName id="EE76530E1C1DFFC4005FFD9EFF7D871D" pageId="21" pageNumber="568">de Chiang Dao</collectorName>
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, grotte
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<collectorName id="EE76530E1C1DFFC40309FD85FD9A871D" box="[311,522,541,566]" pageId="21" pageNumber="568">de Tham Tab Tao, L</collectorName>
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. Deharveng,
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<date id="373D10181C1DFFC400B2FD85FF6B8766" pageId="21" pageNumber="568" value="1985-07-11">
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<collectingDate id="2779E9F01C1DFFC400B2FD85FF6B8766" pageId="21" pageNumber="568" value="1985-07-11">11. VII.1985</collectingDate>
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</date>
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,
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<emphasis id="71F7EACA1C1DFFC40337FDADFE818764" box="[265,273,566,589]" italics="true" pageId="21" pageNumber="568">(</emphasis>
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||
<collectionCode id="2592AE1D1C1DFFC4032CFDACFEFA8766" box="[274,362,567,591]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="568">MNHN</collectionCode>
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);
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<typeStatus id="9C38887A1C1DFFC403B7FDACFE7C8766" box="[393,492,567,591]" pageId="21" pageNumber="568" type="paratype">paratypes</typeStatus>
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1, 1, même provenance (
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<collectionCode id="2592AE1D1C1DFFC4032AFDCAFEFC8740" box="[276,364,593,617]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="568">MNHN</collectionCode>
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).
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||
</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0B9965531C1DFFC402A9FDEEFE9D80AD" pageId="21" pageNumber="568" type="distribution">
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||
<paragraph id="433C36D81C1DFFC402A9FDEEFD06878E" blockId="21.[151,684,411,679]" pageId="21" pageNumber="568">
|
||
DISTRIBUTION. —
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<emphasis id="71F7EACA1C1DFFC40366FDEFFDF087A7" bold="true" box="[344,608,628,654]" pageId="21" pageNumber="568">
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||
<collectingCountry id="3B9476481C1DFFC40366FDEFFE5087A5" box="[344,448,628,652]" name="Thailand" pageId="21" pageNumber="568">Thaïlande</collectingCountry>
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septentrionale.
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</emphasis>
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Grotte de Tham Tab Tao, seule est connue la série typique.
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</paragraph>
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<paragraph id="433C36D81C1DFFC402A9FD72FE998628" blockId="21.[151,684,745,1412]" box="[151,265,745,771]" pageId="21" pageNumber="568">DIAGNOSE</paragraph>
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<paragraph id="433C36D81C1DFFC402A9FC91FE9D80AD" blockId="21.[151,684,745,1412]" pageId="21" pageNumber="568">
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Espèce assez grande et robuste (long.
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||
<quantity id="847B9B3D1C1DFFC4001BFC91FDEE860A" box="[549,638,777,804]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="21" pageNumber="568" unit="mm" value="3.8">3,8 mm</quantity>
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), les antennes et les pattes un peu allongées, fines. Coloris d’un jaune testacé assez sombre, la tête à peine plus foncée, les élytres sans rembrunissement. Yeux bien développés, nettement plus convexes que les tempes. Pronotum nettement transverse (lp/Lp = 1,55), les gouttières latérales creusées, les côtés denticulés, sinués avant les angles basaux qui sont subdroits, petits mais vifs. Édéage (
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<figureCitation id="DBB82A5D1C1DFFC40362FBB1FE24816D" box="[348,436,1066,1092]" captionStart="FIG" captionStartId="11.[151,162,1585,1602]" captionTargetBox="[171,1234,194,1543]" captionTargetId="figure-107@11.[125,1301,124,1581]" captionTargetPageId="11" captionText="FIG. 5. — Eustra Schmidt-Goebel, 1846, édéages des mâles, face latérale du lobe médian et paramère gauche; A, E. japonica Bates, 1892, paratype; B, E. hammondi n. sp., holotype; C, E. gomyi n. sp., holotype; D, E. lebretoni Deuve, 1987, paratype; E, E. leclerci (Deuve, 1986), holotype; F, E. deharvengi (Deuve, 1986), holotype; G, E. chinensis Bänninger, 1949, de Taiwan, « Taihorin ». Abréviation: gc, gonocaulis. Échelle: 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.5391035" httpUri="https://zenodo.org/record/5391035/files/figure.png" pageId="21" pageNumber="568">Figs 5D</figureCitation>
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;
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<figureCitation id="DBB82A5D1C1DFFC403F4FBB1FE7A816D" box="[458,490,1066,1092]" captionStart="FIG" captionStartId="13.[151,162,1455,1472]" captionTargetBox="[160,1239,256,1415]" captionTargetPageId="13" captionText="FIG. 7. — Eustra Schmidt-Goebel, 1846, enveloppes tégumentaires des parties ectodermiques des pièces génitales; A, E. lebretoni Deuve, 1987, édéage, montrant la morphologie des phanères de l’endophallus. Le gonocaulis est visible; B, E. leclerci (Deuve, 1986), segment abdominal IX de la femelle, gonopodes sétifères et pièces sternales intercoxales; C, E. lebretoni Deuve, 1987, vue interne de la face ventrale des segments VIII et IX montrant les gonopodes, les formations sternales IX en position intergonopodiale, la plaque coxosternale VIII divisée en deux parties par la présence du protovagin membraneux, l’entonnoir périgonoporal; D, mêmes formations chez E. troglophila Deuve, 1987 avec seulement quelques différences de forme concernant la longueur des gonopodes et les replis en bordure du protovagin. Abréviations: gc, gonocaulis; cxst.VIII, coxosternite VIII; épl.VIII, épl.IX, épipleurite VIII et IX; gpd.IX, gonopode IX; prtvg, protovagin; spm, spermathèque; st.IX, sternite IX. Échelle: 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391043" httpUri="https://zenodo.org/record/5391043/files/figure.png" pageId="21" pageNumber="568">7A</figureCitation>
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) court mais large, le sommet rétréci en lobe à peine redressé. Endophallus avec à l’apex seulement deux sclérites dentiformes et un troisième beaucoup plus large, subtriangulaire. Paramère gauche très particulier, son extrémité bilobée. Genitalia femelles orthotopiques (
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<figureCitation id="DBB82A5D1C1DFFC4033BFB71FECE802D" box="[261,350,1258,1284]" captionStart="FIG" captionStartId="13.[151,162,1455,1472]" captionTargetBox="[160,1239,256,1415]" captionTargetPageId="13" captionText="FIG. 7. — Eustra Schmidt-Goebel, 1846, enveloppes tégumentaires des parties ectodermiques des pièces génitales; A, E. lebretoni Deuve, 1987, édéage, montrant la morphologie des phanères de l’endophallus. Le gonocaulis est visible; B, E. leclerci (Deuve, 1986), segment abdominal IX de la femelle, gonopodes sétifères et pièces sternales intercoxales; C, E. lebretoni Deuve, 1987, vue interne de la face ventrale des segments VIII et IX montrant les gonopodes, les formations sternales IX en position intergonopodiale, la plaque coxosternale VIII divisée en deux parties par la présence du protovagin membraneux, l’entonnoir périgonoporal; D, mêmes formations chez E. troglophila Deuve, 1987 avec seulement quelques différences de forme concernant la longueur des gonopodes et les replis en bordure du protovagin. Abréviations: gc, gonocaulis; cxst.VIII, coxosternite VIII; épl.VIII, épl.IX, épipleurite VIII et IX; gpd.IX, gonopode IX; prtvg, protovagin; spm, spermathèque; st.IX, sternite IX. Échelle: 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391043" httpUri="https://zenodo.org/record/5391043/files/figure.png" pageId="21" pageNumber="568">Fig. 7C</figureCitation>
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), le gonopode fusiforme mais court, sans gonostyle distinct à son extrémité apicale (
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<figureCitation id="DBB82A5D1C1DFFC402EEFAB1FEB3806D" box="[208,291,1322,1348]" captionStart="FIG" captionStartId="20.[151,162,1564,1581]" captionTargetBox="[195,1155,205,1526]" captionTargetId="figure-147@20.[227,1183,178,1544]" captionTargetPageId="20" captionText="FIG. 9. — Eustra Schmidt-Goebel, 1846. Coxostyles (gonopodes IX) des femelles. Noter la présence du gonostyle (sty), à l’exception de B-D, qui correspondent à des espèces cavernicoles du nord de la Thaïlande; A, E. japonica Bates, 1892; B, E. lebretoni Deuve, 1987; C, E. troglophila Deuve, 1987; D, E. leclerci (Deuve, 1986); E, E. honchongensis Deuve, 1996; F, E. matanga matanga Andrewes, 1919; G, E. taiwanica n. sp; H, E. chinensis Bänninger, 1949; I, E. plagiata peguensis n. subsp; J, E. crucifera Uéno, 1964; K, E. storki n. sp; L, E. andrewesiana n. sp. Échelle: 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391049" httpUri="https://zenodo.org/record/5391049/files/figure.png" pageId="21" pageNumber="568">Fig. 9B</figureCitation>
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); la spermathèque tubulaire, membraneuse, sans replis visibles, recourbée eu U (
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<figureCitation id="DBB82A5D1C1DFFC402A1FAF1FE9080AD" box="[159,256,1386,1412]" captionStart="FIG" captionStartId="25.[151,162,1001,1018]" captionTargetBox="[151,1260,185,991]" captionTargetId="figure-91@25.[152,1287,153,1019]" captionTargetPageId="25" captionText="FIG. 11. — Eustra Schmidt-Goebel, 1846, spermathèque des femelles; A, E. japonica Bates, 1892; B, E. lebretoni Deuve, 1987; C, E. honchongensis Deuve,1996; D, E. chinensis Bänninger, 1949; E, E. taiwanica n. sp; F, E. matanga matanga Andrewes, 1919; G, E. plagiata peguensis n. subsp; H, E. storki n. sp. Échelle: 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.5391055" httpUri="https://zenodo.org/record/5391055/files/figure.png" pageId="21" pageNumber="568">Fig. 11B</figureCitation>
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).
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</paragraph>
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||
</subSubSection>
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||
</treatment>
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||
</document> |