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<document id="C38165788D46154B017546087A39C84D" ID-CLB-Dataset="24430" ID-DOI="10.11646/zootaxa.4718.2.5" ID-GBIF-Dataset="a6efb27e-a61f-4900-9425-b94fd9948169" ID-ISSN="1175-5326" ID-Zenodo-Dep="3602376" ID-ZooBank="51D7C92E-9018-48F4-B195-EE61F0F7AA5E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1578564670572" checkinUser="plazi" docAuthor="Porch, Nick, Smith, Tessa R. &amp; Greig, Kelly" docDate="2020" docId="9C0087DEFFCA2934FF25FB3BFA96FCB7" docLanguage="en" docName="zootaxa.4718.2.5.pdf" docOrigin="Zootaxa 4718 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pycnomerus raivavae Porch &amp; Smith &amp; Greig 2020, sp. nov." docType="treatment" docUuid="7BFAA473-91DF-4C34-84F8-0494D5747CDA" docUuidSource="ZooBank" docVersion="6" lastPageNumber="243" masterDocId="6039FFA6FFC82930FFB2FFCFFFEAFFA1" masterDocTitle="Five new Pycnomerus Erichson (Coleoptera: Zopheridae: Pycnomerini) from Raivavae, French Polynesia" masterLastPageNumber="250" masterPageNumber="239" pageNumber="241" updateTime="1698776543820" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="C63E85942B02CCE92E831200459DA606" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="C9D5F61B32FDE07E0F7F1E45B9B2130E">Five new Pycnomerus Erichson (Coleoptera: Zopheridae: Pycnomerini) from Raivavae, French Polynesia</mods:title>
</mods:titleInfo>
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<mods:role id="7C6DBC3949C7405BC1E432DE54D4F527">
<mods:roleTerm id="F5B076D2526B6A8CAD19EC0BCBCFFE71">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EA74B0066EF880C9E0BC6703844EC088">Porch, Nick</mods:namePart>
</mods:name>
<mods:name id="FB20991B5387BEC7DCD0E70AB215E2DD" type="personal">
<mods:role id="39EE3EA890610D1B3801097C348EEEE5">
<mods:roleTerm id="2BD474C9655E97CF19858E2604F80EDF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F2F62A5D25601724A61BD2610E40CDD1">Smith, Tessa R.</mods:namePart>
</mods:name>
<mods:name id="290133A0C8CAB017B2088233D206367C" type="personal">
<mods:role id="A10C94BB8122A49E8F6A3F576595E53E">
<mods:roleTerm id="E1CB150D7B4665AEBD3F81F620DE5F3A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AA3E6C4680DFFB7290ECC35BD0C2472A">Greig, Kelly</mods:namePart>
</mods:name>
<mods:typeOfResource id="3C2FAAB5D4FE225CB136170F5D26D6BB">text</mods:typeOfResource>
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<mods:title id="56806D9D945EC798FCD5165F86F2EA85">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A3FE7E248DED2F01189273869878C429">
<mods:date id="904470A4E01BDC97BB07C9289BBEA2FD">2020</mods:date>
<mods:detail id="50B63C0FAFFC517325424CBD68834440" type="pubDate">
<mods:number id="C99E3541EFEBFB17A7CD92BAE75D234D">2020-01-07</mods:number>
</mods:detail>
<mods:detail id="C5CD6DF8D84B63E0AA6C08E75FDA8820" type="volume">
<mods:number id="9275FE0EB8ED413C60651E58759E6066">4718</mods:number>
</mods:detail>
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<mods:number id="A27C886BF2A717B46F16FD10EA8B0F23">2</mods:number>
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<mods:classification id="4362AAFC59AFB71611374AC81E840BA3">journal article</mods:classification>
<mods:identifier id="DFB942C0FAB0B62D5DAF2DD2743BB02B" type="CLB-Dataset">24430</mods:identifier>
<mods:identifier id="183E38DF67DA6E2DA468E2DC7C1F323B" type="DOI">10.11646/zootaxa.4718.2.5</mods:identifier>
<mods:identifier id="12004C87098992CE6441ACD36402417D" type="GBIF-Dataset">a6efb27e-a61f-4900-9425-b94fd9948169</mods:identifier>
<mods:identifier id="FD4DB0D38C06196B74F47686875F4D22" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8130223A862CF87105353A3D884C9310" type="Zenodo-Dep">3602376</mods:identifier>
<mods:identifier id="3BEB7497243596BD499432D00250D546" type="ZooBank">51D7C92E-9018-48F4-B195-EE61F0F7AA5E</mods:identifier>
</mods:mods>
<treatment id="9C0087DEFFCA2934FF25FB3BFA96FCB7" ID-GBIF-Taxon="161670836" LSID="urn:lsid:zoobank.org:act:7BFAA473-91DF-4C34-84F8-0494D5747CDA" httpUri="http://treatment.plazi.org/id/9C0087DEFFCA2934FF25FB3BFA96FCB7" lastPageId="4" lastPageNumber="243" pageId="2" pageNumber="241">
<subSubSection id="5CB36543FFCA2932FF25FB3BFD2AFAAC" box="[151,704,1267,1294]" pageId="2" pageNumber="241" type="nomenclature">
<paragraph id="141636C8FFCA2932FF25FB3BFD2AFAAC" blockId="2.[151,704,1267,1330]" box="[151,704,1267,1294]" pageId="2" pageNumber="241">
<heading id="4F5E81A4FFCA2932FF25FB3BFD2AFAAC" bold="true" box="[151,704,1267,1294]" fontSize="11" level="1" pageId="2" pageNumber="241" reason="1">
<emphasis id="26DDEADAFFCA2932FF25FB3BFD2AFAAC" bold="true" box="[151,704,1267,1294]" pageId="2" pageNumber="241">
<taxonomicName id="D3A94D4BFFCA2932FF25FB3BFE71FAAF" authority="Porch &amp; Smith &amp; Greig, 2020" authorityName="Porch &amp; Smith &amp; Greig" authorityYear="2020" box="[151,411,1268,1294]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="241" phylum="Arthropoda" rank="species" species="raivavae" status="sp. nov.">
<emphasis id="26DDEADAFFCA2932FF25FB3BFE71FAAF" bold="true" box="[151,411,1268,1294]" italics="true" pageId="2" pageNumber="241">Pycnomerus raivavae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3DEE57A1FFCA2932FE10FB3BFDEAFAAF" box="[418,512,1268,1294]" pageId="2" pageNumber="241" rank="species">sp. nov.</taxonomicNameLabel>
Porch &amp; Greig
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5CB36543FFCA2932FF25FAD8FEEBFA93" box="[151,257,1303,1330]" pageId="2" pageNumber="241" type="description">
<paragraph id="141636C8FFCA2932FF25FAD8FEEBFA93" blockId="2.[151,704,1267,1330]" box="[151,257,1303,1330]" pageId="2" pageNumber="241">
(
<figureCitation id="8C922A4DFFCA2932FF12FAD8FF37FA90" box="[160,221,1303,1329]" captionStart="FIGURE 1" captionStartId="3.[151,250,1125,1150]" captionTargetBox="[162,1427,460,1092]" captionTargetId="figure@3.[155,1433,450,1098]" captionTargetPageId="3" captionText="FIGURE 1. Holotypes. Pycnomerus raivavae articulated prothorax and head, dorsal (top) and ventral (bottom)—(French Polynesia, Austral archipelago, Raivavae, Raʹirua swamp, RAI16-2 2016 cores, 150–200 cm depth). Pycnomerus rairua prothorax, dorsal (top) and ventral (bottom)—(French Polynesia, Austral archipelago, Raivavae, Raʹirua swamp, DA4 cores, 180–185 cm depth). Pycnomerus vavitu prothorax, dorsal (top) and ventral (bottom)—(French Polynesia, Austral archipelago, Raivavae, Raʹirua Swamp, DA4 2008 core, 170–175 cm depth). Pycnomerus mahanatoa prothorax, dorsal (top) and ventral (bottom)—(French Polynesia, Austral archipelago, Raivavae, Raʹirua Swamp, DA4 2008 core, 205–210 cm depth). Pycnomerus taralewisae prothorax, dorsal (top) and ventral (bottom)—(French Polynesia, Austral archipelago, Raivavae, Raʹirua swamp, DA4 cores, 245–250 cm depth). Scale bars = 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.3602378" httpUri="https://zenodo.org/record/3602378/files/figure.png" pageId="2" pageNumber="241">Fig.1</figureCitation>
,
<figureCitation id="8C922A4DFFCA2932FF5BFAD7FF12FA93" box="[233,248,1304,1330]" captionStart="FIGURE 2" captionStartId="4.[151,250,1516,1541]" captionTargetBox="[164,1427,819,1477]" captionTargetId="figure@4.[152,1436,812,1490]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal views of holotype and paratype prothoraces of P. raivavae; Holotype retains head indicated with ‘HT’. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3602380" httpUri="https://zenodo.org/record/3602380/files/figure.png" pageId="2" pageNumber="241">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5CB36543FFCA2932FF25FAAFFB97F9D4" pageId="2" pageNumber="241" type="materials_examined">
<paragraph id="141636C8FFCA2932FF25FAAFFBA5FADB" blockId="2.[151,1437,1375,2014]" box="[151,1103,1375,1402]" pageId="2" pageNumber="241">
<emphasis id="26DDEADAFFCA2932FF25FAAFFEAEFADB" bold="true" box="[151,324,1375,1402]" pageId="2" pageNumber="241">Type Locality:</emphasis>
Raʹirua Swamp,
<quantity id="D3519B2DFFCA2932FDB7FAAFFDB9FAD8" box="[517,595,1375,1402]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="241" unit="km" value="0.8">0.8 km</quantity>
E. of Raʹirua, at
<geoCoordinate id="719D500FFFCA2932FCA3FA90FC77FADB" box="[785,925,1375,1402]" degrees="23" direction="south" minutes="52.115" orientation="latitude" pageId="2" pageNumber="241" precision="1" value="-23.868584">23°52.115’S</geoCoordinate>
<geoCoordinate id="719D500FFFCA2932FC16FAAFFBA1FADB" box="[932,1099,1376,1402]" degrees="147" direction="west" minutes="40.718" orientation="longitude" pageId="2" pageNumber="241" precision="1" value="-147.67863">147°40.718’W</geoCoordinate>
.
</paragraph>
<paragraph id="141636C8FFCA2932FF75FA4BFE8EFA44" blockId="2.[151,1437,1375,2014]" pageId="2" pageNumber="241">
<emphasis id="26DDEADAFFCA2932FF75FA4BFE92FA3F" bold="true" box="[199,376,1411,1438]" pageId="2" pageNumber="241">Type material.</emphasis>
<materialsCitation id="A4C13C95FFCA2932FE33FA4BFE8EFA44" ID-GBIF-Occurrence="2545448277" collectingDate="2016-02-16" collectionCode="BPBM" country="French Polynesia" county="Ra'irua" location="Austral" municipality="Raivavae" pageId="2" pageNumber="241" specimenCount="1" typeStatus="holotype">
<emphasis id="26DDEADAFFCA2932FE33FA4BFE1DFA3F" bold="true" box="[385,503,1412,1438]" pageId="2" pageNumber="241">
<typeStatus id="CB12886AFFCA2932FE33FA4BFE18FA3F" box="[385,498,1412,1438]" pageId="2" pageNumber="241" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCountry id="6CBE7658FFCA2932FDB3FA4CFD22FA3C" box="[513,712,1411,1437]" name="French Polynesia" pageId="2" pageNumber="241">French Polynesia</collectingCountry>
,
<location id="11766013FFCA2932FD66FA4CFCCCFA3C" LSID="urn:lsid:plazi:treatment:9C0087DEFFCA2934FF25FB3BFA96FCB7:11766013FFCA2932FD66FA4CFCCCFA3C" box="[724,806,1411,1437]" country="French Polynesia" county="Ra'irua" municipality="Raivavae" name="Austral" pageId="2" pageNumber="241">Austral</location>
archipelago,
<collectingMunicipality id="F472ACB2FFCA2932FC76FA4CFBC5FA3C" box="[964,1071,1411,1437]" pageId="2" pageNumber="241">Raivavae</collectingMunicipality>
,
<collectingCounty id="FD774E44FFCA2932FB8FFA4CFB64FA3F" box="[1085,1166,1411,1438]" pageId="2" pageNumber="241">Raʹirua</collectingCounty>
swamp, RAI
<date id="60171008FFCA2932FA99FA4CFA76FA3C" box="[1323,1436,1411,1437]" pageId="2" pageNumber="241" value="2016-02-16">
<collectingDate id="7053E9E0FFCA2932FA99FA4CFA76FA3C" box="[1323,1436,1411,1437]" pageId="2" pageNumber="241" value="2016-02-16">16-2 2016</collectingDate>
</date>
cores,
<quantity id="D3519B2DFFCA2932FF52FA68FE86FA63" box="[224,364,1447,1474]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="2" pageNumber="241" unit="cm" value="175.0" valueMax="200.0" valueMin="150.0">150–200 cm</quantity>
depth—articulated head and prothorax, dorsally and ventrally complete except for appendages.
<collectionCode id="72B8AE0DFFCA2932FF25FA03FF02FA47" box="[151,232,1484,1510]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="2" pageNumber="241" type="Museum">BPBM</collectionCode>
, Honolulu.
</materialsCitation>
</paragraph>
<paragraph id="141636C8FFCA2932FF75FA3FFB97F9D4" blockId="2.[151,1437,1375,2014]" pageId="2" pageNumber="241">
<materialsCitation id="A4C13C95FFCA2932FF75FA3FFE92F98F" ID-GBIF-Occurrence="2545448267" country="French Polynesia" county="Ra'irua" location="Austral" municipality="Raivavae" pageId="2" pageNumber="241" specimenCount="1" typeStatus="paratype">
<emphasis id="26DDEADAFFCA2932FF75FA3FFE9AF9AB" bold="true" box="[199,368,1519,1546]" pageId="2" pageNumber="241">
<typeStatus id="CB12886AFFCA2932FF75FA3FFED5F9AB" box="[199,319,1520,1546]" pageId="2" pageNumber="241" type="paratype">Paratypes</typeStatus>
(8):
</emphasis>
<collectingCountry id="6CBE7658FFCA2932FEC5FA20FDD6F9A8" box="[375,572,1519,1545]" name="French Polynesia" pageId="2" pageNumber="241">French Polynesia</collectingCountry>
,
<location id="11766013FFCA2932FDF4FA20FD72F9A8" LSID="urn:lsid:plazi:treatment:9C0087DEFFCA2934FF25FB3BFA96FCB7:11766013FFCA2932FDF4FA20FD72F9A8" box="[582,664,1519,1545]" country="French Polynesia" county="Ra'irua" municipality="Raivavae" name="Austral" pageId="2" pageNumber="241">Austral</location>
archipelago,
<collectingMunicipality id="F472ACB2FFCA2932FC82FA20FC71F9A8" box="[816,923,1519,1545]" pageId="2" pageNumber="241">Raivavae</collectingMunicipality>
,
<collectingCounty id="FD774E44FFCA2932FC15FA20FC12F9AB" box="[935,1016,1519,1546]" pageId="2" pageNumber="241">Raʹirua</collectingCounty>
swamp, DA4 2008 core:
<quantity id="D3519B2DFFCA2932FAA2FA20FA76F9AB" box="[1296,1436,1519,1546]" metricMagnitude="0" metricUnit="m" metricValue="1.625" metricValueMax="1.65" metricValueMin="1.6" pageId="2" pageNumber="241" unit="cm" value="162.5" valueMax="165.0" valueMin="160.0">160–165 cm</quantity>
depth (1 prothorax)
</materialsCitation>
;
<materialsCitation id="A4C13C95FFCA2932FE36F9DBFC62F98F" ID-GBIF-Occurrence="2545448274" box="[388,904,1555,1582]" country="French Polynesia" county="Ra'irua" location="Austral" municipality="Raivavae" pageId="2" pageNumber="241" specimenCount="1" typeStatus="paratype">
<emphasis id="26DDEADAFFCA2932FE36F9DBFE5DF98F" box="[388,439,1556,1582]" italics="true" pageId="2" pageNumber="241">ditto</emphasis>
except
<quantity id="D3519B2DFFCA2932FDA0F9DCFD75F98F" box="[530,671,1555,1582]" metricMagnitude="0" metricUnit="m" metricValue="1.825" metricValueMax="1.85" metricValueMin="1.8" pageId="2" pageNumber="241" unit="cm" value="182.5" valueMax="185.0" valueMin="180.0">180–185 cm</quantity>
depth (1 prothorax)
</materialsCitation>
;
<materialsCitation id="A4C13C95FFCA2932FC27F9DBFA73F98F" ID-GBIF-Occurrence="2545448279" box="[917,1433,1555,1582]" country="French Polynesia" county="Ra'irua" location="Austral" municipality="Raivavae" pageId="2" pageNumber="241" specimenCount="1" typeStatus="paratype">
<emphasis id="26DDEADAFFCA2932FC27F9DBFC22F98F" box="[917,968,1556,1582]" italics="true" pageId="2" pageNumber="241">ditto</emphasis>
except
<quantity id="D3519B2DFFCA2932FB90F9DCFB45F98F" box="[1058,1199,1555,1582]" metricMagnitude="0" metricUnit="m" metricValue="1.925" metricValueMax="1.95" metricValueMin="1.9" pageId="2" pageNumber="241" unit="cm" value="192.5" valueMax="195.0" valueMin="190.0">190–195 cm</quantity>
depth (1 prothorax)
</materialsCitation>
;
<materialsCitation id="A4C13C95FFCA2932FF25F9F7FB93F9D4" ID-GBIF-Occurrence="2545448275" collectingDate="2016-02-16" collectionCode="BPBM" country="French Polynesia" county="Ra'irua" location="Austral" municipality="Raivavae" pageId="2" pageNumber="241" specimenCount="1" typeStatus="paratype">
<emphasis id="26DDEADAFFCA2932FF25F9F7FF20F9F3" box="[151,202,1592,1618]" italics="true" pageId="2" pageNumber="241">ditto</emphasis>
except
<quantity id="D3519B2DFFCA2932FE93F9F8FE46F9F3" box="[289,428,1591,1618]" metricMagnitude="0" metricUnit="m" metricValue="2.425" metricValueMax="2.45" metricValueMin="2.4" pageId="2" pageNumber="241" unit="cm" value="242.5" valueMax="245.0" valueMin="240.0">240–245 cm</quantity>
depth (1 incomplete prothorax); Raivavae, Raʹirua swamp, RAI
<date id="60171008FFCA2932FBCEF9F8FB01F9F0" box="[1148,1259,1591,1617]" pageId="2" pageNumber="241" value="2016-02-16">
<collectingDate id="7053E9E0FFCA2932FBCEF9F8FB01F9F0" box="[1148,1259,1591,1617]" pageId="2" pageNumber="241" value="2016-02-16">16-2 2016</collectingDate>
</date>
cores,
<quantity id="D3519B2DFFCA2932FA88F9F8FF53F9D7" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="2" pageNumber="241" unit="cm" value="175.0" valueMax="200.0" valueMin="150.0">150–200 cm</quantity>
depth (4 prothoraces: 1 incomplete).
<collectionCode id="72B8AE0DFFCA2932FDECF993FD45F9D7" box="[606,687,1628,1654]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="2" pageNumber="241" type="Museum">BPBM</collectionCode>
, Honolulu and Porch research collection
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5CB36543FFCA2932FF75F9B0FD5EF940" pageId="2" pageNumber="241" type="diagnosis">
<paragraph id="141636C8FFCA2932FF75F9B0FD5EF940" blockId="2.[151,1437,1375,2014]" pageId="2" pageNumber="241">
<emphasis id="26DDEADAFFCA2932FF75F9B0FEA8F93B" bold="true" box="[199,322,1663,1690]" pageId="2" pageNumber="241">Diagnosis.</emphasis>
Distinguished from co-occurring species and those from Rimatara (
<bibRefCitation id="70384B39FFCA2932FBF4F9B0FADFF93B" author="Porch, N. &amp; Smith, T. R." box="[1094,1333,1663,1690]" pageId="2" pageNumber="241" pagination="154 - 166" refId="ref7804" refString="Porch, N. &amp; Smith, T. R. (2017) New Pycnomerus Erichson (Coleoptera: Zopheridae: Pycnomerini) from Rimatara, French Polynesia. Zootaxa, 4237 (1), 154 - 166. https: // doi. org / 10.11646 / zootaxa. 4237.1.8" type="journal article" year="2017">Porch &amp; Smith 2017</bibRefCitation>
) and Tubuai (Greig &amp; Porch, unpubl. data) by its distinctive rounded shape, very sparse punctuation, largely effaced basal sulcus, and thickened, simple, pronotal margins.
</paragraph>
</subSubSection>
<subSubSection id="5CB36543FFCA2934FF75F924FB10FDC3" lastPageId="4" lastPageNumber="243" pageId="2" pageNumber="241" type="description">
<paragraph id="141636C8FFCA2932FF75F924FD04F8A7" blockId="2.[151,1437,1375,2014]" box="[199,750,1771,1798]" pageId="2" pageNumber="241">
<emphasis id="26DDEADAFFCA2932FF75F924FEB3F8A7" bold="true" box="[199,345,1771,1798]" pageId="2" pageNumber="241">Description.</emphasis>
Based on
<typeStatus id="CB12886AFFCA2932FE7CF924FDDAF8A4" box="[462,560,1771,1797]" pageId="2" pageNumber="241" type="holotype">holotype</typeStatus>
and
<specimenCount id="02AFFD41FFCA2932FDD4F923FD00F8A7" box="[614,746,1772,1798]" pageId="2" pageNumber="241" type="generic" typeStatus="paratypes">8 paratypes</specimenCount>
.
</paragraph>
<paragraph id="141636C8FFCA2932FF75F8C0FDE4F888" blockId="2.[151,1437,1375,2014]" box="[199,526,1807,1834]" pageId="2" pageNumber="241">
<emphasis id="26DDEADAFFCA2932FF75F8C0FECEF88B" bold="true" box="[199,292,1807,1834]" pageId="2" pageNumber="241">Colour:</emphasis>
Dark piceous-black.
</paragraph>
<paragraph id="141636C8FFCA2933FF75F8FBFB78FE0E" blockId="2.[151,1437,1375,2014]" lastBlockId="3.[151,1437,153,431]" lastPageId="3" lastPageNumber="242" pageId="2" pageNumber="241">
<emphasis id="26DDEADAFFCA2932FF75F8FBFEABF8EF" bold="true" box="[199,321,1844,1870]" pageId="2" pageNumber="241">Prothorax</emphasis>
approximately as wide as long, the greatest width at about 1/3—1/2 of pronotal length. Lateral margins arcuate and very weakly sinuate just before posterior angles. Lateral margin simple, with a marginal sulcus adjacent to a raised border, narrowest in middle and thickened anteriorly and especially posteriorly to hind angles. Anterior angles rounded and slightly produced. Anterior margin rounded to almost straight in middle, sinuate to anterior angles and unbordered. Posterior angles obtuse yet distinct. Posterior margin arcuate, with an almost effaced marginal channel, this depressed rather than sulcate and most well-developed inward of hind angles where it forms a small, shallow to moderately developed longitudinal fovea; in middle mostly effaced or as a slight transversely elongate depression. Basal border thick but indistinctly developed and with row of very small seta-bearing punctures.
<emphasis id="26DDEADAFFCB2933FF25FECAFED3FEBE" bold="true" box="[151,313,260,287]" pageId="3" pageNumber="242">Pronotal disk</emphasis>
elevated in front of middle, almost flat to weakly raised in medial third, declivous to all, especially lateral, margins. Disk not impressed. Punctures circular to mostly longitudinally ovate, separated by several to many times their length, bearing small, very pale yellowish setae (where preserved), shorter than puncture length. Punctures less abundant along middle, densest and most well developed laterally and more widely spaced posteriorly. Surface between punctures without obvious microsculpture, weakly to moderately shining.
</paragraph>
<caption id="40D66640FFCB2933FF25FBAAFD58FADB" ID-DOI="http://doi.org/10.5281/zenodo.3602378" ID-Zenodo-Dep="3602378" httpUri="https://zenodo.org/record/3602378/files/figure.png" pageId="3" pageNumber="242" startId="3.[151,250,1125,1150]" targetBox="[162,1427,460,1092]" targetPageId="3">
<paragraph id="141636C8FFCB2933FF25FBAAFD58FADB" blockId="3.[151,1437,1125,1402]" pageId="3" pageNumber="242">
<emphasis id="26DDEADAFFCB2933FF25FBAAFEF9FBDF" bold="true" box="[151,275,1125,1150]" pageId="3" pageNumber="242">FIGURE 1.</emphasis>
Holotypes.
<taxonomicName id="D3A94D4BFFCB2933FE3FFBA8FD83FBDF" authorityName="Porch &amp; Smith &amp; Greig" authorityYear="2020" box="[397,617,1127,1150]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="242" phylum="Arthropoda" rank="species" species="raivavae">
<emphasis id="26DDEADAFFCB2933FE3FFBA8FD83FBDF" box="[397,617,1127,1150]" italics="true" pageId="3" pageNumber="242">Pycnomerus raivavae</emphasis>
</taxonomicName>
articulated prothorax and head, dorsal (top) and ventral (bottom)—(French Polynesia, Austral archipelago, Raivavae, Raʹirua swamp, RAI16-2 2016 cores, 150–200 cm depth).
<taxonomicName id="D3A94D4BFFCB2933FBDAFB44FAC7FB03" authorityName="Porch &amp; Smith &amp; Greig" authorityYear="2020" box="[1128,1325,1163,1186]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="242" phylum="Arthropoda" rank="species" species="rairua">
<emphasis id="26DDEADAFFCB2933FBDAFB44FAC7FB03" box="[1128,1325,1163,1186]" italics="true" pageId="3" pageNumber="242">Pycnomerus rairua</emphasis>
</taxonomicName>
prothorax, dorsal (top) and ventral (bottom)—(French Polynesia, Austral archipelago, Raivavae, Raʹirua swamp, DA4 cores, 180–185 cm depth).
<taxonomicName id="D3A94D4BFFCB2933FEBCFB1CFE38FB4B" authorityName="Porch &amp; Smith &amp; Greig" authorityYear="2020" box="[270,466,1235,1258]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="242" phylum="Arthropoda" rank="species" species="vavitu">
<emphasis id="26DDEADAFFCB2933FEBCFB1CFE38FB4B" box="[270,466,1235,1258]" italics="true" pageId="3" pageNumber="242">Pycnomerus vavitu</emphasis>
</taxonomicName>
prothorax, dorsal (top) and ventral (bottom)—(French Polynesia, Austral archipelago, Raivavae, Raʹirua Swamp, DA4 2008 core, 170–175 cm depth).
<taxonomicName id="D3A94D4BFFCB2933FD4FFB38FC1FFAAF" authorityName="Porch &amp; Smith &amp; Greig" authorityYear="2020" box="[765,1013,1270,1294]" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="242" phylum="Arthropoda" rank="species" species="mahanatoa">
<emphasis id="26DDEADAFFCB2933FD4FFB38FC1FFAAF" box="[765,1013,1270,1294]" italics="true" pageId="3" pageNumber="242">Pycnomerus mahanatoa</emphasis>
</taxonomicName>
prothorax, dorsal (top) and ventral (bottom)—(French Polynesia, Austral archipelago, Raivavae, Raʹirua Swamp, DA4 2008 core, 205–210 cm depth).
<taxonomicName id="D3A94D4BFFCB2933FAADFAD4FEE7FAF7" authorityName="Porch &amp; Smith &amp; Greig" authorityYear="2020" class="Insecta" family="Zopheridae" genus="Pycnomerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="242" phylum="Arthropoda" rank="species" species="taralewisae">
<emphasis id="26DDEADAFFCB2933FAADFAD4FEE7FAF7" italics="true" pageId="3" pageNumber="242">Pycnomerus taralewisae</emphasis>
</taxonomicName>
prothorax, dorsal (top) and ventral (bottom)—(French Polynesia, Austral archipelago, Raivavae, Raʹirua swamp, DA4 cores, 245–250 cm depth). Scale bars = 0.5 mm
</paragraph>
</caption>
<paragraph id="141636C8FFCB2933FF75FA66FBCAF93A" blockId="3.[151,1437,1448,2015]" pageId="3" pageNumber="242">
<emphasis id="26DDEADAFFCB2933FF75FA66FEBCFA62" bold="true" box="[199,342,1449,1475]" pageId="3" pageNumber="242">Prosternum</emphasis>
moderately punctate with rounded punctures usually separated by about their length or more, becoming closer and much smaller anteriorly and coalescing on hind margin of prosternal process. Medial punctures with distinct anteriorly-oriented, elongated emargination, becoming weaker away from middle. Anterior with weak impunctate margin that extends across much of prosternum, defined by an indistinct ridge in middle half, with a fringe of short pale yellow setae in well-preserved specimens. Prosternal process very wide (3 three times coxal width) and parallel to very slightly divergent adjacent to coxae, until abrupt postcoxal expansion.Apex of prosternal process weakly angulate emarginate to truncate. Hypomera moderately punctate.
</paragraph>
<paragraph id="141636C8FFCB2933FF75F96AFAD3F8A7" blockId="3.[151,1437,1448,2015]" pageId="3" pageNumber="242">
<emphasis id="26DDEADAFFCB2933FF75F96AFEEDF91E" bold="true" box="[199,263,1701,1727]" pageId="3" pageNumber="242">Head</emphasis>
(based on
<typeStatus id="CB12886AFFCB2933FE33F96BFE09F91F" box="[385,483,1700,1726]" pageId="3" pageNumber="242" type="holotype">holotype</typeStatus>
specimen, head slightly pushed into prothorax). Almost completely impunctate anteriorly but weakly punctate behind antennal insertions. Eyes very small. Frontal margins weakly sinuate, narrowed and rounded to slightly medially protruded, anterior margin. Surface dull. Submentum densely punctured.
</paragraph>
<paragraph id="141636C8FFCB2933FF75F8DFFDD5F88A" blockId="3.[151,1437,1448,2015]" box="[199,575,1808,1835]" pageId="3" pageNumber="242">
<emphasis id="26DDEADAFFCB2933FF75F8DFFDD5F88A" bold="true" box="[199,575,1808,1835]" pageId="3" pageNumber="242">Physical dimensions and Ratios</emphasis>
</paragraph>
<paragraph id="141636C8FFCB2933FF75F8FAFC9EF81A" blockId="3.[151,1437,1448,2015]" pageId="3" pageNumber="242">
<emphasis id="26DDEADAFFCB2933FF75F8FAFEAAF8EE" bold="true" box="[199,320,1845,1871]" pageId="3" pageNumber="242">Pronotum</emphasis>
measurements based on
<specimenCount id="02AFFD41FFCB2933FDEBF8FAFD0DF8EF" box="[601,743,1844,1871]" pageId="3" pageNumber="242" type="generic">9 specimens</specimenCount>
. Measurements of
<typeStatus id="CB12886AFFCB2933FC09F8FBFBC9F8EF" box="[955,1059,1844,1870]" pageId="3" pageNumber="242" type="holotype">Holotype</typeStatus>
in parentheses. Pronotum length: 0.68–0.89 (0.75) mm. Pronotum width: 0.64–0.90 (0.76) mm. Anterior width of pronotum: 0.49–0.66 (0.57) mm. Medial width of pronotum: 0.65–0.92 (0.76) mm. Base of pronotum: 0.56–0.82 (0.65) mm. Lateral length of pronotum: 0.57–0.80 (0.64) mm. Length/Width ratio: 0.96–1.05 (1.00).
</paragraph>
<paragraph id="141636C8FFCB2934FF75F80AFBB7FF5A" blockId="3.[151,1437,1448,2015]" lastBlockId="4.[151,1437,152,791]" lastPageId="4" lastPageNumber="243" pageId="3" pageNumber="242">
<emphasis id="26DDEADAFFCB2933FF75F80AFEBCF87E" bold="true" box="[199,342,1989,2015]" pageId="3" pageNumber="242">Prosternum</emphasis>
measurements based on
<specimenCount id="02AFFD41FFCB2933FDC6F80AFCE8F87F" box="[628,770,1988,2015]" pageId="3" pageNumber="242" type="generic">4 specimens</specimenCount>
. Prosternum length: 0.44–0.55 (0.48) mm. Length proster- num in front of coxae: 0.28–0.32 (0.29) mm. Anterior foramen: 0.44–0.53 (0.51) mm. Basal foramen: 0.44–0.54 (0.44) mm. Prosternal process intercoxal width: 0.23–0.29 (0.27) mm. Maximum prosternal process width: 0.27– 0.32 (0.32) mm. Ratio of Prosternal length/Length in front of coxae: 1.56–1.67 (1.66).
</paragraph>
<paragraph id="141636C8FFCC2934FF75FECAFDF7FEE2" blockId="4.[151,1437,152,791]" pageId="4" pageNumber="243">
<emphasis id="26DDEADAFFCC2934FF75FECAFEEDFEBE" bold="true" box="[199,263,261,287]" pageId="4" pageNumber="243">Head</emphasis>
measurements based on
<typeStatus id="CB12886AFFCC2934FDAEFECBFD94FEBF" box="[540,638,260,286]" pageId="4" pageNumber="243" type="holotype">holotype</typeStatus>
with head partly inside prothorax. Clypeus width: (
<quantity id="D3519B2DFFCC2934FB08FECBFAC9FEBE" box="[1210,1315,260,287]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="4" pageNumber="243" unit="mm" value="0.46">0.46 mm</quantity>
). Interocular width (ventral): (est.
<quantity id="D3519B2DFFCC2934FE1AFEE6FDFAFEE2" box="[424,528,297,323]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="4" pageNumber="243" unit="mm" value="0.44">0.44 mm</quantity>
).
</paragraph>
<paragraph id="141636C8FFCC2934FF75FE83FB10FDC3" blockId="4.[151,1437,152,791]" pageId="4" pageNumber="243">
<emphasis id="26DDEADAFFCC2934FF75FE83FDD4FEC6" bold="true" box="[199,574,332,359]" pageId="4" pageNumber="243">Associated (Non-type) material</emphasis>
(sclerites other than prothorax or articulated head and prothorax). One distinctive elytral
<typeStatus id="CB12886AFFCC2934FEF0FEBEFE98FE2A" box="[322,370,369,395]" pageId="4" pageNumber="243">type</typeStatus>
almost certainly belongs to this species: it is the most common
<typeStatus id="CB12886AFFCC2934FBF5FEBEFB90FE2A" box="[1095,1146,369,395]" pageId="4" pageNumber="243">type</typeStatus>
, it is robust and wide and has similar sculpture to the prothorax:
<collectingCountry id="6CBE7658FFCC2934FDF4FE5BFCE0FE0F" box="[582,778,404,430]" name="French Polynesia" pageId="4" pageNumber="243">French Polynesia</collectingCountry>
, Austral archipelago, Raivavae, Raʹirua swamp, DA4 2008 excavation,
<quantity id="D3519B2DFFCC2934FEACFE77FE43FE72" box="[286,425,440,467]" metricMagnitude="0" metricUnit="m" metricValue="1.725" metricValueMax="1.75" metricValueMin="1.7" pageId="4" pageNumber="243" unit="cm" value="172.5" valueMax="175.0" valueMin="170.0">170–175 cm</quantity>
depth—1 complete left elytron, 1 complete right elytron;
<materialsCitation id="A4C13C95FFCC2934FB81FE77FD5FFE57" ID-GBIF-Occurrence="2545448278" pageId="4" pageNumber="243" specimenCount="1">
<emphasis id="26DDEADAFFCC2934FB81FE77FB8CFE73" box="[1075,1126,440,466]" italics="true" pageId="4" pageNumber="243">ditto</emphasis>
, except
<quantity id="D3519B2DFFCC2934FB77FE77FABAFE72" box="[1221,1360,440,467]" metricMagnitude="0" metricUnit="m" metricValue="1.675" metricValueMax="1.7" metricValueMin="1.65" pageId="4" pageNumber="243" unit="cm" value="167.5" valueMax="170.0" valueMin="165.0">165–170 cm</quantity>
depth, 1 complete left elytron, 1 complete right elytron
</materialsCitation>
;
<materialsCitation id="A4C13C95FFCC2934FD72FE13FABDFE57" ID-GBIF-Occurrence="2545448266" box="[704,1367,476,503]" pageId="4" pageNumber="243" specimenCount="1">
<emphasis id="26DDEADAFFCC2934FD72FE13FD19FE57" box="[704,755,476,502]" italics="true" pageId="4" pageNumber="243">ditto</emphasis>
, except
<quantity id="D3519B2DFFCC2934FCE3FE13FC36FE56" box="[849,988,476,503]" metricMagnitude="0" metricUnit="m" metricValue="1.425" metricValueMax="1.45" metricValueMin="1.4" pageId="4" pageNumber="243" unit="cm" value="142.5" valueMax="145.0" valueMin="140.0">140–145 cm</quantity>
depth, 1 incomplete right elytron
</materialsCitation>
;
<materialsCitation id="A4C13C95FFCC2934FAD0FE13FD0FFDBB" ID-GBIF-Occurrence="2545448276" pageId="4" pageNumber="243" specimenCount="1">
<emphasis id="26DDEADAFFCC2934FAD0FE13FA7FFE57" box="[1378,1429,476,502]" italics="true" pageId="4" pageNumber="243">ditto</emphasis>
, except
<quantity id="D3519B2DFFCC2934FF5BFDCFFE9CFDBA" box="[233,374,512,539]" metricMagnitude="0" metricUnit="m" metricValue="1.925" metricValueMax="1.95" metricValueMin="1.9" pageId="4" pageNumber="243" unit="cm" value="192.5" valueMax="195.0" valueMin="190.0">190–195 cm</quantity>
depth, 1 complete right elytron
</materialsCitation>
;
<materialsCitation id="A4C13C95FFCC2934FD40FDCFFB1CFDC3" ID-GBIF-Occurrence="2545448268" collectionCode="BPBM" country="French Polynesia" county="Representative" location="Austral" municipality="Ra'irua" pageId="4" pageNumber="243" specimenCount="1">
<emphasis id="26DDEADAFFCC2934FD40FDCFFCCFFDBB" box="[754,805,512,538]" italics="true" pageId="4" pageNumber="243">ditto</emphasis>
, except
<quantity id="D3519B2DFFCC2934FC35FDCFFBFEFDBA" box="[903,1044,512,539]" metricMagnitude="0" metricUnit="m" metricValue="1.525" metricValueMax="1.55" metricValueMin="1.5" pageId="4" pageNumber="243" unit="cm" value="152.5" valueMax="155.0" valueMin="150.0">150–155 cm</quantity>
depth, 1 incomplete right elytron.
<collectingCountry id="6CBE7658FFCC2934FF25FDEBFEB4FD9F" box="[151,350,548,574]" name="French Polynesia" pageId="4" pageNumber="243">French Polynesia</collectingCountry>
,
<location id="11766013FFCC2934FED8FDEBFE56FD9F" LSID="urn:lsid:plazi:treatment:9C0087DEFFCA2934FF25FB3BFA96FCB7:11766013FFCC2934FED8FDEBFE56FD9F" box="[362,444,548,574]" country="French Polynesia" county="Representative" municipality="Ra'irua" name="Austral" pageId="4" pageNumber="243">Austral</location>
archipelago,
<location id="11766013FFCC2934FDEAFDEBFD29FD9F" LSID="urn:lsid:plazi:treatment:9C0087DEFFCA2934FF25FB3BFA96FCB7:11766013FFCC2934FDEAFDEBFD29FD9F" box="[600,707,548,574]" country="French Polynesia" county="Representative" municipality="Ra'irua" name="Raivavae" pageId="4" pageNumber="243">Raivavae</location>
,
<collectingMunicipality id="F472ACB2FFCC2934FD62FDEBFCCBFD9E" box="[720,801,548,575]" pageId="4" pageNumber="243">Raʹirua</collectingMunicipality>
swamp, RAI16.2 2016 excavation,
<quantity id="D3519B2DFFCC2934FB0CFDEBFAA1FD9E" box="[1214,1355,548,575]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="4" pageNumber="243" unit="cm" value="175.0" valueMax="200.0" valueMin="150.0">150–200 cm</quantity>
deptharticulated left and right elytra.
<collectingCounty id="FD774E44FFCC2934FE48FD87FD4AFDC3" box="[506,672,584,610]" pageId="4" pageNumber="243">Representative</collectingCounty>
specimens in
<collectionCode id="72B8AE0DFFCC2934FCF2FD86FC7BFDC2" box="[832,913,585,611]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="4" pageNumber="243" type="Museum">BPBM</collectionCode>
, Honolulu and Porch collection
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5CB36543FFCC2934FF75FDA3FD13FD53" pageId="4" pageNumber="243" type="distribution">
<paragraph id="141636C8FFCC2934FF75FDA3FD13FD53" blockId="4.[151,1437,152,791]" pageId="4" pageNumber="243">
<emphasis id="26DDEADAFFCC2934FF75FDA3FE8BFD26" bold="true" box="[199,353,620,647]" pageId="4" pageNumber="243">Distribution.</emphasis>
Known only from Raʹirua swamp, Raivavae, and likely to be a Raivavae endemic. Subfossil and modern material from Tubuai and Rimatara does not include any material remotely close in appearance to this species. The fauna of nearby Rapa is poorly known, however the known modern collections and subfossil species present do not include this species. Potentially extinct.
</paragraph>
</subSubSection>
<subSubSection id="5CB36543FFCC2934FF75FD32FA96FCB7" box="[199,1404,764,791]" pageId="4" pageNumber="243" type="etymology">
<paragraph id="141636C8FFCC2934FF75FD32FA96FCB7" blockId="4.[151,1437,152,791]" box="[199,1404,764,791]" pageId="4" pageNumber="243">
<emphasis id="26DDEADAFFCC2934FF75FD32FEA4FCB6" bold="true" box="[199,334,765,791]" pageId="4" pageNumber="243">Etymology.</emphasis>
Named for its occurrence and apparent endemism on the island of Raivavae,
<collectingCountry id="6CBE7658FFCC2934FB01FD33FA93FCB7" box="[1203,1401,764,790]" name="French Polynesia" pageId="4" pageNumber="243">French Polynesia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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