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<document id="E746882549CF7375D7EB538E734AEE0D" ID-DOI="10.11646/zootaxa.3852.4.1" ID-GBIF-Dataset="15e7f80c-042d-4537-8b2c-696136c4f427" ID-ISSN="1175-5326" ID-Zenodo-Dep="225360" ID-ZooBank="350DD9AE-B559-4DE6-94C6-EDCB90F4EAB4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460884861189" checkinUser="plazi" docAuthor="Bogorodsky, Sergey V., Alpermann, Tilman J., Mal, Ahmad O. &amp; Gabr, Mohamed H." docDate="2014" docId="B37C1D6EFFBF4845AEF2FBFDFB3B9CAA" docLanguage="en" docName="zt03852p437.pdf" docOrigin="Zootaxa 3852 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pomadasys maculatus Bloch 1793" docType="treatment" docVersion="10" lastPageNumber="425" masterDocId="4F456516FFA8485DAE65FF83FFCC9962" masterDocTitle="Survey of demersal fishes from southern Saudi Arabia, with five new records for the Red Sea" masterLastPageNumber="437" masterPageNumber="401" pageNumber="424" updateTime="1698358827164" updateUser="plazi">
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<mods:title id="2170552F9988D29DFD2AC3A936C04538">Survey of demersal fishes from southern Saudi Arabia, with five new records for the Red Sea</mods:title>
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<mods:namePart id="9DF66F396A2AC0487C8E50E856547386">Bogorodsky, Sergey V.</mods:namePart>
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<mods:namePart id="605E600D6596420490DF19161C8CB00D">Alpermann, Tilman J.</mods:namePart>
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<mods:namePart id="23020E21E14D381285A572409A5B5ED5">Mal, Ahmad O.</mods:namePart>
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<mods:namePart id="53D2E329358879F29D031029BEB8D5B1">Gabr, Mohamed H.</mods:namePart>
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<mods:title id="D7CDA312AE4A73A450D918580E4E4B9D">Zootaxa</mods:title>
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<mods:date id="FB3476B711E799023F15B03E65E03E27">2014</mods:date>
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<mods:number id="137BA3EFC9658BEB28D9A72B78222124">3852</mods:number>
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<treatment id="B37C1D6EFFBF4845AEF2FBFDFB3B9CAA" ID-DOI="http://doi.org/10.5281/zenodo.6125314" ID-GBIF-Taxon="119585961" ID-Zenodo-Dep="6125314" LSID="urn:lsid:plazi:treatment:B37C1D6EFFBF4845AEF2FBFDFB3B9CAA" httpUri="http://treatment.plazi.org/id/B37C1D6EFFBF4845AEF2FBFDFB3B9CAA" lastPageId="24" lastPageNumber="425" pageId="23" pageNumber="424">
<subSubSection id="73CFFFF3FFBF484AAEF2FBFDFF349DD8" pageId="23" pageNumber="424" type="nomenclature">
<paragraph id="3B6AAC78FFBF484AAEF2FBFDFD829DFA" blockId="23.[151,590,1150,1210]" box="[151,590,1150,1176]" pageId="23" pageNumber="424">
<heading id="60221B14FFBF484AAEF2FBFDFD829DFA" bold="true" box="[151,590,1150,1176]" fontSize="11" level="1" pageId="23" pageNumber="424" reason="1">
<taxonomicName id="FCD5D7FBFFBF484AAEF2FBFDFD829DFA" ID-CoL="4LQLP" authority="Bloch, 1793" authorityName="Bloch" authorityYear="1793" box="[151,590,1150,1176]" class="Actinopterygii" family="Haemulidae" genus="Pomadasys" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="09A1706AFFBF484AAEF2FBFDFD829DFA" bold="true" box="[151,590,1150,1176]" pageId="23" pageNumber="424">
<emphasis id="09A1706AFFBF484AAEF2FBFDFE6E9DFA" bold="true" box="[151,418,1150,1176]" italics="true" pageId="23" pageNumber="424">Pomadasys maculatus</emphasis>
(Bloch, 1793)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="3B6AAC78FFBF484AAEF2FB22FF349DD8" blockId="23.[151,590,1150,1210]" box="[151,248,1185,1210]" pageId="23" pageNumber="424">
(
<figureCitation id="A3EEB0FDFFBF484AAEFAFB22FF3D9DD8" box="[159,241,1185,1210]" captionStart="FIGURE 10" captionStartId="24.[151,250,1070,1092]" captionTargetBox="[151,1434,193,1047]" captionTargetId="figure@24.[151,1435,193,1049]" captionTargetPageId="24" captionText="FIGURE 10. Pomadasys maculatus, SMF 35117, 18.8 cm SL, Jizan, Saudi Arabia." httpUri="https://zenodo.org/record/225370/files/figure.png" pageId="23" pageNumber="424">Fig. 10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="73CFFFF3FFBF484AAEF2FB6AFF149C28" pageId="23" pageNumber="424" type="materials_examined">
<paragraph id="3B6AAC78FFBF484AAEF2FB6AFF149C28" blockId="23.[151,1437,1257,2038]" pageId="23" pageNumber="424">
<emphasis id="09A1706AFFBF484AAEF2FB6AFE4E9C60" bold="true" box="[151,386,1257,1282]" pageId="23" pageNumber="424">Material examined.</emphasis>
<collectionCode id="5DC434BDFFBF484AAFEEFB69FE0F9C60" LSID="urn:lsid:biocol.org:col:34838" box="[395,451,1258,1282]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="23" pageNumber="424">SMF</collectionCode>
35114 (1:
<quantity id="FC2D019DFFBF484AAC24FB69FD5F9C60" box="[577,659,1258,1282]" metricMagnitude="-2" metricUnit="m" metricValue="9.3" pageId="23" pageNumber="424" unit="cm" value="9.3">9.3 cm</quantity>
);
<collectionCode id="5DC434BDFFBF484AACCCFB69FD2D9C60" LSID="urn:lsid:biocol.org:col:34838" box="[681,737,1258,1282]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="23" pageNumber="424">SMF</collectionCode>
35115 (4: 8.0
<quantity id="FC2D019DFFBF484AADF4FB69FC2F9C60" box="[913,995,1258,1282]" metricMagnitude="-2" metricUnit="m" metricValue="9.4" pageId="23" pageNumber="424" unit="cm" value="9.4">9.4 cm</quantity>
);
<collectionCode id="5DC434BDFFBF484AAD9CFB69FBAB9C60" box="[1017,1127,1258,1282]" pageId="23" pageNumber="424">KAUMM</collectionCode>
156 (5:
<quantity id="FC2D019DFFBF484AAAAEFB69FA829C60" box="[1227,1358,1258,1282]" metricMagnitude="-2" metricUnit="m" metricValue="8.1" metricValueMax="8.4" metricValueMin="7.8" pageId="23" pageNumber="424" unit="cm" value="8.1" valueMax="8.4" valueMin="7.8">7.88.4 cm</quantity>
);
<collectionCode id="5DC434BDFFBF484AAB01FB69FA509C60" LSID="urn:lsid:biocol.org:col:34838" box="[1380,1436,1258,1282]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="23" pageNumber="424">SMF</collectionCode>
35117 (1:
<quantity id="FC2D019DFFBF484AAF69FA8DFEA09C44" box="[268,364,1294,1318]" metricMagnitude="-1" metricUnit="m" metricValue="1.88" pageId="23" pageNumber="424" unit="cm" value="18.8">18.8 cm</quantity>
), obtained from local fishermen at the Jizan fish market, collected in the near surroundings of Jizan.
</paragraph>
</subSubSection>
<subSubSection id="73CFFFF3FFBF484AAEA2FAD6FE3F9EE8" pageId="23" pageNumber="424" type="description">
<paragraph id="3B6AAC78FFBF484AAEA2FAD6FD499FEC" blockId="23.[151,1437,1257,2038]" pageId="23" pageNumber="424">
<emphasis id="09A1706AFFBF484AAEA2FAD6FE959C0C" bold="true" box="[199,345,1365,1390]" pageId="23" pageNumber="424">Description.</emphasis>
Dorsal-fin rays XII,14 or 15 (14: 10; 15: 1); anal-fin rays III,7; pectoral-fin rays 17; lateral-line scales 5052 (50: 2; 51: 3; 52: 6); scales between origin of dorsal fin and lateral line 9; gill rakers 5 or 6 + 13 or 14; body depth
<quantity id="FC2D019DFFBF484AAF7AFA1EFE5E9CD4" box="[287,402,1437,1462]" metricMagnitude="-2" metricUnit="m" metricValue="6.985" metricValueMax="7.366" metricValueMin="6.604" pageId="23" pageNumber="424" unit="in" value="2.75" valueMax="2.9" valueMin="2.6">2.62.9 in</quantity>
standard length; head length
<quantity id="FC2D019DFFBF484AAC82FA1DFC969CD4" box="[743,858,1437,1462]" metricMagnitude="-2" metricUnit="m" metricValue="7.747" metricValueMax="8.128" metricValueMin="7.366" pageId="23" pageNumber="424" unit="in" value="3.05" valueMax="3.2" valueMin="2.9">2.93.2 in</quantity>
standard length; snout short, the length
<quantity id="FC2D019DFFBF484AAB4CFA1DFA509CD4" box="[1321,1436,1437,1462]" metricMagnitude="-2" metricUnit="m" metricValue="9.143999999999998" metricValueMax="9.906" metricValueMin="8.382" pageId="23" pageNumber="424" unit="in" value="3.5999999999999996" valueMax="3.9" valueMin="3.3">3.33.9 in</quantity>
head length; chin with a median pit and two pores; mouth small, upper jaw just reaching to below anterior margin of eye; dorsal profile of head smoothly convex; orbit subequal to snout length, the diameter 3.34.0 in head length; preorbital depth 4.64.9 (
<quantity id="FC2D019DFFBF484AAFA0F989FDC19F40" box="[453,525,1545,1570]" metricMagnitude="-1" metricUnit="m" metricValue="1.0921999999999998" pageId="23" pageNumber="424" unit="in" value="4.3">4.3 in</quantity>
specimen SMF 35117) in head length; interorbital width 4.15.0 in (
<quantity id="FC2D019DFFBF484AAB30F989FA509F40" box="[1365,1436,1545,1570]" metricMagnitude="-2" metricUnit="m" metricValue="8.128" pageId="23" pageNumber="424" unit="in" value="3.2">3.2 in</quantity>
specimen SMF 35117) head length; fourth dorsal spine longest, the length 1.72.0 in head length; last two dorsal spines subequal; longest anal-fin spine 2.0
<quantity id="FC2D019DFFBF484AACE1F9D1FD089F08" box="[644,708,1617,1642]" metricMagnitude="-2" metricUnit="m" metricValue="5.588" pageId="23" pageNumber="424" unit="in" value="2.2">2.2 in</quantity>
head length; caudal peduncle depth
<quantity id="FC2D019DFFBF484AAA07F9D1FB189F08" box="[1122,1236,1617,1642]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="5.588" metricValueMin="4.572" pageId="23" pageNumber="424" unit="in" value="2.0" valueMax="2.2" valueMin="1.8">1.82.2 in</quantity>
its length; caudal fin slightly forked, the length
<quantity id="FC2D019DFFBF484AAF80F9F6FD9A9FEC" box="[485,598,1653,1678]" metricMagnitude="-1" metricUnit="m" metricValue="1.1683999999999999" metricValueMax="1.2953999999999999" metricValueMin="1.0413999999999999" pageId="23" pageNumber="424" unit="in" value="4.6" valueMax="5.1" valueMin="4.1">4.15.1 in</quantity>
SL.
</paragraph>
<paragraph id="3B6AAC78FFBF484AAEA2F91AFE3F9EE8" blockId="23.[151,1437,1257,2038]" pageId="23" pageNumber="424">
Color (when fresh): silvery with three yellowish brown bars, first broad, curved, from nape to below anterior part of lateral line, second curved bar below spinous dorsal fin on upper half of body, the bar is broken in three specimens (8.4, 9.3, and
<quantity id="FC2D019DFFBF484AAFD0F961FDD99F98" box="[437,533,1762,1786]" metricMagnitude="-1" metricUnit="m" metricValue="1.88" pageId="23" pageNumber="424" unit="cm" value="18.8">18.8 cm</quantity>
), the third broken bar below last spines of dorsal fin, two small blotches along back below middle of soft dorsal fin and end of fin respectively, and spot on lateral line below end of dorsal fin; upper part of opercular membrane dark yellow; a large black blotch in dorsal fin between third and sixth or seventh spines; caudal fin greyish; pelvic and anal fins semi-translucent with whitish spines; pectoral fins translucent, sometimes with yellowish hue.
</paragraph>
</subSubSection>
<subSubSection id="73CFFFF3FFBF484AAEA2F816FF289E94" pageId="23" pageNumber="424" type="distribution">
<paragraph id="3B6AAC78FFBF484AAEA2F816FF289E94" blockId="23.[151,1437,1257,2038]" pageId="23" pageNumber="424">
<emphasis id="09A1706AFFBF484AAEA2F816FEAC9ECC" bold="true" box="[199,352,1941,1966]" pageId="23" pageNumber="424">Distribution.</emphasis>
Throughout the Indian Ocean to eastern
<collectingCountry id="43C2ECE8FFBF484AAD4EF816FC5E9ECC" box="[811,914,1941,1966]" name="Australia" pageId="23" pageNumber="424">Australia</collectingCountry>
and southern
<collectingCountry id="43C2ECE8FFBF484AAA4AF815FBBE9ECF" box="[1071,1138,1942,1965]" name="Japan" pageId="23" pageNumber="424">Japan</collectingCountry>
; in the northern part of the western Indian Ocean reported from
<collectingCountry id="43C2ECE8FFBF484AAC54F839FDBA9EB0" box="[561,630,1978,2002]" name="Oman" pageId="23" pageNumber="424">Oman</collectingCountry>
(
<bibRefCitation id="5F44D189FFBF484AACE1F83AFCD09EB0" author="Randall" box="[644,796,1977,2002]" pageId="23" pageNumber="424" refString="Randall, J. E. (1995) Coastal fishes of Oman. Crowford House Publishing Pty Ltd, Bathurst, Australia, 439 pp." type="book" year="1995">Randall 1995</bibRefCitation>
) and
<collectingCountry id="43C2ECE8FFBF484AAD3CF83AFC799EB0" box="[857,949,1977,2002]" name="Somalia" pageId="23" pageNumber="424">Somalia</collectingCountry>
(
<bibRefCitation id="5F44D189FFBF484AADA6F839FB6D9EB0" author="Sommer" box="[963,1185,1977,2002]" pageId="23" pageNumber="424" refString="Sommer, C., Schneider, W. &amp; Poutiers, J. - M. (1996) The living marine resources of Somalia. FAO species identification field guide for fishery purposes, FAO, Rome, XXXII + 376 pp." type="book" year="1996">
Sommer
<emphasis id="09A1706AFFBF484AAA4FF839FBAE9EB0" box="[1066,1122,1977,2002]" italics="true" pageId="23" pageNumber="424">et al.</emphasis>
1996
</bibRefCitation>
) and the Red Sea (this study).
</paragraph>
</subSubSection>
<caption id="6FAAFCF0FFB04845AEF2FBADFC2B9D26" box="[151,999,1070,1092]" httpUri="https://zenodo.org/record/225370/files/figure.png" pageId="24" pageNumber="425" targetBox="[151,1434,193,1047]" targetPageId="24">
<paragraph id="3B6AAC78FFB04845AEF2FBADFC2B9D26" blockId="24.[151,999,1070,1092]" box="[151,999,1070,1092]" pageId="24" pageNumber="425">
<emphasis id="09A1706AFFB04845AEF2FBADFED39D21" bold="true" box="[151,287,1070,1092]" pageId="24" pageNumber="425">FIGURE 10.</emphasis>
<taxonomicName id="FCD5D7FBFFB04845AF40FBADFDCB9D26" box="[293,519,1070,1092]" class="Actinopterygii" family="Haemulidae" genus="Pomadasys" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="425" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="09A1706AFFB04845AF40FBADFDCB9D26" box="[293,519,1070,1092]" italics="true" pageId="24" pageNumber="425">Pomadasys maculatus</emphasis>
</taxonomicName>
, SMF 35117, 18.8 cm SL, Jizan, Saudi Arabia.
</paragraph>
</caption>
<subSubSection id="73CFFFF3FFB04845AEA2FBE8FB3B9CAA" pageId="24" pageNumber="425" type="discussion">
<paragraph id="3B6AAC78FFB04845AEA2FBE8FB3B9CAA" blockId="24.[151,1436,1131,1480]" pageId="24" pageNumber="425">
<emphasis id="09A1706AFFB04845AEA2FBE8FEF79DE6" bold="true" box="[199,315,1131,1156]" pageId="24" pageNumber="425">Remarks.</emphasis>
The species is found in coastal inshore waters on sandy substrata; the Red Sea record is confirmed on the basis of a dozen specimens collected by trawl off Jizan,
<collectingCountry id="43C2ECE8FFB04845AD1BFB0CFBDB9DCA" box="[894,1047,1167,1192]" name="Saudi Arabia" pageId="24" pageNumber="425">Saudi Arabia</collectingCountry>
, from a depth range of
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21
<specimenCount id="2DD367F1FFB04845AB3DFB13FA549DC5" box="[1368,1432,1168,1192]" pageId="24" pageNumber="425" type="generic">24 m</specimenCount>
</quantity>
.
<bibRefCitation id="5F44D189FFB04845AEF2FB30FE269DAE" author="Fowler" box="[151,490,1203,1228]" pageId="24" pageNumber="425" refString="Fowler, H. W. &amp; Steinitz, H. (1956) Fishes from Cyprus, Iran, Iraq, Israel and Oman. Bulletin of the Research Council of Israel, 5 B (3 / 4), 260 - 292." type="journal article" year="1956">Fowler &amp; Steinitz (1956: 276)</bibRefCitation>
listed a single specimen of
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SL from Eilat without a description (at present this specimen is lost, D. Golani, pers. comm.);
<bibRefCitation id="5F44D189FFB04845ACFCFB54FC3E9D92" author="Golani" box="[665,1010,1239,1264]" pageId="24" pageNumber="425" refString="Golani, D. &amp; Bogorodsky, S. V. (2010) The fishes of the Red Sea - reappraisal and updated checklist. Zootaxa, 2463, 1 - 135." type="journal article" year="2010">Golani &amp; Bogorodsky (2010)</bibRefCitation>
did not include the species in their checklist, and noted that “all previous records (see
<bibRefCitation id="5F44D189FFB04845ACA8FB7FFCF49C76" author="Dor" box="[717,824,1276,1300]" pageId="24" pageNumber="425" refString="Dor, M. (1984) Checklist of the Fishes of the Red Sea. Israel Academy of Sciences and Humanities, Jerusalem, xxi + 427 pp." type="book" year="1984">Dor 1984</bibRefCitation>
) are probably misidentifications of
<taxonomicName id="FCD5D7FBFFB04845AAA0FB78FE8D9C5A" authority="Cuvier, 1830" authorityName="Cuvier" authorityYear="1830" class="Actinopterygii" family="Haemulidae" genus="Pomadasys" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="425" phylum="Chordata" rank="species" species="kaakan">
<emphasis id="09A1706AFFB04845AAA0FB78FA509C76" box="[1221,1436,1275,1300]" italics="true" pageId="24" pageNumber="425">Pomadasys kaakan</emphasis>
(Cuvier, 1830)
</taxonomicName>
”. The largest specimen of
<quantity id="FC2D019DFFB04845AC10FAA3FD1D9C55" box="[629,721,1312,1336]" metricMagnitude="-1" metricUnit="m" metricValue="1.88" pageId="24" pageNumber="425" unit="cm" value="18.8">18.8 cm</quantity>
SL has a deeper preorbital and a broader interorbital space in comparison with smaller specimens; more specimens are required to see the dynamics of changes of these two characters with growth. The species can be distinguished easily from other Red Sea congeners
<taxonomicName id="FCD5D7FBFFB04845AB79FAE4FE049CC6" authority="Forsskal, 1775" authorityName="Forsskal" authorityYear="1775" class="Actinopterygii" family="Haemulidae" genus="Pomadasys" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="425" phylum="Chordata" rank="species" species="argenteus">
<emphasis id="09A1706AFFB04845AB79FAE4FEC89CC6" italics="true" pageId="24" pageNumber="425">Pomadasys argenteus</emphasis>
(Forsskål, 1775)
</taxonomicName>
,
<emphasis id="09A1706AFFB04845AFB3FA0EFD8D9CC6" box="[470,577,1419,1444]" italics="true" pageId="24" pageNumber="425">P. k a ak a n</emphasis>
,
<taxonomicName id="FCD5D7FBFFB04845AC31FA0EFC7D9CC6" authority="Ruppell, 1838" authorityName="Ruppell" authorityYear="1838" box="[596,945,1419,1444]" class="Actinopterygii" family="Haemulidae" genus="Pomadasys" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="425" phylum="Chordata" rank="species" species="punctulatus">
<emphasis id="09A1706AFFB04845AC31FA0EFD389CC6" box="[596,756,1419,1444]" italics="true" pageId="24" pageNumber="425">P. punctulatus</emphasis>
(Rüppell, 1838)
</taxonomicName>
, and
<taxonomicName id="FCD5D7FBFFB04845AD95FA0EFAE49CC6" authority="Forsskal, 1775" authorityName="Forsskal" authorityYear="1775" box="[1008,1320,1419,1444]" class="Actinopterygii" family="Haemulidae" genus="Pomadasys" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="425" phylum="Chordata" rank="species" species="stridens">
<emphasis id="09A1706AFFB04845AD95FA0EFBAB9CC6" box="[1008,1127,1419,1444]" italics="true" pageId="24" pageNumber="425">P. stridens</emphasis>
(Forsskål, 1775)
</taxonomicName>
in having three dark bars dorsally on the body and a large black blotch in the spinous portion of the dorsal fin.
</paragraph>
</subSubSection>
</treatment>
</document>