562 lines
61 KiB
XML
562 lines
61 KiB
XML
<document id="3BC192E857C6FFB3AB0CDC48C8D3EA31" ID-CLB-Dataset="29764" ID-DOI="10.11646/zootaxa.4442.2.5" ID-GBIF-Dataset="0fea0081-d093-4611-89c2-316f9baf6ca4" ID-ISSN="1175-5326" ID-Zenodo-Dep="1303125" ID-ZooBank="8EB2105E-3EAA-444A-A20E-819AD215B3C3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1530533589605" checkinUser="plazi" docAuthor="Khaustov, Alexander A. & Frolov, Andrey V." docDate="2018" docId="A1192263FE56BD36C7E104F3437CF81A" docLanguage="en" docName="zootaxa.4442.2.5.pdf" docOrigin="Zootaxa 4442 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Spatulaphorus enoplotruporum Khaustov & Frolov 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="281" masterDocId="5D205A1BFE57BD32C776016A461FFFCF" masterDocTitle="New taxa of pygmephoroid mites (Acari: Pygmephoroidea: Pygmephoridae, Neopygmephoridae) phoretic on Enoplotrupes sharpi (Coleoptera: Geotrupidae) from Thailand" masterLastPageNumber="292" masterPageNumber="277" pageNumber="278" updateTime="1698552142957" updateUser="ExternalLinkService">
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<mods:titleInfo id="39493F39FD54379FCD12EF9D2DA2A410">
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<mods:title id="A2DC57230584D30FD1AE9B7230C4EEED">New taxa of pygmephoroid mites (Acari: Pygmephoroidea: Pygmephoridae, Neopygmephoridae) phoretic on Enoplotrupes sharpi (Coleoptera: Geotrupidae) from Thailand</mods:title>
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<mods:namePart id="625688A9E6338E19C31970F5E33EC1CF">Khaustov, Alexander A.</mods:namePart>
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<mods:name id="78DD41A88248010EF21F5D6969473ABA" type="personal">
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<mods:namePart id="F98331003808B03BC0B289D4C2BA4CFF">Frolov, Andrey V.</mods:namePart>
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<mods:titleInfo id="6155753C3D85674E5EC47C41916E7D02">
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<mods:title id="CA6D97BF073797ECFE3C5A680C54187B">Zootaxa</mods:title>
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<mods:part id="88A3093F4D52AEE48F759863F6AB9D30">
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<mods:date id="586DAF724DE2CD007C18855D51F2FD9D">2018</mods:date>
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<mods:number id="FCADB3C8D326167446439866D78FDF47">2018-07-02</mods:number>
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<mods:number id="EBC0089681F6AB9F11552BAB38133EE7">4442</mods:number>
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<mods:identifier id="132CD7B6F12039DF947F9146CCF0134D" type="DOI">10.11646/zootaxa.4442.2.5</mods:identifier>
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<treatment id="A1192263FE56BD36C7E104F3437CF81A" ID-DOI="http://doi.org/10.5281/zenodo.5949607" ID-GBIF-Taxon="146044081" ID-Zenodo-Dep="5949607" LSID="urn:lsid:plazi:treatment:A1192263FE56BD36C7E104F3437CF81A" httpUri="http://treatment.plazi.org/id/A1192263FE56BD36C7E104F3437CF81A" lastPageId="4" lastPageNumber="281" pageId="1" pageNumber="278">
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<subSubSection id="61AAC0FEFE56BD33C7E104F34715FA1A" pageId="1" pageNumber="278" type="nomenclature">
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<paragraph id="290F9375FE56BD33C7E104F34465FA7C" blockId="1.[151,634,1433,1493]" box="[151,634,1433,1459]" pageId="1" pageNumber="278">
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<heading id="72472419FE56BD33C7E104F34465FA7C" bold="true" box="[151,634,1433,1459]" fontSize="11" level="1" pageId="1" pageNumber="278" reason="1">
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<emphasis id="1BC44F67FE56BD33C7E104F34465FA7C" bold="true" box="[151,634,1433,1459]" pageId="1" pageNumber="278">
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<taxonomicName id="EEB0E8F6FE56BD33C7E104F3440AFA7C" authority="Khaustov & Frolov, 2018" authorityName="Khaustov & Frolov" authorityYear="2018" box="[151,533,1433,1459]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="278" phylum="Arthropoda" rank="species" species="enoplotruporum" status="sp. nov.">
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<emphasis id="1BC44F67FE56BD33C7E104F3440AFA7C" bold="true" box="[151,533,1433,1459]" italics="true" pageId="1" pageNumber="278">Spatulaphorus enoplotruporum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="00F7F21CFE56BD33C56A04F34465FA7C" box="[540,634,1433,1459]" pageId="1" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="290F9375FE56BD33C7E104D64715FA1A" blockId="1.[151,634,1433,1493]" box="[151,266,1468,1493]" pageId="1" pageNumber="278">
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(
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<figureCitation id="B18B8FF0FE56BD33C7E904D6471EFA1A" box="[159,257,1468,1493]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1393,1415]" captionStartId-1="3.[151,250,1954,1976]" captionStartId-2="4.[151,250,1330,1352]" captionStartId-3="5.[151,250,1546,1568]" captionStartId-4="7.[151,250,1804,1826]" captionTargetBox-0="[179,1414,369,1343]" captionTargetBox-1="[209,1332,690,1862]" captionTargetBox-2="[160,1411,246,1289]" captionTargetBox-3="[191,1406,208,1529]" captionTargetBox-4="[151,1436,180,1782]" captionTargetId-0="figure@2.[151,1436,312,1369]" captionTargetId-1="figure@3.[181,1405,643,1923]" captionTargetId-2="figure@4.[151,1436,193,1309]" captionTargetId-3="figure@5.[167,1421,193,1533]" captionTargetId-4="figure@7.[151,1436,180,1782]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="7" captionText-0="FIGURE 1. Spatulaphorus enoplotruporum sp. nov., female: A—dorsum of the body, B—venter of the body. Legs omitted." captionText-1="FIGURE 2. Spatulaphorus enoplotruporum sp. nov., female: A—gnathosoma in dorsal view, B—gnathosoma in ventral view, C—pharyngeal pumps." captionText-2="FIGURE 3. Spatulaphorus enoplotruporum sp. nov., female: A—right leg I in dorsal view, B—right leg II in dorsal view." captionText-3="FIGURE 4. Spatulaphorus enoplotruporum sp. nov., female: A—right leg III in dorsal view, B—right leg IV in dorsal view." captionText-4="FIGURE 5. DIC micrographs of Spatulaphorus enoplotruporum sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri-0="https://zenodo.org/record/1303127/files/figure.png" httpUri-1="https://zenodo.org/record/1303129/files/figure.png" httpUri-2="https://zenodo.org/record/1303131/files/figure.png" httpUri-3="https://zenodo.org/record/1303133/files/figure.png" httpUri-4="https://zenodo.org/record/1303135/files/figure.png" pageId="1" pageNumber="278">Figs 1–5</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="61AAC0FEFE56BD31C7E1076E4792FDF0" lastPageId="3" lastPageNumber="280" pageId="1" pageNumber="278" type="description">
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<paragraph id="290F9375FE56BD33C7E1076E45BEF8F3" blockId="1.[151,1437,1540,2039]" pageId="1" pageNumber="278">
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<emphasis id="1BC44F67FE56BD33C7E1076E473EF9D2" bold="true" box="[151,289,1540,1565]" pageId="1" pageNumber="278">Description</emphasis>
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. FEMALE (
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<figureCitation id="B18B8FF0FE56BD33C6C9076E4433F9D2" box="[447,556,1540,1565]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1393,1415]" captionStartId-1="3.[151,250,1954,1976]" captionStartId-2="4.[151,250,1330,1352]" captionStartId-3="5.[151,250,1546,1568]" captionStartId-4="7.[151,250,1804,1826]" captionTargetBox-0="[179,1414,369,1343]" captionTargetBox-1="[209,1332,690,1862]" captionTargetBox-2="[160,1411,246,1289]" captionTargetBox-3="[191,1406,208,1529]" captionTargetBox-4="[151,1436,180,1782]" captionTargetId-0="figure@2.[151,1436,312,1369]" captionTargetId-1="figure@3.[181,1405,643,1923]" captionTargetId-2="figure@4.[151,1436,193,1309]" captionTargetId-3="figure@5.[167,1421,193,1533]" captionTargetId-4="figure@7.[151,1436,180,1782]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="7" captionText-0="FIGURE 1. Spatulaphorus enoplotruporum sp. nov., female: A—dorsum of the body, B—venter of the body. Legs omitted." captionText-1="FIGURE 2. Spatulaphorus enoplotruporum sp. nov., female: A—gnathosoma in dorsal view, B—gnathosoma in ventral view, C—pharyngeal pumps." captionText-2="FIGURE 3. Spatulaphorus enoplotruporum sp. nov., female: A—right leg I in dorsal view, B—right leg II in dorsal view." captionText-3="FIGURE 4. Spatulaphorus enoplotruporum sp. nov., female: A—right leg III in dorsal view, B—right leg IV in dorsal view." captionText-4="FIGURE 5. DIC micrographs of Spatulaphorus enoplotruporum sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri-0="https://zenodo.org/record/1303127/files/figure.png" httpUri-1="https://zenodo.org/record/1303129/files/figure.png" httpUri-2="https://zenodo.org/record/1303131/files/figure.png" httpUri-3="https://zenodo.org/record/1303133/files/figure.png" httpUri-4="https://zenodo.org/record/1303135/files/figure.png" pageId="1" pageNumber="278">Figs 1–5</figureCitation>
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). Body well sclerotized. Length of idiosoma 205 (175–240), width 120 (100–140).
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<emphasis id="1BC44F67FE56BD33C66C074D47B6F98F" box="[282,425,1575,1600]" italics="true" pageId="1" pageNumber="278">Gnathosoma</emphasis>
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(
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<figureCitation id="B18B8FF0FE56BD33C6CF074D47E1F98F" box="[441,510,1575,1600]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[209,1332,690,1862]" captionTargetId="figure@3.[181,1405,643,1923]" captionTargetPageId="3" captionText="FIGURE 2. Spatulaphorus enoplotruporum sp. nov., female: A—gnathosoma in dorsal view, B—gnathosoma in ventral view, C—pharyngeal pumps." httpUri="https://zenodo.org/record/1303129/files/figure.png" pageId="1" pageNumber="278">Fig. 2</figureCitation>
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). Gnathosomal capsule, excluding palps, subrectangular, length 22 (19–24), width 22 (19–24). Dorsum with one pair of needle-like cheliceral setae (
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<emphasis id="1BC44F67FE56BD33C4E6072645A6F9AA" box="[912,953,1612,1637]" italics="true" pageId="1" pageNumber="278">сha</emphasis>
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) 7 (5–7). Dorsal median apodeme absent. Postpalpal setae (
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<emphasis id="1BC44F67FE56BD33C613071B479EF947" box="[357,385,1649,1672]" italics="true" pageId="1" pageNumber="278">pp</emphasis>
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) spine-like, situated laterally to setae
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<emphasis id="1BC44F67FE56BD33C43F0705456EF947" box="[841,881,1647,1672]" italics="true" pageId="1" pageNumber="278">сha</emphasis>
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. Ventral gnathosoma with one pair of smooth, pointed subcapitular setae
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<emphasis id="1BC44F67FE56BD33C6B507FF47C8F963" box="[451,471,1685,1708]" italics="true" pageId="1" pageNumber="278">m</emphasis>
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6 (5–7) and round pits situated posteriad to
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<emphasis id="1BC44F67FE56BD33C4B807FF45FDF963" box="[974,994,1685,1708]" italics="true" pageId="1" pageNumber="278">m</emphasis>
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. Palps with smooth setae
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<emphasis id="1BC44F67FE56BD33C27C07FE4329F962" box="[1290,1334,1684,1709]" italics="true" pageId="1" pageNumber="278">dFe</emphasis>
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and
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<emphasis id="1BC44F67FE56BD33C21B07FE4383F962" box="[1389,1436,1684,1709]" italics="true" pageId="1" pageNumber="278">dGe</emphasis>
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dorsolaterally, setae
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<emphasis id="1BC44F67FE56BD33C6F607DD47B0F91F" box="[384,431,1719,1744]" italics="true" pageId="1" pageNumber="278">dGe</emphasis>
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pointed, about twice longer than needle-like
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<emphasis id="1BC44F67FE56BD33C4C007DD45FDF91F" box="[950,994,1719,1744]" italics="true" pageId="1" pageNumber="278">dFe</emphasis>
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. Palps ventrally each with mushroomlike accessory setigenous structure (
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<emphasis id="1BC44F67FE56BD33C54A07B7447FF93B" box="[572,608,1757,1780]" italics="true" pageId="1" pageNumber="278">ass</emphasis>
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) and tiny solenidion. Palps terminated with small tibial claw and tiny eupathidium-like seta. Pharyngeal pump 2 with weak transverse striation; pharyngeal pump 1 small, butterfly-like; pharyngeal pump 3 oval, slightly smaller than oval pump 2 (
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<figureCitation id="B18B8FF0FE56BD33C44D064E458BF8F3" box="[827,916,1828,1853]" captionStart="FIGURE 2" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[209,1332,690,1862]" captionTargetId="figure@3.[181,1405,643,1923]" captionTargetPageId="3" captionText="FIGURE 2. Spatulaphorus enoplotruporum sp. nov., female: A—gnathosoma in dorsal view, B—gnathosoma in ventral view, C—pharyngeal pumps." httpUri="https://zenodo.org/record/1303129/files/figure.png" pageId="1" pageNumber="278">Fig. 2C</figureCitation>
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).
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</paragraph>
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<paragraph id="290F9375FE56BD30C7B1062D4319FF31" blockId="1.[151,1437,1540,2039]" lastBlockId="2.[151,1435,151,258]" lastPageId="2" lastPageNumber="279" pageId="1" pageNumber="278">
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<emphasis id="1BC44F67FE56BD33C7B1062D478CF8AF" box="[199,403,1863,1888]" italics="true" pageId="1" pageNumber="278">Idiosomal dorsum</emphasis>
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(
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<figureCitation id="B18B8FF0FE56BD33C6D5062D441DF8AF" box="[419,514,1863,1888]" captionStart="FIGURE 1" captionStartId="2.[151,250,1393,1415]" captionTargetBox="[179,1414,369,1343]" captionTargetId="figure@2.[151,1436,312,1369]" captionTargetPageId="2" captionText="FIGURE 1. Spatulaphorus enoplotruporum sp. nov., female: A—dorsum of the body, B—venter of the body. Legs omitted." httpUri="https://zenodo.org/record/1303127/files/figure.png" pageId="1" pageNumber="278">Figs 1A</figureCitation>
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,
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<figureCitation id="B18B8FF0FE56BD33C566062D4483F8AF" box="[528,668,1863,1888]" captionStart="FIGURE 5" captionStartId="7.[151,250,1804,1826]" captionTargetBox="[151,1436,180,1782]" captionTargetId="figure@7.[151,1436,180,1782]" captionTargetPageId="7" captionText="FIGURE 5. DIC micrographs of Spatulaphorus enoplotruporum sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri="https://zenodo.org/record/1303135/files/figure.png" pageId="1" pageNumber="278">5A, C, E, G</figureCitation>
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). All dorsal shields with numerous tiny, barely discernable dimples (
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<figureCitation id="B18B8FF0FE56BD33C7E90606476FF84B" box="[159,368,1900,1925]" captionStart="FIGURE 5" captionStartId="7.[151,250,1804,1826]" captionTargetBox="[151,1436,180,1782]" captionTargetId="figure@7.[151,1436,180,1782]" captionTargetPageId="7" captionText="FIGURE 5. DIC micrographs of Spatulaphorus enoplotruporum sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri="https://zenodo.org/record/1303135/files/figure.png" pageId="1" pageNumber="278">Figs 5A, C, E, G</figureCitation>
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). Prodorsal shield with three pairs of setae (
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<date id="5D0EB5B5FE56BD33C40B0607458DF846" box="[893,914,1901,1929]" pageId="1" pageNumber="278">
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<emphasis id="1BC44F67FE56BD33C40B06074596F84B" box="[893,905,1901,1924]" italics="true" pageId="1" pageNumber="278">v</emphasis>
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1
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</date>
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,
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<date id="5D0EB5B5FE56BD33C4D2060745A7F846" box="[932,952,1901,1929]" pageId="1" pageNumber="278">
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<emphasis id="1BC44F67FE56BD33C4D2060745AFF84B" box="[932,944,1901,1924]" italics="true" pageId="1" pageNumber="278">v</emphasis>
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2
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</date>
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,
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<emphasis id="1BC44F67FE56BD33C4BC060745FEF84B" box="[970,993,1901,1924]" italics="true" pageId="1" pageNumber="278">sс</emphasis>
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2), one pair of weakly barbed capitate trichobothria and one pair of round stigmata (
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<figureCitation id="B18B8FF0FE56BD33C5D606F8451DF864" box="[672,770,1938,1963]" captionStart="FIGURE 5" captionStartId="7.[151,250,1804,1826]" captionTargetBox="[151,1436,180,1782]" captionTargetId="figure@7.[151,1436,180,1782]" captionTargetPageId="7" captionText="FIGURE 5. DIC micrographs of Spatulaphorus enoplotruporum sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri="https://zenodo.org/record/1303135/files/figure.png" pageId="1" pageNumber="278">Fig. 5 C</figureCitation>
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). Stigmata associated with well developed tracheal trunks. All dorsal idiosomal setae blunt-ended and barbed. Cupules
|
||
<emphasis id="1BC44F67FE56BD33C43F06D24540F800" box="[841,863,1976,1999]" italics="true" pageId="1" pageNumber="278">ia</emphasis>
|
||
on tergite D,
|
||
<emphasis id="1BC44F67FE56BD33C37606D24203F800" box="[1024,1052,1976,1999]" italics="true" pageId="1" pageNumber="278">im</emphasis>
|
||
on tergite EF and
|
||
<emphasis id="1BC44F67FE56BD33C38206DC4315F800" box="[1268,1290,1974,1999]" italics="true" pageId="1" pageNumber="278">ih</emphasis>
|
||
on tergite H large, round. Tergites D, EF and H each with one pair of oval porous areas. Lengths of dorsal setae:
|
||
<date id="5D0EB5B5FE56BD33C38B06B6432AF83C" box="[1277,1333,2011,2039]" pageId="1" pageNumber="278" value="1923-05-01">
|
||
<emphasis id="1BC44F67FE56BD33C38B06B64316F83C" box="[1277,1289,2012,2035]" italics="true" pageId="1" pageNumber="278">v</emphasis>
|
||
1 23
|
||
</date>
|
||
(21–23),
|
||
<date id="5D0EB5B5FE55BD30C7E101F346D1FF7F" box="[151,206,152,180]" pageId="2" pageNumber="279" value="2013-05-02">
|
||
<emphasis id="1BC44F67FE55BD30C7E101F346BCFF7F" box="[151,163,153,176]" italics="true" pageId="2" pageNumber="279">v</emphasis>
|
||
2 13
|
||
</date>
|
||
(10–13),
|
||
<emphasis id="1BC44F67FE55BD30C64C01F3474EFF7F" box="[314,337,153,176]" italics="true" pageId="2" pageNumber="279">sс</emphasis>
|
||
2 49 (45–54),
|
||
<emphasis id="1BC44F67FE55BD30C69F01F347EAFF7F" box="[489,501,153,176]" italics="true" pageId="2" pageNumber="279">с</emphasis>
|
||
1 37 (30–38),
|
||
<emphasis id="1BC44F67FE55BD30C5FF01F3448AFF7F" box="[649,661,153,176]" italics="true" pageId="2" pageNumber="279">с</emphasis>
|
||
2 49 (44–54),
|
||
<emphasis id="1BC44F67FE55BD30C45B01FD4524FF7F" box="[813,827,151,176]" italics="true" pageId="2" pageNumber="279">d</emphasis>
|
||
42 (39–48),
|
||
<emphasis id="1BC44F67FE55BD30C4BC01F345C9FF7F" box="[970,982,153,176]" italics="true" pageId="2" pageNumber="279">e</emphasis>
|
||
25 (23–31),
|
||
<emphasis id="1BC44F67FE55BD30C31001F24271FF7F" box="[1126,1134,152,176]" italics="true" pageId="2" pageNumber="279">f</emphasis>
|
||
54 (44–58),
|
||
<emphasis id="1BC44F67FE55BD30C38B01FD4314FF7F" box="[1277,1291,151,176]" italics="true" pageId="2" pageNumber="279">h</emphasis>
|
||
1 51 (45–57),
|
||
<emphasis id="1BC44F67FE55BD30C7E101D446BAFF18" box="[151,165,190,215]" italics="true" pageId="2" pageNumber="279">h</emphasis>
|
||
2 16 (16–21). Distances between setae:
|
||
<date id="5D0EB5B5FE55BD30C51201AA44A0FF18" box="[612,703,191,219]" pageId="2" pageNumber="279" value="1922-05-01" valueMax="1922-05-01" valueMin="1922-05-01">
|
||
<emphasis id="1BC44F67FE55BD30C51201AA446FFF18" box="[612,624,192,215]" italics="true" pageId="2" pageNumber="279">v</emphasis>
|
||
1–
|
||
<emphasis id="1BC44F67FE55BD30C5F001AA448DFF18" box="[646,658,192,215]" italics="true" pageId="2" pageNumber="279">v</emphasis>
|
||
1 22
|
||
</date>
|
||
(20–23),
|
||
<date id="5D0EB5B5FE55BD30C45901AA4595FF18" box="[815,906,191,219]" pageId="2" pageNumber="279" value="1939-05-02" valueMax="1939-05-02" valueMin="1939-05-02">
|
||
<emphasis id="1BC44F67FE55BD30C45901AA4524FF18" box="[815,827,192,215]" italics="true" pageId="2" pageNumber="279">v</emphasis>
|
||
2
|
||
<emphasis id="1BC44F67FE55BD30C43501AA4541FF18" box="[835,862,192,215]" italics="true" pageId="2" pageNumber="279">–v</emphasis>
|
||
2 39
|
||
</date>
|
||
(34–42),
|
||
<emphasis id="1BC44F67FE55BD30C48C01AA420EFF18" box="[1018,1041,192,215]" italics="true" pageId="2" pageNumber="279">sс</emphasis>
|
||
2
|
||
<emphasis id="1BC44F67FE55BD30C36F01AA4221FF18" box="[1049,1086,192,215]" italics="true" pageId="2" pageNumber="279">–sс</emphasis>
|
||
2 47 (39–49),
|
||
<emphasis id="1BC44F67FE55BD30C3AD01AA42F8FF18" box="[1243,1255,192,215]" italics="true" pageId="2" pageNumber="279">с</emphasis>
|
||
1–
|
||
<emphasis id="1BC44F67FE55BD30C38B01AA4316FF18" box="[1277,1289,192,215]" italics="true" pageId="2" pageNumber="279">с</emphasis>
|
||
1 46 (39–47),
|
||
<emphasis id="1BC44F67FE55BD30C7E1018C46BCFF32" box="[151,163,230,253]" italics="true" pageId="2" pageNumber="279">с</emphasis>
|
||
1–
|
||
<emphasis id="1BC44F67FE55BD30C7CF018C46DAFF32" box="[185,197,230,253]" italics="true" pageId="2" pageNumber="279">с</emphasis>
|
||
2 27 (23–29),
|
||
<emphasis id="1BC44F67FE55BD30C628018F4797FF31" box="[350,392,229,254]" italics="true" pageId="2" pageNumber="279">d–d</emphasis>
|
||
68 (55–74),
|
||
<emphasis id="1BC44F67FE55BD30C56E018C4424FF31" box="[536,571,230,254]" italics="true" pageId="2" pageNumber="279">e–f</emphasis>
|
||
17 (14–17),
|
||
<emphasis id="1BC44F67FE55BD30C5BD018C44F6FF31" box="[715,745,230,254]" italics="true" pageId="2" pageNumber="279">f–f</emphasis>
|
||
40 (31–46),
|
||
<emphasis id="1BC44F67FE55BD30C40F018F4598FF31" box="[889,903,229,254]" italics="true" pageId="2" pageNumber="279">h</emphasis>
|
||
1–
|
||
<emphasis id="1BC44F67FE55BD30C4EB018F45B4FF31" box="[925,939,229,254]" italics="true" pageId="2" pageNumber="279">h</emphasis>
|
||
1 26 (22–32),
|
||
<emphasis id="1BC44F67FE55BD30C335018F424EFF31" box="[1091,1105,229,254]" italics="true" pageId="2" pageNumber="279">h</emphasis>
|
||
1–
|
||
<emphasis id="1BC44F67FE55BD30C311018F426AFF31" box="[1127,1141,229,254]" italics="true" pageId="2" pageNumber="279">h</emphasis>
|
||
2 18 (14–20).
|
||
</paragraph>
|
||
<caption id="7DCFC3FDFE55BD30C7E1041B439CFA48" box="[151,1411,1393,1415]" httpUri="https://zenodo.org/record/1303127/files/figure.png" pageId="2" pageNumber="279" startId="2.[151,250,1393,1415]" targetBox="[179,1414,369,1343]" targetPageId="2">
|
||
<paragraph id="290F9375FE55BD30C7E1041B439CFA48" blockId="2.[151,1411,1393,1415]" box="[151,1411,1393,1415]" pageId="2" pageNumber="279">
|
||
<emphasis id="1BC44F67FE55BD30C7E1041B4712FA49" bold="true" box="[151,269,1393,1415]" pageId="2" pageNumber="279">FIGURE 1</emphasis>
|
||
.
|
||
<taxonomicName id="EEB0E8F6FE55BD30C66F041B4447FA48" authority="Khaustov & Frolov, 2018" authorityName="Khaustov & Frolov" authorityYear="2018" box="[281,600,1393,1415]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="enoplotruporum" status="sp. nov.">
|
||
<emphasis id="1BC44F67FE55BD30C66F041B4447FA48" box="[281,600,1393,1415]" italics="true" pageId="2" pageNumber="279">Spatulaphorus enoplotruporum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="00F7F21CFE55BD30C528041B44B1FA49" box="[606,686,1393,1414]" pageId="2" pageNumber="279" rank="species">
|
||
<emphasis id="1BC44F67FE55BD30C528041B44B1FA49" bold="true" box="[606,686,1393,1414]" pageId="2" pageNumber="279">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, female: A—dorsum of the body, B—venter of the body. Legs omitted.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="290F9375FE55BD30C7B104C44515F8FB" blockId="2.[151,1437,1454,2027]" pageId="2" pageNumber="279">
|
||
<emphasis id="1BC44F67FE55BD30C7B104C44799FA09" box="[199,390,1454,1479]" italics="true" pageId="2" pageNumber="279">Idiosomal venter</emphasis>
|
||
(
|
||
<figureCitation id="B18B8FF0FE55BD30C6EE04C447E9FA08" box="[408,502,1454,1479]" captionStart="FIGURE 1" captionStartId="2.[151,250,1393,1415]" captionTargetBox="[179,1414,369,1343]" captionTargetId="figure@2.[151,1436,312,1369]" captionTargetPageId="2" captionText="FIGURE 1. Spatulaphorus enoplotruporum sp. nov., female: A—dorsum of the body, B—venter of the body. Legs omitted." httpUri="https://zenodo.org/record/1303127/files/figure.png" pageId="2" pageNumber="279">Figs 1B</figureCitation>
|
||
,
|
||
<figureCitation id="B18B8FF0FE55BD30C57304C444B5FA08" box="[517,682,1454,1479]" captionStart="FIGURE 5" captionStartId="7.[151,250,1804,1826]" captionTargetBox="[151,1436,180,1782]" captionTargetId="figure@7.[151,1436,180,1782]" captionTargetPageId="7" captionText="FIGURE 5. DIC micrographs of Spatulaphorus enoplotruporum sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri="https://zenodo.org/record/1303135/files/figure.png" pageId="2" pageNumber="279">5B, D, F, H, I</figureCitation>
|
||
). All ventral plates with numerous tiny dimples. Setae 1
|
||
<emphasis id="1BC44F67FE55BD30C24E04C44359FA08" box="[1336,1350,1454,1479]" italics="true" pageId="2" pageNumber="279">b</emphasis>
|
||
and 2
|
||
<emphasis id="1BC44F67FE55BD30C2F904C54384FA09" box="[1423,1435,1455,1478]" italics="true" pageId="2" pageNumber="279">с</emphasis>
|
||
pointed; other ventral setae with slightly rounded tips. Setae 1
|
||
<emphasis id="1BC44F67FE55BD30C40604B84561FA24" box="[880,894,1490,1515]" italics="true" pageId="2" pageNumber="279">b</emphasis>
|
||
and
|
||
<emphasis id="1BC44F67FE55BD30C4CA04B945CAFA25" box="[956,981,1491,1514]" italics="true" pageId="2" pageNumber="279">ps</emphasis>
|
||
3 with a few barbs; setae
|
||
<emphasis id="1BC44F67FE55BD30C27504B94303FA25" box="[1283,1308,1491,1514]" italics="true" pageId="2" pageNumber="279">ps</emphasis>
|
||
1, 2 strongly barbed; other ventral setae smooth. Bases of setae
|
||
<emphasis id="1BC44F67FE55BD30C5A2049044F2F9DE" box="[724,749,1530,1553]" italics="true" pageId="2" pageNumber="279">ps</emphasis>
|
||
3 situated distinctly anteriad bases of setae
|
||
<emphasis id="1BC44F67FE55BD30C3A4049042F5F9DE" box="[1234,1258,1530,1553]" italics="true" pageId="2" pageNumber="279">ps</emphasis>
|
||
1-2. Apodemes 1 (ap1) and apodemes 2 (ap2) well developed and joined with thick prosternal apodeme (appr); sejugal apodeme (apsej) developed only laterally; apodemes 3 (ap3) weak, not reaching to poststernal apodeme (appo); apodemes 4 (ap4) well developed, exceeding beyond bases of setae 3
|
||
<emphasis id="1BC44F67FE55BD30C47A070D4505F94F" box="[780,794,1639,1664]" italics="true" pageId="2" pageNumber="279">b</emphasis>
|
||
. Apodemes 5 (ap5) well developed, joined with appo and reaching beyond setae 4
|
||
<emphasis id="1BC44F67FE55BD30C6D207E747ADF96B" box="[420,434,1677,1700]" italics="true" pageId="2" pageNumber="279">a</emphasis>
|
||
. Anterior genital sclerite (ags) small, cup-like; posterior genital sclerite (pgs) small, oval, with lateral processes (
|
||
<figureCitation id="B18B8FF0FE55BD30C6D507C5440BF907" box="[419,532,1711,1736]" captionStart="FIGURE 5" captionStartId="7.[151,250,1804,1826]" captionTargetBox="[151,1436,180,1782]" captionTargetId="figure@7.[151,1436,180,1782]" captionTargetPageId="7" captionText="FIGURE 5. DIC micrographs of Spatulaphorus enoplotruporum sp. nov., female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites." httpUri="https://zenodo.org/record/1303135/files/figure.png" pageId="2" pageNumber="279">
|
||
Figure
|
||
<date id="5D0EB5B5FE55BD30C68C07C5440BF907" box="[506,532,1711,1736]" pageId="2" pageNumber="279">5I</date>
|
||
</figureCitation>
|
||
). Posterior margins of posterior sternal plate weakly concave in middle part. Posterior margin of aggenital plate rounded. Lengths of ventral setae: 1
|
||
<emphasis id="1BC44F67FE55BD30C4B607BF45D1F923" box="[960,974,1749,1772]" italics="true" pageId="2" pageNumber="279">a</emphasis>
|
||
9 (9–10), 1
|
||
<emphasis id="1BC44F67FE55BD30C32207BE427DF922" box="[1108,1122,1748,1773]" italics="true" pageId="2" pageNumber="279">b</emphasis>
|
||
17 (15–21), 1
|
||
<emphasis id="1BC44F67FE55BD30C27207BF430FF923" box="[1284,1296,1749,1772]" italics="true" pageId="2" pageNumber="279">с</emphasis>
|
||
11 (10–12), 2
|
||
<emphasis id="1BC44F67FE55BD30C7D3079346ACF8DF" box="[165,179,1785,1808]" italics="true" pageId="2" pageNumber="279">a</emphasis>
|
||
10 (10–11), 2
|
||
<emphasis id="1BC44F67FE55BD30C6230793477EF8DF" box="[341,353,1785,1808]" italics="true" pageId="2" pageNumber="279">с</emphasis>
|
||
23 (18–23), 3
|
||
<emphasis id="1BC44F67FE55BD30C5730793440CF8DF" box="[517,531,1785,1808]" italics="true" pageId="2" pageNumber="279">a</emphasis>
|
||
14 (12–16), 3
|
||
<emphasis id="1BC44F67FE55BD30C5C1079D44DAF8DF" box="[695,709,1783,1808]" italics="true" pageId="2" pageNumber="279">b</emphasis>
|
||
10 (9–11), 3
|
||
<emphasis id="1BC44F67FE55BD30C42C07934579F8DF" box="[858,870,1785,1808]" italics="true" pageId="2" pageNumber="279">с</emphasis>
|
||
12 (11–14), 4
|
||
<emphasis id="1BC44F67FE55BD30C37F07934208F8DF" box="[1033,1047,1785,1808]" italics="true" pageId="2" pageNumber="279">a</emphasis>
|
||
11 (10–13), 4
|
||
<emphasis id="1BC44F67FE55BD30C3CF079D42D8F8DF" box="[1209,1223,1783,1808]" italics="true" pageId="2" pageNumber="279">b</emphasis>
|
||
15 (14–17), 4
|
||
<emphasis id="1BC44F67FE55BD30C21D07934368F8DF" box="[1387,1399,1785,1808]" italics="true" pageId="2" pageNumber="279">с</emphasis>
|
||
12 (12–14),
|
||
<emphasis id="1BC44F67FE55BD30C78B06774709F8FB" box="[253,278,1821,1844]" italics="true" pageId="2" pageNumber="279">ps</emphasis>
|
||
1 22 (21–25),
|
||
<emphasis id="1BC44F67FE55BD30C6D9067747D8F8FB" box="[431,455,1821,1844]" italics="true" pageId="2" pageNumber="279">ps</emphasis>
|
||
2 23 (21–25),
|
||
<emphasis id="1BC44F67FE55BD30C51606774466F8FB" box="[608,633,1821,1844]" italics="true" pageId="2" pageNumber="279">ps</emphasis>
|
||
3 18 (17–20).
|
||
</paragraph>
|
||
<paragraph id="290F9375FE55BD31C7B1062E4735FDD4" blockId="2.[151,1437,1454,2027]" lastBlockId="3.[151,1436,151,575]" lastPageId="3" lastPageNumber="280" pageId="2" pageNumber="279">
|
||
<emphasis id="1BC44F67FE55BD30C7B1062E46E3F894" box="[199,252,1860,1883]" italics="true" pageId="2" pageNumber="279">Legs</emphasis>
|
||
(
|
||
<figureCitation id="B18B8FF0FE55BD30C67D0628474CF894" box="[267,339,1858,1883]" captionStart="FIGURE 3" captionStartId="4.[151,250,1330,1352]" captionTargetBox="[160,1411,246,1289]" captionTargetId="figure@4.[151,1436,193,1309]" captionTargetPageId="4" captionText="FIGURE 3. Spatulaphorus enoplotruporum sp. nov., female: A—right leg I in dorsal view, B—right leg II in dorsal view." httpUri="https://zenodo.org/record/1303131/files/figure.png" pageId="2" pageNumber="279">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="B18B8FF0FE55BD30C6160629476EF894" box="[352,369,1859,1883]" captionStart="FIGURE 4" captionStartId="5.[151,250,1546,1568]" captionTargetBox="[191,1406,208,1529]" captionTargetId="figure@5.[167,1421,193,1533]" captionTargetPageId="5" captionText="FIGURE 4. Spatulaphorus enoplotruporum sp. nov., female: A—right leg III in dorsal view, B—right leg IV in dorsal view." httpUri="https://zenodo.org/record/1303133/files/figure.png" pageId="2" pageNumber="279">4</figureCitation>
|
||
). Leg I (
|
||
<figureCitation id="B18B8FF0FE55BD30C6A706284435F894" box="[465,554,1858,1883]" captionStart="FIGURE 3" captionStartId="4.[151,250,1330,1352]" captionTargetBox="[160,1411,246,1289]" captionTargetId="figure@4.[151,1436,193,1309]" captionTargetPageId="4" captionText="FIGURE 3. Spatulaphorus enoplotruporum sp. nov., female: A—right leg I in dorsal view, B—right leg II in dorsal view." httpUri="https://zenodo.org/record/1303131/files/figure.png" pageId="2" pageNumber="279">Fig. 3A</figureCitation>
|
||
) slightly thicker than leg II. Setal formula:
|
||
<date id="5D0EB5B5FE55BD30C36406294298F894" box="[1042,1159,1859,1883]" pageId="2" pageNumber="279" value="2017-01-04" valueMax="2017-01-02" valueMin="2017-01-04">1–4–2–17</date>
|
||
(4). Tibiotarsus enlarged, with large claw. Setae (
|
||
<emphasis id="1BC44F67FE55BD30C6EB060247B4F8B0" box="[413,427,1896,1919]" italics="true" pageId="2" pageNumber="279">u</emphasis>
|
||
) consolidated into a horn-like structure opposing to claw. Tarsal claw blunt-tipped. Seta
|
||
<emphasis id="1BC44F67FE55BD30C2E706024383F8B0" box="[1425,1436,1896,1919]" italics="true" pageId="2" pageNumber="279">s</emphasis>
|
||
distinctly foliate distally; seta
|
||
<emphasis id="1BC44F67FE55BD30C57606E14436F86D" box="[512,553,1931,1954]" italics="true" pageId="2" pageNumber="279">pv”</emphasis>
|
||
slightly foliate distally. Seta
|
||
<emphasis id="1BC44F67FE55BD30C4F806E04583F86C" box="[910,924,1930,1955]" italics="true" pageId="2" pageNumber="279">d</emphasis>
|
||
of femur spatulate, with short subterminal projection. Setae
|
||
<emphasis id="1BC44F67FE55BD30C62806C44770F808" box="[350,367,1966,1991]" italics="true" pageId="2" pageNumber="279">l’</emphasis>
|
||
of femur
|
||
<emphasis id="1BC44F67FE55BD30C69206C4441CF808" box="[484,515,1966,1991]" italics="true" pageId="2" pageNumber="279">pl’</emphasis>
|
||
and
|
||
<emphasis id="1BC44F67FE55BD30C54B06C44456F808" box="[573,585,1966,1991]" italics="true" pageId="2" pageNumber="279">k</emphasis>
|
||
of tibiotarsus smooth; other leg setae (except eupathidia) sparsely barbed. Setae
|
||
<emphasis id="1BC44F67FE55BD30C7AD06B846F2F824" box="[219,237,2002,2027]" italics="true" pageId="2" pageNumber="279">l’</emphasis>
|
||
of femur, (
|
||
<emphasis id="1BC44F67FE55BD30C61A06B8476BF824" box="[364,372,2002,2027]" italics="true" pageId="2" pageNumber="279">l</emphasis>
|
||
) of genu and
|
||
<emphasis id="1BC44F67FE55BD30C56706B84402F824" box="[529,541,2002,2027]" italics="true" pageId="2" pageNumber="279">k</emphasis>
|
||
of tibiotarsus blunt-ended; other leg setae (except eupathidia) pointed. Lengths of solenidia
|
||
<emphasis id="1BC44F67FE54BD31C65101F34724FF7F" box="[295,315,153,176]" italics="true" pageId="3" pageNumber="280">ω</emphasis>
|
||
1 8 (7–9),
|
||
<emphasis id="1BC44F67FE54BD31C6C401F347D9FF7F" box="[434,454,153,176]" italics="true" pageId="3" pageNumber="280">ω</emphasis>
|
||
2 6 (5–7),
|
||
<emphasis id="1BC44F67FE54BD31C54B01F34453FF7F" box="[573,588,153,176]" italics="true" pageId="3" pageNumber="280">φ</emphasis>
|
||
1 8 (8–9),
|
||
<emphasis id="1BC44F67FE54BD31C5B201F344CCFF7F" box="[708,723,153,176]" italics="true" pageId="3" pageNumber="280">φ</emphasis>
|
||
2 6 (5–6); Solenidion
|
||
<emphasis id="1BC44F67FE54BD31C4B801F345C2FF7F" box="[974,989,153,176]" italics="true" pageId="3" pageNumber="280">φ</emphasis>
|
||
1 thick, clavate, other solenidia weakly clavate. Leg II (
|
||
<figureCitation id="B18B8FF0FE54BD31C63C01D447BEFF18" box="[330,417,190,215]" captionStart="FIGURE 3" captionStartId="4.[151,250,1330,1352]" captionTargetBox="[160,1411,246,1289]" captionTargetId="figure@4.[151,1436,193,1309]" captionTargetPageId="4" captionText="FIGURE 3. Spatulaphorus enoplotruporum sp. nov., female: A—right leg I in dorsal view, B—right leg II in dorsal view." httpUri="https://zenodo.org/record/1303131/files/figure.png" pageId="3" pageNumber="280">Fig. 3B</figureCitation>
|
||
). Setal formula: 1–3–1–4–6(1). Tarsus with padded claws and empodium with three lobes, one distal and two lateral. Solenidion
|
||
<emphasis id="1BC44F67FE54BD31C548018E444DFF34" box="[574,594,228,251]" italics="true" pageId="3" pageNumber="280">ω</emphasis>
|
||
6 (5–7), weakly clavate, solenidion
|
||
<emphasis id="1BC44F67FE54BD31C49F018E45E7FF34" box="[1001,1016,228,251]" italics="true" pageId="3" pageNumber="280">φ</emphasis>
|
||
absent. Setae (
|
||
<emphasis id="1BC44F67FE54BD31C3D4018942D9FF35" box="[1186,1222,227,250]" italics="true" pageId="3" pageNumber="280">pv)</emphasis>
|
||
of tarsus distinctly foliate distally; setae
|
||
<emphasis id="1BC44F67FE54BD31C6F2006C478DFED0" box="[388,402,262,287]" italics="true" pageId="3" pageNumber="280">d</emphasis>
|
||
and
|
||
<emphasis id="1BC44F67FE54BD31C6BE006C47C6FED0" box="[456,473,262,287]" italics="true" pageId="3" pageNumber="280">l’</emphasis>
|
||
of femur blunt-ended; other leg setae pointed. Seta
|
||
<emphasis id="1BC44F67FE54BD31C36A006D422CFED1" box="[1052,1075,263,286]" italics="true" pageId="3" pageNumber="280">u’</emphasis>
|
||
of tarsus smooth and thin; other leg setae barbed. Leg III (
|
||
<figureCitation id="B18B8FF0FE54BD31C6CC00404409FE8C" box="[442,534,298,323]" captionStart="FIGURE 4" captionStartId="5.[151,250,1546,1568]" captionTargetBox="[191,1406,208,1529]" captionTargetId="figure@5.[167,1421,193,1533]" captionTargetPageId="5" captionText="FIGURE 4. Spatulaphorus enoplotruporum sp. nov., female: A—right leg III in dorsal view, B—right leg IV in dorsal view." httpUri="https://zenodo.org/record/1303133/files/figure.png" pageId="3" pageNumber="280">Fig. 4A</figureCitation>
|
||
). Femur divided into basi- and telofemur. Setal formula: 1–2–1–4–6. Claws and empodium of same shape as on tarsus II. Setae (
|
||
<emphasis id="1BC44F67FE54BD31C5CF002544C2FEA9" box="[697,733,335,358]" italics="true" pageId="3" pageNumber="280">pv)</emphasis>
|
||
of tarsus distinctly foliate distally; setae
|
||
<emphasis id="1BC44F67FE54BD31C3D8002542DBFEA9" box="[1198,1220,335,358]" italics="true" pageId="3" pageNumber="280">v’</emphasis>
|
||
of trochanter,
|
||
<emphasis id="1BC44F67FE54BD31C21F00244368FEA8" box="[1385,1399,334,359]" italics="true" pageId="3" pageNumber="280">d</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE54BD31C2F000254383FEA9" box="[1414,1436,335,358]" italics="true" pageId="3" pageNumber="280">v’</emphasis>
|
||
of femur and
|
||
<emphasis id="1BC44F67FE54BD31C6470018475DFE44" box="[305,322,370,395]" italics="true" pageId="3" pageNumber="280">l‘</emphasis>
|
||
of genu blunt-ended; other leg setae pointed. Seta
|
||
<emphasis id="1BC44F67FE54BD31C4F700194587FE45" box="[897,920,371,394]" italics="true" pageId="3" pageNumber="280">u’</emphasis>
|
||
of tarsus smooth; other leg setae barbed. Leg IV (
|
||
<figureCitation id="B18B8FF0FE54BD31C7B200FC4708FE60" box="[196,279,406,431]" captionStart="FIGURE 4" captionStartId="5.[151,250,1546,1568]" captionTargetBox="[191,1406,208,1529]" captionTargetId="figure@5.[167,1421,193,1533]" captionTargetPageId="5" captionText="FIGURE 4. Spatulaphorus enoplotruporum sp. nov., female: A—right leg III in dorsal view, B—right leg IV in dorsal view." httpUri="https://zenodo.org/record/1303133/files/figure.png" pageId="3" pageNumber="280">Fig 4B</figureCitation>
|
||
). Femur divided into basi- and telofemur. Setal formula: 1–2–0–4–6. Claws simple, empodium round, stalked. Setae
|
||
<emphasis id="1BC44F67FE54BD31C64F00D14742FE1D" box="[313,349,443,466]" italics="true" pageId="3" pageNumber="280">tс”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE54BD31C61A00D1479CFE1D" box="[364,387,443,466]" italics="true" pageId="3" pageNumber="280">u’</emphasis>
|
||
, and (
|
||
<emphasis id="1BC44F67FE54BD31C6BD00D647FAFE1C" box="[459,485,444,467]" italics="true" pageId="3" pageNumber="280">pv</emphasis>
|
||
) slightly foliate distally (flattened tips difficult to discern in some specimens); setae
|
||
<emphasis id="1BC44F67FE54BD31C7E100B446B7FE38" box="[151,168,478,503]" italics="true" pageId="3" pageNumber="280">l’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE54BD31C7C100B546D2FE39" box="[183,205,479,502]" italics="true" pageId="3" pageNumber="280">v’</emphasis>
|
||
of tibia and t
|
||
<emphasis id="1BC44F67FE54BD31C61300B54764FE39" box="[357,379,479,502]" italics="true" pageId="3" pageNumber="280">с’</emphasis>
|
||
of tarsus pointed; other leg setae blunt-ended. Seta
|
||
<emphasis id="1BC44F67FE54BD31C4B300B445F5FE38" box="[965,1002,478,503]" italics="true" pageId="3" pageNumber="280">pl”</emphasis>
|
||
of tarsus smooth, spiniform; other leg setae barbed.
|
||
</paragraph>
|
||
<paragraph id="290F9375FE54BD31C7B1034D4792FDF0" blockId="3.[151,1436,151,575]" box="[199,397,550,575]" pageId="3" pageNumber="280">MALE unknown.</paragraph>
|
||
</subSubSection>
|
||
<caption id="7DCFC3FDFE54BD31C7E106C8479CF818" httpUri="https://zenodo.org/record/1303129/files/figure.png" pageId="3" pageNumber="280" startId="3.[151,250,1954,1976]" targetBox="[209,1332,690,1862]" targetPageId="3">
|
||
<paragraph id="290F9375FE54BD31C7E106C8479CF818" blockId="3.[151,1436,1954,2007]" pageId="3" pageNumber="280">
|
||
<emphasis id="1BC44F67FE54BD31C7E106C84712F878" bold="true" box="[151,269,1954,1976]" pageId="3" pageNumber="280">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="EEB0E8F6FE54BD31C66F06C84447F877" authority="Khaustov & Frolov, 2018" authorityName="Khaustov & Frolov" authorityYear="2018" box="[281,600,1954,1976]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="3" pageNumber="280" phylum="Arthropoda" rank="species" species="enoplotruporum" status="sp. nov.">
|
||
<emphasis id="1BC44F67FE54BD31C66F06C84447F877" box="[281,600,1954,1976]" italics="true" pageId="3" pageNumber="280">Spatulaphorus enoplotruporum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="00F7F21CFE54BD31C52806C944B1F877" box="[606,686,1955,1976]" pageId="3" pageNumber="280" rank="species">
|
||
<emphasis id="1BC44F67FE54BD31C52806C944B1F877" bold="true" box="[606,686,1955,1976]" pageId="3" pageNumber="280">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, female: A—gnathosoma in dorsal view, B—gnathosoma in ventral view, C—pharyngeal pumps.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="7DCFC3FDFE53BD36C7E104584370FA87" box="[151,1391,1330,1352]" httpUri="https://zenodo.org/record/1303131/files/figure.png" pageId="4" pageNumber="281" startId="4.[151,250,1330,1352]" targetBox="[160,1411,246,1289]" targetPageId="4">
|
||
<paragraph id="290F9375FE53BD36C7E104584370FA87" blockId="4.[151,1391,1330,1352]" box="[151,1391,1330,1352]" pageId="4" pageNumber="281">
|
||
<emphasis id="1BC44F67FE53BD36C7E104584712FA88" bold="true" box="[151,269,1330,1352]" pageId="4" pageNumber="281">FIGURE 3</emphasis>
|
||
.
|
||
<taxonomicName id="EEB0E8F6FE53BD36C66F04584447FA87" authority="Khaustov & Frolov, 2018" authorityName="Khaustov & Frolov" authorityYear="2018" box="[281,600,1330,1352]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="281" phylum="Arthropoda" rank="species" species="enoplotruporum" status="sp. nov.">
|
||
<emphasis id="1BC44F67FE53BD36C66F04584447FA87" box="[281,600,1330,1352]" italics="true" pageId="4" pageNumber="281">Spatulaphorus enoplotruporum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="00F7F21CFE53BD36C528045844B1FA88" box="[606,686,1330,1351]" pageId="4" pageNumber="281" rank="species">
|
||
<emphasis id="1BC44F67FE53BD36C528045844B1FA88" bold="true" box="[606,686,1330,1351]" pageId="4" pageNumber="281">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, female: A—right leg I in dorsal view, B—right leg II in dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="61AAC0FEFE53BD36C7B10405445AF9D7" pageId="4" pageNumber="281" type="materials_examined">
|
||
<paragraph id="290F9375FE53BD36C7B104054258FA63" blockId="4.[151,1436,1391,2006]" pageId="4" pageNumber="281">
|
||
<materialsCitation id="99D89928FE53BD36C7B1040542F0FA47" ID-GBIF-Occurrence="1852084072" box="[199,1263,1391,1416]" collectingDate="1986-10" collectionCode="ZISP" country="Thailand" location="Chiang Mai" pageId="4" pageNumber="281" specimenCount="2" stateProvince="Chiang Mai" typeStatus="holotype">
|
||
<emphasis id="1BC44F67FE53BD36C7B10405476CFA47" bold="true" box="[199,371,1391,1416]" pageId="4" pageNumber="281">
|
||
<typeStatus id="F60B2DD7FE53BD36C7B10405471DFA47" box="[199,258,1391,1416]" pageId="4" pageNumber="281">Type</typeStatus>
|
||
material
|
||
</emphasis>
|
||
. Female
|
||
<typeStatus id="F60B2DD7FE53BD36C69304054455FA47" box="[485,586,1391,1416]" pageId="4" pageNumber="281" type="holotype">holotype</typeStatus>
|
||
, slide
|
||
<collectionCode id="4FA10BB0FE53BD36C5EB041A44C8FA47" box="[669,727,1392,1416]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="4" pageNumber="281">ZISP</collectionCode>
|
||
T-Pygm-001,
|
||
<collectingCountry id="51A7D3E5FE53BD36C4F5040545F6FA47" box="[899,1001,1391,1416]" name="Thailand" pageId="4" pageNumber="281">Thailand</collectingCountry>
|
||
,
|
||
<collectingRegion id="EB745D97FE53BD36C48C04054297FA47" box="[1018,1160,1391,1416]" country="Thailand" name="Chiang Mai" pageId="4" pageNumber="281">Chiang Mai</collectingRegion>
|
||
,
|
||
<collectingDate id="4D4A4C5DFE53BD36C3EF040542F0FA47" box="[1177,1263,1391,1416]" pageId="4" pageNumber="281" value="1986-10">
|
||
<date id="5D0EB5B5FE53BD36C3EF040542F0FA47" box="[1177,1263,1391,1416]" pageId="4" pageNumber="281" value="1986-10">X.1986</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="99D89928FE53BD36C276041A4771FA63" ID-GBIF-Occurrence="1852084069" collectingDate="1986-10" collectionCode="ZISP" country="Thailand" location="Chiang Mai" pageId="4" pageNumber="281" specimenCount="19" specimenCount-female="19" stateProvince="Chiang Mai" typeStatus="paratype">
|
||
<typeStatus id="F60B2DD7FE53BD36C276041A436FFA47" box="[1280,1392,1392,1416]" pageId="4" pageNumber="281" type="paratype">paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="3FB658FCFE53BD36C2F6041A46EEFA63" pageId="4" pageNumber="281" type="female">19 females</specimenCount>
|
||
, same data
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="99D89928FE53BD36C60F04FE4258FA63" ID-GBIF-Occurrence="1852084070" box="[377,1095,1427,1452]" collectingDate="1992-07" collectorName="N. Kagana." country="Thailand" location="Mae Ai" pageId="4" pageNumber="281" specimenCount="4" specimenCount-female="4" stateProvince="Chiang Mai" typeStatus="paratype">
|
||
<specimenCount id="3FB658FCFE53BD36C60F04FE47F7FA63" box="[377,488,1427,1452]" pageId="4" pageNumber="281" type="female">4 females</specimenCount>
|
||
,
|
||
<collectingCountry id="51A7D3E5FE53BD36C68504F94446FA63" box="[499,601,1427,1452]" name="Thailand" pageId="4" pageNumber="281">Thailand</collectingCountry>
|
||
,
|
||
<collectingRegion id="EB745D97FE53BD36C51304F944F2FA63" box="[613,749,1427,1452]" country="Thailand" name="Chiang Mai" pageId="4" pageNumber="281">Chiang Mai</collectingRegion>
|
||
,
|
||
<location id="2C6FC5AEFE53BD36C58E04FE4550FA63" LSID="urn:lsid:plazi:treatment:A1192263FE56BD36C7E104F3437CF81A:2C6FC5AEFE53BD36C58E04FE4550FA63" box="[760,847,1427,1452]" country="Thailand" name="Mae Ai" pageId="4" pageNumber="281" stateProvince="Chiang Mai">Mae Ai</location>
|
||
,
|
||
<collectingDate id="4D4A4C5DFE53BD36C42C04FE45DCFA63" box="[858,963,1428,1452]" pageId="4" pageNumber="281" value="1992-07">
|
||
<date id="5D0EB5B5FE53BD36C42C04FE45DCFA63" box="[858,963,1428,1452]" pageId="4" pageNumber="281" value="1992-07">VII.1992</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="8445F6A3FE53BD36C4B804FE4258FA63" box="[974,1095,1428,1452]" pageId="4" pageNumber="281">N.Kagana.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="290F9375FE53BD36C7B104DD445AF9D7" blockId="4.[151,1436,1391,2006]" pageId="4" pageNumber="281">
|
||
<materialsCitation id="99D89928FE53BD36C7B104DD441FFA3B" ID-GBIF-Occurrence="1852084154" collectionCode="RAS" country="Russia" location="Saint Petersburg" pageId="4" pageNumber="281" specimenCount="3" stateProvince="Saint Petersburg" typeStatus="holotype">
|
||
<emphasis id="1BC44F67FE53BD36C7B104DD4799FA1F" bold="true" box="[199,390,1463,1488]" pageId="4" pageNumber="281">
|
||
<typeStatus id="F60B2DD7FE53BD36C7B104DD471DFA1F" box="[199,258,1463,1488]" pageId="4" pageNumber="281">Type</typeStatus>
|
||
deposition
|
||
</emphasis>
|
||
. The
|
||
<typeStatus id="F60B2DD7FE53BD36C6BA04DD4431FA1F" box="[460,558,1463,1488]" pageId="4" pageNumber="281" type="holotype">holotype</typeStatus>
|
||
and four
|
||
<typeStatus id="F60B2DD7FE53BD36C5D404D24512FA1F" box="[674,781,1464,1488]" pageId="4" pageNumber="281" type="paratype">paratypes</typeStatus>
|
||
are deposited in the collection of Zoological Institute of
|
||
<collectionCode id="4FA10BB0FE53BD36C7E104B646CFFA3B" box="[151,208,1500,1524]" country="Myanmar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14256" name="Union of Burma Applied Research Institute" pageId="4" pageNumber="281">RAS</collectionCode>
|
||
,
|
||
<collectingRegion id="EB745D97FE53BD36C79704B147BEFA3B" box="[225,417,1499,1524]" country="Russia" name="Saint Petersburg" pageId="4" pageNumber="281">Saint Petersburg</collectingRegion>
|
||
,
|
||
<collectingCountry id="51A7D3E5FE53BD36C6C404B1441FFA3B" box="[434,512,1499,1524]" name="Russia" pageId="4" pageNumber="281">Russia</collectingCountry>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="99D89928FE53BD36C56404B6445AF9D7" ID-GBIF-Occurrence="1852084071" country="Russia" location="Tyumen State University Museum of Zoology" pageId="4" pageNumber="281" specimenCount="19" stateProvince="Tyumen" typeStatus="paratype">
|
||
<specimenCount id="3FB658FCFE53BD36C56404B644B9FA3B" box="[530,678,1500,1524]" pageId="4" pageNumber="281" type="generic" typeStatus="paratypes">19 paratypes</specimenCount>
|
||
are deposited in the collection of the
|
||
<location id="2C6FC5AEFE53BD36C30F04B64761F9D7" LSID="urn:lsid:plazi:treatment:A1192263FE56BD36C7E104F3437CF81A:2C6FC5AEFE53BD36C30F04B64761F9D7" country="Russia" name="Tyumen State University Museum of Zoology" pageId="4" pageNumber="281" stateProvince="Tyumen">Tyumen State University Museum of Zoology</location>
|
||
,
|
||
<collectingRegion id="EB745D97FE53BD36C6FF076A47F8F9D7" box="[393,487,1536,1560]" country="Russia" name="Tyumen'" pageId="4" pageNumber="281">Tyumen</collectingRegion>
|
||
,
|
||
<collectingCountry id="51A7D3E5FE53BD36C6850495445EF9D7" box="[499,577,1535,1560]" name="Russia" pageId="4" pageNumber="281">Russia</collectingCountry>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61AAC0FEFE53BD36C7B10749476DF87D" pageId="4" pageNumber="281" type="diagnosis">
|
||
<paragraph id="290F9375FE53BD36C7B10749476DF87D" blockId="4.[151,1436,1391,2006]" pageId="4" pageNumber="281">
|
||
<emphasis id="1BC44F67FE53BD36C7B1074947D7F9F3" bold="true" box="[199,456,1571,1596]" pageId="4" pageNumber="281">Differential diagnosis</emphasis>
|
||
. The new species is most similar to
|
||
<emphasis id="1BC44F67FE53BD36C418074E4563F9F4" box="[878,892,1572,1595]" italics="true" pageId="4" pageNumber="281">S</emphasis>
|
||
.
|
||
<emphasis id="1BC44F67FE53BD36C4FA0749421EF9F3" box="[908,1025,1571,1596]" italics="true" pageId="4" pageNumber="281">сamerikae</emphasis>
|
||
<bibRefCitation id="4D21EE84FE53BD36C37F0749433BF9F3" box="[1033,1316,1571,1596]" pageId="4" pageNumber="281" refId="ref7015" refString="Dastych, H. & Rack, G. (1993) Spatulaphorus gen. n. and three new species of pygmephorid mites associated with scarab beetles in Botswana and Vietnam (Acari: Heterostigmata). Mitteilungen Hamburgisches Zoologisches Museum und Institut, 90, 265 - 84." type="journal article">Dastych and Rack, 1993</bibRefCitation>
|
||
, described from
|
||
<collectingCountry id="51A7D3E5FE53BD36C7AC07224757F990" box="[218,328,1608,1631]" name="Botswana" pageId="4" pageNumber="281">Botswana</collectingCountry>
|
||
(
|
||
<bibRefCitation id="4D21EE84FE53BD36C62A072D4476F9AF" box="[348,617,1607,1632]" pageId="4" pageNumber="281" refId="ref7015" refString="Dastych, H. & Rack, G. (1993) Spatulaphorus gen. n. and three new species of pygmephorid mites associated with scarab beetles in Botswana and Vietnam (Acari: Heterostigmata). Mitteilungen Hamburgisches Zoologisches Museum und Institut, 90, 265 - 84." type="journal article">Dastych & Rack 1993</bibRefCitation>
|
||
), by the similar modifications of setae on tibiotarsus I and tarsus II, presence of apodemes 5, and characteristic position of setae
|
||
<emphasis id="1BC44F67FE53BD36C44907064547F94C" box="[831,856,1644,1667]" italics="true" pageId="4" pageNumber="281">ps</emphasis>
|
||
3 anteriorly to
|
||
<emphasis id="1BC44F67FE53BD36C48F0706420DF94C" box="[1017,1042,1644,1667]" italics="true" pageId="4" pageNumber="281">ps</emphasis>
|
||
1, 2. The new species differs from
|
||
<emphasis id="1BC44F67FE53BD36C2F10706438AF94C" box="[1415,1429,1644,1667]" italics="true" pageId="4" pageNumber="281">S</emphasis>
|
||
.
|
||
<emphasis id="1BC44F67FE53BD36C7E107F84713F964" box="[151,268,1682,1707]" italics="true" pageId="4" pageNumber="281">сamerikae</emphasis>
|
||
by distinctly barbed setae
|
||
<emphasis id="1BC44F67FE53BD36C54907F94447F965" box="[575,600,1683,1706]" italics="true" pageId="4" pageNumber="281">ps</emphasis>
|
||
1, 2 (vs. smooth in
|
||
<emphasis id="1BC44F67FE53BD36C45707F94530F965" box="[801,815,1683,1706]" italics="true" pageId="4" pageNumber="281">S</emphasis>
|
||
.
|
||
<emphasis id="1BC44F67FE53BD36C44807F845ADF964" box="[830,946,1682,1707]" italics="true" pageId="4" pageNumber="281">сamerikae</emphasis>
|
||
), foliate setae on tarsi III and IV (vs. nonmodified in
|
||
<emphasis id="1BC44F67FE53BD36C66907D047B0F91D" box="[287,431,1721,1746]" italics="true" pageId="4" pageNumber="281">S. сamerikae</emphasis>
|
||
) and by distinctly longer dorsal setae
|
||
<emphasis id="1BC44F67FE53BD36C42E07D34579F91D" box="[856,870,1721,1746]" italics="true" pageId="4" pageNumber="281">d</emphasis>
|
||
39–48,
|
||
<emphasis id="1BC44F67FE53BD36C4B607D345D7F91E" box="[960,968,1721,1745]" italics="true" pageId="4" pageNumber="281">f</emphasis>
|
||
44–58,
|
||
<emphasis id="1BC44F67FE53BD36C35507D3422EF91D" box="[1059,1073,1721,1746]" italics="true" pageId="4" pageNumber="281">h</emphasis>
|
||
1 45–57 (vs.
|
||
<emphasis id="1BC44F67FE53BD36C3CA07D342D5F91D" box="[1212,1226,1721,1746]" italics="true" pageId="4" pageNumber="281">d</emphasis>
|
||
25–30,
|
||
<emphasis id="1BC44F67FE53BD36C25207D34333F91E" box="[1316,1324,1721,1745]" italics="true" pageId="4" pageNumber="281">f</emphasis>
|
||
24–31,
|
||
<emphasis id="1BC44F67FE53BD36C2F007D3438BF91D" box="[1414,1428,1721,1746]" italics="true" pageId="4" pageNumber="281">h</emphasis>
|
||
<date id="5D0EB5B5FE53BD36C2E207A246C2F937" pageId="4" pageNumber="281" value="1924-01-31">
|
||
1
|
||
<quantity id="EE483E90FE53BD36C7E1078A46C2F937" box="[151,221,1760,1784]" metricMagnitude="-1" metricUnit="m" metricValue="6.984999999999999" metricValueMax="7.874" metricValueMin="6.096" pageId="4" pageNumber="281" unit="in" value="27.5" valueMax="31.0" valueMin="24.0">24–31</quantity>
|
||
</date>
|
||
in
|
||
<emphasis id="1BC44F67FE53BD36C677078A4710F938" box="[257,271,1760,1783]" italics="true" pageId="4" pageNumber="281">S</emphasis>
|
||
.
|
||
<emphasis id="1BC44F67FE53BD36C66B07B5478EF937" box="[285,401,1759,1784]" italics="true" pageId="4" pageNumber="281">сamerikae</emphasis>
|
||
). The new species also similar to S.
|
||
<taxonomicName id="EEB0E8F6FE53BD36C45A07B5429DF937" authority="Dastych and Rack, 1993" authorityName="Dastych and Rack" authorityYear="1993" box="[812,1154,1759,1784]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="281" phylum="Arthropoda" rank="species" species="langi">
|
||
<emphasis id="1BC44F67FE53BD36C45A07B54579F937" box="[812,870,1759,1784]" italics="true" pageId="4" pageNumber="281">langi</emphasis>
|
||
<bibRefCitation id="4D21EE84FE53BD36C41B07B5429DF937" box="[877,1154,1759,1784]" pageId="4" pageNumber="281" refId="ref7015" refString="Dastych, H. & Rack, G. (1993) Spatulaphorus gen. n. and three new species of pygmephorid mites associated with scarab beetles in Botswana and Vietnam (Acari: Heterostigmata). Mitteilungen Hamburgisches Zoologisches Museum und Institut, 90, 265 - 84." type="journal article">Dastych and Rack, 1993</bibRefCitation>
|
||
</taxonomicName>
|
||
, described from
|
||
<collectingCountry id="51A7D3E5FE53BD36C24B07B54383F937" box="[1341,1436,1759,1784]" name="Vietnam" pageId="4" pageNumber="281">Vietnam</collectingCountry>
|
||
(
|
||
<bibRefCitation id="4D21EE84FE53BD36C7E8066E47B7F8D3" box="[158,424,1796,1821]" pageId="4" pageNumber="281" refId="ref7015" refString="Dastych, H. & Rack, G. (1993) Spatulaphorus gen. n. and three new species of pygmephorid mites associated with scarab beetles in Botswana and Vietnam (Acari: Heterostigmata). Mitteilungen Hamburgisches Zoologisches Museum und Institut, 90, 265 - 84." type="journal article">Dastych & Rack 1993</bibRefCitation>
|
||
), by the similar relative lengths of dorsal idiosomal setae and by the same number of modified foliate setae on tibiotarsus I and tarsi I and II. The new species differs from
|
||
<taxonomicName id="EEB0E8F6FE53BD36C30A064242CBF88F" authorityName="Dastych and Rack" authorityYear="1993" box="[1148,1236,1831,1856]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="281" phylum="Arthropoda" rank="species" species="langi">
|
||
<emphasis id="1BC44F67FE53BD36C30A064242CBF88F" box="[1148,1236,1831,1856]" italics="true" pageId="4" pageNumber="281">S. langi</emphasis>
|
||
</taxonomicName>
|
||
by non-modified setae
|
||
<emphasis id="1BC44F67FE53BD36C7A0062746ECF8AB" box="[214,243,1869,1892]" italics="true" pageId="4" pageNumber="281">tс’</emphasis>
|
||
on tarsi II and III (vs. spiniform in
|
||
<taxonomicName id="EEB0E8F6FE53BD36C5FC062744FEF8AA" authorityName="Dastych and Rack" authorityYear="1993" box="[650,737,1868,1893]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="281" phylum="Arthropoda" rank="species" species="langi">
|
||
<emphasis id="1BC44F67FE53BD36C5FC062744FEF8AA" box="[650,737,1868,1893]" italics="true" pageId="4" pageNumber="281">S. langi</emphasis>
|
||
</taxonomicName>
|
||
), by setae
|
||
<emphasis id="1BC44F67FE53BD36C42A0627456AF8AB" box="[860,885,1869,1892]" italics="true" pageId="4" pageNumber="281">ps</emphasis>
|
||
1, 2 distinctly barbed and subequal to
|
||
<emphasis id="1BC44F67FE53BD36C26406274334F8AB" box="[1298,1323,1869,1892]" italics="true" pageId="4" pageNumber="281">ps</emphasis>
|
||
3 (vs.
|
||
<emphasis id="1BC44F67FE53BD36C21A0627439AF8AB" box="[1388,1413,1869,1892]" italics="true" pageId="4" pageNumber="281">ps</emphasis>
|
||
1, 2 smooth and distinctly shorter than
|
||
<emphasis id="1BC44F67FE53BD36C56E061E442EF844" box="[536,561,1908,1931]" italics="true" pageId="4" pageNumber="281">ps</emphasis>
|
||
<quantity id="EE483E90FE53BD36C54706EB444AF844" box="[561,597,1906,1935]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="281" unit="in" value="3.0">3 in</quantity>
|
||
<taxonomicName id="EEB0E8F6FE53BD36C52D061944AEF844" authorityName="Dastych and Rack" authorityYear="1993" box="[603,689,1906,1931]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="281" phylum="Arthropoda" rank="species" species="langi">
|
||
<emphasis id="1BC44F67FE53BD36C52D061944AEF844" box="[603,689,1906,1931]" italics="true" pageId="4" pageNumber="281">S. langi</emphasis>
|
||
</taxonomicName>
|
||
), and by setae 2
|
||
<emphasis id="1BC44F67FE53BD36C413061E456CF844" box="[869,883,1908,1931]" italics="true" pageId="4" pageNumber="281">a</emphasis>
|
||
distinctly shorter than 2
|
||
<emphasis id="1BC44F67FE53BD36C3F7061E4292F844" box="[1153,1165,1908,1931]" italics="true" pageId="4" pageNumber="281">с</emphasis>
|
||
(vs. 2
|
||
<emphasis id="1BC44F67FE53BD36C3A7061E42C0F844" box="[1233,1247,1908,1931]" italics="true" pageId="4" pageNumber="281">a</emphasis>
|
||
distinctly longer than 2
|
||
<emphasis id="1BC44F67FE53BD36C7AA06F046F7F87E" box="[220,232,1946,1969]" italics="true" pageId="4" pageNumber="281">с</emphasis>
|
||
in
|
||
<taxonomicName id="EEB0E8F6FE53BD36C67A06F0477EF87D" authorityName="Dastych and Rack" authorityYear="1993" box="[268,353,1945,1970]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="281" phylum="Arthropoda" rank="species" species="langi">
|
||
<emphasis id="1BC44F67FE53BD36C67A06F04705F87E" box="[268,282,1946,1969]" italics="true" pageId="4" pageNumber="281">S</emphasis>
|
||
.
|
||
<emphasis id="1BC44F67FE53BD36C65E06F3477EF87D" box="[296,353,1945,1970]" italics="true" pageId="4" pageNumber="281">langi</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61AAC0FEFE53BD36C7B106D7437CF81A" box="[199,1379,1981,2006]" pageId="4" pageNumber="281" type="etymology">
|
||
<paragraph id="290F9375FE53BD36C7B106D7437CF81A" blockId="4.[151,1436,1391,2006]" box="[199,1379,1981,2006]" pageId="4" pageNumber="281">
|
||
<emphasis id="1BC44F67FE53BD36C7B106D74759F819" bold="true" box="[199,326,1981,2006]" pageId="4" pageNumber="281">Etymology</emphasis>
|
||
. The name of the new species refers to its association with beetles of the genus
|
||
<taxonomicName id="EEB0E8F6FE53BD36C3B306D74343F819" authorityName="Lucas" authorityYear="1869" box="[1221,1372,1981,2006]" class="Insecta" family="Geotrupidae" genus="Enoplotrupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="281" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1BC44F67FE53BD36C3B306D74343F819" box="[1221,1372,1981,2006]" italics="true" pageId="4" pageNumber="281">Enoplotrupes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |