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<document id="6ADE24ACFE868FCC4343C4FDFF05DB70" ID-DOI="10.5281/zenodo.275907" ID-GBIF-Dataset="d5d27825-82a5-4aaf-b47d-35d24b33aaaa" ID-ISSN="1175-5326" ID-Zenodo-Dep="275907" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460189977646" checkinUser="plazi" docAuthor="Lott, Derek A." docDate="2010" docId="4D54A843FFA11A7A0694FA49EAFCFA49" docLanguage="en" docName="zt02402p051.pdf" docOrigin="Zootaxa 2402" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Acylophorus lomaensis Levasseur" docType="treatment" docVersion="5" lastPageNumber="28" masterDocId="B16DD03BFFB81A610603FFAFEE40FFEE" masterDocTitle="The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa" masterLastPageNumber="51" masterPageNumber="1" pageNumber="26" updateTime="1698236417163" updateUser="plazi">
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<mods:title id="C3C6846B404738E9C8E982BD6419AA5A">The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa</mods:title>
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<mods:namePart id="D8A2A102AA2896F7FA908B3E708A4BFA">Lott, Derek A.</mods:namePart>
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<mods:date id="996170E1D81E3AB2203012E57A90E355">2010</mods:date>
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<treatment id="4D54A843FFA11A7A0694FA49EAFCFA49" ID-DOI="http://doi.org/10.5281/zenodo.6196048" ID-GBIF-Taxon="127655340" ID-Zenodo-Dep="6196048" LSID="urn:lsid:plazi:treatment:4D54A843FFA11A7A0694FA49EAFCFA49" httpUri="http://treatment.plazi.org/id/4D54A843FFA11A7A0694FA49EAFCFA49" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26">
<subSubSection id="8DE74ADEFFA11A780694FA49EFBAF9CD" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="C5421955FFA11A780694FA49EC76F9EE" blockId="25.[151,566,1510,1571]" box="[151,566,1510,1536]" pageId="25" pageNumber="26">
<heading id="9E0AAE39FFA11A780694FA49EC76F9EE" bold="true" box="[151,566,1510,1536]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<taxonomicName id="02FD62D6FFA11A780694FA49EC76F9EE" ID-CoL="9W87" authority="Levasseur" authorityName="Levasseur" box="[151,566,1510,1536]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="lomaensis">
<emphasis id="F789C547FFA11A780694FA49EC76F9EE" bold="true" box="[151,566,1510,1536]" pageId="25" pageNumber="26">
<emphasis id="F789C547FFA11A780694FA49EFF0F9EE" bold="true" box="[151,432,1510,1536]" italics="true" pageId="25" pageNumber="26">Acylophorus lomaensis</emphasis>
Levasseur
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C5421955FFA11A780694F9A6EFBAF9CD" blockId="25.[151,566,1510,1571]" box="[151,506,1545,1571]" pageId="25" pageNumber="26">
(
<figureCitation id="5DC605D0FFA11A78069CF9A6EEBAF9CD" box="[159,250,1545,1571]" captionStart="FIGURES 18 – 20" captionStartId="26.[151,272,1902,1926]" captionTargetBox="[174,1406,185,1901]" captionTargetId="figure@26.[163,1448,167,1913]" captionTargetPageId="26" captionText="FIGURES 18 – 20. Forebodies of species in the Acylophorus salifi group. 18, A. salifi new species; 19, A. lomaensis Levasseur; 20. A. minor new species. FIGURES 21 – 25. Forebodies of species in the Acylophorus trigonocephalus group. 21, A. trigonocephalus Cameron; 22, A. setiger new species; 23, A. usambarae Bernhauer; 24, A. grandis Bernhauer; 25, A. bafutensis Levasseur." httpUri="https://zenodo.org/record/275910/files/figure.png" pageId="25" pageNumber="26">Figs 19</figureCitation>
,
<figureCitation id="5DC605D0FFA11A780705F9A6EF66F9CD" box="[262,294,1545,1571]" captionStart="FIGURES 42 – 50" captionStartId="35.[151,269,1552,1576]" captionTargetBox="[171,1412,194,1517]" captionTargetId="figure@35.[159,1444,183,1554]" captionTargetPageId="35" captionText="FIGURES 42 – 50. Mandibles of Acylophorus species. 42, A. congoensis Cameron; 43, A. ziloensis Levasseur (Katanga); 44, A. ziloensis Levasseur (Kivu); 45, A. allardi Levasseur; 46, A. salifi new species; 47, A. lomaensis Levasseur; 48, A. mareei Bernhauer; 49, A. bafutensis Levasseur; 50, A. lualabaensis Levasseur." httpUri="https://zenodo.org/record/275913/files/figure.png" pageId="25" pageNumber="26">47</figureCitation>
,
<figureCitation id="5DC605D0FFA11A780731F9A6EF12F9CD" box="[306,338,1545,1571]" captionStart="FIGURES 51 – 76" captionStartId="36.[151,271,1818,1842]" captionTargetBox="[176,1425,305,1773]" captionTargetId="figure@36.[155,1440,293,1797]" captionTargetPageId="36" captionText="FIGURES 51 – 76. Terminal segments of the maxilliary palpi of Acylophorus species. 51, A. orientalis Fauvel; 52, A. nitens new species; 53, A. rossii Bordoni, 54, A. bumbunae Bordoni; 55, A. capensis Cameron; 56, A. dankalensis Bordoni; 57, A. tshuapensis new species; 58, A. makhoreae new species; 59, A. densipennis Bernhauer; 60, A. antennalis Cameron; 61, A. collarti Cameron; 62. A. micans new species; 63, A. allardi Levasseur; 64, A. congoensis Cameron; 65, A. ziloensis Levasseur; 66, A. mareei Bernhauer; 67, A. salifi new species; 68, A. lomaensis Levasseur; 69. A. minor new species; 70, A. trigonocephalus Cameron; 71, A. setiger new species; 72, A. usambarae Bernhauer; 73, A. grandis Bernhauer; 74, A. bafutensis Levasseur; 75, A. kambuiensis Bordoni, 76. A. lualabaensis Levasseur." httpUri="https://zenodo.org/record/275914/files/figure.png" pageId="25" pageNumber="26">68</figureCitation>
,
<figureCitation id="5DC605D0FFA11A78075DF9A6EF3EF9CD" box="[350,382,1545,1571]" captionStart="FIGURES 77 – 96" captionStartId="37.[151,271,1853,1877]" captionTargetBox="[209,1389,189,1828]" captionTargetId="figure@37.[168,1428,165,1848]" captionTargetPageId="37" captionText="FIGURES 77 – 96. Antennae of Acylophorus species. 77, A. orientalis Fauvel; 78, A. nitens new species; 79, A. rossii Bordoni, 80, A. bumbunae Bordoni; 81, A. capensis Cameron; 82, A. dankalensis Bordoni; 83, A. tshuapensis new species; 84, A. makhoreae new species; 85, A. densipennis Bernhauer; 86, A. antennalis Cameron; 87, A. collarti Cameron; 88. A. micans new species; 89, A. allardi Levasseur; 90, A. congoensis Cameron; 91, A. ziloensis Levasseur; 92, A. mareei Bernhauer; 93, A. methneri Bernhauer; 94, A. salifi new species; 95, A. lomaensis Levasseur; 96. A. minor new species." httpUri="https://zenodo.org/record/275915/files/figure.png" pageId="25" pageNumber="26">95</figureCitation>
,
<figureCitation id="5DC605D0FFA11A780789F9A6EFB2F9CD" box="[394,498,1545,1571]" captionStart="FIGURES 126 – 131" captionStartId="41.[151,269,1916,1940]" captionTargetBox="[283,1289,186,1915]" captionTargetId="figure@41.[282,1306,178,1917]" captionTargetPageId="41" captionText="FIGURES 126 – 131. Aedeagi of species in the Acylophorus salifi group. 126, A. salifi new species (Burkina Faso); 127, A. salifi new species (Zimbabwe); 128, A. salifi new species (RD Congo); 129, A. lomaensis Levasseur (Lac Tumba, RD Congo); 130, A. lomaensis Levasseur (R. Lukendu, RD Congo); 131. A. minor new species. FIGURES 132 – 133. Male sternite IX and styles of tergite IX of Acylophorus species. 132, A. densipennis Bernhauer; 133, A. kambuiensis Bordoni." httpUri="https://zenodo.org/record/275919/files/figure.png" pageId="25" pageNumber="26">129, 130</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="8DE74ADEFFA11A780694F9FBEDFDF910" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="C5421955FFA11A780694F9FBED2BF985" blockId="25.[151,875,1620,1643]" box="[151,875,1620,1643]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="E5ED3E7BFFA11A780694F9FBED2BF985" box="[151,875,1620,1643]" pageId="25" pageNumber="26">
<taxonomicName id="02FD62D6FFA11A780694F9FBEC3CF985" ID-CoL="9W87" authority="Levasseur, 1971: 303" authorityName="Levasseur" authorityPageNumber="303" authorityYear="1971" box="[151,636,1620,1643]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="lomaensis">
<emphasis id="F789C547FFA11A780694F9FBEFCFF985" box="[151,399,1620,1643]" italics="true" pageId="25" pageNumber="26">Acylophorus lomaensis</emphasis>
<treatmentCitation id="445C3F44FFA11A780796F9FBEC3CF985" author="Levasseur" box="[405,636,1620,1643]" page="303" pageId="25" pageNumber="26" year="1971">
<bibRefCitation id="A16C64A4FFA11A780796F9FBEC07F985" author="Levasseur" box="[405,583,1620,1643]" pageId="25" pageNumber="51" refString="Levasseur, L. (1971). Coleoptera Staphylinidae. 1 er partie: Steninae, Staphylininae, Tachyporinae et Aleocharinae. In Le Massif des Monts Loma (Sierra Leone) Fascicule I. Recherches geographiques, botaniques et zoologiques. Memoires de l'Institut Fondamental d'Afrique Noire, 86, 291 - 315." type="journal article" year="1971">Levasseur, 1971</bibRefCitation>
: 303
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="445C3F44FFA11A780484F9FBED27F985" author="Herman" box="[647,871,1620,1643]" page="3033" pageId="25" pageNumber="26" year="2001">
<bibRefCitation id="A16C64A4FFA11A780484F9FBED63F985" author="Herman" box="[647,803,1620,1643]" pageId="25" pageNumber="50" refString="Herman, L. H. (2001). Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3). Bulletin of the American Museum of Natural History, 265, 3021 - 3840." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 3033
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C5421955FFA11A780694F937EDFDF910" blockId="25.[151,1436,1688,1864]" pageId="25" pageNumber="26">
<emphasis id="F789C547FFA11A780694F937EFEDF95C" bold="true" box="[151,429,1688,1714]" pageId="25" pageNumber="26">Diagnostic characters.</emphasis>
Levasseur’s description contains illustrations of the aedeagus and antenna in addition to a textual description of morphological features. The following section contains additional diagnostic features as well as variations exhibited by further specimens studied.
</paragraph>
</subSubSection>
<subSubSection id="8DE74ADEFFA11A7A06C7F8A6EBC3FCE9" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26" type="description">
<paragraph id="C5421955FFA11A7806C7F8A6ED99F8A6" blockId="25.[151,1436,1688,1864]" pageId="25" pageNumber="26">
Length varies from
<quantity id="0205B4B0FFA11A7807B3F8A6EC4EF8CD" box="[432,526,1801,1827]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="25" pageNumber="26" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7mm</quantity>
. Body often all brown with head dark brown. Abdominal tergites iridescent. Antennae often all pale with apical segments of antennae almost white.
</paragraph>
<caption id="918249DDFFA21A7B0694F8C1EC7EF81E" httpUri="https://zenodo.org/record/275910/files/figure.png" pageId="26" pageNumber="27" targetBox="[174,1406,185,1901]" targetPageId="26">
<paragraph id="C5421955FFA21A7B0694F8C1EC7EF81E" blockId="26.[151,1436,1902,2032]" pageId="26" pageNumber="27">
<emphasis id="F789C547FFA21A7B0694F8C1EF1CF868" bold="true" box="[151,348,1902,1926]" pageId="26" pageNumber="27">FIGURES 18–20</emphasis>
. Forebodies of species in the
<taxonomicName id="02FD62D6FFA21A7B04ABF8C0ED32F868" box="[680,882,1903,1926]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="salifi">
<emphasis id="F789C547FFA21A7B04ABF8C0ED32F868" box="[680,882,1903,1926]" italics="true" pageId="26" pageNumber="27">Acylophorus salifi</emphasis>
</taxonomicName>
group. 18,
<taxonomicName id="02FD62D6FFA21A7B05F0F8C0EA09F868" box="[1011,1097,1903,1926]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="salifi" status="sp. nov.">
<emphasis id="F789C547FFA21A7B05F0F8C0EA09F868" box="[1011,1097,1903,1926]" italics="true" pageId="26" pageNumber="27">A. salifi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECBA783CFFA21A7B0252F8C0EA99F868" box="[1105,1241,1903,1926]" pageId="26" pageNumber="27" rank="species">new species</taxonomicNameLabel>
; 19,
<taxonomicName id="02FD62D6FFA21A7B030CF8C0EF4BF847" authority="Levasseur" authorityName="Levasseur" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="lomaensis">
<emphasis id="F789C547FFA21A7B030CF8C0EBDCF868" box="[1295,1436,1903,1926]" italics="true" pageId="26" pageNumber="27">A. lomaensis</emphasis>
Levasseur
</taxonomicName>
; 20.
<taxonomicName id="02FD62D6FFA21A7B0742F83DEFE3F847" box="[321,419,1938,1961]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="minor" status="sp. nov.">
<emphasis id="F789C547FFA21A7B0742F83DEFE3F847" box="[321,419,1938,1961]" italics="true" pageId="26" pageNumber="27">A. minor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECBA783CFFA21A7B07A9F83DEC74F847" box="[426,564,1938,1961]" pageId="26" pageNumber="27" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="F789C547FFA21A7B043CF83EED46F847" bold="true" box="[575,774,1937,1961]" pageId="26" pageNumber="27">FIGURES 21–25</emphasis>
. Forebodies of species in the
<taxonomicName id="02FD62D6FFA21A7B0256F83DEBDCF847" box="[1109,1436,1938,1961]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="trigonocephalus">
<emphasis id="F789C547FFA21A7B0256F83DEBDCF847" box="[1109,1436,1938,1961]" italics="true" pageId="26" pageNumber="27">Acylophorus trigonocephalus</emphasis>
</taxonomicName>
group. 21,
<taxonomicName id="02FD62D6FFA21A7B0710F81BEC1FF825" authority="Cameron" authorityName="Cameron" box="[275,607,1972,1995]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="trigonocephalus">
<emphasis id="F789C547FFA21A7B0710F81BEFACF825" box="[275,492,1972,1995]" italics="true" pageId="26" pageNumber="27">A. trigonocephalus</emphasis>
Cameron
</taxonomicName>
; 22,
<taxonomicName id="02FD62D6FFA21A7B0495F81BED43F825" box="[662,771,1972,1995]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="setiger" status="sp. nov.">
<emphasis id="F789C547FFA21A7B0495F81BED43F825" box="[662,771,1972,1995]" italics="true" pageId="26" pageNumber="27">A. setiger</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECBA783CFFA21A7B050FF81BEDD8F825" box="[780,920,1972,1995]" pageId="26" pageNumber="27" rank="species">new species</taxonomicNameLabel>
; 23,
<taxonomicName id="02FD62D6FFA21A7B05D3F81BEAAEF825" authority="Bernhauer" authorityName="Bernhauer" box="[976,1262,1972,1995]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="usambarae">
<emphasis id="F789C547FFA21A7B05D3F81BEA2DF825" box="[976,1133,1972,1995]" italics="true" pageId="26" pageNumber="27">A. usambarae</emphasis>
Bernhauer
</taxonomicName>
; 24,
<taxonomicName id="02FD62D6FFA21A7B0325F81BEF49F81E" authority="Bernhauer" authorityName="Bernhauer" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="F789C547FFA21A7B0325F81BEBDCF825" box="[1318,1436,1972,1995]" italics="true" pageId="26" pageNumber="27">A. grandis</emphasis>
Bernhauer
</taxonomicName>
; 25,
<taxonomicName id="02FD62D6FFA21A7B073FF876EC7EF81E" authority="Levasseur." authorityName="Levasseur." box="[316,574,2009,2032]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="bafutensis">
<emphasis id="F789C547FFA21A7B073FF876EF85F81E" box="[316,453,2009,2032]" italics="true" pageId="26" pageNumber="27">A. bafutensis</emphasis>
Levasseur.
</taxonomicName>
</paragraph>
</caption>
<paragraph id="C5421955FFA31A7A06C7FF37EC07FDEC" blockId="27.[151,1437,152,1447]" pageId="27" pageNumber="28">
Head small (pronotum 1.9x wider than head), more or less as long as wide with temples barely suggested (
<figureCitation id="5DC605D0FFA31A7A06A3FF12EEBDFF39" box="[160,253,189,215]" captionStart="FIGURES 18 – 20" captionStartId="26.[151,272,1902,1926]" captionTargetBox="[174,1406,185,1901]" captionTargetId="figure@26.[163,1448,167,1913]" captionTargetPageId="26" captionText="FIGURES 18 – 20. Forebodies of species in the Acylophorus salifi group. 18, A. salifi new species; 19, A. lomaensis Levasseur; 20. A. minor new species. FIGURES 21 – 25. Forebodies of species in the Acylophorus trigonocephalus group. 21, A. trigonocephalus Cameron; 22, A. setiger new species; 23, A. usambarae Bernhauer; 24, A. grandis Bernhauer; 25, A. bafutensis Levasseur." httpUri="https://zenodo.org/record/275910/files/figure.png" pageId="27" pageNumber="28">Fig. 19</figureCitation>
). Antennae inserted right on front margin. Genae well developed and produced at sides of front margin. Micro-punctures extremely sparse. Short pubescence behind eyes localised and very sparse. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other. Five postocular setae visible from above on each side. No extra seta by hind margin of eye. Underside of head weakly depressed at base. Gular sutures continued to base of head, converging toward base but well separated throughout their length. Mandibles without medial teeth, similar to
<taxonomicName id="02FD62D6FFA31A7A05C6FED7EA63FE7F" box="[965,1059,376,401]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="salifi">
<emphasis id="F789C547FFA31A7A05C6FED7EA63FE7F" box="[965,1059,376,401]" italics="true" pageId="27" pageNumber="28">A. salifi</emphasis>
</taxonomicName>
(
<figureCitation id="5DC605D0FFA31A7A0237FED7EACFFE7C" box="[1076,1167,376,402]" captionStart="FIGURES 42 – 50" captionStartId="35.[151,269,1552,1576]" captionTargetBox="[171,1412,194,1517]" captionTargetId="figure@35.[159,1444,183,1554]" captionTargetPageId="35" captionText="FIGURES 42 – 50. Mandibles of Acylophorus species. 42, A. congoensis Cameron; 43, A. ziloensis Levasseur (Katanga); 44, A. ziloensis Levasseur (Kivu); 45, A. allardi Levasseur; 46, A. salifi new species; 47, A. lomaensis Levasseur; 48, A. mareei Bernhauer; 49, A. bafutensis Levasseur; 50, A. lualabaensis Levasseur." httpUri="https://zenodo.org/record/275913/files/figure.png" pageId="27" pageNumber="28">Fig. 47</figureCitation>
). Maxillary palpi with terminal segment densely pubescent, markedly asymmetric, longer than short, triangular, penultimate segment, which is glabrous (
<figureCitation id="5DC605D0FFA31A7A07E6FE6DEC7DFE32" box="[485,573,450,476]" captionStart="FIGURES 51 – 76" captionStartId="36.[151,271,1818,1842]" captionTargetBox="[176,1425,305,1773]" captionTargetId="figure@36.[155,1440,293,1797]" captionTargetPageId="36" captionText="FIGURES 51 – 76. Terminal segments of the maxilliary palpi of Acylophorus species. 51, A. orientalis Fauvel; 52, A. nitens new species; 53, A. rossii Bordoni, 54, A. bumbunae Bordoni; 55, A. capensis Cameron; 56, A. dankalensis Bordoni; 57, A. tshuapensis new species; 58, A. makhoreae new species; 59, A. densipennis Bernhauer; 60, A. antennalis Cameron; 61, A. collarti Cameron; 62. A. micans new species; 63, A. allardi Levasseur; 64, A. congoensis Cameron; 65, A. ziloensis Levasseur; 66, A. mareei Bernhauer; 67, A. salifi new species; 68, A. lomaensis Levasseur; 69. A. minor new species; 70, A. trigonocephalus Cameron; 71, A. setiger new species; 72, A. usambarae Bernhauer; 73, A. grandis Bernhauer; 74, A. bafutensis Levasseur; 75, A. kambuiensis Bordoni, 76. A. lualabaensis Levasseur." httpUri="https://zenodo.org/record/275914/files/figure.png" pageId="27" pageNumber="28">Fig. 68</figureCitation>
). First segment of antenna nearly as long as next four (
<figureCitation id="5DC605D0FFA31A7A02BDFE6DEB55FE32" box="[1214,1301,450,476]" captionStart="FIGURES 77 – 96" captionStartId="37.[151,271,1853,1877]" captionTargetBox="[209,1389,189,1828]" captionTargetId="figure@37.[168,1428,165,1848]" captionTargetPageId="37" captionText="FIGURES 77 – 96. Antennae of Acylophorus species. 77, A. orientalis Fauvel; 78, A. nitens new species; 79, A. rossii Bordoni, 80, A. bumbunae Bordoni; 81, A. capensis Cameron; 82, A. dankalensis Bordoni; 83, A. tshuapensis new species; 84, A. makhoreae new species; 85, A. densipennis Bernhauer; 86, A. antennalis Cameron; 87, A. collarti Cameron; 88. A. micans new species; 89, A. allardi Levasseur; 90, A. congoensis Cameron; 91, A. ziloensis Levasseur; 92, A. mareei Bernhauer; 93, A. methneri Bernhauer; 94, A. salifi new species; 95, A. lomaensis Levasseur; 96. A. minor new species." httpUri="https://zenodo.org/record/275915/files/figure.png" pageId="27" pageNumber="28">Fig. 95</figureCitation>
). Segments I to V elongate, VII to XI transverse.
</paragraph>
<paragraph id="C5421955FFA31A7A06C7FDA2EC6FFD79" blockId="27.[151,1437,152,1447]" pageId="27" pageNumber="28">
Pronotum less transverse than in
<taxonomicName id="02FD62D6FFA31A7A0451FDA2ECEFFDC8" box="[594,687,525,550]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="salifi">
<emphasis id="F789C547FFA31A7A0451FDA2ECEFFDC8" box="[594,687,525,550]" italics="true" pageId="27" pageNumber="28">A. salifi</emphasis>
</taxonomicName>
(1.2x wider than long) with rounded sides and widest in basal half. Shining with no micro-punctures. Elytra strongly transverse (1.6x wider than long). Fringe of bristles on hind margin longer than the hairs on the rest of the elytra. Asperate punctures on abdominal tergites stronger than on elytra, denser toward base.
</paragraph>
<paragraph id="C5421955FFA31A7A06C7FD0DEBC3FCE9" blockId="27.[151,1437,152,1447]" pageId="27" pageNumber="28">
There is some variation in the form of the aedeagus. In most specimens, including the
<typeStatus id="1A46A7F7FFA31A7A02B7FD0DEB59FD52" box="[1204,1305,674,700]" pageId="27" pageNumber="28" type="paratype">paratype</typeStatus>
that I have seen, each lobe is weakly arched and proximate and the pegs confusedly arranged in apical half of each lobe (
<figureCitation id="5DC605D0FFA31A7A069CFD42EF1CFCE9" box="[159,348,749,775]" captionStart="FIGURES 126 – 131" captionStartId="41.[151,269,1916,1940]" captionTargetBox="[283,1289,186,1915]" captionTargetId="figure@41.[282,1306,178,1917]" captionTargetPageId="41" captionText="FIGURES 126 – 131. Aedeagi of species in the Acylophorus salifi group. 126, A. salifi new species (Burkina Faso); 127, A. salifi new species (Zimbabwe); 128, A. salifi new species (RD Congo); 129, A. lomaensis Levasseur (Lac Tumba, RD Congo); 130, A. lomaensis Levasseur (R. Lukendu, RD Congo); 131. A. minor new species. FIGURES 132 – 133. Male sternite IX and styles of tergite IX of Acylophorus species. 132, A. densipennis Bernhauer; 133, A. kambuiensis Bordoni." httpUri="https://zenodo.org/record/275919/files/figure.png" pageId="27" pageNumber="28">Figs 129 &amp; 130</figureCitation>
). The median lobe is longer than the paramere and expanded at the apex which is rounded.
</paragraph>
</subSubSection>
<subSubSection id="8DE74ADEFFA31A7A06C7FCBDEDC8FB92" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="C5421955FFA31A7A06C7FCBDEF4CFC99" blockId="27.[151,1437,152,1447]" pageId="27" pageNumber="28">
<emphasis id="F789C547FFA31A7A06C7FCBDEFB4FCC2" bold="true" box="[196,500,786,812]" pageId="27" pageNumber="28">
<typeStatus id="1A46A7F7FFA31A7A06C7FCBDEF41FCC2" box="[196,257,786,812]" pageId="27" pageNumber="28">Type</typeStatus>
material.
<typeStatus id="1A46A7F7FFA31A7A0781FCBDEFB4FCC2" box="[386,500,786,812]" pageId="27" pageNumber="28" type="paratype">Paratype</typeStatus>
</emphasis>
3: “Mission ENS-IFAN aux monts
<collectionCode id="A3EC8190FFA31A7A05A2FCBDEDB7FCC2" box="[929,1015,786,812]" pageId="27" pageNumber="28">LOMA</collectionCode>
/ Ravin SE du Camp
<date id="B1433F95FFA31A7A02FAFCBDEB24FCC2" box="[1273,1380,786,812]" bridgedPair="-" pageId="27" pageNumber="28" value="1600-02">
II –
<quantity id="0205B4B0FFA31A7A0329FCBDEB24FCC2" box="[1322,1380,786,812]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="27" pageNumber="28" unit="m" value="1600.0">1600</quantity>
</date>
m
<date id="B1433F95FFA31A7A0387FCBDEEAEFCBC" pageId="27" pageNumber="28" value="1966-01-08">8- 1-1966</date>
/ tamisage bord ruisseau /
<taxonomicName id="02FD62D6FFA31A7A0443FC97ED20FCBF" box="[576,864,824,849]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="lomaensis" status="sp. nov.">
<emphasis id="F789C547FFA31A7A0443FC97ED20FCBF" box="[576,864,824,849]" italics="true" pageId="27" pageNumber="28">Acylophorus lomaensis</emphasis>
</taxonomicName>
<emphasis id="F789C547FFA31A7A056AFC98EDF0FCBF" bold="true" box="[873,944,823,849]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="ECBA783CFFA31A7A056AFC98EDF0FCBF" box="[873,944,823,849]" pageId="27" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
LEVASSEUR DET 1967
<typeStatus id="1A46A7F7FFA31A7A02FBFC97EBD3FCBC" box="[1272,1427,824,850]" pageId="27" pageNumber="28" type="paratype">PARATYPE</typeStatus>
” (
<collectionCode id="A3EC8190FFA31A7A069CFCF2EF40FC99" LSID="urn:lsid:biocol.org:col:34988" box="[159,256,861,887]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="28">MNHN</collectionCode>
).
</paragraph>
<paragraph id="C5421955FFA31A7A06C7FC2DEDC8FB92" blockId="27.[151,1437,152,1447]" pageId="27" pageNumber="28">
<emphasis id="F789C547FFA31A7A06C7FC2DECA9FC72" bold="true" box="[196,745,898,924]" pageId="27" pageNumber="28">
Further material examined. R.D.
<collectingCountry id="BDEA59C5FFA31A7A047BFC2DECA9FC72" box="[632,745,898,924]" name="Republic of the Congo" pageId="27" pageNumber="28">CONGO</collectingCountry>
</emphasis>
:
<emphasis id="F789C547FFA31A7A04F8FC2DEDD1FC72" bold="true" box="[763,913,898,924]" pageId="27" pageNumber="28">
Bas
<collectingCountry id="BDEA59C5FFA31A7A0531FC2DEDCBFC72" box="[818,907,898,924]" name="Republic of the Congo" pageId="27" pageNumber="28">Congo</collectingCountry>
:
</emphasis>
Kisantu, P. Basilewsky, 29–30.
<date id="B1433F95FFA31A7A0310FC2DEBC5FC72" box="[1299,1413,898,924]" pageId="27" pageNumber="28" value="1952-11-01">xi.1952 1</date>
Ƥ (
<collectionCode id="A3EC8190FFA31A7A069CFC07EEBDFC2C" box="[159,253,936,962]" pageId="27" pageNumber="28">ISRNB</collectionCode>
);
<emphasis id="F789C547FFA31A7A0710FC08EFD6FC2F" bold="true" box="[275,406,935,961]" pageId="27" pageNumber="28">
<collectingCountry id="BDEA59C5FFA31A7A0710FC08EFD1FC2F" box="[275,401,935,961]" name="Ecuador" pageId="27" pageNumber="28">Equateur</collectingCountry>
:
</emphasis>
Coquilhatville, N Leleup,
<date id="B1433F95FFA31A7A04D9FC07ED0BFC2C" box="[730,843,936,962]" pageId="27" pageNumber="28" value="1959-10-02">x.1959, 2</date>
Ƥ (
<collectionCode id="A3EC8190FFA31A7A0570FC07ED91FC2C" box="[883,977,936,962]" pageId="27" pageNumber="28">ISRNB</collectionCode>
);
<emphasis id="F789C547FFA31A7A05E4FC08EA6FFC2F" bold="true" box="[999,1071,935,961]" pageId="27" pageNumber="28">Kivu:</emphasis>
Kalehe, Bunyakiri,
<quantity id="0205B4B0FFA31A7A0327FC07EB3AFC2C" box="[1316,1402,936,962]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="27" pageNumber="28" unit="m" value="1050.0">1050m</quantity>
, N Leleup,
<date id="B1433F95FFA31A7A06F4FC62EF10FC09" box="[247,336,973,999]" pageId="27" pageNumber="28" value="1953-09">ix.1953</date>
131Ƥ (
<collectionCode id="A3EC8190FFA31A7A07B2FC62EC4EFC09" box="[433,526,973,999]" pageId="27" pageNumber="28">ISRNB</collectionCode>
); Masisi, Walikali,
<quantity id="0205B4B0FFA31A7A04F2FC62ED77FC09" box="[753,823,973,999]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="27" pageNumber="28" unit="m" value="700.0">700m</quantity>
, N Leleup,
<date id="B1433F95FFA31A7A05C3FC62EA06FC08" box="[960,1094,972,999]" pageId="27" pageNumber="28" value="1953-09-13">ix.1953 13</date>
(
<collectionCode id="A3EC8190FFA31A7A0255FC62EAF3FC09" box="[1110,1203,973,999]" pageId="27" pageNumber="28">ISRNB</collectionCode>
); Masisi, Mutakato,
<quantity id="0205B4B0FFA31A7A0694FC5DEE9FFBE2" box="[151,223,1010,1036]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="27" pageNumber="28" unit="m" value="800.0">800m</quantity>
, N Leleup,
<date id="B1433F95FFA31A7A0772FC5DEF8EFBE2" box="[369,462,1010,1036]" pageId="27" pageNumber="28" value="1953-09">ix.1953</date>
231Ƥ (
<collectionCode id="A3EC8190FFA31A7A0437FC5DECD5FBE2" box="[564,661,1010,1036]" pageId="27" pageNumber="28">ISRNB</collectionCode>
); Uvira, rives basses, Sanghe,
<quantity id="0205B4B0FFA31A7A0213FC5DEA19FBE2" box="[1040,1113,1010,1036]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="27" pageNumber="28" unit="m" value="980.0">980m</quantity>
, N Leleup,
<date id="B1433F95FFA31A7A02E9FC5DEB2EFBE2" box="[1258,1390,1010,1036]" pageId="27" pageNumber="28" value="1951-09-13">13.ix.1951</date>
13 (
<collectionCode id="A3EC8190FFA31A7A069CFBB7EEBFFBDC" box="[159,255,1048,1074]" pageId="27" pageNumber="28">ISRNB</collectionCode>
);
<emphasis id="F789C547FFA31A7A0715FBB8EFCEFBDF" bold="true" box="[278,398,1047,1073]" pageId="27" pageNumber="28">Tshuapa:</emphasis>
Bikoro, Mabali, Lac Tumba, N Leleup,
<date id="B1433F95FFA31A7A0582FBB7EA43FBDC" box="[897,1027,1048,1074]" pageId="27" pageNumber="28" value="1955-09-29">29.ix.1955</date>
–
<date id="B1433F95FFA31A7A0216FBB7EA34FBDC" box="[1045,1140,1048,1074]" pageId="27" pageNumber="28" value="1959-09">ix.1959</date>
, 35 (
<collectionCode id="A3EC8190FFA31A7A02B3FBB7EB50FBDC" box="[1200,1296,1048,1074]" pageId="27" pageNumber="28">ISRNB</collectionCode>
); Ikela, riv. Lukendu, N Leleup,
<date id="B1433F95FFA31A7A0789FB92EC5FFBB8" box="[394,543,1084,1111]" pageId="27" pageNumber="28" value="1959-08-13">viii.1959 13</date>
(
<collectionCode id="A3EC8190FFA31A7A042DFB92ECCBFBB9" box="[558,651,1085,1111]" pageId="27" pageNumber="28">ISRNB</collectionCode>
); Ikela, vill. Besoke, N Leleup,
<date id="B1433F95FFA31A7A05FDFB92EA2CFBB9" box="[1022,1132,1085,1111]" pageId="27" pageNumber="28" value="1959-09-01">ix.1959 1</date>
Ƥ (
<collectionCode id="A3EC8190FFA31A7A0292FB92EAAEFBB9" box="[1169,1262,1085,1111]" pageId="27" pageNumber="28">ISRNB</collectionCode>
).
<emphasis id="F789C547FFA31A7A0302FB92EBD3FBB9" bold="true" box="[1281,1427,1085,1111]" pageId="27" pageNumber="28">
R.
<collectingCountry id="BDEA59C5FFA31A7A0326FB92EBD3FBB9" box="[1317,1427,1085,1111]" name="Republic of the Congo" pageId="27" pageNumber="28">CONGO</collectingCountry>
</emphasis>
: Brazzaville, Forêt Pattes d’Oie, J.P. Adam,
<date id="B1433F95FFA31A7A0490FBCDECBAFB92" box="[659,762,1122,1148]" pageId="27" pageNumber="28" value="1960-10-02">x.1960 2</date>
Ƥ (
<collectionCode id="A3EC8190FFA31A7A051CFBCDED3BFB92" box="[799,891,1122,1148]" pageId="27" pageNumber="28">ISRNB</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="8DE74ADEFFA31A7A06C7FB28EB54FAD9" pageId="27" pageNumber="28" type="distribution">
<paragraph id="C5421955FFA31A7A06C7FB28EEA3FB02" blockId="27.[151,1437,152,1447]" pageId="27" pageNumber="28">
<emphasis id="F789C547FFA31A7A06C7FB28EC60FB4F" bold="true" box="[196,544,1159,1185]" pageId="27" pageNumber="28">Distribution and bionomics.</emphasis>
Apart from the
<typeStatus id="1A46A7F7FFA31A7A04D8FB27ED4EFB4C" box="[731,782,1160,1186]" pageId="27" pageNumber="28">type</typeStatus>
series from
<collectingCountry id="BDEA59C5FFA31A7A059DFB27EA77FB4C" box="[926,1079,1160,1186]" name="Sierra Leone" pageId="27" pageNumber="28">Sierra Leone</collectingCountry>
, all material seen so far comes from the
<collectingCountry id="BDEA59C5FFA31A7A070BFB02EF19FB29" box="[264,345,1197,1223]" name="Republic of the Congo" pageId="27" pageNumber="28">Congo</collectingCountry>
catchment (
<figureCitation id="5DC605D0FFA31A7A07ECFB02EC1CFB29" box="[495,604,1197,1223]" captionStart="FIGURE 146" captionStartId="47.[151,255,898,922]" captionTargetBox="[151,1436,194,870]" captionTargetId="figure@47.[151,1436,194,870]" captionTargetPageId="47" captionText="FIGURE 146. Distribution of species in the Acylophorus salifi group = A. lomaensis Levasseur, = A. minor new species)." httpUri="https://zenodo.org/record/275927/files/figure.png" pageId="27" pageNumber="28">Fig. 146</figureCitation>
). Habitats range from forest humus to clay lake margins and stream banks.
</paragraph>
<paragraph id="C5421955FFA31A7A06C7FB58EB54FAD9" blockId="27.[151,1437,152,1447]" pageId="27" pageNumber="28">
<emphasis id="F789C547FFA31A7A06C7FB58EF81FAFF" bold="true" box="[196,449,1271,1297]" pageId="27" pageNumber="28">Comparative notes.</emphasis>
Often smaller than
<taxonomicName id="02FD62D6FFA31A7A04B4FB57ED57FAFF" box="[695,791,1272,1297]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="salifi">
<emphasis id="F789C547FFA31A7A04B4FB57ED57FAFF" box="[695,791,1272,1297]" italics="true" pageId="27" pageNumber="28">A. salifi</emphasis>
</taxonomicName>
with a less transverse pronotum, smaller eyes, more developed genae and a distinctive aedeagus with narrower, more proximate lobes to the paramere.
</paragraph>
</subSubSection>
<subSubSection id="8DE74ADEFFA31A7A06C7FAEDEAFCFA49" pageId="27" pageNumber="28" type="discussion">
<paragraph id="C5421955FFA31A7A06C7FAEDEAFCFA49" blockId="27.[151,1437,152,1447]" pageId="27" pageNumber="28">
<emphasis id="F789C547FFA31A7A06C7FAEDEF0FFAB2" bold="true" box="[196,335,1346,1372]" pageId="27" pageNumber="28">Discussion.</emphasis>
Specimens from the
<collectingCountry id="BDEA59C5FFA31A7A0444FAEDECD5FAB2" box="[583,661,1346,1372]" name="Republic of the Congo" pageId="27" pageNumber="28">Congo</collectingCountry>
catchment are smaller (
<quantity id="0205B4B0FFA31A7A05AEFAEDEA4AFAB2" box="[941,1034,1346,1372]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="27" pageNumber="28" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6mm</quantity>
) than those from
<collectingCountry id="BDEA59C5FFA31A7A02DBFAEDEB2EFAB2" box="[1240,1390,1346,1372]" name="Sierra Leone" pageId="27" pageNumber="28">Sierra Leone</collectingCountry>
(
<quantity id="0205B4B0FFA31A7A037EFAEDEE94FA6C" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="27" pageNumber="28" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6– 7mm</quantity>
).There are variations in the form of the aedeagus (
<figureCitation id="5DC605D0FFA31A7A052BFAC7EDA6FA6C" box="[808,998,1384,1410]" captionStart="FIGURES 126 – 131" captionStartId="41.[151,269,1916,1940]" captionTargetBox="[283,1289,186,1915]" captionTargetId="figure@41.[282,1306,178,1917]" captionTargetPageId="41" captionText="FIGURES 126 – 131. Aedeagi of species in the Acylophorus salifi group. 126, A. salifi new species (Burkina Faso); 127, A. salifi new species (Zimbabwe); 128, A. salifi new species (RD Congo); 129, A. lomaensis Levasseur (Lac Tumba, RD Congo); 130, A. lomaensis Levasseur (R. Lukendu, RD Congo); 131. A. minor new species. FIGURES 132 – 133. Male sternite IX and styles of tergite IX of Acylophorus species. 132, A. densipennis Bernhauer; 133, A. kambuiensis Bordoni." httpUri="https://zenodo.org/record/275919/files/figure.png" pageId="27" pageNumber="28">Figs 129 &amp; 130</figureCitation>
). Some populations may be referable to distinct taxa once sufficient material from further localities becomes available for study.
</paragraph>
</subSubSection>
</treatment>
</document>
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