105 lines
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105 lines
11 KiB
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<document id="45E7FF136D2433CA78A05350E9B5B0CB" ID-DOI="http://dx.doi.org/10.3897/zookeys.433.8022" ID-GBIF-Dataset="cc89bbf1-736b-4259-b56a-1d94f103f3f0" ID-PMC="PMC4141180" ID-Pensoft-Pub="1313-2970-433-77" ID-PubMed="25152674" ID-ZooBank="065CFC3A4DD84759B55D040FCC3351AA" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-433-77" ModsDocOrigin="ZooKeys 433" ModsDocTitle="A review of the primary types of the Hawaiian stag beetle genus Apterocyclus Waterhouse (Coleoptera, Lucanidae, Lucaninae), with the description of a new species" checkinTime="1451245511645" checkinUser="pensoft" docAuthor="Paulsen, M. J. & Hawks, David C." docDate="2014" docId="0569136ACE567ABFF71AB6AC9B486FFF" docLanguage="en" docName="ZooKeys 433: 77-88" docOrigin="ZooKeys 433" docSource="http://dx.doi.org/10.3897/zookeys.433.8022" docTitle="Apterocyclus waterhousei Sharp 1908" docType="treatment" docVersion="4" lastPageNumber="84" masterDocId="CD526101B053FF86711AFF83BC67FFAB" masterDocTitle="A review of the primary types of the Hawaiian stag beetle genus Apterocyclus Waterhouse (Coleoptera, Lucanidae, Lucaninae), with the description of a new species" masterLastPageNumber="88" masterPageNumber="77" pageNumber="83" updateTime="1701315137992" updateUser="plazi">
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<mods:title id="7BFD37035A7FE31A4A18DF5AB288A7AF">A review of the primary types of the Hawaiian stag beetle genus Apterocyclus Waterhouse (Coleoptera, Lucanidae, Lucaninae), with the description of a new species</mods:title>
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<mods:namePart id="26D554B8BD9214B159F7C2DF63C26925">Paulsen, M. J.</mods:namePart>
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<mods:name id="CC83A532672B142569FA99C6C7DB345A" type="personal">
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<mods:roleTerm id="C31C1D30765475D2F6C336D91C070939">Author</mods:roleTerm>
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<mods:namePart id="1F1105DB8FB12473477CDE6C1B4112EC">Hawks, David C.</mods:namePart>
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<mods:typeOfResource id="A85A160561ED63464B4118163D1A68BA">text</mods:typeOfResource>
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<mods:title id="9BB7C9AEEF889977B309D6B47D70DBB3">ZooKeys</mods:title>
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<mods:part id="776B843A9EE1C388A397DDBB4E6C0565">
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<mods:date id="8A34C1824495CF092BECFF1B4C832B12">2014</mods:date>
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<mods:detail id="226AA34A7049F628BC829FE65A6FC74A" type="volume">
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<mods:number id="B12C74F51550A0A8B8B2279559B47FBB">433</mods:number>
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<mods:start id="4E8FD196C88262484762644C5A427E83">77</mods:start>
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<mods:end id="582F1A9FC0C37E867C58CC4585C7A15C">88</mods:end>
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<mods:url id="C17F04311817093487A514A2F57EC483">http://dx.doi.org/10.3897/zookeys.433.8022</mods:url>
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<mods:classification id="38F1E57CBEE4C23E73994A9CB405E2CE">journal article</mods:classification>
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<mods:identifier id="C3C20A47D0FB53D976897F8D6DEA663A" type="DOI">http://dx.doi.org/10.3897/zookeys.433.8022</mods:identifier>
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<mods:identifier id="9FB215B885E3E409E0872384242031CC" type="Pensoft-Pub">1313-2970-433-77</mods:identifier>
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<treatment id="0569136ACE567ABFF71AB6AC9B486FFF" ID-GBIF-Taxon="152054932" LSID="urn:lsid:plazi:treatment:0569136ACE567ABFF71AB6AC9B486FFF" httpUri="http://treatment.plazi.org/id/0569136ACE567ABFF71AB6AC9B486FFF" lastPageId="7" lastPageNumber="84" pageId="6" pageNumber="83">
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<subSubSection id="402B232E26CB4BE9A6C042367331B5AA" pageId="6" pageNumber="83" type="multiple">
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<paragraph id="565C1BD1E5F07AF36A3D2387573BE656" pageId="6" pageNumber="83">Taxon classification Animalia Coleoptera Lucanidae</paragraph>
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</subSubSection>
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<subSubSection id="67DC744F7D3DFD1F9B5E0B91B7639829" pageId="6" pageNumber="83" type="nomenclature">
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<paragraph id="F7007459A15880656DBD9D0C60A05506" pageId="6" pageNumber="83">
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<taxonomicName id="A0DB9318BFEB5E767A3BCA9D0F9A46CE" ID-CoL="FYKB" authority="Sharp, 1908" authorityName="Sharp" authorityYear="1908" class="Insecta" family="Lucanidae" genus="Apterocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apterocyclus waterhousei" order="Coleoptera" pageId="6" pageNumber="83" phylum="Arthropoda" rank="species" species="waterhousei">Apterocyclus waterhousei Sharp, 1908</taxonomicName>
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Figs 8, 16, 21
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</paragraph>
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</subSubSection>
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<subSubSection id="25279D6FC97933245DB5FE05F50B53F5" pageId="6" pageNumber="83" type="reference_group">
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<paragraph id="2E142173B60A4AA3CC2D6AE896F317E5" pageId="6" pageNumber="83">
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<taxonomicName id="50AB725B8CC8A5D561A5B8949768C51A" class="Insecta" family="Lucanidae" genus="Apterocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apterocyclus waterhousei" order="Coleoptera" pageId="6" pageNumber="83" phylum="Arthropoda" rank="species" species="waterhousei">Apterocyclus waterhousei</taxonomicName>
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Sharp in
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<bibRefCitation id="0AE30706DE388364700A09C63008B533" author="Sharp, D" journalOrPublisher="Fauna Hawaiiensis or the zoology of the Sandwich (Hawaiian) Isles: being results of the explorations instituted by the Joint Committee appointed by British Association for the Advancement of Science and carried on with the assistance of those bodies and of the Trustees of the Bernice Pauahi Bishop Museum at Honolulu" pageId="10" pageNumber="87" pagination="367 - 579" volume="3" year="1908">Sharp and Scott 1908</bibRefCitation>
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: 403. Type material: Holotype male (NHM) labeled: a) handwritten, on cork "Apterocyclus waterhousei / Type D.S./ Kauai, Perkins"; b) "Sandwich Is. / 1912
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<normalizedToken id="03222F74BEF5C3C2787AFD65E1D27701" originalValue="–215”">-215"</normalizedToken>
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; c) red-bordered circular label
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<normalizedToken id="13387CE28E3EF161DFE393A92AF39F93" originalValue="“Type”">"Type"</normalizedToken>
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; d) red paper, "Apterocyclus / waterhousei m /Sharp, 1908 / HOLOTYPE / des. M.J. Paulsen"; e) "Apterocyclus / waterhousei / Sharp, 1908 / det. M.J. Paulsen 2012".
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</paragraph>
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</subSubSection>
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<subSubSection id="842E63510070122019A112E157EE3031" pageId="6" pageNumber="83" type="diagnosis">
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<paragraph id="6BFF8780AF92FFDCBDC423215C0827C3" pageId="6" pageNumber="83">Diagnosis.</paragraph>
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<paragraph id="C1FE4271FF38C46451D4171872E09D23" pageId="6" pageNumber="83">
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The large, median tooth present on the meso- and metatibiae mentioned by Sharp (
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<bibRefCitation id="5CFBC3ACC1D5D5891B5F318A10D37E8C" author="Sharp, D" journalOrPublisher="Fauna Hawaiiensis or the zoology of the Sandwich (Hawaiian) Isles: being results of the explorations instituted by the Joint Committee appointed by British Association for the Advancement of Science and carried on with the assistance of those bodies and of the Trustees of the Bernice Pauahi Bishop Museum at Honolulu" pageId="10" pageNumber="87" pagination="367 - 579" volume="3" year="1908">1908</bibRefCitation>
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) for the holotype is present on all other specimens of both sexes, and not found in any other species. These large teeth, in addition to the broad, serrate protibiae and almost obsolete canthus separate this species from
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<taxonomicName id="45870A47CB0A313E807E179201C60940" class="Insecta" family="Lucanidae" genus="Apterocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apterocyclus honoluluensis" order="Coleoptera" pageId="6" pageNumber="83" phylum="Arthropoda" rank="species" species="honoluluensis">Apterocyclus honoluluensis</taxonomicName>
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(a few
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<taxonomicName id="394F0D93B1F2EAC9E35D6D298D44FD28" class="Insecta" family="Lucanidae" genus="Apterocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apterocyclus honoluluensis" order="Coleoptera" pageId="6" pageNumber="83" phylum="Arthropoda" rank="species" species="honoluluensis">Apterocyclus honoluluensis</taxonomicName>
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have a weakly indicated tooth on the mesotibiae). The species is most similar to
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<taxonomicName id="EC1FBCAAF85BA80CBE05CEEF2A192EFF" class="Insecta" family="Lucanidae" genus="Apterocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apterocyclus munroi" order="Coleoptera" pageId="6" pageNumber="83" phylum="Arthropoda" rank="species" species="munroi">Apterocyclus munroi</taxonomicName>
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, but lacks distinct setae on the elytra and the anteriorly divergent clypeus. Specimens range from 18 to 22 mm in length.
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</paragraph>
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</subSubSection>
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<subSubSection id="DBA131CEBBFD3BD8734A265C865A4616" lastPageId="7" lastPageNumber="84" pageId="6" pageNumber="83" type="distribution">
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<paragraph id="CACEB6935699CA6389FD9531E8E9B2B2" pageId="6" pageNumber="83">Distribution.</paragraph>
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<paragraph id="3A1075EA921724C34744754F316329EF" lastPageId="7" lastPageNumber="84" pageId="6" pageNumber="83">
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Specimens have been examined from Kohua/Mohiki [Mohihi?] Stream, Kaholuamano, and near Waialae River (BPBM).
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<bibRefCitation id="711CF8F75590488ADD3129D4FB8A3447" author="Mizunuma, T" journalOrPublisher="Toshitsugu Endo, Tokyo" pageId="10" pageNumber="87" title="Endless Collection" year="2000">Mizunuma (2000)</bibRefCitation>
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illustrated
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<pageBreakToken id="EF7ECBBDF0543068A286BE95AB239BA1" pageId="7" pageNumber="84" start="start">specimens</pageBreakToken>
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from
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<normalizedToken id="2EAC7B1EB2C97E8A0BAA0AA98CFC0D0E" originalValue="Po’omau">Po'omau</normalizedToken>
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Canyon, and
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<bibRefCitation id="58D8E6A167D962F308B092E9653EC5FC" author="Fujita, H" journalOrPublisher="Mushi-Sha, Tokyo" pageId="10" pageNumber="87" title="The Lucanid Beetles of the World." year="2010">Fujita (2010)</bibRefCitation>
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included specimens from Kohua Ridge and
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<normalizedToken id="BA500C44D8246A2E3C04ACEF1F2EBA30" originalValue="Po’omau">Po'omau</normalizedToken>
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Canyon.
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</paragraph>
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</subSubSection>
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<subSubSection id="31D8B9BF6DAAEC6E44F48CC53C0249AA" pageId="7" pageNumber="84" type="remarks">
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<paragraph id="02E0F70FE240B0B8E1D3913111DE481D" pageId="7" pageNumber="84">Remarks.</paragraph>
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<paragraph id="B561786609D8423EAD9F3F10F9355C50" pageId="7" pageNumber="84">
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Described from a single male, Sharp (
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<bibRefCitation id="40EFDE46D832AB10BB492936158229BA" author="Sharp, D" journalOrPublisher="Fauna Hawaiiensis or the zoology of the Sandwich (Hawaiian) Isles: being results of the explorations instituted by the Joint Committee appointed by British Association for the Advancement of Science and carried on with the assistance of those bodies and of the Trustees of the Bernice Pauahi Bishop Museum at Honolulu" pageId="10" pageNumber="87" pagination="367 - 579" volume="3" year="1908">1908</bibRefCitation>
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) noted the thick legs,
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<normalizedToken id="64472ADDD888E3B972765A2B97621A6F" originalValue="‘crenulate’">'crenulate'</normalizedToken>
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protibiae, and pointed elytral apices (Fig. 8). No other specimens have been seen that exhibit the pointed elytral apices of the holotype, and we consider the character to be possibly aberrant and not of specific-level importance.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |