138 lines
12 KiB
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138 lines
12 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.531.6097" ID-GBIF-Dataset="70702787-a6a8-4af7-9702-738f344a5cbe" ID-PMC="PMC4668911" ID-Pensoft-Pub="1313-2970-531-1" ID-PubMed="26692803" ID-ZBK="C845838EC9124BD8AB4E07980F91959E" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-531-1" ModsDocOrigin="ZooKeys 531" ModsDocTitle="Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species" checkinTime="1451243834596" checkinUser="pensoft" docAuthor="Vermeulen, Jaap J., Liew, Thor-Seng & Schilthuizen, Menno" docDate="2015" docId="123C6BFE24C0F89EBA03373F6DEB70B4" docLanguage="en" docName="ZooKeys 531: 1-139" docOrigin="ZooKeys 531" docSource="http://dx.doi.org/10.3897/zookeys.531.6097" docTitle="Ditropopsis tyloacron Vermeulen, Liew & Schilthuizen, sp. n." docType="treatment" docUuid="4BF26EBE-935F-4FF1-9BAD-0F62543E18AF" docUuidSource="ZooBank" docVersion="10" lastPageNumber="15" masterDocId="1527DF5A6645C540FFF1FF85AF4AE358" masterDocTitle="Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species" masterLastPageNumber="139" masterPageNumber="1" pageNumber="13" updateTime="1668160884494" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liew, Thor-Seng</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schilthuizen, Menno</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>531</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>139</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.531.6097</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.531.6097</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-531-1</mods:identifier>
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<mods:identifier type="ZBK">C845838EC9124BD8AB4E07980F91959E</mods:identifier>
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<mods:identifier type="ZooBank">C845838EC9124BD8AB4E07980F91959E</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="144057495" LSID="urn:lsid:zoobank.org:act:4BF26EBE-935F-4FF1-9BAD-0F62543E18AF" httpUri="http://treatment.plazi.org/id/123C6BFE24C0F89EBA03373F6DEB70B4" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
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<subSubSection pageId="12" pageNumber="13" type="multiple">
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<paragraph pageId="12" pageNumber="13">Taxon classification Animalia Architaenioglossa Cyclophoridae</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="13" type="nomenclature">
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<paragraph pageId="12" pageNumber="13">
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<taxonomicName LSID="http://zoobank.org/4BF26EBE-935F-4FF1-9BAD-0F62543E18AF" authority="Vermeulen, Liew & Schilthuizen" class="Gastropoda" family="Cyclophoridae" genus="Ditropopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ditropopsis tyloacron" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="tyloacron">Ditropopsis tyloacron Vermeulen, Liew & Schilthuizen</taxonomicName>
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<taxonomicNameLabel pageId="12" pageNumber="13">sp. n.</taxonomicNameLabel>
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Figure 12
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="reference_group">
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<paragraph pageId="13" pageNumber="14">
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<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Ditropopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ditropopsis" order="Architaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
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<pageBreakToken pageId="13" pageNumber="14" start="start">Ditropopsis</pageBreakToken>
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</taxonomicName>
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sp.
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<normalizedToken originalValue="“BO-02”" pageId="13" pageNumber="14">"BO-02"</normalizedToken>
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,
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<bibRefCitation author="Schilthuizen, M" journalOrPublisher="Journal of Molluscan Studies" pageId="13" pageNumber="14" pagination="255 - 258" title="Microsnails at microscales in Borneo: distributions of Prosobranchia versus Pulmonata." url="10.1093/mollus/68.3.255" volume="68" year="2002">Schilthuizen et al. 2002</bibRefCitation>
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: 257-258.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="holotype">
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<paragraph pageId="13" pageNumber="14">
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<materialsCitation collectingDate="2015-01-01" collectingDateMax="2015-12-31" collectingDateMin="2015-01-01" collectionCode="RMNH" country="Malaysia" httpUri="http://data.biodiversitydata.nl/naturalis/specimen/RMNH.5003920" location="Tawau Province" municipality="Sabah" specimenCode="RMNH 5003920" specimenCount="1" typeStatus="holotype">
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<typeStatus>Holotype</typeStatus>
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.
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<collectingCountry>Malaysia</collectingCountry>
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,
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<collectingMunicipality>Sabah</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:123C6BFE24C0F89EBA03373F6DEB70B4:74EAB6AD4F0375B12F4815F8B0DD8D3B" country="Malaysia" municipality="Sabah" name="Tawau Province">Tawau Province</location>
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,
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<location LSID="urn:lsid:plazi:treatment:123C6BFE24C0F89EBA03373F6DEB70B4:F89F642C064B137C51C333A278AA5BCA" country="Malaysia" municipality="Sabah" name="Danum Valley Conservation Area">Danum Valley Conservation Area</location>
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(
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<specimenCode>RMNH5003920</specimenCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="materials_examined">
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<paragraph pageId="13" pageNumber="14">Examined material from Sabah.</paragraph>
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<paragraph pageId="13" pageNumber="14">Tawau Province. Danum Valley Conservation Area (leg. UMS students, V 9880, BOR/MOL 313).</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="description">
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<paragraph pageId="13" pageNumber="14">Description.</paragraph>
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<paragraph pageId="13" pageNumber="14">Shell very small, rather thick, somewhat translucent, white to pale (yellowish) green. Surface shiny or glossy. Spire almost flat, but apex protruding, slightly oblique. Sculpture. Radial sculpture: fine growth lines, locally grading into fine, densely placed riblets. Spiral sculpture on the last whorl: 6 very distinct cords: 1 supraperipheral and 1 peripheral, the latter widest and widely projecting, 1 basal and 3 umbilical; next to these a fine spiral striation on the lower surface. Sculpture continuing up to the peristome, but close to the peristome distorted and partly obliterated by densely placed, low radial riblets and increasingly coarse spiral threads. Aperture. Peristome simple, not expanded, distinctly constricted in fully adult shells, parietal side attached to the penultimate whorl in fully adult shells, basal side angular, slightly to distinctly drawn out. Dimensions. Height 1.4-1.65 mm; width 2.5-2.8 mm; h/w 0.52-0.56; umbilicus measured over the basal spiral cord 33-37 % of the shell width; number of whorls c. 4; height aperture 0.7-0.9 mm; width aperture 0.8-0.9 mm.</paragraph>
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<caption pageId="13" pageNumber="14">
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<paragraph pageId="13" pageNumber="14">
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Figure 12-13. 12
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<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Ditropopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ditropopsis tyloacron" order="Architaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="tyloacron">Ditropopsis tyloacron</taxonomicName>
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sp. n. 12A Frontal view, shell 1.6 mm high 12B Frontal view, shell 1.5 mm high 12C Umbilical view (12A, 12C Malaysia, Sabah, Tawau Province, Danum Valley Conservation Area, RMNH.5003920, holotype; 12B Indonesia, Kalimantan Selatan, Meratus Mountains, c. 18 Km East of Barabai, Nateh near Batu Tangga, V 3003) 13
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<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Ditropopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ditropopsis constricta" order="Architaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="constricta">Ditropopsis constricta</taxonomicName>
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sp. n. 13A Frontal view, shell 1.5 mm high 13B Umbilical view (Malaysia, Sabah, Interior Province, Sepulut valley, Gua Sanaron, RMNH.5003919, holotype).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="habitat in sabah and distribution">
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<paragraph pageId="13" pageNumber="14">Habitat in Sabah and distribution.</paragraph>
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<paragraph pageId="13" pageNumber="14">Primary forest on limestone soil, rarely on sandstone, 0-600 m alt. Sabah: S.E. part. Also in Kalimantan, East part. Endemic to Borneo.</paragraph>
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</subSubSection>
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<subSubSection pageId="14" pageNumber="15" type="cross diagnosis">
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<paragraph pageId="14" pageNumber="15">
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<pageBreakToken pageId="14" pageNumber="15" start="start">Cross</pageBreakToken>
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diagnosis.
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</paragraph>
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<paragraph pageId="14" pageNumber="15">
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Well characterized among Sabah
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<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Ditropopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ditropopsis" order="Architaenioglossa" pageId="14" pageNumber="15" phylum="Mollusca" rank="genus">Ditropopsis</taxonomicName>
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by the low, almost flat spire with only the apex protruding. In Kalimantan the Sabah form (illustrated) locally grades into series with relatively smooth, large shells and with the supraperipheral and umbilical ridges partly or entirely missing. Such shells look a little like
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<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Ditropopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ditropopsis imadatei" order="Architaenioglossa" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="imadatei">Ditropopsis imadatei</taxonomicName>
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and
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<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Ditropopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ditropopsis cincta" order="Architaenioglossa" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="cincta">Ditropopsis cincta</taxonomicName>
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, but have a flatter spire and a wider umbilicus.
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</paragraph>
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<paragraph pageId="14" pageNumber="15">
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Elsewhere,
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<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Ditropopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ditropopsis moellendorfii" order="Architaenioglossa" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="moellendorfii">Ditropopsis moellendorfii</taxonomicName>
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Boettger, 1891 and
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<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Ditropopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ditropopsis papuana" order="Architaenioglossa" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="papuana">Ditropopsis papuana</taxonomicName>
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E.A. Smith, 1897 have similarly flat shells, but the first has a double peristome, and the second a more rounded spire.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="14" pageNumber="15" type="etymology">
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<paragraph pageId="14" pageNumber="15">Etymology.</paragraph>
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<paragraph pageId="14" pageNumber="15">The name refers to the shape of the apex [tulos (Gr.) = knob; akros (Gr.) = at the top; a knobhead, therefore].</paragraph>
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</subSubSection>
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</treatment>
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</document> |