144 lines
16 KiB
XML
144 lines
16 KiB
XML
<document id="32A23CD56349989B2B41663D9B404234" ID-DOI="10.5281/zenodo.275907" ID-GBIF-Dataset="d5d27825-82a5-4aaf-b47d-35d24b33aaaa" ID-ISSN="1175-5326" ID-Zenodo-Dep="275907" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460189977646" checkinUser="plazi" docAuthor="Lott, Derek A." docDate="2010" docId="4D54A843FFA61A7F0694FDDDEF4AF951" docLanguage="en" docName="zt02402p051.pdf" docOrigin="Zootaxa 2402" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Acylophorus usambarae Bernhauer" docType="treatment" docVersion="5" lastPageNumber="31" masterDocId="B16DD03BFFB81A610603FFAFEE40FFEE" masterDocTitle="The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa" masterLastPageNumber="51" masterPageNumber="1" pageNumber="31" updateTime="1698236417163" updateUser="plazi">
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<mods:title id="35B1A3F8B99E40DEB0844749A514A984">The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa</mods:title>
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<mods:namePart id="E089C27D8DAC8340C1382E602E3F36C3">Lott, Derek A.</mods:namePart>
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<mods:title id="33267CA04A91C9FFFBF01A6B1AF21D93">Zootaxa</mods:title>
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<mods:part id="7E77FB12E653767F52E7B63FA40C4FFB">
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<mods:date id="EB4E8004231D2BC4FFD68EBDE0BDAA15">2010</mods:date>
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<mods:detail id="D83E0D4EA9BF8E353046464E663A8AA8" type="volume">
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<mods:number id="DAC70F29FEA21E898B68F6B451B6BCED">2402</mods:number>
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<mods:identifier id="36EDC57BCE497B743BFA054361B99B37" type="DOI">10.5281/zenodo.275907</mods:identifier>
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<mods:identifier id="D666B3DB40F400AC67A824A36878AF91" type="ISSN">1175-5326</mods:identifier>
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<treatment id="4D54A843FFA61A7F0694FDDDEF4AF951" ID-DOI="http://doi.org/10.5281/zenodo.6196059" ID-GBIF-Taxon="127655335" ID-Zenodo-Dep="6196059" LSID="urn:lsid:plazi:treatment:4D54A843FFA61A7F0694FDDDEF4AF951" httpUri="http://treatment.plazi.org/id/4D54A843FFA61A7F0694FDDDEF4AF951" lastPageNumber="31" pageId="30" pageNumber="31">
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<subSubSection id="8DE74ADEFFA61A7F0694FDDDEF19FD41" pageId="30" pageNumber="31" type="nomenclature">
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<paragraph id="C5421955FFA61A7F0694FDDDEC0BFD62" blockId="30.[151,587,626,687]" box="[151,587,626,652]" pageId="30" pageNumber="31">
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<heading id="9E0AAE39FFA61A7F0694FDDDEC0BFD62" bold="true" box="[151,587,626,652]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
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<taxonomicName id="02FD62D6FFA61A7F0694FDDDEC0BFD62" ID-CoL="65399" authority="Bernhauer" authorityName="Bernhauer" box="[151,587,626,652]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="usambarae">
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<emphasis id="F789C547FFA61A7F0694FDDDEC0BFD62" bold="true" box="[151,587,626,652]" pageId="30" pageNumber="31">
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<emphasis id="F789C547FFA61A7F0694FDDDEFFDFD62" bold="true" box="[151,445,626,652]" italics="true" pageId="30" pageNumber="31">Acylophorus usambarae</emphasis>
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Bernhauer
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="C5421955FFA61A7F0694FD3AEF19FD41" blockId="30.[151,587,626,687]" box="[151,345,661,687]" pageId="30" pageNumber="31">(Figs 23, 72, 99)</paragraph>
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</subSubSection>
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<subSubSection id="8DE74ADEFFA61A7F0694FD4FEF35FC8A" pageId="30" pageNumber="31" type="reference_group">
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<paragraph id="C5421955FFA61A7F0694FD4FEA3CFD19" blockId="30.[151,1148,736,759]" box="[151,1148,736,759]" pageId="30" pageNumber="31">
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<treatmentCitationGroup id="E5ED3E7BFFA61A7F0694FD4FEA3CFD19" box="[151,1148,736,759]" pageId="30" pageNumber="31">
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<taxonomicName id="02FD62D6FFA61A7F0694FD4FEC3EFD19" ID-CoL="65399" authority="Bernhauer, 1917: 48" authorityName="Bernhauer" authorityPageNumber="48" authorityYear="1917" box="[151,638,736,759]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="usambarae">
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<emphasis id="F789C547FFA61A7F0694FD4FEFDAFD19" box="[151,410,736,759]" italics="true" pageId="30" pageNumber="31">Acylophorus usambarae</emphasis>
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<treatmentCitation id="445C3F44FFA61A7F07A2FD4FEC3EFD19" author="Bernhauer" box="[417,638,736,759]" page="48" pageId="30" pageNumber="31" year="1917">
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<bibRefCitation id="A16C64A4FFA61A7F07A2FD4FEC15FD19" author="Bernhauer" box="[417,597,736,759]" pageId="30" pageNumber="50" refString="Bernhauer, M. (1917). Beitrag zur Staphylinidenfauna des tropischen Afrika. Entomologische Blatter, 13, 46 - 49." type="journal article" year="1917">Bernhauer, 1917</bibRefCitation>
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: 48
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</treatmentCitation>
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</taxonomicName>
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;
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<treatmentCitation id="445C3F44FFA61A7F048AFD4FEDCEFD19" author="Scheerpeltz" box="[649,910,736,759]" page="1467" pageId="30" pageNumber="31" year="1933">
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<bibRefCitation id="A16C64A4FFA61A7F048AFD4FED0BFD19" author="Scheerpeltz" box="[649,843,736,759]" pageId="30" pageNumber="51" refString="Scheerpeltz, O. (1933). Staphylinidae VII. In: S. Schenkling (ed.), Coleopterorum Catalogus, 6 (129). Junk, Berlin, pp. 989 - 1500." type="journal article" year="1933">Scheerpeltz, 1933</bibRefCitation>
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: 1467
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</treatmentCitation>
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;
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<treatmentCitation id="445C3F44FFA61A7F059AFD4FEA39FD19" author="Herman" box="[921,1145,736,759]" page="3038" pageId="30" pageNumber="31" year="2001">
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<bibRefCitation id="A16C64A4FFA61A7F059AFD4FEA75FD19" author="Herman" box="[921,1077,736,759]" pageId="30" pageNumber="50" refString="Herman, L. H. (2001). Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3). Bulletin of the American Museum of Natural History, 265, 3021 - 3840." type="journal article" year="2001">Herman, 2001</bibRefCitation>
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: 3038
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="C5421955FFA61A7F0694FC8AEF35FC8A" blockId="30.[151,1437,805,1727]" pageId="30" pageNumber="31">
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<emphasis id="F789C547FFA61A7F0694FC8AEF0DFCD1" bold="true" box="[151,333,805,831]" pageId="30" pageNumber="31">Redescription.</emphasis>
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Length
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<quantity id="0205B4B0FFA61A7F07B3FC8AEFB0FCD1" box="[432,496,805,831]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="30" pageNumber="31" unit="mm" value="7.0">7mm</quantity>
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. Body brown with head darker. Legs and palpi pale, antennae brown with paler terminal segments.
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</paragraph>
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</subSubSection>
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<subSubSection id="8DE74ADEFFA61A7F06C7FCDFEFD9FA7A" pageId="30" pageNumber="31" type="description">
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<paragraph id="C5421955FFA61A7F06C7FCDFEFC3FB84" blockId="30.[151,1437,805,1727]" pageId="30" pageNumber="31">
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Head large (pronotum 1.5x wider than long) slightly transverse (1.1x wider than long), temples well developed (Fig. 23). Area in front of antennal insertion pigmented. Micro-punctures very sparse. Short dense pubescence behind eyes. Three pairs of interocular setae and a line of five postocular setae visible from above on each side. Maxillary palpi with last two segments densely pubescent, terminal segment much more rounded on outer margin than inner margin, so markedly asymmetric, longer than penultimate segment which is also elongate (
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<figureCitation id="5DC605D0FFA61A7F075EFB85EFF7FBAA" box="[349,439,1066,1092]" captionStart="FIGURES 51 – 76" captionStartId="36.[151,271,1818,1842]" captionTargetBox="[176,1425,305,1773]" captionTargetId="figure@36.[155,1440,293,1797]" captionTargetPageId="36" captionText="FIGURES 51 – 76. Terminal segments of the maxilliary palpi of Acylophorus species. 51, A. orientalis Fauvel; 52, A. nitens new species; 53, A. rossii Bordoni, 54, A. bumbunae Bordoni; 55, A. capensis Cameron; 56, A. dankalensis Bordoni; 57, A. tshuapensis new species; 58, A. makhoreae new species; 59, A. densipennis Bernhauer; 60, A. antennalis Cameron; 61, A. collarti Cameron; 62. A. micans new species; 63, A. allardi Levasseur; 64, A. congoensis Cameron; 65, A. ziloensis Levasseur; 66, A. mareei Bernhauer; 67, A. salifi new species; 68, A. lomaensis Levasseur; 69. A. minor new species; 70, A. trigonocephalus Cameron; 71, A. setiger new species; 72, A. usambarae Bernhauer; 73, A. grandis Bernhauer; 74, A. bafutensis Levasseur; 75, A. kambuiensis Bordoni, 76. A. lualabaensis Levasseur." httpUri="https://zenodo.org/record/275914/files/figure.png" pageId="30" pageNumber="31">Fig. 72</figureCitation>
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). First segment of antenna as long as next five. Segments I to V elongate, VIII to XI transverse (
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<figureCitation id="5DC605D0FFA61A7F071EFBFFEF35FB84" box="[285,373,1104,1130]" captionStart="FIGURES 97 – 103" captionStartId="38.[151,270,1806,1830]" captionTargetBox="[178,1406,194,1766]" captionTargetId="figure@38.[161,1418,178,1785]" captionTargetPageId="38" captionText="FIGURES 97 – 103. Antennae of Acylophorus species. 97, A. trigonocephalus Cameron; 98, A. setiger new species; 99, A. usambarae Bernhauer; 100, A. grandis Bernhauer; 101, A. bafutensis Levasseur; 102, A. kambuiensis Bordoni, 103. A. lualabaensis Levasseur. FIGURES 104 – 107. Details of Acylophorus delphinus Fauvel. 104, forebody; 105, terminal segments of maxilliary palpi; 106, antenna; 107, aedeagus. FIGURE 108. Lateral view of head of Acylophorus salifi new species." httpUri="https://zenodo.org/record/275916/files/figure.png" pageId="30" pageNumber="31">Fig. 99</figureCitation>
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).
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</paragraph>
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<paragraph id="C5421955FFA61A7F06C7FBDAEFD9FA7A" blockId="30.[151,1437,805,1727]" pageId="30" pageNumber="31">
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Pronotum transverse (1.2x wider than long) with rounded sides and widest at centre. Shining with no micro-punctures. One pair of dorsal and one pair of lateral setae. Marginal setae long and more numerous than in
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<taxonomicName id="02FD62D6FFA61A7F06B5FB6FEF0BFB37" box="[182,331,1216,1241]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="orientalis">
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<emphasis id="F789C547FFA61A7F06B5FB6FEF0BFB37" box="[182,331,1216,1241]" italics="true" pageId="30" pageNumber="31">A. orientalis</emphasis>
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</taxonomicName>
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. Elytra transverse (1.6x wider than long) with long pubescence, asperate punctures stronger but less dense than in
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<taxonomicName id="02FD62D6FFA61A7F079CFB4AEC71FB10" box="[415,561,1253,1278]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="orientalis">
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<emphasis id="F789C547FFA61A7F079CFB4AEC71FB10" box="[415,561,1253,1278]" italics="true" pageId="30" pageNumber="31">A. orientalis</emphasis>
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</taxonomicName>
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. Apical fringe of bristles much longer than hairs on the rest of the elytra. Scutellum markedly less strongly punctured than elytra. Abdominal tergites with very long and semi-erect pubescence, asperate punctures much weaker and sparser than on elytra. Punctures on apical tergites much sparser than on proximal segments. Apical fringe of bristles on each tergite of equal length. The form of the aedeagus is unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="8DE74ADEFFA61A7F06C7FA30ED61F9C4" pageId="30" pageNumber="31" type="materials_examined">
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<paragraph id="C5421955FFA61A7F06C7FA30ED61F9C4" blockId="30.[151,1437,805,1727]" pageId="30" pageNumber="31">
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<emphasis id="F789C547FFA61A7F06C7FA30EF3BFA57" bold="true" box="[196,379,1439,1465]" pageId="30" pageNumber="31">
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<typeStatus id="1A46A7F7FFA61A7F06C7FA30EF40FA57" box="[196,256,1439,1465]" pageId="30" pageNumber="31">Type</typeStatus>
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material.
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</emphasis>
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Bernhauer described this species from a single specimen, which has been located. This specimen is the
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<typeStatus id="1A46A7F7FFA61A7F0750FA6AEFFAFA31" box="[339,442,1477,1503]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
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by monotypy.
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<emphasis id="F789C547FFA61A7F0469FA6AEC9CFA31" bold="true" box="[618,732,1477,1503]" pageId="30" pageNumber="31">
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<typeStatus id="1A46A7F7FFA61A7F0469FA6AEC9CFA31" box="[618,732,1477,1503]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
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</emphasis>
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Ƥ: “West Usambara;
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<date id="B1433F95FFA61A7F05DDFA6AEA56FA31" box="[990,1046,1477,1503]" pageId="30" pageNumber="31" value="1912-02">II 12</date>
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Methner / D. Ostafrika; Methner /
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<taxonomicName id="02FD62D6FFA61A7F06ABFA45EF71F9EA" box="[168,305,1514,1540]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="48" phylum="Arthropoda" rank="species" species="usambarae">Usambarae</taxonomicName>
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Bernh.
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<typeStatus id="1A46A7F7FFA61A7F0796FA45EF9FF9EA" box="[405,479,1514,1540]" pageId="30" pageNumber="31">Typus</typeStatus>
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unic / Chicago NHMus M. Bernhauer Collection /
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<typeStatus id="1A46A7F7FFA61A7F0259FA45EABAF9EA" box="[1114,1274,1514,1540]" pageId="30" pageNumber="31" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="02FD62D6FFA61A7F0301FA44EFF8F9C7" authority="Bernhauer P" authorityName="Bernhauer P" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="usambarae">
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<emphasis id="F789C547FFA61A7F0301FA44EF59F9C7" italics="true" pageId="30" pageNumber="31">Acylophorus usambarae</emphasis>
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Bernhauer Ƥ
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</taxonomicName>
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det. DA Lott, 2009” (
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<collectionCode id="A3EC8190FFA61A7F04B9F9BFED56F9C4" LSID="urn:lsid:biocol.org:col:34795" box="[698,790,1552,1578]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="30" pageNumber="31">FMNH</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="8DE74ADEFFA61A7F06C7F99AEF4AF951" pageId="30" pageNumber="31" type="distribution">
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<paragraph id="C5421955FFA61A7F06C7F99AEFC4F99A" blockId="30.[151,1437,805,1727]" pageId="30" pageNumber="31">
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<emphasis id="F789C547FFA61A7F06C7F99AEC69F9A1" bold="true" box="[196,553,1589,1615]" pageId="30" pageNumber="31">Distribution and bionomics.</emphasis>
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Only known from the unique
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<typeStatus id="1A46A7F7FFA61A7F0594F99AED8BF9A1" box="[919,971,1589,1615]" pageId="30" pageNumber="31">type</typeStatus>
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collected in the Usambara Mountains,
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<collectingCountry id="BDEA59C5FFA61A7F0694F9F5EF40F99A" box="[151,256,1626,1652]" name="Tanzania" pageId="30" pageNumber="31">Tanzania</collectingCountry>
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(
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<figureCitation id="5DC605D0FFA61A7F0713F9F5EF37F99A" box="[272,375,1626,1652]" captionStart="FIGURE 147" captionStartId="47.[151,255,1866,1890]" captionTargetBox="[151,1436,1004,1838]" captionTargetId="figure@47.[151,1436,1004,1838]" captionTargetPageId="47" captionText="FIGURE 147. Distribution of species in the Acylophorus trigonocephalus group = A. trigonocephalus Cameron, = A. setiger new species, = A. usambarae Bernhauer, = A. grandis Bernhauer; = A. bafutensis Levasseur)." httpUri="https://zenodo.org/record/275928/files/figure.png" pageId="30" pageNumber="31">Fig. 147</figureCitation>
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).
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</paragraph>
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<paragraph id="C5421955FFA61A7F06C7F9D0EF4AF951" blockId="30.[151,1437,805,1727]" pageId="30" pageNumber="31">
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<emphasis id="F789C547FFA61A7F06C7F9D0EFF7F977" bold="true" box="[196,439,1663,1689]" pageId="30" pageNumber="31">Comparative notes.</emphasis>
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Easily recognisable within the species group by the long and outstanding hairs on the abdomen.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |