treatments-xml/data/4D/35/EF/4D35EF104F9773D4213F1DBEB46CFEC7.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.768.24423" ID-PMC="PMC6019434" ID-Pensoft-Pub="1313-2970-768-113" ID-Pensoft-UUID="CB01CA30FFB4F96DFFA2FF86FFAEFFB6" ID-PubMed="29955215" ID-Zenodo-Dep="1298776" ID-ZooBank="399BCC3E9D6F4231870E05C79B9FD4B0" ModsDocID="1313-2970-768-113" checkinTime="1529468775230" checkinUser="pensoft" docAuthor="Giron, Jennifer C. &amp; Short, Andrew Edward Z." docDate="2018" docId="4D35EF104F9773D4213F1DBEB46CFEC7" docLanguage="en" docName="ZooKeys 768: 113-158" docOrigin="ZooKeys 768" docPubDate="2018-06-19" docSource="http://dx.doi.org/10.3897/zookeys.768.24423" docTitle="Katasophistes Girón &amp; Short 2018, gen. n." docType="treatment" docUuid="3EB28C66-5B17-4540-8E86-DB99504C7227" docUuidSource="ZooBank" docVersion="6" id="CB01CA30FFB4F96DFFA2FF86FFAEFFB6" lastPageNumber="128" masterDocId="CB01CA30FFB4F96DFFA2FF86FFAEFFB6" masterDocTitle="Three new genera of acidocerine water scavenger beetles from tropical South America (Coleoptera, Hydrophilidae, Acidocerinae)" masterLastPageNumber="158" masterPageNumber="113" pageNumber="126" updateTime="1668165917053" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Three new genera of acidocerine water scavenger beetles from tropical South America (Coleoptera, Hydrophilidae, Acidocerinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Giron, Jennifer C.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0851-6883</mods:nameIdentifier>
<mods:affiliation>Department of Ecology &amp; Evolutionary Biology, and Division of Entomology, Biodiversity Institute, University of Kansas, Lawrence, KS 66045, USA</mods:affiliation>
<mods:nameIdentifier type="email">entiminae@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Short, Andrew Edward Z.</mods:namePart>
<mods:affiliation>Department of Ecology &amp; Evolutionary Biology, and Division of Entomology, Biodiversity Institute, University of Kansas, Lawrence, KS 66045, USA</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2018</mods:date>
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<mods:number>2018-06-19</mods:number>
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<mods:detail type="volume">
<mods:number>768</mods:number>
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<mods:start>113</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.768.24423</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.768.24423</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-768-113</mods:identifier>
<mods:identifier type="ZooBank">399BCC3E9D6F4231870E05C79B9FD4B0</mods:identifier>
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<treatment ID-GBIF-Taxon="146002394" LSID="urn:lsid:zoobank.org:act:3EB28C66-5B17-4540-8E86-DB99504C7227" httpUri="http://treatment.plazi.org/id/4D35EF104F9773D4213F1DBEB46CFEC7" lastPageId="15" lastPageNumber="128" pageId="13" pageNumber="126">
<subSubSection pageId="13" pageNumber="126" type="nomenclature">
<paragraph pageId="13" pageNumber="126">
<taxonomicName LSID="http://zoobank.org/3EB28C66-5B17-4540-8E86-DB99504C7227" authority="Girón &amp; Short, 2018" authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Katasophistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Katasophistes" order="Coleoptera" pageId="13" pageNumber="126" phylum="Arthropoda" rank="genus" status="gen. n.">Katasophistes</taxonomicName>
<taxonomicNameLabel pageId="13" pageNumber="126">gen. n.</taxonomicNameLabel>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus of Katasophistes spp.: A-E Katasophistes charynae: A dorsal view B lateral view C ventral view D head E-H Katasophistes cuzco: E dorsal view F lateral view G ventral view H head. Scale bars: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure9" httpUri="https://binary.pensoft.net/fig/211220" pageId="13" pageNumber="126">Figs 9</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Katasophistes spp.: A-E Katasophistes merida: A dorsal view B lateral view C ventral view D head E-H Katasophistes superficialis: E dorsal view F lateral view G ventral view H head. Scale bar: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure10" httpUri="https://binary.pensoft.net/fig/211221" pageId="13" pageNumber="126">, 10</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Thorax and abdomen of Katasophistes spp.: A-C Katasophistes merida: A ventral view of mesoventrite (white arrow pointing transverse ridge) B posterior view of metafurca: (at) anterior tendon, (c) crux, (ds) dorsal sheath, (fa) furcal arm, (h) hemiductus, (ls) lateral sheath, (s) stalk C fifth abdominal ventrite D Katasophistes charynae, fifth abdominal ventrite; E Katasophistes cuzco, fifth abdominal ventrite." figureDoi="10.3897/zookeys.768.24423.figure11" httpUri="https://binary.pensoft.net/fig/211223" pageId="13" pageNumber="126">, 11</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Aedeagus of Katasophistes spp.: A-B Katasophistes charynae: A &quot; colln 32 &quot; B &quot; colln 42 a &quot; C Katasophistes cuzco; D Katasophistes merida E Katasophistes superficialis. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.768.24423.figure12" httpUri="https://binary.pensoft.net/fig/211224" pageId="13" pageNumber="126">, 12</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Distribution of Katasophistes species." figureDoi="10.3897/zookeys.768.24423.figure13" httpUri="https://binary.pensoft.net/fig/211225" pageId="13" pageNumber="126">, 13</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Habitat of Katasophistes spp. A-B Type locality and habitat of K. merida: Venezuela: Merida State, Collecting Event VZ 12 - 0122 - 03 A C-D Habitat of K. merida: Venezuela: Merida State, Collecting Event VZ 12 - 0124 - 01 A E-F Type locality and habitat of K. superficialis: Ecuador: Pastaza Province, Collecting Event AS- 08 - 08 b." figureDoi="10.3897/zookeys.768.24423.figure14" httpUri="https://binary.pensoft.net/fig/211227" pageId="13" pageNumber="126">, 14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="126" type="type species">
<paragraph pageId="13" pageNumber="126">Type species.</paragraph>
<paragraph pageId="13" pageNumber="126">
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Katasophistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Katasophistes merida" order="Coleoptera" pageId="13" pageNumber="126" phylum="Arthropoda" rank="species" species="merida">
<emphasis italics="true" pageId="13" pageNumber="126">Katasophistes merida</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="126" type="differential diagnosis">
<paragraph pageId="13" pageNumber="126">Differential diagnosis.</paragraph>
<paragraph pageId="13" pageNumber="126">
Medium to small beetles, total body length 2.7-4.5 mm, width 1.7-2.4 mm. Color orange brown to dark brown, rather uniform along body regions. Body shape oval to elongated in dorsal view; moderately convex in lateral view (see Figs 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus of Katasophistes spp.: A-E Katasophistes charynae: A dorsal view B lateral view C ventral view D head E-H Katasophistes cuzco: E dorsal view F lateral view G ventral view H head. Scale bars: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure9" httpUri="https://binary.pensoft.net/fig/211220" pageId="13" pageNumber="126">9</figureCitation>
- 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Katasophistes spp.: A-E Katasophistes merida: A dorsal view B lateral view C ventral view D head E-H Katasophistes superficialis: E dorsal view F lateral view G ventral view H head. Scale bar: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure10" httpUri="https://binary.pensoft.net/fig/211221" pageId="13" pageNumber="126">10</figureCitation>
). Antennae with nine antennomeres (e.g., Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Katasophistes spp.: A-E Katasophistes merida: A dorsal view B lateral view C ventral view D head E-H Katasophistes superficialis: E dorsal view F lateral view G ventral view H head. Scale bar: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure10" httpUri="https://binary.pensoft.net/fig/211221" pageId="13" pageNumber="126">10C</figureCitation>
). Maxillary palps curved inward, moderately long to long (e.g., Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Katasophistes spp.: A-E Katasophistes merida: A dorsal view B lateral view C ventral view D head E-H Katasophistes superficialis: E dorsal view F lateral view G ventral view H head. Scale bar: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure10" httpUri="https://binary.pensoft.net/fig/211221" pageId="13" pageNumber="126">10D, H</figureCitation>
). Each elytron with five rows of deep/large systematic punctures (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Katasophistes spp.: A-E Katasophistes merida: A dorsal view B lateral view C ventral view D head E-H Katasophistes superficialis: E dorsal view F lateral view G ventral view H head. Scale bar: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure10" httpUri="https://binary.pensoft.net/fig/211221" pageId="13" pageNumber="126">10A</figureCitation>
; not as evident in 
<taxonomicName lsidName="K. charynae" pageId="13" pageNumber="126" rank="species" species="charynae">
<emphasis italics="true" pageId="13" pageNumber="126">K. charynae</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="K. cuzco" pageId="13" pageNumber="126" rank="species" species="cuzco">
<emphasis italics="true" pageId="13" pageNumber="126">K. cuzco</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="K. superficialis" pageId="13" pageNumber="126" rank="species" species="superficialis">
<emphasis italics="true" pageId="13" pageNumber="126">K. superficialis</emphasis>
</taxonomicName>
); elytra without sutural striae, with outer margins slightly flared; serial punctures not evident. Posterior elevation of mesoventrite, with a well-defined, curved transverse ridge (Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Thorax and abdomen of Katasophistes spp.: A-C Katasophistes merida: A ventral view of mesoventrite (white arrow pointing transverse ridge) B posterior view of metafurca: (at) anterior tendon, (c) crux, (ds) dorsal sheath, (fa) furcal arm, (h) hemiductus, (ls) lateral sheath, (s) stalk C fifth abdominal ventrite D Katasophistes charynae, fifth abdominal ventrite; E Katasophistes cuzco, fifth abdominal ventrite." figureDoi="10.3897/zookeys.768.24423.figure11" httpUri="https://binary.pensoft.net/fig/211223" pageId="13" pageNumber="126">11A</figureCitation>
). Posterior femora glabrous at most along apical third. Fifth abdominal ventrite apically truncate to slightly emarginate, with stout setae (see Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Thorax and abdomen of Katasophistes spp.: A-C Katasophistes merida: A ventral view of mesoventrite (white arrow pointing transverse ridge) B posterior view of metafurca: (at) anterior tendon, (c) crux, (ds) dorsal sheath, (fa) furcal arm, (h) hemiductus, (ls) lateral sheath, (s) stalk C fifth abdominal ventrite D Katasophistes charynae, fifth abdominal ventrite; E Katasophistes cuzco, fifth abdominal ventrite." figureDoi="10.3897/zookeys.768.24423.figure11" httpUri="https://binary.pensoft.net/fig/211223" pageId="13" pageNumber="126">11C-E</figureCitation>
).
</paragraph>
<paragraph pageId="13" pageNumber="126">
At first glance the genus may appear similar to some species of 
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chasmogenus" order="Coleoptera" pageId="13" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="13" pageNumber="126">Chasmogenus</emphasis>
</taxonomicName>
, however the lack of sutural striae easily separates the two. The enlargement of the rows of elytral systematic punctures is also rare within the 
<taxonomicName baseAuthorityName="Zaitzev" baseAuthorityYear="1908" lsidName="" pageId="13" pageNumber="126" rank="subFamily" subFamily="Acidocerinae">Acidocerinae</taxonomicName>
(found in some 
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chasmogenus" order="Coleoptera" pageId="13" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="13" pageNumber="126">Chasmogenus</emphasis>
</taxonomicName>
and 
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Agraphydrus" order="Coleoptera" pageId="13" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="13" pageNumber="126">Agraphydrus</emphasis>
</taxonomicName>
) and will separate it from other New World 
<taxonomicName class="Hexapoda" family="Hydrophilidae" genus="Helochares" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helochares" order="Coleoptera" pageId="13" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="13" pageNumber="126">Helochares</emphasis>
</taxonomicName>
, with which it may also be confused.
</paragraph>
</subSubSection>
<subSubSection lastPageId="15" lastPageNumber="128" pageId="13" pageNumber="126" type="description">
<paragraph pageId="13" pageNumber="126">Description.</paragraph>
<paragraph lastPageId="15" lastPageNumber="128" pageId="13" pageNumber="126">
Body oval to elongate, weakly to moderately convex (e.g., Figs 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus of Katasophistes spp.: A-E Katasophistes charynae: A dorsal view B lateral view C ventral view D head E-H Katasophistes cuzco: E dorsal view F lateral view G ventral view H head. Scale bars: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure9" httpUri="https://binary.pensoft.net/fig/211220" pageId="13" pageNumber="126">9F</figureCitation>
, 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Katasophistes spp.: A-E Katasophistes merida: A dorsal view B lateral view C ventral view D head E-H Katasophistes superficialis: E dorsal view F lateral view G ventral view H head. Scale bar: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure10" httpUri="https://binary.pensoft.net/fig/211221" pageId="13" pageNumber="126">10F</figureCitation>
), coloration rather uniform along body regions, orange brown to dark brown, often with maxillary palps slightly paler. 
<emphasis bold="true" pageId="13" pageNumber="126">Head.</emphasis>
Frons and clypeus (e.g., Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus of Katasophistes spp.: A-E Katasophistes charynae: A dorsal view B lateral view C ventral view D head E-H Katasophistes cuzco: E dorsal view F lateral view G ventral view H head. Scale bars: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure9" httpUri="https://binary.pensoft.net/fig/211220" pageId="13" pageNumber="126">9D</figureCitation>
) with moderately marked ground punctures, irregularly and rather densely distributed over the surface, accompanied by seta-bearing systematic punctures, particularly noticeable on frons behind frontoclypeal suture and around inner margins of eyes, and on lateral areas of clypeus; surface between punctures smooth and shiny; anterior corners of clypeus roundly angulated; anterior margin of clypeus widely roundly emarginate. Eyes subquadrate in dorsal view (e.g., Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Katasophistes spp.: A-E Katasophistes merida: A dorsal view B lateral view C ventral view D head E-H Katasophistes superficialis: E dorsal view F lateral view G ventral view H head. Scale bar: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure10" httpUri="https://binary.pensoft.net/fig/211221" pageId="13" pageNumber="126">10H</figureCitation>
). Labrum wide, fully exposed, nearly half as long, and collinear to clypeus (e.g., Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus of Katasophistes spp.: A-E Katasophistes charynae: A dorsal view B lateral view C ventral view D head E-H Katasophistes cuzco: E dorsal view F lateral view G ventral view H head. Scale bars: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure9" httpUri="https://binary.pensoft.net/fig/211220" pageId="13" pageNumber="126">9D</figureCitation>
); dorsal surface slightly convex, with scattered fine punctures; anterior margin mesally slightly emarginate and barely bent inwards; lateral and anterior margins fringed by rather long setae. Temporae densely covered by very short and fine setae (hydrofuge pubescence). Mentum parallel sided, with lateral oblique longitudinal ridges; anterior margin with wide, deep, concave median impression, not demarcated by a transverse carina. Submentum rather flat and pubescent at base, glabrous, shiny and ascending at apex, with sparse rather long setae near limit of pubescent/glabrous line; ocular ridge well-developed (e.g., Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus of Katasophistes spp.: A-E Katasophistes charynae: A dorsal view B lateral view C ventral view D head E-H Katasophistes cuzco: E dorsal view F lateral view G ventral view H head. Scale bars: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure9" httpUri="https://binary.pensoft.net/fig/211220" pageId="13" pageNumber="126">9C</figureCitation>
). Maxilla (e.g., Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus of Katasophistes spp.: A-E Katasophistes charynae: A dorsal view B lateral view C ventral view D head E-H Katasophistes cuzco: E dorsal view F lateral view G ventral view H head. Scale bars: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure9" httpUri="https://binary.pensoft.net/fig/211220" pageId="13" pageNumber="126">9C</figureCitation>
) with ventral surface of cardo and stipes smooth and shiny, with scattered and shallow punctures (basal outer margin of cardo with a fringe of fine and rather long setae in 
<taxonomicName lsidName="K. merida" pageId="13" pageNumber="126" rank="species" species="merida">
<emphasis italics="true" pageId="13" pageNumber="126">K. merida</emphasis>
</taxonomicName>
); outer dorsal margin of palpifer with a row of stiff decumbent spiniform setae; limit between cardo and stipes oblique; maxillary palps brown to orange, slender, longer than antennae (slightly so and stout in 
<taxonomicName lsidName="K. merida" pageId="13" pageNumber="126" rank="species" species="merida">
<emphasis italics="true" pageId="13" pageNumber="126">K. merida</emphasis>
</taxonomicName>
); apex of palpomere 3 bearing sensilla. Mandibles with apex bifid (observed in 
<taxonomicName lsidName="K. merida" pageId="13" pageNumber="126" rank="species" species="merida">
<emphasis italics="true" pageId="13" pageNumber="126">K. merida</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="K. charynae" pageId="13" pageNumber="126" rank="species" species="charynae">
<emphasis italics="true" pageId="13" pageNumber="126">K. charynae</emphasis>
</taxonomicName>
). Labial palps yellowish, nearly as long as mentum, dorsoventrally flattened; palpomere 2 with inner and outer margin convex apicad of midpoint; palpomere 3 digitiform, with a long subapical seta on outer corner. Antennae (e.g., Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Katasophistes spp.: A-E Katasophistes merida: A dorsal view B lateral view C ventral view D head E-H Katasophistes superficialis: E dorsal view F lateral view G ventral view H head. Scale bar: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure10" httpUri="https://binary.pensoft.net/fig/211221" pageId="13" pageNumber="126">10C</figureCitation>
) with nine antennomeres, either similar or paler than general coloration of head; antennomere 1 anteriorly projected near base, almost reaching midpoint of ventral surface of eye (not reaching cardo-stipes joint), nearly 1.4-1.8-times longer than antennomere 2; antennomere 2 nearly as long as antennomeres 3-4 combined; antennomere 6 forming a well differentiated, only slightly asymmetric cupule; antennomeres 7-9 similar in size (8 shortest, 9 longest) and shape, slightly flattened (less so in 
<taxonomicName lsidName="K. merida" pageId="13" pageNumber="126" rank="species" species="merida">
<emphasis italics="true" pageId="13" pageNumber="126">K. merida</emphasis>
</taxonomicName>
), forming a loosely articulated, pubescent club; apex of antennomeres 7-9 with few scattered setae longer than general pubescence of club. 
<emphasis bold="true" pageId="13" pageNumber="126">Thorax.</emphasis>
Pronotum widest at base, narrowed anteriorly, with anterior and posterior margins only slightly sinuate; surface evenly convex, ground punctation moderate, uniformly dense, with surface between punctures smooth and shiny; seta bearing systematic punctures forming paired anterolateral semicircles. Scutellar shield of moderate size, triangular, nearly as long as wide, with punctation as in pronotum. Prosternum (e.g., Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus of Katasophistes spp.: A-E Katasophistes charynae: A dorsal view B lateral view C ventral view D head E-H Katasophistes cuzco: E dorsal view F lateral view G ventral view H head. Scale bars: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure9" httpUri="https://binary.pensoft.net/fig/211220" pageId="13" pageNumber="126">9C</figureCitation>
) 
<pageBreakToken pageId="14" pageNumber="127" start="start">nearly</pageBreakToken>
as long as 2/3 of length of a coxa, weakly convex to only slightly ascending longitudinally on central area; anterior margin of prosternum mesally projected as a wide, roundly pointed triangle, usually with a low carina along midline of projection; surface of prosternum covered by rather long, fine, scattered setae; intercoxal process projected from posterior margin of procoxal cavities, rectangular shaped in outline, with pyramidal surface. Mesoventrite not fused to mesepisterna, with anterior margin 0.2-times as wide as anterior margin of mesepisternum; anterior rib of mesoventrite bearing a medial semitriangular, pearlescent macula; posterior elevation of mesoventrite with a well-defined, curved transverse ridge, sometimes swollen at midpoint (as in 
<taxonomicName lsidName="K. cuzco" pageId="14" pageNumber="127" rank="species" species="cuzco">
<emphasis italics="true" pageId="14" pageNumber="127">K. cuzco</emphasis>
</taxonomicName>
); surface of mesoventrite reticulated for the most part; mesepisternum obliquely widely concave, with reticulated surface; mesepimeron trapezoid, with reticulate and pubescent surface. Mesofurca (examined in 
<taxonomicName lsidName="K. merida" pageId="14" pageNumber="127" rank="species" species="merida">
<emphasis italics="true" pageId="14" pageNumber="127">K. merida</emphasis>
</taxonomicName>
) with short arms, 0.6-times length of mesocoxae; apical half of arms free irregularly explanate, with outer corners sharply pointed. Metaventrite posteromesally elevated, with elevation rather flat; surface of metaventrite reticulate and pubescent, except for postero median and glabrous patch. Metepisterna 3-4-times longer than wide, only slightly narrowing posteriorly. Metepimeron triangular and acute. Metafurca (examined in 
<taxonomicName lsidName="K. merida" pageId="14" pageNumber="127" rank="species" species="merida">
<emphasis italics="true" pageId="14" pageNumber="127">K. merida</emphasis>
</taxonomicName>
, Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Thorax and abdomen of Katasophistes spp.: A-C Katasophistes merida: A ventral view of mesoventrite (white arrow pointing transverse ridge) B posterior view of metafurca: (at) anterior tendon, (c) crux, (ds) dorsal sheath, (fa) furcal arm, (h) hemiductus, (ls) lateral sheath, (s) stalk C fifth abdominal ventrite D Katasophistes charynae, fifth abdominal ventrite; E Katasophistes cuzco, fifth abdominal ventrite." figureDoi="10.3897/zookeys.768.24423.figure11" httpUri="https://binary.pensoft.net/fig/211223" pageId="14" pageNumber="127">11B</figureCitation>
) 1.6-times wider than long, with furcal arms (fa) slightly longer than stalk (s); stalk triangular (wider near the crux (c), gradually narrowing ventrally), with paired longitudinal keels extending along basal third of posterior face, and a well-developed median keel on anterior face extending to anterior margin of dorsal sheets (ds); outer margins of stalk diverging from base towards proximal third of furcal arms; furcal arms somewhat trapezoid, with apex (hemiductus (h)) only slightly explanate, obliquely positioned; anterior tendons (at) inserted at basal third of dorsal edge of furcal arms; dorsal sheaths well-developed, slightly wider than widest point of lateral sheaths (ls). 
<emphasis bold="true" pageId="14" pageNumber="127">Elytra.</emphasis>
Surface even (without elevations or depressions), serial punctures not clearly defined (e.g., Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus of Katasophistes spp.: A-E Katasophistes charynae: A dorsal view B lateral view C ventral view D head E-H Katasophistes cuzco: E dorsal view F lateral view G ventral view H head. Scale bars: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure9" httpUri="https://binary.pensoft.net/fig/211220" pageId="14" pageNumber="127">9E</figureCitation>
), not impressed into striae; seta bearing systematic punctures rather enlarged and deep (less strikingly so in 
<taxonomicName lsidName="K. charynae" pageId="14" pageNumber="127" rank="species" species="charynae">
<emphasis italics="true" pageId="14" pageNumber="127">K. charynae</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="K. cuzco" pageId="14" pageNumber="127" rank="species" species="cuzco">
<emphasis italics="true" pageId="14" pageNumber="127">K. cuzco</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="K. superficialis" pageId="14" pageNumber="127" rank="species" species="superficialis">
<emphasis italics="true" pageId="14" pageNumber="127">K. superficialis</emphasis>
</taxonomicName>
), forming five longitudinal rows, fifth row very close to outer margin of elytron; elytral margins slightly flared. Epipleura well-developed, slightly oblique, with sparse setae, anteriorly wide, gradually narrowing posteriorly, extending up to line of posterior margin of first abdominal ventrite; pseudepipleura well-developed, perpendicularly positioned, nearly half as wide as anterior portion of epipleura, extending along entire outer margin of elytra. Hind wings well-developed. 
<emphasis bold="true" pageId="14" pageNumber="127">Legs.</emphasis>
All femora with dense pubescence, at least along basal two thirds, remainder of surface glabrous, shiny and slightly reticulated; all femora antero-posteriorly flattened; metafemora with rather weak tibial grooves. Tibiae slender, weakly flattened, with well-developed spines; protibiae with a median longitudinal row of long spathulate setae along anterior surface. All tarsi with five tarsomeres, bearing long apical hair-like setae on dorsal face, and two lateral rows of spines and/or hair-like spines on ventral face of tarsomeres 2-4; protarsomeres 1-4 similar in size and shape; meso- and meta tarsomeres with tarsomere 2 nearly as long as tarsomere 5; tarsomere 5 approximately as long as tarsomeres 3-4 combined, without spines on ventral face; claws rather large, curved; empodium well-developed, bearing 
<pageBreakToken pageId="15" pageNumber="128" start="start">a</pageBreakToken>
pair of long, curved apical setae. 
<emphasis bold="true" pageId="15" pageNumber="128">Abdomen.</emphasis>
Abdomen with five ventrites, medially weakly convex; all ventrites reticulated, with uniform, dense, fine pubescence; posterior margin of fifth ventrite truncate to mesally weakly emarginated, set with a row of thick, flat spine-like setae (see Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Thorax and abdomen of Katasophistes spp.: A-C Katasophistes merida: A ventral view of mesoventrite (white arrow pointing transverse ridge) B posterior view of metafurca: (at) anterior tendon, (c) crux, (ds) dorsal sheath, (fa) furcal arm, (h) hemiductus, (ls) lateral sheath, (s) stalk C fifth abdominal ventrite D Katasophistes charynae, fifth abdominal ventrite; E Katasophistes cuzco, fifth abdominal ventrite." figureDoi="10.3897/zookeys.768.24423.figure11" httpUri="https://binary.pensoft.net/fig/211223" pageId="15" pageNumber="128">11C-E</figureCitation>
). Aedeagus (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Aedeagus of Katasophistes spp.: A-B Katasophistes charynae: A &quot; colln 32 &quot; B &quot; colln 42 a &quot; C Katasophistes cuzco; D Katasophistes merida E Katasophistes superficialis. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.768.24423.figure12" httpUri="https://binary.pensoft.net/fig/211224" pageId="15" pageNumber="128">12</figureCitation>
) nearly parallel sided, with basal piece between 0.5 and 1.1-times the length of parameres; median lobe wider than each paramere, gradually narrowing apically, with a conspicuous median longitudinal sclerotization, and well-developed lateral basal apodemes; apex of median lobe acute; parameres nearly as long as median lobe, with apical setae; gonopore preapically situated.
</paragraph>
<caption ID-Zenodo-Dep="1298850" doi="10.3897/zookeys.768.24423.figure8" httpUri="https://binary.pensoft.net/fig/211219" pageId="15" pageNumber="128" start="Figure 8" startId="F8">
<paragraph pageId="15" pageNumber="128">
<emphasis bold="true" pageId="15" pageNumber="128">Figure 8.</emphasis>
Habitat of 
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Crucisternum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crucisternum" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="15" pageNumber="128">Crucisternum</emphasis>
</taxonomicName>
spp. 
<emphasis bold="true" pageId="15" pageNumber="128">A-B</emphasis>
Habitat for 
<taxonomicName lsidName="C. ouboteri" pageId="15" pageNumber="128" rank="species" species="ouboteri">
<emphasis italics="true" pageId="15" pageNumber="128">C. ouboteri</emphasis>
</taxonomicName>
Suriname, Raleighvallen, collecting event SR12-0727-03A 
<emphasis bold="true" pageId="15" pageNumber="128">C</emphasis>
Type locality and habitat for 
<taxonomicName lsidName="C. ouboteri" pageId="15" pageNumber="128" rank="species" species="ouboteri">
<emphasis italics="true" pageId="15" pageNumber="128">C. ouboteri</emphasis>
</taxonomicName>
, Suriname, Brownsberg Nature Park, collecting event SR12-0804-01A 
<emphasis bold="true" pageId="15" pageNumber="128">D</emphasis>
Type locality and habitat for 
<taxonomicName lsidName="C. vanessae" pageId="15" pageNumber="128" rank="species" species="vanessae">
<emphasis italics="true" pageId="15" pageNumber="128">C. vanessae</emphasis>
</taxonomicName>
, Suriname, Tafelberg Summit, Caiman Creek, collecting event SR13-0819-05D 
<emphasis bold="true" pageId="15" pageNumber="128">E-F</emphasis>
Type locality and habitat for 
<taxonomicName lsidName="C. escalera" pageId="15" pageNumber="128" rank="species" species="escalera">
<emphasis italics="true" pageId="15" pageNumber="128">C. escalera</emphasis>
</taxonomicName>
, Venezuela, Bolivar State, along La Escalera, collecting event AS-08-058.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="1298852" doi="10.3897/zookeys.768.24423.figure9" httpUri="https://binary.pensoft.net/fig/211220" pageId="15" pageNumber="128" start="Figure 9" startId="F9">
<paragraph pageId="15" pageNumber="128">
<emphasis bold="true" pageId="15" pageNumber="128">Figure 9.</emphasis>
Habitus of 
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Katasophistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Katasophistes" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="15" pageNumber="128">Katasophistes</emphasis>
</taxonomicName>
spp.: 
<emphasis bold="true" pageId="15" pageNumber="128">A-E</emphasis>
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Katasophistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Katasophistes charynae" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="species" species="charynae">
<emphasis italics="true" pageId="15" pageNumber="128">Katasophistes charynae</emphasis>
</taxonomicName>
: 
<emphasis bold="true" pageId="15" pageNumber="128">A</emphasis>
dorsal view 
<emphasis bold="true" pageId="15" pageNumber="128">B</emphasis>
lateral view 
<emphasis bold="true" pageId="15" pageNumber="128">C</emphasis>
ventral view 
<emphasis bold="true" pageId="15" pageNumber="128">D</emphasis>
head 
<emphasis bold="true" pageId="15" pageNumber="128">E-H</emphasis>
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Katasophistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Katasophistes cuzco" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="species" species="cuzco">
<emphasis italics="true" pageId="15" pageNumber="128">Katasophistes cuzco</emphasis>
</taxonomicName>
: 
<emphasis bold="true" pageId="15" pageNumber="128">E</emphasis>
dorsal view 
<emphasis bold="true" pageId="15" pageNumber="128">F</emphasis>
lateral view 
<emphasis bold="true" pageId="15" pageNumber="128">G</emphasis>
ventral view 
<emphasis bold="true" pageId="15" pageNumber="128">H</emphasis>
head. Scale bars: 1 mm.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="1298814" doi="10.3897/zookeys.768.24423.figure10" httpUri="https://binary.pensoft.net/fig/211221" pageId="15" pageNumber="128" start="Figure 10" startId="F10">
<paragraph pageId="15" pageNumber="128">
<emphasis bold="true" pageId="15" pageNumber="128">Figure 10.</emphasis>
Habitus of 
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Katasophistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Katasophistes" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="15" pageNumber="128">Katasophistes</emphasis>
</taxonomicName>
spp.: 
<emphasis bold="true" pageId="15" pageNumber="128">A-E</emphasis>
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Katasophistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Katasophistes merida" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="species" species="merida">
<emphasis italics="true" pageId="15" pageNumber="128">Katasophistes merida</emphasis>
</taxonomicName>
: 
<emphasis bold="true" pageId="15" pageNumber="128">A</emphasis>
dorsal view 
<emphasis bold="true" pageId="15" pageNumber="128">B</emphasis>
lateral view 
<emphasis bold="true" pageId="15" pageNumber="128">C</emphasis>
ventral view 
<emphasis bold="true" pageId="15" pageNumber="128">D</emphasis>
head 
<emphasis bold="true" pageId="15" pageNumber="128">E-H</emphasis>
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Katasophistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Katasophistes superficialis" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="species" species="superficialis">
<emphasis italics="true" pageId="15" pageNumber="128">Katasophistes superficialis</emphasis>
</taxonomicName>
: 
<emphasis bold="true" pageId="15" pageNumber="128">E</emphasis>
dorsal view 
<emphasis bold="true" pageId="15" pageNumber="128">F</emphasis>
lateral view 
<emphasis bold="true" pageId="15" pageNumber="128">G</emphasis>
ventral view 
<emphasis bold="true" pageId="15" pageNumber="128">H</emphasis>
head. Scale bar: 1 mm.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="1298816" doi="10.3897/zookeys.768.24423.figure11" httpUri="https://binary.pensoft.net/fig/211223" pageId="15" pageNumber="128" start="Figure 11" startId="F11">
<paragraph pageId="15" pageNumber="128">
<emphasis bold="true" pageId="15" pageNumber="128">Figure 11.</emphasis>
Thorax and abdomen of 
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Katasophistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Katasophistes" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="15" pageNumber="128">Katasophistes</emphasis>
</taxonomicName>
spp.: 
<emphasis bold="true" pageId="15" pageNumber="128">A-C</emphasis>
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Katasophistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Katasophistes merida" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="species" species="merida">
<emphasis italics="true" pageId="15" pageNumber="128">Katasophistes merida</emphasis>
</taxonomicName>
: 
<emphasis bold="true" pageId="15" pageNumber="128">A</emphasis>
ventral view of mesoventrite (white arrow pointing transverse ridge) 
<emphasis bold="true" pageId="15" pageNumber="128">B</emphasis>
posterior view of metafurca: (at) anterior tendon, (c) crux, (ds) dorsal sheath, (fa) furcal arm, (h) hemiductus, (ls) lateral sheath, (s) stalk 
<emphasis bold="true" pageId="15" pageNumber="128">C</emphasis>
fifth abdominal ventrite 
<emphasis bold="true" pageId="15" pageNumber="128">D</emphasis>
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Katasophistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Katasophistes charynae" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="species" species="charynae">
<emphasis italics="true" pageId="15" pageNumber="128">Katasophistes charynae</emphasis>
</taxonomicName>
, fifth abdominal ventrite; 
<emphasis bold="true" pageId="15" pageNumber="128">E</emphasis>
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Katasophistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Katasophistes cuzco" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="species" species="cuzco">
<emphasis italics="true" pageId="15" pageNumber="128">Katasophistes cuzco</emphasis>
</taxonomicName>
, fifth abdominal ventrite.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="15" pageNumber="128" type="larvae">
<paragraph pageId="15" pageNumber="128">Larvae.</paragraph>
<paragraph pageId="15" pageNumber="128">The immature stages are unknown.</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="128" type="etymology">
<paragraph pageId="15" pageNumber="128">Etymology.</paragraph>
<paragraph pageId="15" pageNumber="128">
Named from the Greek word 
<emphasis italics="true" pageId="15" pageNumber="128">katasophistes</emphasis>
, meaning trickster, in reference to the disparity of the general appearance of some of the known species. Name to be treated as masculine.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="128" type="distribution">
<paragraph pageId="15" pageNumber="128">Distribution.</paragraph>
<paragraph pageId="15" pageNumber="128">
Venezuela (
<normalizedToken originalValue="Mérida">Merida</normalizedToken>
), Ecuador (Pastaza), Peru (Cuzco, Madre de Dios). See Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Distribution of Katasophistes species." figureDoi="10.3897/zookeys.768.24423.figure13" httpUri="https://binary.pensoft.net/fig/211225" pageId="15" pageNumber="128">13</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="128" type="biology_ecology">
<paragraph pageId="15" pageNumber="128">Biology.</paragraph>
<paragraph pageId="15" pageNumber="128">
Species in this genus exhibit a combination of ecologies: 
<taxonomicName lsidName="K. merida" pageId="15" pageNumber="128" rank="species" species="merida">
<emphasis italics="true" pageId="15" pageNumber="128">K. merida</emphasis>
</taxonomicName>
is restricted to seepage habitats, while the type series of 
<taxonomicName lsidName="K. superficialis" pageId="15" pageNumber="128" rank="species" species="superficialis">
<emphasis italics="true" pageId="15" pageNumber="128">K. superficialis</emphasis>
</taxonomicName>
was collected from forested stream pools with abundant detritus. See Fig. 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Habitat of Katasophistes spp. A-B Type locality and habitat of K. merida: Venezuela: Merida State, Collecting Event VZ 12 - 0122 - 03 A C-D Habitat of K. merida: Venezuela: Merida State, Collecting Event VZ 12 - 0124 - 01 A E-F Type locality and habitat of K. superficialis: Ecuador: Pastaza Province, Collecting Event AS- 08 - 08 b." figureDoi="10.3897/zookeys.768.24423.figure14" httpUri="https://binary.pensoft.net/fig/211227" pageId="15" pageNumber="128">14</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="128" type="description">
<paragraph pageId="15" pageNumber="128">
Characters of taxonomic importance for 
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Katasophistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Katasophistes" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="15" pageNumber="128">Katasophistes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="15" pageNumber="128">
With the exception of the enlarged elytral systematic punctures, there is nothing particularly remarkable about the external morphology of 
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Katasophistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Katasophistes" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="15" pageNumber="128">Katasophistes</emphasis>
</taxonomicName>
. Careful examination of the elytral systematic punctures is needed in order to recognize 
<taxonomicName lsidName="K. charynae" pageId="15" pageNumber="128" rank="species" species="charynae">
<emphasis italics="true" pageId="15" pageNumber="128">K. charynae</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="K. cuzco" pageId="15" pageNumber="128" rank="species" species="cuzco">
<emphasis italics="true" pageId="15" pageNumber="128">K. cuzco</emphasis>
</taxonomicName>
, and 
<taxonomicName lsidName="K. superficialis" pageId="15" pageNumber="128" rank="species" species="superficialis">
<emphasis italics="true" pageId="15" pageNumber="128">K. superficialis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="15" pageNumber="128">
<emphasis bold="true" pageId="15" pageNumber="128">
<emphasis italics="true" pageId="15" pageNumber="128">Punctation</emphasis>
.
</emphasis>
The most prominent feature of 
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Katasophistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Katasophistes" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="15" pageNumber="128">Katasophistes</emphasis>
</taxonomicName>
is the enlargement of the elytral systematic punctures, which is evident in 
<taxonomicName lsidName="K. merida" pageId="15" pageNumber="128" rank="species" species="merida">
<emphasis italics="true" pageId="15" pageNumber="128">K. merida</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Katasophistes spp.: A-E Katasophistes merida: A dorsal view B lateral view C ventral view D head E-H Katasophistes superficialis: E dorsal view F lateral view G ventral view H head. Scale bar: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure10" httpUri="https://binary.pensoft.net/fig/211221" pageId="15" pageNumber="128">10A, B</figureCitation>
), but much less so in 
<taxonomicName lsidName="K. charynae" pageId="15" pageNumber="128" rank="species" species="charynae">
<emphasis italics="true" pageId="15" pageNumber="128">K. charynae</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="K. cuzco" pageId="15" pageNumber="128" rank="species" species="cuzco">
<emphasis italics="true" pageId="15" pageNumber="128">K. cuzco</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="K. superficialis" pageId="15" pageNumber="128" rank="species" species="superficialis">
<emphasis italics="true" pageId="15" pageNumber="128">K. superficialis</emphasis>
</taxonomicName>
. One way to recognize the enlarged elytral systematic punctures would be by spotting the long setae that systematic punctures bear.
</paragraph>
<paragraph pageId="15" pageNumber="128">
<emphasis bold="true" pageId="15" pageNumber="128">
<emphasis italics="true" pageId="15" pageNumber="128">Posterior elevation of mesoventrite</emphasis>
.
</emphasis>
It is usually well-defined and shaped as a curved transverse ridge. Only in 
<taxonomicName lsidName="K. cuzco" pageId="15" pageNumber="128" rank="species" species="cuzco">
<emphasis italics="true" pageId="15" pageNumber="128">K. cuzco</emphasis>
</taxonomicName>
is this transverse ridge additionally medially swollen, as to form a low bump.
</paragraph>
<paragraph pageId="15" pageNumber="128">
<emphasis bold="true" pageId="15" pageNumber="128">
<emphasis italics="true" pageId="15" pageNumber="128">Aedeagus</emphasis>
.
</emphasis>
In 
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Katasophistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Katasophistes" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="15" pageNumber="128">Katasophistes</emphasis>
</taxonomicName>
the aedeagus exhibits the same general shape of the median lobe, combined with a wide variety of shapes of the parameres (see Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Aedeagus of Katasophistes spp.: A-B Katasophistes charynae: A &quot; colln 32 &quot; B &quot; colln 42 a &quot; C Katasophistes cuzco; D Katasophistes merida E Katasophistes superficialis. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.768.24423.figure12" httpUri="https://binary.pensoft.net/fig/211224" pageId="15" pageNumber="128">12</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="128" type="key">
<paragraph pageId="15" pageNumber="128">
Key to the species of 
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Katasophistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Katasophistes" order="Coleoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="15" pageNumber="128">Katasophistes</emphasis>
</taxonomicName>
</paragraph>
<paragraph pageId="15" pageNumber="128">
<table inLine="true" pageId="15" pageNumber="128">
<tr pageId="15" pageNumber="128">
<td colspan="1" pageId="15" pageNumber="128" rowspan="1">1</td>
<td colspan="1" pageId="15" pageNumber="128" rowspan="1">
Elytra with strikingly large and deeply impressed systematic punctures (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Katasophistes spp.: A-E Katasophistes merida: A dorsal view B lateral view C ventral view D head E-H Katasophistes superficialis: E dorsal view F lateral view G ventral view H head. Scale bar: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure10" httpUri="https://binary.pensoft.net/fig/211221" pageId="15" pageNumber="128">10A, B</figureCitation>
); maxillary palps relatively short and stout (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Katasophistes spp.: A-E Katasophistes merida: A dorsal view B lateral view C ventral view D head E-H Katasophistes superficialis: E dorsal view F lateral view G ventral view H head. Scale bar: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure10" httpUri="https://binary.pensoft.net/fig/211221" pageId="15" pageNumber="128">10D</figureCitation>
)
</td>
<td colspan="1" pageId="15" pageNumber="128" rowspan="1">
<taxonomicName lsidName="K. merida" pageId="15" pageNumber="128" rank="species" species="merida">
<emphasis bold="true" italics="true" pageId="15" pageNumber="128">K. merida</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="15" pageNumber="128">
<td colspan="1" pageId="15" pageNumber="128" rowspan="1">-</td>
<td colspan="1" pageId="15" pageNumber="128" rowspan="1">
Elytra with slightly enlarged systematic punctures which are shallowly impressed; maxillary palps relatively long and slender (e.g., Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Katasophistes spp.: A-E Katasophistes merida: A dorsal view B lateral view C ventral view D head E-H Katasophistes superficialis: E dorsal view F lateral view G ventral view H head. Scale bar: 1 mm." figureDoi="10.3897/zookeys.768.24423.figure10" httpUri="https://binary.pensoft.net/fig/211221" pageId="15" pageNumber="128">10H</figureCitation>
)
</td>
<td colspan="1" pageId="15" pageNumber="128" rowspan="1">
<emphasis bold="true" pageId="15" pageNumber="128">2</emphasis>
</td>
</tr>
<tr pageId="15" pageNumber="128">
<td colspan="1" pageId="15" pageNumber="128" rowspan="1">2</td>
<td colspan="1" pageId="15" pageNumber="128" rowspan="1">
Body length around 4.5 mm; basal piece of aedeagus nearly as long as parameres (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Aedeagus of Katasophistes spp.: A-B Katasophistes charynae: A &quot; colln 32 &quot; B &quot; colln 42 a &quot; C Katasophistes cuzco; D Katasophistes merida E Katasophistes superficialis. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.768.24423.figure12" httpUri="https://binary.pensoft.net/fig/211224" pageId="15" pageNumber="128">12E</figureCitation>
)
</td>
<td colspan="1" pageId="15" pageNumber="128" rowspan="1">
<taxonomicName lsidName="K. superficialis" pageId="15" pageNumber="128" rank="species" species="superficialis">
<emphasis bold="true" italics="true" pageId="15" pageNumber="128">K. superficialis</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="15" pageNumber="128">
<td colspan="1" pageId="15" pageNumber="128" rowspan="1">-</td>
<td colspan="1" pageId="15" pageNumber="128" rowspan="1">
Body length less than 4 mm; basal piece of aedeagus nearly half as long as parameres (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Aedeagus of Katasophistes spp.: A-B Katasophistes charynae: A &quot; colln 32 &quot; B &quot; colln 42 a &quot; C Katasophistes cuzco; D Katasophistes merida E Katasophistes superficialis. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.768.24423.figure12" httpUri="https://binary.pensoft.net/fig/211224" pageId="15" pageNumber="128">12A-C</figureCitation>
)
</td>
<td colspan="1" pageId="15" pageNumber="128" rowspan="1">
<emphasis bold="true" pageId="15" pageNumber="128">3</emphasis>
</td>
</tr>
<tr pageId="15" pageNumber="128">
<td colspan="1" pageId="15" pageNumber="128" rowspan="1">3</td>
<td colspan="1" pageId="15" pageNumber="128" rowspan="1">
Transverse ridge on posterior elevation of mesoventrite uniformly wide; parameres with widest basal point clearly wider than pre-apical width (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Aedeagus of Katasophistes spp.: A-B Katasophistes charynae: A &quot; colln 32 &quot; B &quot; colln 42 a &quot; C Katasophistes cuzco; D Katasophistes merida E Katasophistes superficialis. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.768.24423.figure12" httpUri="https://binary.pensoft.net/fig/211224" pageId="15" pageNumber="128">12A, B</figureCitation>
)
</td>
<td colspan="1" pageId="15" pageNumber="128" rowspan="1">
<taxonomicName lsidName="K. charynae" pageId="15" pageNumber="128" rank="species" species="charynae">
<emphasis bold="true" italics="true" pageId="15" pageNumber="128">K. charynae</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="15" pageNumber="128">
<td colspan="1" pageId="15" pageNumber="128" rowspan="1">-</td>
<td colspan="1" pageId="15" pageNumber="128" rowspan="1">
Transverse ridge on posterior elevation of mesoventrite swollen at midpoint; parameres with widest basal point nearly as wide as pre-apical width (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Aedeagus of Katasophistes spp.: A-B Katasophistes charynae: A &quot; colln 32 &quot; B &quot; colln 42 a &quot; C Katasophistes cuzco; D Katasophistes merida E Katasophistes superficialis. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.768.24423.figure12" httpUri="https://binary.pensoft.net/fig/211224" pageId="15" pageNumber="128">12C</figureCitation>
)
</td>
<td colspan="1" pageId="15" pageNumber="128" rowspan="1">
<taxonomicName lsidName="K. cuzco" pageId="15" pageNumber="128" rank="species" species="cuzco">
<emphasis bold="true" italics="true" pageId="15" pageNumber="128">K. cuzco</emphasis>
</taxonomicName>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>