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<document ID-DOI="http://dx.doi.org/10.3897/evolsyst.4.59893" ID-GBIF-Dataset="62c58258-c691-427e-9bf3-081a946616ee" ID-GBIF-Taxon="169091723" ID-Pensoft-Pub="2535-0730-2-71" ID-Pensoft-UUID="C2F8869E8A18583F93087CC85413B251" ID-ZooBank="4EEF928329EF47AEA1A5ED43ECF87299" LSID="urn:lsid:zoobank.org:act:D4C3AC48-60D0-4E02-9C37-FF09BA7099EA" ModsDocID="2535-0730-2-71" checkinTime="1605571711922" checkinUser="pensoft" docAuthor="Pellinen, Markku J., Zahiri, Reza &amp; Sihvonen, Pasi" docDate="2020" docId="878A3589AF665E3C82FC27A3DABF80D4" docLanguage="en" docName="EvolutSyst 4(2): 71-77" docOrigin="Evolutionary Systematics 4 (2)" docSource="http://dx.doi.org/10.3897/evolsyst.4.59893" docTitle="Sacada chaehomensis Pellinen &amp; Zahiri 2020, sp. nov." docType="treatment" docUuid="D4C3AC48-60D0-4E02-9C37-FF09BA7099EA" docUuidSource="ZooBank" docVersion="3" id="C2F8869E8A18583F93087CC85413B251" lastPageNumber="71" masterDocId="C2F8869E8A18583F93087CC85413B251" masterDocTitle="A new species of Sacada Walker, 1862 from Thailand (Lepidoptera, Pyralidae, Pyralinae)" masterLastPageNumber="77" masterPageNumber="71" pageId="0" pageNumber="71" updateTime="1643451100217" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Sacada Walker, 1862 from Thailand (Lepidoptera, Pyralidae, Pyralinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pellinen, Markku J.</mods:namePart>
<mods:affiliation>175, M 1 Muban Phichai, Mueang Lampang, Lampang, 52000, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">markku.j.pellinen@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zahiri, Reza</mods:namePart>
<mods:affiliation>Universitaet Hamburg, Centrum fuer Naturkunde, Martin-Luther-King Platz 3, 20146, Hamburg, Germany &amp; Canadian Food Inspection Agency, Ottawa Plant Laboratory, Entomology Unit, Bldg. 18, 960 Carling Ave., Ottawa, Ontario, K 1 A 0 C 6 Canada</mods:affiliation>
<mods:nameIdentifier type="email">reza.zahiri@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sihvonen, Pasi</mods:namePart>
<mods:affiliation>Finnish Museum of Natural History, Pohjoinen Rautatiekatu 13, FI- 00014, University of Helsinki, Helsinki, Finland</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-2237-9325</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Evolutionary Systematics</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>4</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>71</mods:start>
<mods:end>77</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/evolsyst.4.59893</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/evolsyst.4.59893</mods:identifier>
<mods:identifier type="Pensoft-Pub">2535-0730-2-71</mods:identifier>
<mods:identifier type="ZooBank">4EEF928329EF47AEA1A5ED43ECF87299</mods:identifier>
<mods:identifier type="Pensoft-UUID">C2F8869E8A18583F93087CC85413B251</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="169091723" LSID="urn:lsid:zoobank.org:act:D4C3AC48-60D0-4E02-9C37-FF09BA7099EA" httpUri="http://treatment.plazi.org/id/878A3589AF665E3C82FC27A3DABF80D4" lastPageNumber="71" pageId="0" pageNumber="71">
<subSubSection pageId="0" pageNumber="71" type="nomenclature">
<paragraph pageId="0" pageNumber="71">
<taxonomicName LSID="878A3589-AF66-5E3C-82FC-27A3DABF80D4" authority="Pellinen &amp; Zahiri" authorityName="Pellinen &amp; Zahiri" authorityYear="2020" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada chaehomensis" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chaehomensis" status="sp. nov.">Sacada chaehomensis Pellinen &amp; Zahiri</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 2–4" captionStartId="F2" captionText="Figures 2 - 4. Adults of Sacada spp. with diagnostic characters indicated 2 S. dzonguensis Singh, Kirti &amp; Ranjan (male) 3 S. umtasorensis Singh, Kirti &amp; Ranjan (male) 4 a S. chaehomensis sp. nov. (male, dorsal view) 4 b S. chaehomensis sp. nov. (male, ventral view). Figs 2, 3 based on Singh et al. (2020), used with permission." httpUri="https://binary.pensoft.net/fig/475843" pageId="0" pageNumber="71">Figs 4</figureCitation>
<figureCitation captionStart="Figures 5–7" captionStartId="F3" captionText="Figures 5 - 7. Male genitalia of Sacada with diagnostic characters indicated 5 S. dzonguensis Singh, Kirti &amp; Ranjan (holotype) 5 a aedeagus 5 b genitalia 6 S. umtasorensis Singh, Kirti &amp; Ranjan (holotype) 6 a aedeagus 6 b genitalia 7 S. chaehomensis sp. nov. (holotype, slide Sihvonen 2828) 7 a aedeagus with vesica everted 7 b aedeagus 7 c genitalia. Figs 5, 6 based on Singh et al. (2020), used with permission." httpUri="https://binary.pensoft.net/fig/475844" pageId="0" pageNumber="71">, 7</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F4" captionText="Figure 8. Descaled male abdomen of Sacada chaehomensis sp. nov. (holotype) 8 a abdomen 8 b 8 th tergite 8 c tympanal organs (slide Sihvonen 2828)." httpUri="https://binary.pensoft.net/fig/475845" pageId="0" pageNumber="71">, 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♂ (label data given verbatim,
<normalizedToken originalValue="“/”">&quot;/&quot;</normalizedToken>
indicates new line,
<normalizedToken originalValue="“;”">&quot;;&quot;</normalizedToken>
indicates new label]: Thailand Lampang/ Chae Hom 340 M/
<geoCoordinate degrees="18" direction="north" minutes="43" orientation="latitude" precision="15" seconds="19" value="18.721945">18°43'19&quot;N</geoCoordinate>
,
<geoCoordinate degrees="99" direction="east" minutes="33" orientation="longitude" precision="15" seconds="11" value="99.553055">99°33'11&quot;E</geoCoordinate>
/ 23.5.2020/ Leg. Pellinen, Markku [white label]; MJPT0020 [green label, indicates M. J. Pellinen DNA tissue sample]; ZMH-DNA0199 [indicates
<normalizedToken originalValue="Universität">Universitaet</normalizedToken>
Hamburg DNA process number]; 0146 [pink label, indicates M. J. Pellinen photograph number]; Pasi Sihvonen,/prep. number 2828 [blue label]; http://id.luomus.fi/GBT.1;
<emphasis italics="true" pageId="0" pageNumber="71">
<taxonomicName authorityName="Walker" authorityYear="1862" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Sacada</taxonomicName>
/
<taxonomicName authorityName="Pellinen &amp; Zahiri" authorityYear="2020" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="chaehomensis" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chaehomensis">chaehomensis</taxonomicName>
</emphasis>
/ Pellinen &amp; Zahiri/ HOLOTYPE [red label] (in coll. Finnish Museum of Natural History, Helsinki, Finland).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="materials_examined">
<paragraph pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="71">Only holotype male is known.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Pellinen &amp; Zahiri" authorityYear="2020" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada chaehomensis" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chaehomensis">
<emphasis italics="true" pageId="0" pageNumber="71">Sacada chaehomensis</emphasis>
</taxonomicName>
sp. nov. is morphologically similar to
<taxonomicName genus="S." lsidName="S. dzonguensis" pageId="0" pageNumber="71" rank="species" species="dzonguensis">
<emphasis italics="true" pageId="0" pageNumber="71">S. dzonguensis</emphasis>
</taxonomicName>
Singh, Kirti &amp; Ranjan, 2020 and
<taxonomicName genus="S." lsidName="S. umtasorensis" pageId="0" pageNumber="71" rank="species" species="umtasorensis">
<emphasis italics="true" pageId="0" pageNumber="71">S. umtasorensis</emphasis>
</taxonomicName>
Singh, Kirti &amp; Ranjan, 2020. Diagnostic male characters of these three species are indicated on Figs
<figureCitation captionStart="Figures 2–4" captionStartId="F2" captionText="Figures 2 - 4. Adults of Sacada spp. with diagnostic characters indicated 2 S. dzonguensis Singh, Kirti &amp; Ranjan (male) 3 S. umtasorensis Singh, Kirti &amp; Ranjan (male) 4 a S. chaehomensis sp. nov. (male, dorsal view) 4 b S. chaehomensis sp. nov. (male, ventral view). Figs 2, 3 based on Singh et al. (2020), used with permission." httpUri="https://binary.pensoft.net/fig/475843" pageId="0" pageNumber="71">2</figureCitation>
-
<figureCitation captionStart="Figures 5–7" captionStartId="F3" captionText="Figures 5 - 7. Male genitalia of Sacada with diagnostic characters indicated 5 S. dzonguensis Singh, Kirti &amp; Ranjan (holotype) 5 a aedeagus 5 b genitalia 6 S. umtasorensis Singh, Kirti &amp; Ranjan (holotype) 6 a aedeagus 6 b genitalia 7 S. chaehomensis sp. nov. (holotype, slide Sihvonen 2828) 7 a aedeagus with vesica everted 7 b aedeagus 7 c genitalia. Figs 5, 6 based on Singh et al. (2020), used with permission." httpUri="https://binary.pensoft.net/fig/475844" pageId="0" pageNumber="71">7</figureCitation>
(females of these three species are unknown). Forewing postmedial line is straight on costa and medial line is straight in
<taxonomicName genus="S." lsidName="S. chaehomensis" pageId="0" pageNumber="71" rank="species" species="chaehomensis">
<emphasis italics="true" pageId="0" pageNumber="71">S. chaehomensis</emphasis>
</taxonomicName>
sp. nov. (postmedial line curved and medial line angled in
<taxonomicName genus="S." lsidName="S. dzonguensis" pageId="0" pageNumber="71" rank="species" species="dzonguensis">
<emphasis italics="true" pageId="0" pageNumber="71">S. dzonguensis</emphasis>
</taxonomicName>
; postmedial line straight and medial line angled in
<taxonomicName genus="S." lsidName="S. umtasorensis" pageId="0" pageNumber="71" rank="species" species="umtasorensis">
<emphasis italics="true" pageId="0" pageNumber="71">S. umtasorensis</emphasis>
</taxonomicName>
). With regard to diagnostic male genitalia characters, see Figs
<figureCitation captionStart="Figures 5–7" captionStartId="F3" captionText="Figures 5 - 7. Male genitalia of Sacada with diagnostic characters indicated 5 S. dzonguensis Singh, Kirti &amp; Ranjan (holotype) 5 a aedeagus 5 b genitalia 6 S. umtasorensis Singh, Kirti &amp; Ranjan (holotype) 6 a aedeagus 6 b genitalia 7 S. chaehomensis sp. nov. (holotype, slide Sihvonen 2828) 7 a aedeagus with vesica everted 7 b aedeagus 7 c genitalia. Figs 5, 6 based on Singh et al. (2020), used with permission." httpUri="https://binary.pensoft.net/fig/475844" pageId="0" pageNumber="71">5-7</figureCitation>
.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/475843" pageId="0" pageNumber="71" start="Figures 2–4" startId="F2">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figures 2-4.</emphasis>
Adults of
<taxonomicName authorityName="Walker" authorityYear="1862" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Sacada</emphasis>
</taxonomicName>
spp. with diagnostic characters indicated
<emphasis bold="true" pageId="0" pageNumber="71">2</emphasis>
<taxonomicName genus="S." lsidName="S. dzonguensis" pageId="0" pageNumber="71" rank="species" species="dzonguensis">
<emphasis italics="true" pageId="0" pageNumber="71">S. dzonguensis</emphasis>
</taxonomicName>
Singh, Kirti &amp; Ranjan (male)
<emphasis bold="true" pageId="0" pageNumber="71">3</emphasis>
<taxonomicName genus="S." lsidName="S. umtasorensis" pageId="0" pageNumber="71" rank="species" species="umtasorensis">
<emphasis italics="true" pageId="0" pageNumber="71">S. umtasorensis</emphasis>
</taxonomicName>
Singh, Kirti &amp; Ranjan (male)
<emphasis bold="true" pageId="0" pageNumber="71">4a</emphasis>
<taxonomicName genus="S." lsidName="S. chaehomensis" pageId="0" pageNumber="71" rank="species" species="chaehomensis">
<emphasis italics="true" pageId="0" pageNumber="71">S. chaehomensis</emphasis>
</taxonomicName>
sp. nov. (male, dorsal view)
<emphasis bold="true" pageId="0" pageNumber="71">4b</emphasis>
<taxonomicName genus="S." lsidName="S. chaehomensis" pageId="0" pageNumber="71" rank="species" species="chaehomensis">
<emphasis italics="true" pageId="0" pageNumber="71">S. chaehomensis</emphasis>
</taxonomicName>
sp. nov. (male, ventral view). Figs
<emphasis bold="true" pageId="0" pageNumber="71">2, 3</emphasis>
based on
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.962.51194" author="Singh, N" journalOrPublisher="ZooKeys" pageId="0" pageNumber="71" pagination="139 - 163" refId="B12" refString="Singh, N, Kriti, JS, Ranjan, R, Chandra, K, Speidel, W, 2020. On the taxonomy of the genus Sacada Walker, 1862 from India, with descriptions of a new genus and two new species (Pyralinae, Pyralidae, Lepidoptera). ZooKeys 962: 139 - 163, DOI: https://doi.org/10.3897/zookeys.962.51194" title="On the taxonomy of the genus Sacada Walker, 1862 from India, with descriptions of a new genus and two new species (Pyralinae, Pyralidae, Lepidoptera)." url="https://doi.org/10.3897/zookeys.962.51194" volume="962" year="2020">Singh et al. (2020)</bibRefCitation>
, used with permission.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/475844" pageId="0" pageNumber="71" start="Figures 5–7" startId="F3">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figures 5-7.</emphasis>
Male genitalia of
<taxonomicName authorityName="Walker" authorityYear="1862" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Sacada</emphasis>
</taxonomicName>
with diagnostic characters indicated
<emphasis bold="true" pageId="0" pageNumber="71">5</emphasis>
<taxonomicName genus="S." lsidName="S. dzonguensis" pageId="0" pageNumber="71" rank="species" species="dzonguensis">
<emphasis italics="true" pageId="0" pageNumber="71">S. dzonguensis</emphasis>
</taxonomicName>
Singh, Kirti &amp; Ranjan (holotype)
<emphasis bold="true" pageId="0" pageNumber="71">5a</emphasis>
aedeagus
<emphasis bold="true" pageId="0" pageNumber="71">5b</emphasis>
genitalia
<emphasis bold="true" pageId="0" pageNumber="71">6</emphasis>
<taxonomicName genus="S." lsidName="S. umtasorensis" pageId="0" pageNumber="71" rank="species" species="umtasorensis">
<emphasis italics="true" pageId="0" pageNumber="71">S. umtasorensis</emphasis>
</taxonomicName>
Singh, Kirti &amp; Ranjan (holotype)
<emphasis bold="true" pageId="0" pageNumber="71">6a</emphasis>
aedeagus
<emphasis bold="true" pageId="0" pageNumber="71">6b</emphasis>
genitalia
<emphasis bold="true" pageId="0" pageNumber="71">7</emphasis>
<taxonomicName genus="S." lsidName="S. chaehomensis" pageId="0" pageNumber="71" rank="species" species="chaehomensis">
<emphasis italics="true" pageId="0" pageNumber="71">S. chaehomensis</emphasis>
</taxonomicName>
sp. nov. (holotype, slide Sihvonen 2828)
<emphasis bold="true" pageId="0" pageNumber="71">7a</emphasis>
aedeagus with vesica everted
<emphasis bold="true" pageId="0" pageNumber="71">7b</emphasis>
aedeagus
<emphasis bold="true" pageId="0" pageNumber="71">7c</emphasis>
genitalia. Figs
<emphasis bold="true" pageId="0" pageNumber="71">5, 6</emphasis>
based on
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.962.51194" author="Singh, N" journalOrPublisher="ZooKeys" pageId="0" pageNumber="71" pagination="139 - 163" refId="B12" refString="Singh, N, Kriti, JS, Ranjan, R, Chandra, K, Speidel, W, 2020. On the taxonomy of the genus Sacada Walker, 1862 from India, with descriptions of a new genus and two new species (Pyralinae, Pyralidae, Lepidoptera). ZooKeys 962: 139 - 163, DOI: https://doi.org/10.3897/zookeys.962.51194" title="On the taxonomy of the genus Sacada Walker, 1862 from India, with descriptions of a new genus and two new species (Pyralinae, Pyralidae, Lepidoptera)." url="https://doi.org/10.3897/zookeys.962.51194" volume="962" year="2020">Singh et al. (2020)</bibRefCitation>
, used with permission.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">Description.</paragraph>
<paragraph pageId="0" pageNumber="71">
Description is based on male, female is unknown. Wingspan 40 mm. Antennae with basal two thirds bipectinate, apical third fasciculate. Head, thorax and abdomen rufous. Tegula with iridescent scales of purplish-grey hue. Labial palps short. Base of proboscis covered densely with scales. Fore- and midleg tibia with massive hair tuft, spur formula of legs 0-2-4. Forewing narrow, reddish brown, medial, postmedial and terminal lines dry straw colored (fuscous). Medial line angled, postmedial line straight, forming triangular area on midwing. Terminal line narrow, fringes concolorous with wings. Hind wing wide, grey, weakly rufous near margin, veins darker. Terminal line fuscous, fringes concolorous with wings. Forewing underside rufous grey-brown, medially darker and inner margin pale. Hindwing underside rufous grey-brown on costa, otherwise paler. Postmedial line weakly visible on both wings. Tympanal organs large, semi-circular, fornix tympani separate but connected via narrow sclerotised ridge (in Fig.
<figureCitation captionStart="Figures 5–7" captionStartId="F3" captionText="Figures 5 - 7. Male genitalia of Sacada with diagnostic characters indicated 5 S. dzonguensis Singh, Kirti &amp; Ranjan (holotype) 5 a aedeagus 5 b genitalia 6 S. umtasorensis Singh, Kirti &amp; Ranjan (holotype) 6 a aedeagus 6 b genitalia 7 S. chaehomensis sp. nov. (holotype, slide Sihvonen 2828) 7 a aedeagus with vesica everted 7 b aedeagus 7 c genitalia. Figs 5, 6 based on Singh et al. (2020), used with permission." httpUri="https://binary.pensoft.net/fig/475844" pageId="0" pageNumber="71">7c</figureCitation>
metathorax sclerotisations and metafurca are visible between tympanal organs). Abdominal segments weakly sclerotised, 8th tergite with two posterior lobes, 8th sternite with round posterior margin.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Male genitalia.</emphasis>
Uncus hood-shaped, curved, densely setose, with rounded lobes basolaterally. Gnathos sclerotised, arms forming narrow bands, arms fused medially forming upwards directed weakly dentate hook; basolateral parts curved inwards, lobe-shaped. Tegumen formed of two narrow plates. Valva simple, setose, pointing weakly outwards; costa straight. Transtilla large, sclerotised, plate-like, with distinct elongated, upwards directed projection medially, its apex blunt. Juxta broad, weakly constricted laterally, anterior margin even, posterior margin with two heavily sclerotised arms bearing few spikes. Vinculum narrow. Saccus elongated, apex round. Aedeagus stout and weakly undulating, caecum round, apical apodeme ventrally with small group of dentate spikes. Vesica large, simple, without distinct diverticula, basal part wider, apical part narrow and weakly curved; base covered with minute spikes, followed by minute sclerotisations, those become fewer towards vesica apex.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="etymology">
<paragraph pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="71">The species is named after its type locality, Chae Hom, in Lampang province, Thailand.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="biology_ecology">
<paragraph pageId="0" pageNumber="71">Biology.</paragraph>
<paragraph pageId="0" pageNumber="71">
The single known male was collected in May 2020 at 340 m altitude; it was attracted to light in forest with diverse tree species, interspersed with small vegetable plantations (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F5" captionText="Figure 9. Collecting site of Sacada chaehomensis sp. nov. Thailand, Lampang, Chae Hom, 8.9.2013, photo by Markku Pellinen." httpUri="https://binary.pensoft.net/fig/475846" pageId="0" pageNumber="71">9</figureCitation>
). Immature stages are unknown.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/475845" pageId="0" pageNumber="71" start="Figure 8" startId="F4">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 8.</emphasis>
Descaled male abdomen of
<taxonomicName authorityName="Pellinen &amp; Zahiri" authorityYear="2020" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada chaehomensis" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chaehomensis">
<emphasis italics="true" pageId="0" pageNumber="71">Sacada chaehomensis</emphasis>
</taxonomicName>
sp. nov. (holotype)
<emphasis bold="true" pageId="0" pageNumber="71">8a</emphasis>
abdomen
<emphasis bold="true" pageId="0" pageNumber="71">8b</emphasis>
8th tergite
<emphasis bold="true" pageId="0" pageNumber="71">8c</emphasis>
tympanal organs (slide Sihvonen 2828).
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/475846" pageId="0" pageNumber="71" start="Figure 9" startId="F5">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 9.</emphasis>
Collecting site of
<taxonomicName authorityName="Pellinen &amp; Zahiri" authorityYear="2020" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada chaehomensis" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chaehomensis">
<emphasis italics="true" pageId="0" pageNumber="71">Sacada chaehomensis</emphasis>
</taxonomicName>
sp. nov. Thailand, Lampang, Chae Hom, 8.9.2013, photo by Markku Pellinen.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">The only specimen was collected in Lampang (Chae Hom) in northern Thailand.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="barcode analysis">
<paragraph pageId="0" pageNumber="71">Barcode analysis.</paragraph>
<paragraph pageId="0" pageNumber="71">
A 642 bp barcode of
<taxonomicName authorityName="Pellinen &amp; Zahiri" authorityYear="2020" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada chaehomensis" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chaehomensis">
<emphasis italics="true" pageId="0" pageNumber="71">Sacada chaehomensis</emphasis>
</taxonomicName>
sp. nov. was submitted to the identification engine on BOLD. Genetically nearest neighbors are unidentified pyralids from Malaysia, Sabah (North of Borneo) with a minimum divergence of 6.17% and another from Malaysia, North of Kuala Lumpur with 6.50%. The other pre-existing data for
<taxonomicName authorityName="Walker" authorityYear="1862" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Sacada</emphasis>
</taxonomicName>
on BOLD were
<taxonomicName authorityName="Snellen" authorityYear="1892" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada ragonotalis" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ragonotalis">
<emphasis italics="true" pageId="0" pageNumber="71">Sacada ragonotalis</emphasis>
</taxonomicName>
(6.54%) and
<taxonomicName authorityName="Hampson" authorityYear="1917" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada albioculalis" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="albioculalis">
<emphasis italics="true" pageId="0" pageNumber="71">Sacada albioculalis</emphasis>
</taxonomicName>
(8.93%), thus indicating that the unidentified pyralids may be
<taxonomicName authorityName="Walker" authorityYear="1862" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Sacada</emphasis>
</taxonomicName>
sp. The species morphologically most similar (i.e.,
<taxonomicName genus="S." lsidName="S. dzonguensis" pageId="0" pageNumber="71" rank="species" species="dzonguensis">
<emphasis italics="true" pageId="0" pageNumber="71">S. dzonguensis</emphasis>
</taxonomicName>
and
<taxonomicName genus="S." lsidName="S. umtasorensis" pageId="0" pageNumber="71" rank="species" species="umtasorensis">
<emphasis italics="true" pageId="0" pageNumber="71">S. umtasorensis</emphasis>
</taxonomicName>
) to the new taxon lacked representations on the BOLD DNA library. The tree topology generated by the BI analysis (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F6" captionText="Figure 10. The Bayesian tree from analysis of the COI 5 P sequences of Sacada chaehomensis sp. nov. and 17 public Sacada DNA sequences. The nodes are scored by posterior probabilities (PP)." httpUri="https://binary.pensoft.net/fig/475847" pageId="0" pageNumber="71">10</figureCitation>
) was very similar to the NJ-tree confirming the association (PP = 0.87) of the new species (ZMH199) with the sister pair of unidentified pyralids from Malaysia, Borneo (LPMLY526-14+ LPMLY208-14) and Malaysia, North of Kuala Lumpur (LPMAL244-14+LEPML271-13). The relationships between analyzed
<taxonomicName authorityName="Walker" authorityYear="1862" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Sacada</emphasis>
</taxonomicName>
must be treated with caution, because those are based on short DNA barcode sequences only, and are likely to change with addition of more molecular data and expanding taxon sampling.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/475847" pageId="0" pageNumber="71" start="Figure 10" startId="F6">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 10.</emphasis>
The Bayesian tree from analysis of the COI5P sequences of
<taxonomicName authorityName="Pellinen &amp; Zahiri" authorityYear="2020" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada chaehomensis" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="chaehomensis">
<emphasis italics="true" pageId="0" pageNumber="71">Sacada chaehomensis</emphasis>
</taxonomicName>
sp. nov. and 17 public
<taxonomicName authorityName="Walker" authorityYear="1862" class="Insecta" family="Pyralidae" genus="Sacada" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sacada" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Sacada</emphasis>
</taxonomicName>
DNA sequences. The nodes are scored by posterior probabilities (PP).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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