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<document id="3ABF7444F10363918C980830C6BE6111" ID-DOI="10.5281/zenodo.275907" ID-GBIF-Dataset="d5d27825-82a5-4aaf-b47d-35d24b33aaaa" ID-ISSN="1175-5326" ID-Zenodo-Dep="275907" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460189977646" checkinUser="plazi" docAuthor="Lott, Derek A." docDate="2010" docId="4D54A843FFB61A6E0694FE3DEC8AFF39" docLanguage="en" docName="zt02402p051.pdf" docOrigin="Zootaxa 2402" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Acylophorus makhoreae Lott, new species" docType="treatment" docVersion="5" lastPageNumber="15" masterDocId="B16DD03BFFB81A610603FFAFEE40FFEE" masterDocTitle="The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa" masterLastPageNumber="51" masterPageNumber="1" pageNumber="15" updateTime="1698236417163" updateUser="plazi">
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<mods:title id="8672B1B190E3E7C550C7F3026FB26F90">The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa</mods:title>
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<mods:namePart id="1AF8FDC143312743864227EBD7E99EA9">Lott, Derek A.</mods:namePart>
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<mods:date id="5898C789553A3BFE515947E7AE341387">2010</mods:date>
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<treatment id="4D54A843FFB61A6E0694FE3DEC8AFF39" ID-DOI="http://doi.org/10.5281/zenodo.6196019" ID-GBIF-Taxon="127655319" ID-Zenodo-Dep="6196019" LSID="urn:lsid:plazi:treatment:4D54A843FFB61A6E0694FE3DEC8AFF39" httpUri="http://treatment.plazi.org/id/4D54A843FFB61A6E0694FE3DEC8AFF39" lastPageId="15" lastPageNumber="15" pageId="14" pageNumber="15">
<subSubSection id="8DE74ADEFFB61A6F0694FE3DEFF0FE21" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="C5421955FFB61A6F0694FE3DECDAFE42" blockId="14.[151,666,402,463]" box="[151,666,402,428]" pageId="14" pageNumber="15">
<heading id="9E0AAE39FFB61A6F0694FE3DECDAFE42" bold="true" box="[151,666,402,428]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="F789C547FFB61A6F0694FE3DECDAFE42" bold="true" box="[151,666,402,428]" pageId="14" pageNumber="15">
<taxonomicName id="02FD62D6FFB61A6F0694FE3DEFBDFE42" ID-CoL="9W8F" authority="Lott" authorityName="Lott" box="[151,509,402,428]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="makhoreae" status="sp. nov.">
<emphasis id="F789C547FFB61A6F0694FE3DEFFEFE42" bold="true" box="[151,446,402,428]" italics="true" pageId="14" pageNumber="15">Acylophorus makhoreae</emphasis>
Lott
</taxonomicName>
,
<taxonomicNameLabel id="ECBA783CFFB61A6F0409FE3DECDAFE42" box="[522,666,402,428]" pageId="14" pageNumber="15" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<paragraph id="C5421955FFB61A6F0694FE1AEFF0FE21" blockId="14.[151,666,402,463]" box="[151,432,437,463]" pageId="14" pageNumber="15">
(
<figureCitation id="5DC605D0FFB61A6F069CFE1AEEAAFE21" box="[159,234,437,463]" captionStart="FIGURES 1 8" captionStartId="5.[151,272,1933,1957]" captionTargetBox="[171,1408,165,1908]" captionTargetId="figure@5.[156,1441,151,1941]" captionTargetPageId="5" captionText="FIGURES 1 8. Forebodies of species in the Acylophorus orientalis group. 1, A. orientalis Fauvel; 2, A. nitens new species; 3, A. rossii Bordoni, 4. A. bumbunae Bordoni; 5, A. capensis Cameron; 6, A. dankalensis Bordoni; 7, A. tshuapensis new species; 8, A. makhoreae new species." httpUri="https://zenodo.org/record/275908/files/figure.png" pageId="14" pageNumber="15">Figs 8</figureCitation>
,
<figureCitation id="5DC605D0FFB61A6F06FBFE1AEF58FE21" box="[248,280,437,463]" captionStart="FIGURES 33 41" captionStartId="34.[151,270,1501,1525]" captionTargetBox="[173,1411,192,1467]" captionTargetId="figure@34.[155,1440,183,1495]" captionTargetPageId="34" captionText="FIGURES 33 41. Mandibles of Acylophorus species. 33, A. orientalis Fauvel; 34, A. nitens new species; 35, A. rossii Bordoni, 36, A. capensis Cameron; 37, A. dankalensis Bordoni; 38, A. makhoreae new species; 39, A. antennalis Cameron (RD Congo); 40, A. antennalis Cameron (R Congo); 41, A. collarti Cameron." httpUri="https://zenodo.org/record/275912/files/figure.png" pageId="14" pageNumber="15">38</figureCitation>
,
<figureCitation id="5DC605D0FFB61A6F0720FE1AEF03FE21" box="[291,323,437,463]" captionStart="FIGURES 51 76" captionStartId="36.[151,271,1818,1842]" captionTargetBox="[176,1425,305,1773]" captionTargetId="figure@36.[155,1440,293,1797]" captionTargetPageId="36" captionText="FIGURES 51 76. Terminal segments of the maxilliary palpi of Acylophorus species. 51, A. orientalis Fauvel; 52, A. nitens new species; 53, A. rossii Bordoni, 54, A. bumbunae Bordoni; 55, A. capensis Cameron; 56, A. dankalensis Bordoni; 57, A. tshuapensis new species; 58, A. makhoreae new species; 59, A. densipennis Bernhauer; 60, A. antennalis Cameron; 61, A. collarti Cameron; 62. A. micans new species; 63, A. allardi Levasseur; 64, A. congoensis Cameron; 65, A. ziloensis Levasseur; 66, A. mareei Bernhauer; 67, A. salifi new species; 68, A. lomaensis Levasseur; 69. A. minor new species; 70, A. trigonocephalus Cameron; 71, A. setiger new species; 72, A. usambarae Bernhauer; 73, A. grandis Bernhauer; 74, A. bafutensis Levasseur; 75, A. kambuiensis Bordoni, 76. A. lualabaensis Levasseur." httpUri="https://zenodo.org/record/275914/files/figure.png" pageId="14" pageNumber="15">58</figureCitation>
,
<figureCitation id="5DC605D0FFB61A6F0753FE1AEF30FE21" box="[336,368,437,463]" captionStart="FIGURES 77 96" captionStartId="37.[151,271,1853,1877]" captionTargetBox="[209,1389,189,1828]" captionTargetId="figure@37.[168,1428,165,1848]" captionTargetPageId="37" captionText="FIGURES 77 96. Antennae of Acylophorus species. 77, A. orientalis Fauvel; 78, A. nitens new species; 79, A. rossii Bordoni, 80, A. bumbunae Bordoni; 81, A. capensis Cameron; 82, A. dankalensis Bordoni; 83, A. tshuapensis new species; 84, A. makhoreae new species; 85, A. densipennis Bernhauer; 86, A. antennalis Cameron; 87, A. collarti Cameron; 88. A. micans new species; 89, A. allardi Levasseur; 90, A. congoensis Cameron; 91, A. ziloensis Levasseur; 92, A. mareei Bernhauer; 93, A. methneri Bernhauer; 94, A. salifi new species; 95, A. lomaensis Levasseur; 96. A. minor new species." httpUri="https://zenodo.org/record/275915/files/figure.png" pageId="14" pageNumber="15">84</figureCitation>
,
<figureCitation id="5DC605D0FFB61A6F0778FE1AEFE7FE21" box="[379,423,437,463]" captionStart="FIGURES 109 116" captionStartId="39.[151,271,1945,1969]" captionTargetBox="[226,1362,196,1910]" captionTargetId="figure@39.[207,1395,159,1942]" captionTargetPageId="39" captionText="FIGURES 109 116. Aedeagi of species in the Acylophorus orientalis group. 109, A. orientalis Fauvel; 110, A. nitens new species; 111, A. rossii Bordoni, 112. A. bumbunae Bordoni; 113, A. capensis Cameron; 114, A. dankalensis Bordoni; 115, A. tshuapensis new species; 116, A. makhoreae new species." httpUri="https://zenodo.org/record/275917/files/figure.png" pageId="14" pageNumber="15">116</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="8DE74ADEFFB61A6F0694FE50ED15FAEA" pageId="14" pageNumber="15" type="description">
<paragraph id="C5421955FFB61A6F0694FE50ECFDFD64" blockId="14.[151,1437,511,2031]" pageId="14" pageNumber="15">
<emphasis id="F789C547FFB61A6F0694FE50EF6FFDF7" bold="true" box="[151,303,511,537]" pageId="14" pageNumber="15">Description.</emphasis>
Length
<quantity id="0205B4B0FFB61A6F0792FDAFEF91FDF4" box="[401,465,512,538]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="15" unit="mm" value="7.0">7mm</quantity>
. Body colour very variable ranging from pale brown to black. Pronotum yellow to red-brown with a dark, diffuse mark covering the disc and sometimes suffused almost to the edges. Abdomen iridescent when dark. Appendages similarly variable in colour, but the terminal segment of the maxillary palpi is always darker than the penultimate segment.
</paragraph>
<paragraph id="C5421955FFB61A6F06C7FD3AEACBFC11" blockId="14.[151,1437,511,2031]" pageId="14" pageNumber="15">
Head of average size (pronotum 1.75x wider than head), more or less as long as wide with rounded temples not very evident (
<figureCitation id="5DC605D0FFB61A6F07C6FD15EC4CFD3A" box="[453,524,698,724]" captionStart="FIGURES 1 8" captionStartId="5.[151,272,1933,1957]" captionTargetBox="[171,1408,165,1908]" captionTargetId="figure@5.[156,1441,151,1941]" captionTargetPageId="5" captionText="FIGURES 1 8. Forebodies of species in the Acylophorus orientalis group. 1, A. orientalis Fauvel; 2, A. nitens new species; 3, A. rossii Bordoni, 4. A. bumbunae Bordoni; 5, A. capensis Cameron; 6, A. dankalensis Bordoni; 7, A. tshuapensis new species; 8, A. makhoreae new species." httpUri="https://zenodo.org/record/275908/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
). Pigmented area of head extending well in front of antennal insertion. Micropunctures extending over much of head, but very sparse away from the front of the head and next to the eyes. Dense short pubescence behind eyes. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other. Only four postocular setae visible from above on each side. Underside of head sparsely pubescent, depressed at base with gular sutures separate, but very proximate toward base. Right mandible with one sharp median tooth; left mandible lacking sharp tooth (
<figureCitation id="5DC605D0FFB61A6F022FFCDAEAC8FC61" box="[1068,1160,885,911]" captionStart="FIGURES 33 41" captionStartId="34.[151,270,1501,1525]" captionTargetBox="[173,1411,192,1467]" captionTargetId="figure@34.[155,1440,183,1495]" captionTargetPageId="34" captionText="FIGURES 33 41. Mandibles of Acylophorus species. 33, A. orientalis Fauvel; 34, A. nitens new species; 35, A. rossii Bordoni, 36, A. capensis Cameron; 37, A. dankalensis Bordoni; 38, A. makhoreae new species; 39, A. antennalis Cameron (RD Congo); 40, A. antennalis Cameron (R Congo); 41, A. collarti Cameron." httpUri="https://zenodo.org/record/275912/files/figure.png" pageId="14" pageNumber="15">Fig. 38</figureCitation>
). Maxillary palpi with terminal segment densely pubescent, with rounded angle on outer margin, asymmetric and less elongate than
<emphasis id="F789C547FFB61A6F0694FC6FEF70FC37" box="[151,304,960,985]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="02FD62D6FFB61A6F0694FC6FEF6BFC37" box="[151,299,960,985]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="orientalis">A. orientalis</taxonomicName>
,
</emphasis>
longer than glabrous penultimate segment which is slightly elongate (
<figureCitation id="5DC605D0FFB61A6F026EFC6FEA86FC34" box="[1133,1222,960,986]" captionStart="FIGURES 51 76" captionStartId="36.[151,271,1818,1842]" captionTargetBox="[176,1425,305,1773]" captionTargetId="figure@36.[155,1440,293,1797]" captionTargetPageId="36" captionText="FIGURES 51 76. Terminal segments of the maxilliary palpi of Acylophorus species. 51, A. orientalis Fauvel; 52, A. nitens new species; 53, A. rossii Bordoni, 54, A. bumbunae Bordoni; 55, A. capensis Cameron; 56, A. dankalensis Bordoni; 57, A. tshuapensis new species; 58, A. makhoreae new species; 59, A. densipennis Bernhauer; 60, A. antennalis Cameron; 61, A. collarti Cameron; 62. A. micans new species; 63, A. allardi Levasseur; 64, A. congoensis Cameron; 65, A. ziloensis Levasseur; 66, A. mareei Bernhauer; 67, A. salifi new species; 68, A. lomaensis Levasseur; 69. A. minor new species; 70, A. trigonocephalus Cameron; 71, A. setiger new species; 72, A. usambarae Bernhauer; 73, A. grandis Bernhauer; 74, A. bafutensis Levasseur; 75, A. kambuiensis Bordoni, 76. A. lualabaensis Levasseur." httpUri="https://zenodo.org/record/275914/files/figure.png" pageId="14" pageNumber="15">Fig. 58</figureCitation>
). First segment of antenna as long as next five. Segments I to IV elongate, VII to XI transverse (
<figureCitation id="5DC605D0FFB61A6F0226FC4AEA3DFC11" box="[1061,1149,997,1023]" captionStart="FIGURES 77 96" captionStartId="37.[151,271,1853,1877]" captionTargetBox="[209,1389,189,1828]" captionTargetId="figure@37.[168,1428,165,1848]" captionTargetPageId="37" captionText="FIGURES 77 96. Antennae of Acylophorus species. 77, A. orientalis Fauvel; 78, A. nitens new species; 79, A. rossii Bordoni, 80, A. bumbunae Bordoni; 81, A. capensis Cameron; 82, A. dankalensis Bordoni; 83, A. tshuapensis new species; 84, A. makhoreae new species; 85, A. densipennis Bernhauer; 86, A. antennalis Cameron; 87, A. collarti Cameron; 88. A. micans new species; 89, A. allardi Levasseur; 90, A. congoensis Cameron; 91, A. ziloensis Levasseur; 92, A. mareei Bernhauer; 93, A. methneri Bernhauer; 94, A. salifi new species; 95, A. lomaensis Levasseur; 96. A. minor new species." httpUri="https://zenodo.org/record/275915/files/figure.png" pageId="14" pageNumber="15">Fig. 84</figureCitation>
).
</paragraph>
<paragraph id="C5421955FFB61A6F06C7FBA5EF38FB54" blockId="14.[151,1437,511,2031]" pageId="14" pageNumber="15">
Pronotum only slightly transverse (1.1x wider than long) with rounded sides and widest in basal half. Shining with no micro-punctures. Dorsal, lateral and marginal setae shorter than in
<taxonomicName id="02FD62D6FFB61A6F02A5FB9FEB01FBA7" box="[1190,1345,1072,1097]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="F789C547FFB61A6F02A5FB9FEB01FBA7" box="[1190,1345,1072,1097]" italics="true" pageId="14" pageNumber="15">A. orientalis</emphasis>
</taxonomicName>
. Elytra transverse (1.7x wider than long) with pubescence arising from asperate punctures. Apical fringe of bristles longer than the hairs on the rest of the elytra. Abdominal tergites with evenly spaced, relatively sparse asperate punctures.
</paragraph>
<paragraph id="C5421955FFB61A6F06C7FB6AED15FAEA" blockId="14.[151,1437,511,2031]" pageId="14" pageNumber="15">
Paramere bilobed, each lobe fairly flat, at least toward apex, pegs concentrated in dense mass at apex (
<figureCitation id="5DC605D0FFB61A6F036FFB6AEE86FAEA" captionStart="FIGURES 109 116" captionStartId="39.[151,271,1945,1969]" captionTargetBox="[226,1362,196,1910]" captionTargetId="figure@39.[207,1395,159,1942]" captionTargetPageId="39" captionText="FIGURES 109 116. Aedeagi of species in the Acylophorus orientalis group. 109, A. orientalis Fauvel; 110, A. nitens new species; 111, A. rossii Bordoni, 112. A. bumbunae Bordoni; 113, A. capensis Cameron; 114, A. dankalensis Bordoni; 115, A. tshuapensis new species; 116, A. makhoreae new species." httpUri="https://zenodo.org/record/275917/files/figure.png" pageId="14" pageNumber="15">Fig. 116</figureCitation>
). Median lobe longer than paramere with truncate apex.
</paragraph>
</subSubSection>
<subSubSection id="8DE74ADEFFB61A6F06C7FAA0EA42F9C1" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="C5421955FFB61A6F06C7FAA0EA42F9C1" blockId="14.[151,1437,511,2031]" pageId="14" pageNumber="15">
<emphasis id="F789C547FFB61A6F06C7FAA0EFB2FAC7" bold="true" box="[196,498,1295,1321]" pageId="14" pageNumber="15">
<typeStatus id="1A46A7F7FFB61A6F06C7FAA0EF41FAC7" box="[196,257,1295,1321]" pageId="14" pageNumber="15">Type</typeStatus>
material.
<typeStatus id="1A46A7F7FFB61A6F0783FAA0EFB2FAC7" box="[384,498,1295,1321]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
</emphasis>
3: “Under plants at a stream edge /
<collectingCountry id="BDEA59C5FFB61A6F0594FABFEA63FAC4" box="[919,1059,1296,1322]" name="Ethiopia" pageId="14" pageNumber="15">ETHIOPIA</collectingCountry>
: Kaffa
<quantity id="0205B4B0FFB61A6F0275FABFEAFDFAC4" box="[1142,1213,1296,1322]" metricMagnitude="4" metricUnit="m" metricValue="2.8" pageId="14" pageNumber="15" unit="km" value="28.0">28km</quantity>
. S. of Jimma 2,000 m
<date id="B1433F95FFB61A6F06C2FA9AEF5DFAA1" box="[193,285,1333,1359]" pageId="14" pageNumber="15" value="1971-11">xi.1971</date>
/ R.
<collectingCountry id="BDEA59C5FFB61A6F0756FA9AEF2FFAA1" box="[341,367,1333,1359]" name="Iceland" pageId="14" pageNumber="15">O</collectingCountry>
.S. Clark B.M. 1973-450 /
<typeStatus id="1A46A7F7FFB61A6F04B0FA9AED12FAA1" box="[691,850,1333,1359]" pageId="14" pageNumber="15" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="02FD62D6FFB61A6F0559FA9AEAC1FAA0" box="[858,1153,1333,1358]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="makhoreae" status="sp. nov.">
<emphasis id="F789C547FFB61A6F0559FA9AEAC1FAA0" box="[858,1153,1333,1358]" italics="true" pageId="14" pageNumber="15">Acylophorus makhoreae</emphasis>
</taxonomicName>
<emphasis id="F789C547FFB61A6F0288FA9AEA91FAA1" bold="true" box="[1163,1233,1333,1359]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="ECBA783CFFB61A6F0288FA9AEA91FAA1" box="[1163,1233,1333,1359]" pageId="14" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
3 det. DA Lott, 2009” (
<collectionCode id="A3EC8190FFB61A6F06EEFAF5EF0CFA9A" box="[237,332,1370,1396]" pageId="14" pageNumber="15">BMNH</collectionCode>
).
<emphasis id="F789C547FFB61A6F075DFAF5EF9BFA9A" bold="true" box="[350,475,1370,1396]" pageId="14" pageNumber="15">
<typeStatus id="1A46A7F7FFB61A6F075DFAF5EF9BFA9A" box="[350,475,1370,1396]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
</emphasis>
13 1Ƥ: “
<collectingCountry id="BDEA59C5FFB61A6F0448FAF5EC9AFA9A" box="[587,730,1370,1396]" name="Ethiopia" pageId="14" pageNumber="15">ETHIOPIA</collectingCountry>
: ILUBADOR
<quantity id="0205B4B0FFB61A6F0582FAF5ED88FA9A" box="[897,968,1370,1396]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="15" unit="km" value="10.0">10km</quantity>
. W. Bedelle 0825N 3618E
<date id="B1433F95FFB61A6F0300FAF5EB12FA9A" box="[1283,1362,1370,1396]" pageId="14" pageNumber="15" value="1972-10">x.1972</date>
<quantity id="0205B4B0FFB61A6F0359FAF5EEF1FA74" metricMagnitude="0" metricUnit="m" metricValue="1.8" pageId="14" pageNumber="15" unit="m" value="1.8">1,800 m</quantity>
. / R.
<collectingCountry id="BDEA59C5FFB61A6F06EBFA2FEF41FA74" box="[232,257,1408,1434]" name="Iceland" pageId="14" pageNumber="15">O</collectingCountry>
.S. Clark B.M. 1973-450 /
<typeStatus id="1A46A7F7FFB61A6F0430FA2FEC84FA74" box="[563,708,1408,1434]" pageId="14" pageNumber="15" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="02FD62D6FFB61A6F04C8FA2FEDA7FA77" box="[715,999,1408,1433]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="makhoreae" status="sp. nov.">
<emphasis id="F789C547FFB61A6F04C8FA2FEDA7FA77" box="[715,999,1408,1433]" italics="true" pageId="14" pageNumber="15">Acylophorus makhoreae</emphasis>
</taxonomicName>
<emphasis id="F789C547FFB61A6F05EDFAD0EA70FA77" bold="true" box="[1006,1072,1407,1433]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="ECBA783CFFB61A6F05EDFAD0EA70FA77" box="[1006,1072,1407,1433]" pageId="14" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
det. DA Lott, 2009” (
<collectionCode id="A3EC8190FFB61A6F0332FA2FEBD0FA74" box="[1329,1424,1408,1434]" pageId="14" pageNumber="15">BMNH</collectionCode>
); 4: “Under plants &amp; stones at edge stream /
<collectingCountry id="BDEA59C5FFB61A6F049FFA0AED69FA51" box="[668,809,1445,1471]" name="Ethiopia" pageId="14" pageNumber="15">ETHIOPIA</collectingCountry>
: Kaffa Belleta F.
<quantity id="0205B4B0FFB61A6F05F9FA0AEA1CFA51" box="[1018,1116,1445,1471]" metricMagnitude="0" metricUnit="m" metricValue="2.1" pageId="14" pageNumber="15" unit="m" value="2.1">2,100 m</quantity>
<quantity id="0205B4B0FFB61A6F0266FA0AEAEDFA51" box="[1125,1197,1445,1471]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="15" unit="km" value="40.0">40km</quantity>
. S.W. Jimma 1971 / R.
<collectingCountry id="BDEA59C5FFB61A6F06B0FA65EE8CFA0A" box="[179,204,1482,1508]" name="Iceland" pageId="14" pageNumber="15">O</collectingCountry>
.S. Clark B.M. 1973-450 /
<typeStatus id="1A46A7F7FFB61A6F07FCFA65ECD0FA0A" box="[511,656,1482,1508]" pageId="14" pageNumber="15" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="02FD62D6FFB61A6F0494FA64EDF3FA0A" box="[663,947,1483,1508]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="makhoreae" status="sp. nov.">
<emphasis id="F789C547FFB61A6F0494FA64EDF3FA0A" box="[663,947,1483,1508]" italics="true" pageId="14" pageNumber="15">Acylophorus makhoreae</emphasis>
</taxonomicName>
<emphasis id="F789C547FFB61A6F05B8FA65EDBCFA0A" bold="true" box="[955,1020,1482,1508]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="ECBA783CFFB61A6F05B8FA65EDBCFA0A" box="[955,1020,1482,1508]" pageId="14" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
det. DA Lott, 2009” (
<collectionCode id="A3EC8190FFB61A6F02FEFA65EB1CFA0A" box="[1277,1372,1482,1508]" pageId="14" pageNumber="15">BMNH</collectionCode>
); 1Ƥ: “ih red ref. /
<collectingCountry id="BDEA59C5FFB61A6F073AFA5FEF8BF9E4" box="[313,459,1520,1546]" name="Ethiopia" pageId="14" pageNumber="15">ETHIOPIA</collectingCountry>
: Kaffa Jimma to Addis Rd. 0749-370
<date id="B1433F95FFB61A6F05A7FA5FEA47F9E4" box="[932,1031,1520,1546]" pageId="14" pageNumber="15" value="1972-03">iii.1972</date>
. / R.
<collectingCountry id="BDEA59C5FFB61A6F0247FA5FEA1EF9E4" box="[1092,1118,1520,1546]" name="Iceland" pageId="14" pageNumber="15">O</collectingCountry>
.S. Clark B.M. 1973-450 /
<typeStatus id="1A46A7F7FFB61A6F0694F9BAEF69F9C1" box="[151,297,1557,1583]" pageId="14" pageNumber="15" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="02FD62D6FFB61A6F072CF9BAEC0CF9C0" box="[303,588,1557,1582]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="makhoreae" status="sp. nov.">
<emphasis id="F789C547FFB61A6F072CF9BAEC0CF9C0" box="[303,588,1557,1582]" italics="true" pageId="14" pageNumber="15">Acylophorus makhoreae</emphasis>
</taxonomicName>
<emphasis id="F789C547FFB61A6F0450F9BAECD5F9C1" bold="true" box="[595,661,1557,1583]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="ECBA783CFFB61A6F0450F9BAECD5F9C1" box="[595,661,1557,1583]" pageId="14" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
det. DA Lott, 2009” (
<collectionCode id="A3EC8190FFB61A6F0594F9BAEDB6F9C1" box="[919,1014,1557,1583]" pageId="14" pageNumber="15">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="8DE74ADEFFB61A6F06C7F995EC06F8DA" pageId="14" pageNumber="15" type="distribution">
<paragraph id="C5421955FFB61A6F06C7F995EA8CF971" blockId="14.[151,1437,511,2031]" pageId="14" pageNumber="15">
<emphasis id="F789C547FFB61A6F06C7F995EC61F9BA" bold="true" box="[196,545,1594,1620]" pageId="14" pageNumber="15">Distribution and bionomics.</emphasis>
All material seen so far comes from the south-west of
<collectingCountry id="BDEA59C5FFB61A6F02A7F995EB47F9BA" box="[1188,1287,1594,1620]" name="Ethiopia" pageId="14" pageNumber="15">Ethiopia</collectingCountry>
(
<figureCitation id="5DC605D0FFB61A6F0314F995EB3DF9BA" box="[1303,1405,1594,1620]" captionStart="FIGURE 142" captionStartId="45.[151,260,891,915]" captionTargetBox="[153,1436,186,864]" captionTargetId="figure@45.[151,1436,186,864]" captionTargetPageId="45" captionText="FIGURE 142. Distribution of species in the Acylophorus orientalis group = A. bumbunae Bordoni, = A. tshuapensis new species, = A. makhoreae new species)." httpUri="https://zenodo.org/record/275923/files/figure.png" pageId="14" pageNumber="15">Fig. 142</figureCitation>
). I have also seen a female specimen collected at “8,000” feet from the Djem-Djem Forest to the west of Addis Ababa that may belong to this species. Most of the specimens were collected along streams.
</paragraph>
<paragraph id="C5421955FFB61A6F06C7F905EC06F8DA" blockId="14.[151,1437,511,2031]" pageId="14" pageNumber="15">
<emphasis id="F789C547FFB61A6F06C7F905EFF9F92A" bold="true" box="[196,441,1706,1732]" pageId="14" pageNumber="15">Comparative notes.</emphasis>
Fairly distinct within the
<taxonomicName id="02FD62D6FFB61A6F04EDF904EDC0F92A" box="[750,896,1707,1732]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="F789C547FFB61A6F04EDF904EDC0F92A" box="[750,896,1707,1732]" italics="true" pageId="14" pageNumber="15">A. orientalis</emphasis>
</taxonomicName>
species group by virtue of the position of the antennal insertion, the broader, more asymmetric terminal segment of the maxillary palpi and the more transverse medial antennal segments. The form of the aedeagus and the arrangement of medial teeth on the mandibles are also useful characters.
</paragraph>
</subSubSection>
<subSubSection id="8DE74ADEFFB61A6F06C7F890EE9EF84A" pageId="14" pageNumber="15" type="etymology">
<paragraph id="C5421955FFB61A6F06C7F890EE9EF84A" blockId="14.[151,1437,511,2031]" pageId="14" pageNumber="15">
<emphasis id="F789C547FFB61A6F06C7F890EF11F8B7" bold="true" box="[196,337,1855,1881]" pageId="14" pageNumber="15">Etymology.</emphasis>
Makhore was a legendary queen, who founded the kingdom of Jimma in the area where many of the
<typeStatus id="1A46A7F7FFB61A6F06E5F8CAEF5AF891" box="[230,282,1893,1919]" pageId="14" pageNumber="15">type</typeStatus>
specimens were collected. The species name is the genitive case of a Latinised version of her name.
</paragraph>
</subSubSection>
<subSubSection id="8DE74ADEFFB61A6E06C7F800EC8AFF39" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="discussion">
<paragraph id="C5421955FFB61A6E06C7F800EC8AFF39" blockId="14.[151,1437,511,2031]" lastBlockId="15.[151,1436,152,215]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="F789C547FFB61A6F06C7F800EF16F827" bold="true" box="[196,342,1967,1993]" pageId="14" pageNumber="15">Discussion.</emphasis>
<taxonomicName id="02FD62D6FFB61A6F0763F81FEC4BF827" box="[352,523,1968,1993]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="makhoreae">
<emphasis id="F789C547FFB61A6F0763F81FEC4BF827" box="[352,523,1968,1993]" italics="true" pageId="14" pageNumber="15">A. makhoreae</emphasis>
</taxonomicName>
has been placed in the
<taxonomicName id="02FD62D6FFB61A6F0532F81FED8CF827" box="[817,972,1968,1993]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="F789C547FFB61A6F0532F81FED8CF827" box="[817,972,1968,1993]" italics="true" pageId="14" pageNumber="15">A. orientalis</emphasis>
</taxonomicName>
group, because it lacks dense micropunctures on the head and pronotum. However, the form of the maxillary palpi, the mandibles and the aedeagus and the position of the antennal insertion point all suggest that it may be more closely related to some of the species in the
<taxonomicName id="02FD62D6FFB71A6E07CFFF12EC37FF38" box="[460,631,189,214]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="densipennis">
<emphasis id="F789C547FFB71A6E07CFFF12EC37FF38" box="[460,631,189,214]" italics="true" pageId="15" pageNumber="16">A. densipennis</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
</treatment>
</document>