treatments-xml/data/2E/2F/87/2E2F87C1FFCA272CFE3FFC592018FAF0.xml
2024-06-21 12:32:37 +02:00

266 lines
30 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="F70508B73A117B241C4A17FD0C173383" ID-DOI="10.1080/00222933.2017.1355500" ID-ISSN="1464-5262" ID-Zenodo-Dep="5182722" ID-ZooBank="4E0A6295-A2FA-43BB-993D-BB00AFF54406" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628707176855" checkinUser="felipe" docAuthor="Csuzdi, Csaba, Pearlson, Oren &amp; Pavlíček, Tomás" docDate="2017" docId="2E2F87C1FFCA272CFE3FFC592018FAF0" docLanguage="en" docName="JNatHist.51.31-32.1899-1912.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 51 (31 - 32)" docSource="http://dx.doi.org/10.1080/00222933.2017.1355500" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Acanthodrilus barrierei Csuzdi &amp; Pearlson &amp; Pavlíček 2017, sp. nov." docType="treatment" docVersion="4" lastPageNumber="1902" masterDocId="D216FFB9FFC82728FFEEFFEF2423FFE6" masterDocTitle="New Acanthodrilus species from New Caledonia (Clitellata, Megadrili, Acanthodrilidae)" masterLastPageNumber="1912" masterPageNumber="1899" pageNumber="1900" updateTime="1699050311385" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="681706F0C7BCF50222D10E8A77C57D59" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B8FBE245B6DE613D1F1B673B40990085">
<mods:title id="DB3A35C621C79A2862149B4CD4A39ABC">New Acanthodrilus species from New Caledonia (Clitellata, Megadrili, Acanthodrilidae)</mods:title>
</mods:titleInfo>
<mods:name id="58BD959C73F451348E60E991C0E375B8" type="personal">
<mods:role id="FD4829EBC2B68E42E0628BA26BC0302D">
<mods:roleTerm id="6B7770B49C275D752AC1F3D897950412">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A1FDACC4274DB65ACEC7B81B1BCC9041">Csuzdi, Csaba</mods:namePart>
</mods:name>
<mods:name id="853385E731836F6FBB556FFCD84CF79E" type="personal">
<mods:role id="384CE4EA6E81A15C684C76ADE4215C2E">
<mods:roleTerm id="709D2422C199357CD79A33BE5DAB58DA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F4E5438AE0107B75E04D47C50FBB00CC">Pearlson, Oren</mods:namePart>
</mods:name>
<mods:name id="2982EDACFD9A8A026B66BC4C54721793" type="personal">
<mods:role id="E1EBC0DF7D65DF38D54120B13BB8A98F">
<mods:roleTerm id="AEFF7E9BD3462CDA59D04FC900AEE843">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="88DC78DED9D67D8E4D01747C466F3290">Pavlíček, Tomás</mods:namePart>
</mods:name>
<mods:typeOfResource id="0267D3EEE3B4E803E408C5B069F97CFD">text</mods:typeOfResource>
<mods:relatedItem id="2BB2F8F6B72CD5607F725C24741FE28F" type="host">
<mods:titleInfo id="E7DCE43F74EA788AB9227725DBD302D0">
<mods:title id="C233D003238F740CF4B37F38562EF604">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="3D971B273B48D4C5D3DA181E67060CBA">
<mods:date id="3079315715B5779ABBB79DD741F9442A">2017</mods:date>
<mods:detail id="FEEDA83F97CAA8CA8B477A7AD12B5516" type="series">
<mods:title id="6BACBBF6C4724BB7BE51DD1764418716">J. Nat. Hist.</mods:title>
</mods:detail>
<mods:detail id="4800D2C5281E222A7B877BF49F36FE92" type="pubDate">
<mods:number id="81F0B6E73693DA7477BB36CFBE240FCB">2017-08-15</mods:number>
</mods:detail>
<mods:detail id="EF2AC2E6C58B63F22B5AA004281E3DAC" type="volume">
<mods:number id="BA8753B4C813A9CA928EADC198ADEF5A">51</mods:number>
</mods:detail>
<mods:detail id="891871AD4FAE3BF293B61699DC9893AA" type="issue">
<mods:number id="4327C80BA1A39EDB3327272D3BFADDFD">31 - 32</mods:number>
</mods:detail>
<mods:extent id="EC99B095E00CF241EE4830BD30E20420" unit="page">
<mods:start id="1F884F7D5F816005F38E0AC8F6EF693E">1899</mods:start>
<mods:end id="8B2479BD43764C106FF1841FD5A8EDF4">1912</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="501B666B64AA8EA8D5C60A0FC488321D">
<mods:url id="3312F9C7D6411AB2D4333E7A2CF870E7">http://dx.doi.org/10.1080/00222933.2017.1355500</mods:url>
</mods:location>
<mods:classification id="6504CF10975D51C0B193B00C1BB3443C">journal article</mods:classification>
<mods:identifier id="108604502056EFFADE9069E7284A88A3" type="DOI">10.1080/00222933.2017.1355500</mods:identifier>
<mods:identifier id="F23C31B283EDEA36E2B89A6FB6C7CF0E" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="0210E416CE0DD0CC36E1230FFCA3464D" type="Zenodo-Dep">5182722</mods:identifier>
<mods:identifier id="99EB68F422F449FE3CA37D1B5F8ECA0C" type="ZooBank">4E0A6295-A2FA-43BB-993D-BB00AFF54406</mods:identifier>
</mods:mods>
<treatment id="2E2F87C1FFCA272CFE3FFC592018FAF0" ID-DOI="http://doi.org/10.5281/zenodo.5191710" ID-Zenodo-Dep="5191710" LSID="urn:lsid:plazi:treatment:2E2F87C1FFCA272CFE3FFC592018FAF0" httpUri="http://treatment.plazi.org/id/2E2F87C1FFCA272CFE3FFC592018FAF0" lastPageId="4" lastPageNumber="1902" pageId="2" pageNumber="1900">
<subSubSection id="EE9C655CFFCA272AFE3FFC592770FC36" box="[465,851,950,976]" pageId="2" pageNumber="1900" type="nomenclature">
<paragraph id="A63936D7FFCA272AFE3FFC592770FC36" blockId="2.[419,896,882,1011]" box="[465,851,950,976]" pageId="2" pageNumber="1900">
<heading id="FD7181BBFFCA272AFE3FFC592770FC36" box="[465,851,950,976]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="1900" reason="2">
<taxonomicName id="61864D54FFCA272AFE3FFC5926CEFC36" authority="Csuzdi &amp; Pearlson &amp; Pavlíček, 2017" authorityName="Csuzdi &amp; Pearlson &amp; Pavlíček" authorityYear="2017" box="[465,749,950,976]" class="Clitellata" family="Ocnerodrilidae" genus="Acanthodrilus" kingdom="Animalia" order="Haplotaxida" pageId="2" pageNumber="1900" phylum="Annelida" rank="species" species="barrierei" status="sp. nov.">
<emphasis id="94F2EAC5FFCA272AFE3FFC5926CEFC36" bold="true" box="[465,749,950,976]" italics="true" pageId="2" pageNumber="1900">Acanthodrilus barrierei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8FC157BEFFCA272AFD18FC582770FC36" box="[758,851,951,976]" pageId="2" pageNumber="1900" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EE9C655CFFCA272AFDB7FC3526E8FC15" box="[601,715,986,1011]" pageId="2" pageNumber="1900" type="description">
<paragraph id="A63936D7FFCA272AFDB7FC3526E8FC15" blockId="2.[419,896,882,1011]" box="[601,715,986,1011]" pageId="2" pageNumber="1900">
(
<figureCitation id="3EBD2A52FFCA272AFD8FFC3526E1FC15" box="[609,706,986,1011]" captionStart="Figure 1" captionStartId="3.[160,225,1634,1657]" captionTargetBox="[205,1115,142,1603]" captionTargetId="figure-1@3.[205,1116,142,1603]" captionTargetPageId="3" captionText="Figure 1. Acanthodrilus barrieri sp. n. (a) ventral view of the clitellar region; Fp, female pore; Gp, genital papillae; Gs, genital setal papilla; Mp, male pore; Prp, prostate pores; St, spermathecal pores; scale bar, 1 mm. (b) nephridial bladder from the postclitellar segments. (c) penial seta; scale bar, 0.05 mm. (d) genital seta; scale bar, 0.03 mm. (e) spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5182724" httpUri="https://zenodo.org/record/5182724/files/figure.png" pageId="2" pageNumber="1900">Figure 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="EE9C655CFFCA272AFF4EFBC4207BFB4A" pageId="2" pageNumber="1900" type="materials_examined">
<paragraph id="A63936D7FFCA272AFF4EFBC42566FBA2" blockId="2.[160,1156,1067,1196]" box="[160,325,1067,1092]" pageId="2" pageNumber="1900">
<emphasis id="94F2EAC5FFCA272AFF4EFBC42566FBA2" bold="true" box="[160,325,1067,1092]" italics="true" pageId="2" pageNumber="1900">Type material</emphasis>
</paragraph>
<paragraph id="A63936D7FFCA272AFF4EFBA1207BFB4A" blockId="2.[160,1156,1067,1196]" pageId="2" pageNumber="1900">
<materialsCitation id="16EE3C8AFFCA272AFF4EFBA120A3FB6F" collectingDate="2014-01-31" collectionCode="HNHM, T" collectorName="Pavli C" country="New Caledonia" county="River Provincial Park" latitude="-22.15" location="Blue River Provincial Park" longLatPrecision="756" longitude="166.73" municipality="Blue River" pageId="2" pageNumber="1900" specimenCode="AF5683" specimenCount="1" typeStatus="holotype">
<emphasis id="94F2EAC5FFCA272AFF4EFBA12532FB80" bold="true" box="[160,273,1102,1127]" italics="true" pageId="2" pageNumber="1900">
<typeStatus id="793D8875FFCA272AFF4EFBA12528FB81" box="[160,267,1102,1127]" pageId="2" pageNumber="1900" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="C097AE12FFCA272AFECFFBA1254FFB80" box="[289,364,1102,1126]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="2" pageNumber="1900">HNHM</collectionCode>
/
<specimenCode id="F6209EACFFCA272AFE9DFBA125E8FB80" box="[371,459,1102,1126]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="2" pageNumber="1900">AF5683</specimenCode>
, clitellate adult, complete specimen.
<collectingCountry id="DE917647FFCA272AFC60FBA12062FB80" box="[910,1089,1102,1126]" name="New Caledonia" pageId="2" pageNumber="1900">New Caledonia</collectingCountry>
,
<location id="A359600CFFCA272AFBBCFBA125A9FB6F" LSID="urn:lsid:plazi:treatment:2E2F87C1FFCA272CFE3FFC592018FAF0:A359600CFFCA272AFBBCFBA125A9FB6F" country="New Caledonia" county="River Provincial Park" latitude="-22.15" longLatPrecision="756" longitude="166.73" municipality="Blue River" name="Blue River Provincial Park" pageId="2" pageNumber="1900">
<collectingMunicipality id="465DACADFFCA272AFBBCFBA124F4FB6F" pageId="2" pageNumber="1900">
Blue
<collectingCounty id="4F584E5BFFCA272AFF4EFB9E24F4FB6F" box="[160,215,1137,1161]" pageId="2" pageNumber="1900">River</collectingCounty>
</collectingMunicipality>
Provincial Park
</location>
(
<geoCoordinate id="C3B25010FFCA272AFE74FB9E25D8FB6F" box="[410,507,1137,1161]" degrees="22.15" direction="south" orientation="latitude" pageId="2" pageNumber="1900" precision="555" value="-22.15">22.15° S</geoCoordinate>
,
<geoCoordinate id="C3B25010FFCA272AFDE7FB9E2655FB6F" box="[521,630,1137,1161]" degrees="166.73" direction="east" orientation="longitude" pageId="2" pageNumber="1900" precision="555" value="166.73">166.73° E</geoCoordinate>
), rainforest,
<date id="D2381017FFCA272AFCEBFB9E27E6FB6F" box="[773,965,1137,1161]" pageId="2" pageNumber="1900" value="2014-01-31">
<collectingDate id="C27CE9FFFFCA272AFCEBFB9E27E6FB6F" box="[773,965,1137,1161]" pageId="2" pageNumber="1900" value="2014-01-31">31 January 2014</collectingDate>
</date>
, leg.
<collectionCode id="C097AE12FFCA272AFBE8FB9E2035FB6F" box="[1030,1046,1137,1161]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="1900">T</collectionCode>
.
<collectorName id="0B735301FFCA272AFBCDFB9E2047FB6F" box="[1059,1124,1137,1161]" pageId="2" pageNumber="1900">
Pavlí
<emphasis id="94F2EAC5FFCA272AFBB6FB9E2047FB6F" bold="true" box="[1112,1124,1137,1161]" pageId="2" pageNumber="1900">Č</emphasis>
</collectorName>
ek
</materialsCitation>
.
<materialsCitation id="16EE3C8AFFCA272AFF4EFB7B26EFFB4A" box="[160,716,1172,1196]" collectionCode="HNHM" country="New Caledonia" pageId="2" pageNumber="1900" specimenCode="AF5684" specimenCount="1" typeStatus="paratype">
<typeStatus id="793D8875FFCA272AFF4EFB7B2533FB4A" box="[160,272,1172,1196]" pageId="2" pageNumber="1900" type="paratype">Paratypes</typeStatus>
:
<collectionCode id="C097AE12FFCA272AFEF5FB7B2546FB4A" box="[283,357,1172,1196]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="2" pageNumber="1900">HNHM</collectionCode>
/
<specimenCode id="F6209EACFFCA272AFE83FB7B25E6FB4A" box="[365,453,1172,1196]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="2" pageNumber="1900">AF5684</specimenCode>
, two clitellate adult ex
</materialsCitation>
., locality same as that of
<typeStatus id="793D8875FFCA272AFC00FB7B2076FB4A" box="[1006,1109,1172,1196]" pageId="2" pageNumber="1900" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="EE9C655CFFCA272AFF4EFB3326BAFADC" pageId="2" pageNumber="1900" type="etymology">
<paragraph id="A63936D7FFCA272AFF4EFB33253EFB13" blockId="2.[160,1156,1244,1338]" box="[160,285,1244,1269]" pageId="2" pageNumber="1900">
<emphasis id="94F2EAC5FFCA272AFF4EFB33253EFB13" bold="true" box="[160,285,1244,1269]" italics="true" pageId="2" pageNumber="1900">Etymology</emphasis>
</paragraph>
<paragraph id="A63936D7FFCA272AFF4EFAEF26BAFADC" blockId="2.[160,1156,1244,1338]" pageId="2" pageNumber="1900">
The new species is named in honour of Frédérique Alain-Barrière for her generous help during the collecting trip to
<collectingCountry id="DE917647FFCA272AFE09FACD26B7FADC" box="[487,660,1314,1338]" name="New Caledonia" pageId="2" pageNumber="1900">New Caledonia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="EE9C655CFFCA272AFF4EFA842073F9B5" pageId="2" pageNumber="1900" type="diagnosis">
<paragraph id="A63936D7FFCA272AFF4EFA842537FA62" blockId="2.[160,1156,1387,1619]" box="[160,276,1387,1412]" pageId="2" pageNumber="1900">
<emphasis id="94F2EAC5FFCA272AFF4EFA842537FA62" bold="true" box="[160,276,1387,1412]" italics="true" pageId="2" pageNumber="1900">Diagnosis</emphasis>
</paragraph>
<paragraph id="A63936D7FFCA272AFF4EFA612073F9B5" blockId="2.[160,1156,1387,1619]" pageId="2" pageNumber="1900">
L.
<quantity id="617E9B32FFCA272AFF52FA61252CFA40" box="[188,271,1422,1446]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="1900" unit="mm" value="90.0">90 mm</quantity>
. D.
<quantity id="617E9B32FFCA272AFED0FA6125A7FA40" box="[318,388,1422,1446]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="1900" unit="mm" value="2.0">2 mm</quantity>
. Average weight
<quantity id="617E9B32FFCA272AFDA2FA6126A4FA40" box="[588,647,1422,1446]" metricMagnitude="-4" metricUnit="kg" metricValue="6.0" pageId="2" pageNumber="1900" unit="g" value="0.6">0.6 g</quantity>
. Colour yellowish. First dorsal pore in 12/13. Clitellum ½13½17, circular. Prostatic pores paired in 17, 19. Penial setae L.
<quantity id="617E9B32FFCA272AFBE8FA5E207DFA2F" box="[1030,1118,1457,1481]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="1900" unit="mm" value="3.5">3.5 mm</quantity>
, D.
<quantity id="617E9B32FFCA272AFF4EFA3C2520FA0D" box="[160,259,1491,1515]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="2" pageNumber="1900" unit="mm" value="0.07">0.07 mm</quantity>
, ornamentation group of spines. Spermathecal pores 7/8, 8/9
<emphasis id="94F2EAC5FFCA272AFC41FA3D279EFA0D" box="[943,957,1490,1515]" italics="true" pageId="2" pageNumber="1900">b</emphasis>
. Muscular gizzard in 6, calciferous glands lacking. Last pair of hearts in 13. Spermathecae with long narrow duct and a somewhat shorter oval ampoule joining the duct at a right angle. The duct entally bears a small diverticulum filled with sperm-chambers. Genital setae present.
</paragraph>
</subSubSection>
<subSubSection id="EE9C655CFFCA272AFF4EF96B2505F97B" box="[160,294,1668,1693]" pageId="2" pageNumber="1900" type="description">
<paragraph id="A63936D7FFCA272AFF4EF96B2505F97B" blockId="2.[160,736,1668,1727]" box="[160,294,1668,1693]" pageId="2" pageNumber="1900">
<emphasis id="94F2EAC5FFCA272AFF4EF96B2505F97B" bold="true" box="[160,294,1668,1693]" italics="true" pageId="2" pageNumber="1900">Description</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="EE9C655CFFCA272CFF4EF949202EFBDF" lastPageId="4" lastPageNumber="1902" pageId="2" pageNumber="1900" type="materials_examined">
<paragraph id="A63936D7FFCA272AFF4EF94926C3F959" blockId="2.[160,736,1668,1727]" box="[160,736,1702,1727]" pageId="2" pageNumber="1900">
<emphasis id="94F2EAC5FFCA272AFF4EF9492532F959" bold="true" box="[160,273,1702,1727]" italics="true" pageId="2" pageNumber="1900">
<typeStatus id="793D8875FFCA272AFF4EF9492528F959" box="[160,267,1702,1727]" pageId="2" pageNumber="1900" type="holotype">Holotype</typeStatus>
.
</emphasis>
L.
<quantity id="617E9B32FFCA272AFED2F94825ACF959" box="[316,399,1703,1727]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="1900" unit="mm" value="90.0">90 mm</quantity>
, D.
<quantity id="617E9B32FFCA272AFE55F9482623F959" box="[443,512,1703,1727]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="1900" unit="mm" value="2.0">2 mm</quantity>
, No. segments 165.
</paragraph>
<caption id="F2F9665FFFCB272BFF4EF98D276FF937" ID-DOI="http://doi.org/10.5281/zenodo.5182724" ID-Zenodo-Dep="5182724" httpUri="https://zenodo.org/record/5182724/files/figure.png" pageId="3" pageNumber="1901" startId="3.[160,225,1634,1657]" targetBox="[205,1115,142,1603]" targetPageId="3">
<paragraph id="A63936D7FFCB272BFF4EF98D276FF937" blockId="3.[160,1162,1633,1745]" pageId="3" pageNumber="1901">
<emphasis id="94F2EAC5FFCB272BFF4EF98D2522F99F" bold="true" box="[160,257,1634,1657]" pageId="3" pageNumber="1901">Figure 1.</emphasis>
<taxonomicName id="61864D54FFCB272BFEE7F98E25FDF99E" authority="Csuzdi &amp; Pearlson &amp; Pavlíček, 2017" authorityName="Csuzdi &amp; Pearlson &amp; Pavlíček" authorityYear="2017" box="[265,478,1633,1656]" class="Clitellata" family="Ocnerodrilidae" genus="Acanthodrilus" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="1901" phylum="Annelida" rank="species" species="barrieri" status="sp. nov.">
<emphasis id="94F2EAC5FFCB272BFEE7F98E25FDF99E" box="[265,478,1633,1656]" italics="true" pageId="3" pageNumber="1901">Acanthodrilus barrieri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8FC157BEFFCB272BFE02F98D2604F99F" box="[492,551,1634,1657]" pageId="3" pageNumber="1901" rank="species">sp. n.</taxonomicNameLabel>
(a) ventral view of the clitellar region;
<emphasis id="94F2EAC5FFCB272BFC33F98E27D7F99E" box="[989,1012,1633,1656]" italics="true" pageId="3" pageNumber="1901">Fp</emphasis>
, female pore;
<emphasis id="94F2EAC5FFCB272BFF4EF9912498F973" box="[160,187,1662,1685]" italics="true" pageId="3" pageNumber="1901">Gp</emphasis>
, genital papillae;
<emphasis id="94F2EAC5FFCB272BFE9AF99125A8F973" box="[372,395,1662,1685]" italics="true" pageId="3" pageNumber="1901">Gs</emphasis>
, genital setal papilla;
<emphasis id="94F2EAC5FFCB272BFD9EF99126ACF973" box="[624,655,1662,1685]" italics="true" pageId="3" pageNumber="1901">Mp</emphasis>
, male pore;
<emphasis id="94F2EAC5FFCB272BFCFAF9912717F973" box="[788,820,1662,1685]" italics="true" pageId="3" pageNumber="1901">Prp</emphasis>
, prostate pores;
<emphasis id="94F2EAC5FFCB272BFC0DF99127D6F973" box="[995,1013,1662,1685]" italics="true" pageId="3" pageNumber="1901">St</emphasis>
, spermathecal pores; scale bar, 1 mm. (b) nephridial bladder from the postclitellar segments. (c) penial seta; scale bar, 0.05 mm. (d) genital seta; scale bar, 0.03 mm. (e) spermatheca.
</paragraph>
</caption>
<paragraph id="A63936D7FFCC272CFF4EFF6126B9FDE4" blockId="4.[160,1156,142,514]" pageId="4" pageNumber="1902">
<emphasis id="94F2EAC5FFCC272CFF4EFF612538FF41" bold="true" box="[160,283,142,167]" italics="true" pageId="4" pageNumber="1902">
<typeStatus id="793D8875FFCC272CFF4EFF612536FF41" box="[160,277,142,167]" pageId="4" pageNumber="1902" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<quantity id="617E9B32FFCC272CFEC2FF602587FF41" box="[300,420,143,167]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="4.6" metricValueMin="4.4" pageId="4" pageNumber="1902" unit="mm" value="45.0" valueMax="46.0" valueMin="44.0">4446 mm</quantity>
in length,
<quantity id="617E9B32FFCC272CFDCFFF602640FF41" box="[545,611,143,167]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="1902" unit="mm" value="2.0">2 mm</quantity>
in diameter, No. segments 92, 117, tail missing. Colour yellowish, pigmentation lacking. Prostomium tanylobous, the first dorsal pore occurs in the intersegmental furrow 17/18. Setae closely paired, setal formula after the clitellum aa:ab:bc:cd:dd = 8.9:1.1:6.4:1:28.6. Setae
<emphasis id="94F2EAC5FFCC272CFD35FF1926D4FEE9" box="[731,759,246,271]" italics="true" pageId="4" pageNumber="1902">ab</emphasis>
of segment 9 modified to genital setae. The clitellum extends over segments ½1317, clearly saddle-shaped, reaching ventrally the setal line
<emphasis id="94F2EAC5FFCC272CFE46FED42595FEB2" box="[424,438,315,340]" italics="true" pageId="4" pageNumber="1902">a</emphasis>
(
<figureCitation id="3EBD2A52FFCC272CFE26FED32610FEB2" box="[456,563,316,340]" captionStart="Figure 1" captionStartId="3.[160,225,1634,1657]" captionTargetBox="[205,1115,142,1603]" captionTargetId="figure-1@3.[205,1116,142,1603]" captionTargetPageId="3" captionText="Figure 1. Acanthodrilus barrieri sp. n. (a) ventral view of the clitellar region; Fp, female pore; Gp, genital papillae; Gs, genital setal papilla; Mp, male pore; Prp, prostate pores; St, spermathecal pores; scale bar, 1 mm. (b) nephridial bladder from the postclitellar segments. (c) penial seta; scale bar, 0.05 mm. (d) genital seta; scale bar, 0.03 mm. (e) spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5182724" httpUri="https://zenodo.org/record/5182724/files/figure.png" pageId="4" pageNumber="1902">Figure 1a</figureCitation>
). Prostatic pores paired on small papillae in 17, 19. Male pores externally clearly visible on segment 18, in the curly bracket-shaped seminal grooves. Female pores paired on 14, small slits anterior to setae
<emphasis id="94F2EAC5FFCC272CFC9DFE6E27B4FE7C" box="[883,919,385,410]" italics="true" pageId="4" pageNumber="1902">b-b</emphasis>
. Spermathecal pores large in furrows 7/8, 8/
<quantity id="617E9B32FFCC272CFE44FE4B25F4FE5A" box="[426,471,420,444]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="4" pageNumber="1902" unit="in" value="9.0">9 in</quantity>
setal line
<emphasis id="94F2EAC5FFCC272CFDBCFE4C2643FE5A" box="[594,608,419,444]" italics="true" pageId="4" pageNumber="1902">b</emphasis>
. Nephridiopores aligned in setal line
<emphasis id="94F2EAC5FFCC272CFBFAFE4C200EFE5A" box="[1044,1069,419,444]" italics="true" pageId="4" pageNumber="1902">cd</emphasis>
. Paired oval porophores in 9/10, 10/11 aligned with setae
<emphasis id="94F2EAC5FFCC272CFD11FE292738FE39" box="[767,795,454,479]" italics="true" pageId="4" pageNumber="1902">ab</emphasis>
, and unpaired ventral median porophores in segment 1719, 21/2222/23.
</paragraph>
<paragraph id="A63936D7FFCC272CFF4EFDD425A2FDB2" blockId="4.[160,1156,571,1081]" box="[160,385,571,596]" pageId="4" pageNumber="1902">
<emphasis id="94F2EAC5FFCC272CFF4EFDD425A2FDB2" bold="true" box="[160,385,571,596]" italics="true" pageId="4" pageNumber="1902">Internal characters</emphasis>
</paragraph>
<paragraph id="A63936D7FFCC272CFF4EFDB0202EFBDF" blockId="4.[160,1156,571,1081]" pageId="4" pageNumber="1902">
All septa membranous. Oesophageal gizzard large in 6. Calciferous glands lacking, the intestine commences from segment 23. Typhlosole lamellar, moderately sized, beginning around segment 30. Excretory system holoic, nephridial bladders sack-shaped (
<figureCitation id="3EBD2A52FFCC272CFBE3FD4B2054FD5A" box="[1037,1143,676,700]" captionStart="Figure 1" captionStartId="3.[160,225,1634,1657]" captionTargetBox="[205,1115,142,1603]" captionTargetId="figure-1@3.[205,1116,142,1603]" captionTargetPageId="3" captionText="Figure 1. Acanthodrilus barrieri sp. n. (a) ventral view of the clitellar region; Fp, female pore; Gp, genital papillae; Gs, genital setal papilla; Mp, male pore; Prp, prostate pores; St, spermathecal pores; scale bar, 1 mm. (b) nephridial bladder from the postclitellar segments. (c) penial seta; scale bar, 0.05 mm. (d) genital seta; scale bar, 0.03 mm. (e) spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5182724" httpUri="https://zenodo.org/record/5182724/files/figure.png" pageId="4" pageNumber="1902">Figure 1b</figureCitation>
). Paired hearts present in segments 1013. Testes are in 10, 11 free. Two pairs of seminal vesicles present in 11, 12. A pair of small ovaries in segment 13. Two pairs of prostates present in 17, 19. The glands are moderately long, reaching up to segment 22. Each prostate accompanied by a penial setal sac containing an adult and several replacement setae. The adult seta
<quantity id="617E9B32FFCC272CFEFBFCBE2549FC8F" box="[277,362,849,873]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="1902" unit="mm" value="1.5">1.5 mm</quantity>
in length and the middle
<quantity id="617E9B32FFCC272CFD7CFCBE26D7FC8F" box="[658,756,849,873]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="1902" unit="mm" value="0.05">0.05 mm</quantity>
wide. The seta forms almost a halfcircle, tip pointed, ornamentation consists of small serrated bars (
<figureCitation id="3EBD2A52FFCC272CFC8FFC9B27E6FC6A" box="[865,965,884,908]" captionStart="Figure 1" captionStartId="3.[160,225,1634,1657]" captionTargetBox="[205,1115,142,1603]" captionTargetId="figure-1@3.[205,1116,142,1603]" captionTargetPageId="3" captionText="Figure 1. Acanthodrilus barrieri sp. n. (a) ventral view of the clitellar region; Fp, female pore; Gp, genital papillae; Gs, genital setal papilla; Mp, male pore; Prp, prostate pores; St, spermathecal pores; scale bar, 1 mm. (b) nephridial bladder from the postclitellar segments. (c) penial seta; scale bar, 0.05 mm. (d) genital seta; scale bar, 0.03 mm. (e) spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5182724" httpUri="https://zenodo.org/record/5182724/files/figure.png" pageId="4" pageNumber="1902">Figure 1c</figureCitation>
). The genital seta is almost straight,
<quantity id="617E9B32FFCC272CFE95FC7825F7FC49" box="[379,468,919,943]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="1902" unit="mm" value="1.1">1.1 mm</quantity>
long and in the middle
<quantity id="617E9B32FFCC272CFCEBFC78275AFC49" box="[773,889,919,943]" metricMagnitude="-5" metricUnit="m" metricValue="3.7" pageId="4" pageNumber="1902" unit="mm" value="0.037">0.037 mm</quantity>
wide. Its tip rounded, ornamentation consists of small serrated scales (
<figureCitation id="3EBD2A52FFCC272CFD25FC56271BFC37" box="[715,824,953,977]" captionStart="Figure 1" captionStartId="3.[160,225,1634,1657]" captionTargetBox="[205,1115,142,1603]" captionTargetId="figure-1@3.[205,1116,142,1603]" captionTargetPageId="3" captionText="Figure 1. Acanthodrilus barrieri sp. n. (a) ventral view of the clitellar region; Fp, female pore; Gp, genital papillae; Gs, genital setal papilla; Mp, male pore; Prp, prostate pores; St, spermathecal pores; scale bar, 1 mm. (b) nephridial bladder from the postclitellar segments. (c) penial seta; scale bar, 0.05 mm. (d) genital seta; scale bar, 0.03 mm. (e) spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5182724" httpUri="https://zenodo.org/record/5182724/files/figure.png" pageId="4" pageNumber="1902">Figure 1d</figureCitation>
). Two pairs of spermathecae present in segments 8 and 9. Each consists of a narrow and duct and an equally long, oblong ampoule. The ampoule joins the duct almost in a right angle. The ental part of the duct forms a protruding elbow containing several small sperm chambers (
<figureCitation id="3EBD2A52FFCC272CFC77FBCE2023FBDF" box="[921,1024,1057,1081]" captionStart="Figure 1" captionStartId="3.[160,225,1634,1657]" captionTargetBox="[205,1115,142,1603]" captionTargetId="figure-1@3.[205,1116,142,1603]" captionTargetPageId="3" captionText="Figure 1. Acanthodrilus barrieri sp. n. (a) ventral view of the clitellar region; Fp, female pore; Gp, genital papillae; Gs, genital setal papilla; Mp, male pore; Prp, prostate pores; St, spermathecal pores; scale bar, 1 mm. (b) nephridial bladder from the postclitellar segments. (c) penial seta; scale bar, 0.05 mm. (d) genital seta; scale bar, 0.03 mm. (e) spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5182724" httpUri="https://zenodo.org/record/5182724/files/figure.png" pageId="4" pageNumber="1902">Figure 1e</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="EE9C655CFFCC272CFF4EFB9C2018FAF0" pageId="4" pageNumber="1902" type="discussion">
<paragraph id="A63936D7FFCC272CFF4EFB9C2525FB6A" blockId="4.[160,1155,1139,1302]" box="[160,262,1139,1164]" pageId="4" pageNumber="1902">
<emphasis id="94F2EAC5FFCC272CFF4EFB9C2525FB6A" bold="true" box="[160,262,1139,1164]" italics="true" pageId="4" pageNumber="1902">Remarks</emphasis>
</paragraph>
<paragraph id="A63936D7FFCC272CFF4EFB782018FAF0" blockId="4.[160,1155,1139,1302]" pageId="4" pageNumber="1902">
The new species with its medium size, lack of pigmentation and vesicles in 11, 12 seems to be most close to
<taxonomicName id="61864D54FFCC272CFE6DFB572690FB37" authority="Michaelsen, 1913" authorityName="Michaelsen" authorityYear="1913" box="[387,691,1208,1233]" class="Clitellata" family="Ocnerodrilidae" genus="Acanthodrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="1902" phylum="Annelida" rank="species" species="foanus">
<emphasis id="94F2EAC5FFCC272CFE6DFB5725C9FB37" box="[387,490,1208,1233]" italics="true" pageId="4" pageNumber="1902">A. foanus</emphasis>
<bibRefCitation id="C2174B26FFCC272CFE1FFB562690FB37" author="Michaelsen W" box="[497,691,1209,1233]" pageId="4" pageNumber="1902" pagination="173 - 280" refId="ref6164" refString="Michaelsen W. 1913. Die Oligochaten von Neu-Caledonien und den benachbarten Inselgruppen. Nova Caledonia Zool. 1 (3): 173 - 280." type="journal article" year="1913">Michaelsen, 1913</bibRefCitation>
</taxonomicName>
. However, it differs from the latter in the presence of genital setae, in the size and ornamentation of the penial setae (
<quantity id="617E9B32FFCC272CFBC4FB3320A0FB12" box="[1066,1155,1244,1268]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="1902" unit="mm" value="1.5">1.5 mm</quantity>
with pointed tip vs.
<quantity id="617E9B32FFCC272CFE66FB1125FDFAF0" box="[392,478,1278,1302]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="1902" unit="mm" value="4.5">4.5 mm</quantity>
with a blunt tip) and the form of the spermathecae.
</paragraph>
</subSubSection>
</treatment>
</document>