treatments-xml/data/05/59/87/055987E28B237360FF7DD5DDFD1F889A.xml
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<document id="AC21C1212B4938D16D64BC5A68401F88" ID-DOI="10.11646/zootaxa.5375.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10184224" ID-ZooBank="F3D0E008-556C-4FAD-BF51-4F1A714325DA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700642529072" checkinUser="plazi" docAuthor="Sokolov, Igor M." docDate="2023" docId="055987E28B237360FF7DD5DDFD1F889A" docLanguage="en" docName="zootaxa.5375.2.1.pdf" docOrigin="Zootaxa 5375 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5375.2.1/52321" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nesamblyops hobbit Sokolov 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="169" masterDocId="F960FF9A8B327372FFEAD113FFB08B1B" masterDocTitle="Fourteen new species of the genus Nesamblyops Jeannel (Coleoptera: Carabidae: Anillini) from the South Island of New Zealand with redescription of the genus and description of a new subtribe" masterLastPageNumber="192" masterPageNumber="151" pageNumber="168" updateTime="1700643183691" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="02FD27670729DC9AD2C2763867D384A2">Fourteen new species of the genus Nesamblyops Jeannel (Coleoptera: Carabidae: Anillini) from the South Island of New Zealand with redescription of the genus and description of a new subtribe</mods:title>
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<mods:namePart id="65933E04C7F14B525DF63DC75D411710">Sokolov, Igor M.</mods:namePart>
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<mods:title id="89360682946F15AE54C8B3FB4F3E09F0">Zootaxa</mods:title>
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<mods:date id="F134E5A847994E3DF90A3AC87FF69503">2023</mods:date>
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<mods:number id="053F45CBFC5D374DC1564FE304DC7332">2023-11-22</mods:number>
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<mods:identifier id="9BD0F0813340414FF336C4E7CC3514ED" type="DOI">10.11646/zootaxa.5375.2.1</mods:identifier>
<mods:identifier id="D608ADF2F41A6A9AD1343BE33B2EA432" type="ISSN">1175-5326</mods:identifier>
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<treatment id="055987E28B237360FF7DD5DDFD1F889A" LSID="urn:lsid:plazi:treatment:055987E28B237360FF7DD5DDFD1F889A" httpUri="http://treatment.plazi.org/id/055987E28B237360FF7DD5DDFD1F889A" lastPageId="18" lastPageNumber="169" pageId="17" pageNumber="168">
<subSubSection id="C5EA657F8B237363FF7DD5DDFE478FF3" box="[151,503,1230,1257]" pageId="17" pageNumber="168" type="nomenclature">
<paragraph id="8D4F36F48B237363FF7DD5DDFE478FF3" blockId="17.[151,531,1230,1293]" box="[151,503,1230,1257]" pageId="17" pageNumber="168">
<heading id="D60781988B237363FF7DD5DDFE478FF3" bold="true" box="[151,503,1230,1257]" fontSize="11" level="1" pageId="17" pageNumber="168" reason="1">
<emphasis id="BF84EAE68B237363FF7DD5DDFE478FF3" bold="true" box="[151,503,1230,1257]" pageId="17" pageNumber="168">
<taxonomicName id="4AF04D778B237363FF7DD5DDFE3B8FF2" authority="Sokolov, 2023" authorityName="Sokolov" authorityYear="2023" box="[151,395,1230,1257]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="hobbit" status="sp. nov.">
<emphasis id="BF84EAE68B237363FF7DD5DDFE3B8FF2" bold="true" box="[151,395,1230,1257]" italics="true" pageId="17" pageNumber="168">Nesamblyops hobbit</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A4B7579D8B237363FE73D5DDFE478FF3" box="[409,503,1230,1256]" pageId="17" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C5EA657F8B237363FF7DD5E1FDA38E17" box="[151,531,1266,1293]" pageId="17" pageNumber="168" type="description">
<paragraph id="8D4F36F48B237363FF7DD5E1FDA38E17" blockId="17.[151,531,1230,1293]" box="[151,531,1266,1293]" pageId="17" pageNumber="168">
<figureCitation id="15CB2A718B237363FF7DD5E1FEAC8E17" box="[151,284,1266,1292]" captionStart="FIGURE 9" captionStartId="15.[151,250,1649,1673]" captionTargetBox="[155,1436,181,1621]" captionTargetId="figure-15@15.[151,1436,181,1625]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect.A—N. canaanensis (Canaan, Nelson, SO), B—N. disjunctus (Fletchers Creek, Buller, SO), C—N. hobbit (9mi. N Karamea, Nelson, SO), D—N. karamea (Karamea River Gorge, Nelson, SO), E—N. kuscheli (Flora Hut, MountArthur, Nelson, SO), F—N. magnificus (Allison Reserve,Akatore, Dunedin, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184245" httpUri="https://zenodo.org/record/10184245/files/figure.png" pageId="17" pageNumber="168">Figures. 9C</figureCitation>
,
<figureCitation id="15CB2A718B237363FEC2D5E1FEE98E17" box="[296,345,1266,1292]" captionStart="FIGURE 12" captionStartId="26.[151,250,1215,1239]" captionTargetBox="[157,1430,181,1192]" captionTargetId="figure-32@26.[157,1430,181,1192]" captionTargetPageId="26" captionText="FIGURE 12. Digital images of pronota of New Zealand Nesamblyops species, dorsal aspect. A—N. canaanensis (Canaan, Nelson, SO), B—N. disjunctus (Fletchers Creek, Buller, SO), C—N. hobbit (9mi. N Karamea, Nelson, SO), D—N. karamea (Karamea River Gorge, Nelson, SO), E—N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO), F—N. magnificus (Allison Reserve, Akatore, Dunedin, SO), G—N. montanus (Lake Sylvester, Kahurangi National Park, Nelson, SO), H—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), I—N. ovipennis (Mount Arthur, Nelson, SO). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10184251" httpUri="https://zenodo.org/record/10184251/files/figure.png" pageId="17" pageNumber="168">12C</figureCitation>
,
<figureCitation id="15CB2A718B237363FE8FD5E1FE1F8E16" box="[357,431,1266,1293]" captionStart="FIGURE 14" captionStartId="30.[151,250,1365,1389]" captionTargetBox="[157,1433,187,1336]" captionTargetId="figure-32@30.[151,1436,181,1341]" captionTargetPageId="30" captionText="FIGURE 14. Line drawings of male genitalia of New Zealand Nesamblyops species. N. canaanensis (Canaan, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. disjunctus (Fletchers Creek, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. hobbit (Mount Domett, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. karamea (Karamea River Gorge, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. magnificus (Allison Reserve, Akatore, Dunedin, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Legend: rCs—reversed C-contour; Vs—V-contour. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184255" httpUri="https://zenodo.org/record/10184255/files/figure.png" pageId="17" pageNumber="168">14GI</figureCitation>
,
<figureCitation id="15CB2A718B237363FE50D5E1FE5C8E17" box="[442,492,1266,1292]" captionStart="FIGURE 17" captionStartId="34.[151,250,1191,1215]" captionTargetBox="[163,1421,187,1168]" captionTargetId="figure-32@34.[151,1436,181,1168]" captionTargetPageId="34" captionText="FIGURE 17. Line drawings of ring sclerite of New Zealand Nesamblyops species, male genitalia, dorsal aspect. A—N. canaanensis (Canaan, Nelson, SO), B—N. disjunctus (Fletchers Creek, Buller, SO), C—N. hobbit (Mount Domett, Nelson, SO), D—N. karamea (Karamea River Gorge, Nelson, SO), E—N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO), F—N. montanus (Relax Shelter, Mount Robert Ridge, Nelson, SO), G—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), H—N. ovipennis (Mount Arthur, Nelson, SO), I—N. rotundicollis (Onekaka, Nelson, SO), J—N. subcaecus (Margarets Tarn Lake, Arthurs Pass, Mid Canterbury, SO), K—N. viator (Secretary Island, Grono Bay, Fjordland, SO), L—N. victoriae (Capleston, Victoria Range, Buller, SO). Scale = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10184261" httpUri="https://zenodo.org/record/10184261/files/figure.png" pageId="17" pageNumber="168">17C</figureCitation>
,
<figureCitation id="15CB2A718B237363FE1DD5E1FDA38E17" box="[503,531,1266,1292]" captionStart="FIGURE 19" captionStartId="36.[151,250,1381,1405]" captionTargetBox="[151,1436,181,1357]" captionTargetId="figure-32@36.[151,1436,181,1357]" captionTargetPageId="36" captionText="FIGURE 19. Distributional records for the Nesamblyops species and images of male median lobe, obtained from specimens, collected in the line-connected with images localities. Green circles—N. canaanensis; black circles—N. hobbit; blue circles— N. ovipennis; yellow circles—N. karamea; red circles—N. kuscheli. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10184267" httpUri="https://zenodo.org/record/10184267/files/figure.png" pageId="17" pageNumber="168">19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B237363FF7DD429FB4D8D37" pageId="17" pageNumber="168" type="materials_examined">
<paragraph id="8D4F36F48B237363FF7DD429FB4D8D37" blockId="17.[151,1436,1338,2013]" pageId="17" pageNumber="168">
<emphasis id="BF84EAE68B237363FF7DD429FEF58E4F" bold="true" box="[151,325,1338,1364]" pageId="17" pageNumber="168">Type material.</emphasis>
<materialsCitation id="3D983CA98B237363FEA6D429FE218E62" collectionCode="NZAC, DSIR" collectorName="Dec &amp; G. W. Ramsay &amp; Litter &amp; No" country="New Zealand" elevation="1000" location="Mt. Domett" pageId="17" pageNumber="168" specimenCount="1" typeStatus="holotype">
<emphasis id="BF84EAE68B237363FEA6D429FE518E4F" bold="true" box="[332,481,1338,1364]" italics="true" pageId="17" pageNumber="168">
<typeStatus id="524B88568B237363FEA6D429FE518E4F" box="[332,481,1338,1364]" pageId="17" pageNumber="168" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
, male, in
<collectionCode id="EBE1AE318B237363FDBBD429FD108E4F" box="[593,672,1338,1364]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="17" pageNumber="168" type="Other institutional type">NZAC</collectionCode>
, labeled: \
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<quantity id="4A089B118B237363FC77D429FC5A8E4F" box="[925,1002,1338,1364]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="168" unit="m" value="1000.0">
<elevation id="06DDD1C78B237363FC77D429FC5A8E4F" box="[925,1002,1338,1364]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="168" unit="m" value="1000.0">1000m</elevation>
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1
<collectorName id="200553228B237363FBEDD428FB848E4E" box="[1031,1076,1339,1365]" pageId="17" pageNumber="168">Dec</collectorName>
71
<collectorName id="200553228B237363FBB5D429FB438E4E" box="[1119,1267,1338,1365]" pageId="17" pageNumber="168">G.W.Ramsay</collectorName>
\
<collectorName id="200553228B237363FAE3D429FAF78E4F" box="[1289,1351,1338,1364]" pageId="17" pageNumber="168">Litter</collectorName>
71/176 \
<collectionCode id="EBE1AE318B237363FF42D44DFF588E63" box="[168,232,1374,1400]" country="New Zealand" httpUri="http://grbio.org/cool/6qmb-xjj9" name="Department of Scientific and Industrial Research" pageId="17" pageNumber="168">DSIR</collectionCode>
\
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\
<collectingCountry id="F5E776648B237363FEAAD44CFED58E62" box="[320,357,1375,1401]" name="New Zealand" pageId="17" pageNumber="168">NZ</collectingCountry>
PB
</materialsCitation>
\.
<emphasis id="BF84EAE68B237363FE58D44DFDFD8E63" bold="true" box="[434,589,1374,1400]" italics="true" pageId="17" pageNumber="168">
<typeStatus id="524B88568B237363FE58D44DFDFD8E63" box="[434,589,1374,1400]" pageId="17" pageNumber="168" type="paratype">PARATYPES</typeStatus>
</emphasis>
(
<specimenCount id="9BF6FD7D8B237363FDB5D44DFD4E8E63" box="[607,766,1374,1400]" count="30" pageId="17" pageNumber="168" type="generic">30 specimens</specimenCount>
, dissected 5 exx.),
<specimenCount id="9BF6FD7D8B237363FC36D44DFB848E63" box="[988,1076,1374,1400]" count="6" pageId="17" pageNumber="168" type="male">6 males</specimenCount>
and
<specimenCount id="9BF6FD7D8B237363FB9AD44DFB6E8E63" box="[1136,1246,1374,1400]" count="6" pageId="17" pageNumber="168" type="female">6 females</specimenCount>
labeled same as
<typeStatus id="524B88568B237363FF7DD491FF4D8E87" box="[151,253,1410,1436]" pageId="17" pageNumber="168" type="holotype">holotype</typeStatus>
;
<materialsCitation id="3D983CA98B237363FEE3D491FEAF8EDA" collectionCode="DSIR" collectorName="Dec &amp; J. S. Dugdale" country="New Zealand" elevation="1463" location="Mt. Domett" municipality="No" pageId="17" pageNumber="168" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9BF6FD7D8B237363FEE3D491FEE68E87" box="[265,342,1410,1436]" count="1" pageId="17" pageNumber="168" type="male">1 male</specimenCount>
and
<specimenCount id="9BF6FD7D8B237363FE65D491FE4D8E87" box="[399,509,1410,1436]" count="3" pageId="17" pageNumber="168" type="female">3 females</specimenCount>
labeled: \
<location id="882F602F8B237363FD9CD490FD428E86" LSID="urn:lsid:plazi:treatment:055987E28B237360FF7DD5DDFD1F889A:882F602F8B237363FD9CD490FD428E86" box="[630,754,1411,1437]" country="New Zealand" municipality="No" name="Mt. Domett" pageId="17" pageNumber="168">Mt.Domett</location>
<quantity id="4A089B118B237363FD10D491FCF88E87" box="[762,840,1410,1436]" metricMagnitude="3" metricUnit="m" metricValue="1.463" pageId="17" pageNumber="168" unit="m" value="1463.0">
<elevation id="06DDD1C78B237363FD10D491FCF88E87" box="[762,840,1410,1436]" metricMagnitude="3" metricUnit="m" metricValue="1.463" pageId="17" pageNumber="168" unit="m" value="1463.0">1463m</elevation>
</quantity>
1
<collectorName id="200553228B237363FC8DD490FC248E86" box="[871,916,1411,1437]" pageId="17" pageNumber="168">Dec</collectorName>
71
<collectorName id="200553228B237363FC2AD491FBE08E87" box="[960,1104,1410,1436]" pageId="17" pageNumber="168">J.S. Dugdale</collectorName>
\ mat plants 71/181 \
<collectionCode id="EBE1AE318B237363FAA6D491FA3C8E87" box="[1356,1420,1410,1436]" country="New Zealand" httpUri="http://grbio.org/cool/6qmb-xjj9" name="Department of Scientific and Industrial Research" pageId="17" pageNumber="168">DSIR</collectionCode>
\
<collectingMunicipality id="6D2BAC8E8B237363FF7DD4B4FF098EDA" box="[151,185,1447,1473]" pageId="17" pageNumber="168">No</collectingMunicipality>
\
<collectingCountry id="F5E776648B237363FF3AD4B4FF458EDA" box="[208,245,1447,1473]" name="New Zealand" pageId="17" pageNumber="168">NZ</collectingCountry>
PB
</materialsCitation>
\;
<materialsCitation id="3D983CA98B237363FED4D4B5FEAC8EFE" collectionCode="DSIR" collectorName="G. Kuschel" country="New Zealand" county="No" elevation="1250" location="Mt. Domett" municipality="Nov-Dec" pageId="17" pageNumber="168" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9BF6FD7D8B237363FED4D4B5FE248EDB" box="[318,404,1446,1472]" count="2" pageId="17" pageNumber="168" type="male">2 males</specimenCount>
and
<specimenCount id="9BF6FD7D8B237363FE26D4B5FD898EDB" box="[460,569,1446,1472]" count="2" pageId="17" pageNumber="168" type="female">2 females</specimenCount>
labeled: \
<location id="882F602F8B237363FD5AD4B4FC9B8EDA" LSID="urn:lsid:plazi:treatment:055987E28B237360FF7DD5DDFD1F889A:882F602F8B237363FD5AD4B4FC9B8EDA" box="[688,811,1447,1473]" country="New Zealand" county="No" municipality="Nov-Dec" name="Mt. Domett" pageId="17" pageNumber="168">Mt.Domett</location>
<quantity id="4A089B118B237363FCD9D4B5FC308EDA" box="[819,896,1446,1473]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="17" pageNumber="168" unit="m" value="1250.0">
<elevation id="06DDD1C78B237363FCD9D4B5FC308EDA" box="[819,896,1446,1473]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="17" pageNumber="168" unit="m" value="1250.0">1250m</elevation>
</quantity>
<collectingMunicipality id="6D2BAC8E8B237363FC62D4B4FC5F8EDA" box="[904,1007,1447,1473]" pageId="17" pageNumber="168">Nov-Dec</collectingMunicipality>
71
<collectorName id="200553228B237363FBF0D4B5FB218EDB" box="[1050,1169,1446,1473]" pageId="17" pageNumber="168">G.Kuschel</collectorName>
\ moss 71/169 \
<collectionCode id="EBE1AE318B237363FAA7D4B5FA3D8EDB" box="[1357,1421,1446,1472]" country="New Zealand" httpUri="http://grbio.org/cool/6qmb-xjj9" name="Department of Scientific and Industrial Research" pageId="17" pageNumber="168">DSIR</collectionCode>
\
<collectingCounty id="642E4E788B237363FF7DD4D8FF098EFE" box="[151,185,1483,1509]" pageId="17" pageNumber="168">No</collectingCounty>
\
<collectingCountry id="F5E776648B237363FF24D4D8FF438EFE" box="[206,243,1483,1509]" name="New Zealand" pageId="17" pageNumber="168">NZ</collectingCountry>
PB
</materialsCitation>
\;
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labeled: \
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PB
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\;
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<specimenCount id="9BF6FD7D8B237363FABBD4D9FA2C8EFF" box="[1361,1436,1482,1508]" count="1" pageId="17" pageNumber="168" type="male">1 male</specimenCount>
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NN
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71 \
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\ moss 71/165 \
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and
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labeled: \
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<taxonomicName id="4AF04D778B237363FE2CD700FDFE8D36" authorityName="Sokolov" authorityYear="2023" box="[454,590,1555,1581]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="187" phylum="Arthropoda" rank="species" species="karamea">N. Karamea</taxonomicName>
\ 1200 20.vi.
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\ litter 67/217 \
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\.
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B237363FF2DD725FB2D8D6F" pageId="17" pageNumber="168" type="etymology">
<paragraph id="8D4F36F48B237363FF2DD725FB2D8D6F" blockId="17.[151,1436,1338,2013]" pageId="17" pageNumber="168">
<emphasis id="BF84EAE68B237363FF2DD725FEFE8D4B" bold="true" box="[199,334,1590,1616]" pageId="17" pageNumber="168">Etymology.</emphasis>
The specific epithet is a noun in apposition in the nominative case based on the imaginary race of people in the novel of J. R. R. Tolkien “The Hobbit”, and refers to this speciess way of life.
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B237363FF2DD76DFE488C1F" pageId="17" pageNumber="168" type="materials_examined">
<paragraph id="8D4F36F48B237363FF2DD76DFB028D82" blockId="17.[151,1436,1338,2013]" box="[199,1202,1662,1689]" pageId="17" pageNumber="168">
<materialsCitation id="3D983CA98B237363FF2DD76DFB1E8D82" box="[199,1198,1662,1689]" country="New Zealand" county="South Island" location="Mt Domett" municipality="Kahurangi National Park" pageId="17" pageNumber="168" specimenCount="1" stateProvince="Nelson" typeStatus="holotype">
<emphasis id="BF84EAE68B237363FF2DD76DFED68D83" bold="true" box="[199,358,1662,1688]" pageId="17" pageNumber="168">
<typeStatus id="524B88568B237363FF2DD76DFEB28D83" box="[199,258,1662,1688]" pageId="17" pageNumber="168">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F5E776648B237363FE86D76CFDB58D83" box="[364,517,1662,1689]" name="New Zealand" pageId="17" pageNumber="168">New Zealand</collectingCountry>
,
<collectingCounty id="642E4E788B237363FDFAD76DFD108D83" box="[528,672,1662,1688]" pageId="17" pageNumber="168">South Island</collectingCounty>
,
<collectingRegion id="4F34F8168B237363FD41D76DFD4D8D83" box="[683,765,1662,1688]" country="New Zealand" name="Nelson" pageId="17" pageNumber="168">Nelson</collectingRegion>
,
<collectingMunicipality id="6D2BAC8E8B237363FCE2D76DFB938D83" box="[776,1059,1662,1688]" pageId="17" pageNumber="168">Kahurangi National Park</collectingMunicipality>
,
<location id="882F602F8B237363FBC4D76CFB1E8D82" LSID="urn:lsid:plazi:treatment:055987E28B237360FF7DD5DDFD1F889A:882F602F8B237363FBC4D76CFB1E8D82" box="[1070,1198,1663,1689]" country="New Zealand" county="South Island" municipality="Kahurangi National Park" name="Mt Domett" pageId="17" pageNumber="168" stateProvince="Nelson">Mt Domett</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8D4F36F48B237363FF2DD7B1FE488C1F" blockId="17.[151,1436,1338,2013]" pageId="17" pageNumber="168">
<emphasis id="BF84EAE68B237363FF2DD7B1FEED8DA7" bold="true" box="[199,349,1698,1724]" pageId="17" pageNumber="168">Recognition.</emphasis>
Adults of this species (
<figureCitation id="15CB2A718B237363FD6ED7B1FD508DA7" box="[644,736,1698,1724]" captionStart="FIGURE 9" captionStartId="15.[151,250,1649,1673]" captionTargetBox="[155,1436,181,1621]" captionTargetId="figure-15@15.[151,1436,181,1625]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect.A—N. canaanensis (Canaan, Nelson, SO), B—N. disjunctus (Fletchers Creek, Buller, SO), C—N. hobbit (9mi. N Karamea, Nelson, SO), D—N. karamea (Karamea River Gorge, Nelson, SO), E—N. kuscheli (Flora Hut, MountArthur, Nelson, SO), F—N. magnificus (Allison Reserve,Akatore, Dunedin, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184245" httpUri="https://zenodo.org/record/10184245/files/figure.png" pageId="17" pageNumber="168">Fig. 9C</figureCitation>
) are practically indistinguishable from the adults of many
<taxonomicName id="4AF04D778B237363FF7DD7D5FE388DFB" box="[151,392,1734,1760]" pageId="17" pageNumber="168">
<emphasis id="BF84EAE68B237363FF7DD7D5FE9C8DFB" box="[151,300,1734,1760]" italics="true" pageId="17" pageNumber="168">Nesamblyops</emphasis>
species
</taxonomicName>
based on external characters (e.g.,
<figureCitation id="15CB2A718B237363FCCED7D5FC338DFB" box="[804,899,1734,1760]" captionStart="FIGURE 9" captionStartId="15.[151,250,1649,1673]" captionTargetBox="[155,1436,181,1621]" captionTargetId="figure-15@15.[151,1436,181,1625]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect.A—N. canaanensis (Canaan, Nelson, SO), B—N. disjunctus (Fletchers Creek, Buller, SO), C—N. hobbit (9mi. N Karamea, Nelson, SO), D—N. karamea (Karamea River Gorge, Nelson, SO), E—N. kuscheli (Flora Hut, MountArthur, Nelson, SO), F—N. magnificus (Allison Reserve,Akatore, Dunedin, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184245" httpUri="https://zenodo.org/record/10184245/files/figure.png" pageId="17" pageNumber="168">Figs 9A</figureCitation>
,
<figureCitation id="15CB2A718B237363FC7BD7D5FC718DFB" box="[913,961,1734,1760]" captionStart="FIGURE 10" captionStartId="21.[151,250,1533,1557]" captionTargetBox="[155,1431,188,1505]" captionTargetId="figure-15@21.[154,1434,181,1509]" captionTargetPageId="21" captionText="FIGURE 10. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. montanus (Lake Sylvester, Kahurangi National Park, Nelson, SO), B—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), C—N. ovipennis (Mount Arthur, Nelson, SO), D—N. rotundicollis (Onekaka, Nelson, SO), E—N. solitarius (Italian Creek, Capleston, Buller, SO), F—N. subcaecus (Sharp) (Greymouth, Buller, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184247" httpUri="https://zenodo.org/record/10184247/files/figure.png" pageId="17" pageNumber="168">10C</figureCitation>
) and are distinguished from them by the structures of the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B237360FF2DD61DFD1F889A" lastPageId="18" lastPageNumber="169" pageId="17" pageNumber="168" type="description">
<paragraph id="8D4F36F48B237363FF2DD61DFB2E8C33" blockId="17.[151,1436,1338,2013]" box="[199,1182,1806,1833]" pageId="17" pageNumber="168">
<emphasis id="BF84EAE68B237363FF2DD61DFEE98C33" bold="true" box="[199,345,1806,1832]" pageId="17" pageNumber="168">Description.</emphasis>
Large for genus (SBL range
<quantity id="4A089B118B237363FD75D61DFCF48C32" box="[671,836,1806,1833]" metricMagnitude="-3" metricUnit="m" metricValue="2.08" metricValueMax="2.19" metricValueMin="1.97" pageId="17" pageNumber="168" unit="mm" value="2.08" valueMax="2.19" valueMin="1.97">1.972.19 mm</quantity>
, mean 2.09±
<quantity id="4A089B118B237363FC39D61DFBF88C32" box="[979,1096,1806,1833]" metricMagnitude="-5" metricUnit="m" metricValue="8.2" pageId="17" pageNumber="168" unit="mm" value="0.082">0.082 mm</quantity>
, n=16).
</paragraph>
<paragraph id="8D4F36F48B237363FF2DD621FDAB8C8E" blockId="17.[151,1436,1338,2013]" pageId="17" pageNumber="168">
<emphasis id="BF84EAE68B237363FF2DD621FE978C57" box="[199,295,1842,1868]" italics="true" pageId="17" pageNumber="168">Habitus.</emphasis>
Body form (
<figureCitation id="15CB2A718B237363FE44D621FDB18C57" box="[430,513,1842,1868]" captionStart="FIGURE 9" captionStartId="15.[151,250,1649,1673]" captionTargetBox="[155,1436,181,1621]" captionTargetId="figure-15@15.[151,1436,181,1625]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect.A—N. canaanensis (Canaan, Nelson, SO), B—N. disjunctus (Fletchers Creek, Buller, SO), C—N. hobbit (9mi. N Karamea, Nelson, SO), D—N. karamea (Karamea River Gorge, Nelson, SO), E—N. kuscheli (Flora Hut, MountArthur, Nelson, SO), F—N. magnificus (Allison Reserve,Akatore, Dunedin, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184245" httpUri="https://zenodo.org/record/10184245/files/figure.png" pageId="17" pageNumber="168">Fig.9C</figureCitation>
) markedly convex, elongate ovoid, general proportions wide (WE/SBL0.40±0.009), head narrow relative to pronotum (WH/WPm 0.66±0.028), proportions of pronotum in comparison to elytra average for genus (WPm/WE 0.75±0.018).
</paragraph>
<paragraph id="8D4F36F48B237363FF2DD68DFCFD8CA2" blockId="17.[151,1436,1338,2013]" box="[199,845,1950,1977]" pageId="17" pageNumber="168">
<emphasis id="BF84EAE68B237363FF2DD68DFEBD8CA3" box="[199,269,1950,1976]" italics="true" pageId="17" pageNumber="168">Color.</emphasis>
Body color brunneorufous, appendages testaceous.
</paragraph>
<paragraph id="8D4F36F48B237360FF2DD6D1FA3B8BE2" blockId="17.[151,1436,1338,2013]" lastBlockId="18.[151,1437,150,897]" lastPageId="18" lastPageNumber="169" pageId="17" pageNumber="168">
<emphasis id="BF84EAE68B237363FF2DD6D1FEF18CC7" box="[199,321,1986,2012]" italics="true" pageId="17" pageNumber="168">Prothorax.</emphasis>
Pronotum (
<figureCitation id="15CB2A718B237363FE2CD6D1FD9A8CC7" box="[454,554,1986,2012]" captionStart="FIGURE 12" captionStartId="26.[151,250,1215,1239]" captionTargetBox="[157,1430,181,1192]" captionTargetId="figure-32@26.[157,1430,181,1192]" captionTargetPageId="26" captionText="FIGURE 12. Digital images of pronota of New Zealand Nesamblyops species, dorsal aspect. A—N. canaanensis (Canaan, Nelson, SO), B—N. disjunctus (Fletchers Creek, Buller, SO), C—N. hobbit (9mi. N Karamea, Nelson, SO), D—N. karamea (Karamea River Gorge, Nelson, SO), E—N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO), F—N. magnificus (Allison Reserve, Akatore, Dunedin, SO), G—N. montanus (Lake Sylvester, Kahurangi National Park, Nelson, SO), H—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), I—N. ovipennis (Mount Arthur, Nelson, SO). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10184251" httpUri="https://zenodo.org/record/10184251/files/figure.png" pageId="17" pageNumber="168">Fig. 12C</figureCitation>
) moderately long in comparison to elytra (LP/LE 0.42±0.010) and moderately transverse (WPm/LP 1.22±0.035), with lateral margins arcuately constricted posteriorly (WPm/WPp 1.30±0.026). Anterior angles indistinct, posterior angles obtuse (109117°), rounded. Width between posterior angles greater than between anterior angles (WPa/WPp 0.86±0.029). Basal margin almost rectilinear, slightly convex at middle.
</paragraph>
<paragraph id="8D4F36F48B207360FF2DD010FD578A7E" blockId="18.[151,1437,150,897]" pageId="18" pageNumber="169">
<emphasis id="BF84EAE68B207360FF2DD010FEA58A06" box="[199,277,259,285]" italics="true" pageId="18" pageNumber="169">Elytra.</emphasis>
Ovoid, narrowly depressed along suture, comparatively long (LE/SBL 0.59±0.005) and moderately wide (WE/LE 0.68±0.015). Humeri completely rounded. Lateral margins slightly divergent at basal half, subparallel at middle and evenly rounded to apex in apical third.
</paragraph>
<paragraph id="8D4F36F48B207360FF2DD07CFBCD897A" blockId="18.[151,1437,150,897]" pageId="18" pageNumber="169">
<emphasis id="BF84EAE68B207360FF2DD07CFEC18A92" box="[199,369,367,393]" italics="true" pageId="18" pageNumber="169">Male genitalia.</emphasis>
Median lobe of aedeagus (
<figureCitation id="15CB2A718B207360FD7FD07CFD5E8A92" box="[661,750,367,393]" captionStart="FIGURE 14" captionStartId="30.[151,250,1365,1389]" captionTargetBox="[157,1433,187,1336]" captionTargetId="figure-32@30.[151,1436,181,1341]" captionTargetPageId="30" captionText="FIGURE 14. Line drawings of male genitalia of New Zealand Nesamblyops species. N. canaanensis (Canaan, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. disjunctus (Fletchers Creek, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. hobbit (Mount Domett, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. karamea (Karamea River Gorge, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. magnificus (Allison Reserve, Akatore, Dunedin, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Legend: rCs—reversed C-contour; Vs—V-contour. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184255" httpUri="https://zenodo.org/record/10184255/files/figure.png" pageId="18" pageNumber="169">Fig. 14I</figureCitation>
) moderately arcuate and moderately twisted. Shaft subparallel basally, tapering in apical half.Apex almost straight with narrowly rounded tip. Apical orifice long, occupies almost half of the shaft length. Ventral margin of median lobe straight. Walls of shaft with a number of poriferous canals scattered along ventral margin. Dorsal copulatory sclerites with well-developed rC-sclerite and reduced V-contour with short dorsal branch slightly protruding toward basal orifice (
<figureCitation id="15CB2A718B207360FC87D0ECFC798902" box="[877,969,511,537]" captionStart="FIGURE 14" captionStartId="30.[151,250,1365,1389]" captionTargetBox="[157,1433,187,1336]" captionTargetId="figure-32@30.[151,1436,181,1341]" captionTargetPageId="30" captionText="FIGURE 14. Line drawings of male genitalia of New Zealand Nesamblyops species. N. canaanensis (Canaan, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. disjunctus (Fletchers Creek, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. hobbit (Mount Domett, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. karamea (Karamea River Gorge, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. magnificus (Allison Reserve, Akatore, Dunedin, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Legend: rCs—reversed C-contour; Vs—V-contour. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184255" httpUri="https://zenodo.org/record/10184255/files/figure.png" pageId="18" pageNumber="169">Fig. 14I</figureCitation>
). Left paramere (
<figureCitation id="15CB2A718B207360FB62D0ECFB5C8902" box="[1160,1260,511,537]" captionStart="FIGURE 14" captionStartId="30.[151,250,1365,1389]" captionTargetBox="[157,1433,187,1336]" captionTargetId="figure-32@30.[151,1436,181,1341]" captionTargetPageId="30" captionText="FIGURE 14. Line drawings of male genitalia of New Zealand Nesamblyops species. N. canaanensis (Canaan, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. disjunctus (Fletchers Creek, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. hobbit (Mount Domett, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. karamea (Karamea River Gorge, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. magnificus (Allison Reserve, Akatore, Dunedin, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Legend: rCs—reversed C-contour; Vs—V-contour. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184255" httpUri="https://zenodo.org/record/10184255/files/figure.png" pageId="18" pageNumber="169">Fig. 14G</figureCitation>
) comparatively wide, with attenuate apex, bearing three long setae. Right paramere (
<figureCitation id="15CB2A718B207360FC72D330FC4E8926" box="[920,1022,547,573]" captionStart="FIGURE 14" captionStartId="30.[151,250,1365,1389]" captionTargetBox="[157,1433,187,1336]" captionTargetId="figure-32@30.[151,1436,181,1341]" captionTargetPageId="30" captionText="FIGURE 14. Line drawings of male genitalia of New Zealand Nesamblyops species. N. canaanensis (Canaan, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. disjunctus (Fletchers Creek, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. hobbit (Mount Domett, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. karamea (Karamea River Gorge, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. magnificus (Allison Reserve, Akatore, Dunedin, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Legend: rCs—reversed C-contour; Vs—V-contour. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184255" httpUri="https://zenodo.org/record/10184255/files/figure.png" pageId="18" pageNumber="169">Fig. 14H</figureCitation>
) long and narrow, bearing three long setae, which are slightly shorter than the length of paramere. Ring sclerite as in
<figureCitation id="15CB2A718B207360FBFED354FBC9897A" box="[1044,1145,583,609]" captionStart="FIGURE 17" captionStartId="34.[151,250,1191,1215]" captionTargetBox="[163,1421,187,1168]" captionTargetId="figure-32@34.[151,1436,181,1168]" captionTargetPageId="34" captionText="FIGURE 17. Line drawings of ring sclerite of New Zealand Nesamblyops species, male genitalia, dorsal aspect. A—N. canaanensis (Canaan, Nelson, SO), B—N. disjunctus (Fletchers Creek, Buller, SO), C—N. hobbit (Mount Domett, Nelson, SO), D—N. karamea (Karamea River Gorge, Nelson, SO), E—N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO), F—N. montanus (Relax Shelter, Mount Robert Ridge, Nelson, SO), G—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), H—N. ovipennis (Mount Arthur, Nelson, SO), I—N. rotundicollis (Onekaka, Nelson, SO), J—N. subcaecus (Margarets Tarn Lake, Arthurs Pass, Mid Canterbury, SO), K—N. viator (Secretary Island, Grono Bay, Fjordland, SO), L—N. victoriae (Capleston, Victoria Range, Buller, SO). Scale = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10184261" httpUri="https://zenodo.org/record/10184261/files/figure.png" pageId="18" pageNumber="169">Fig. 17C</figureCitation>
.
</paragraph>
<paragraph id="8D4F36F48B207360FF2DD378FD29899E" blockId="18.[151,1437,150,897]" box="[199,665,619,645]" pageId="18" pageNumber="169">
<emphasis id="BF84EAE68B207360FF2DD378FE5D899E" box="[199,493,619,645]" italics="true" pageId="18" pageNumber="169">Female internal genitalia.</emphasis>
Not examined.
</paragraph>
<paragraph id="8D4F36F48B207360FF2DD39DFD6889EA" blockId="18.[151,1437,150,897]" pageId="18" pageNumber="169">
<emphasis id="BF84EAE68B207360FF2DD39DFDB589B3" bold="true" box="[199,517,654,680]" pageId="18" pageNumber="169">Geographical distribution.</emphasis>
The species inhabits the central-western part of the Kahurangi National Park with a range stretching from the catchment of the Aorere River at the northeast to the northernmost parts of the
<collectingRegion id="4F34F8168B207360FA8FD3A0FE8589EA" country="New Zealand" name="West Coast" pageId="18" pageNumber="169">West Coast District</collectingRegion>
, South Island (
<figureCitation id="15CB2A718B207360FE36D3C4FD9E89EA" box="[476,558,727,753]" captionStart="FIGURE 19" captionStartId="36.[151,250,1381,1405]" captionTargetBox="[151,1436,181,1357]" captionTargetId="figure-32@36.[151,1436,181,1357]" captionTargetPageId="36" captionText="FIGURE 19. Distributional records for the Nesamblyops species and images of male median lobe, obtained from specimens, collected in the line-connected with images localities. Green circles—N. canaanensis; black circles—N. hobbit; blue circles— N. ovipennis; yellow circles—N. karamea; red circles—N. kuscheli. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10184267" httpUri="https://zenodo.org/record/10184267/files/figure.png" pageId="18" pageNumber="169">Fig. 19</figureCitation>
, black circles).
</paragraph>
<paragraph id="8D4F36F48B207360FF2DD3E9FB9F880E" blockId="18.[151,1437,150,897]" box="[199,1071,762,789]" pageId="18" pageNumber="169">
<emphasis id="BF84EAE68B207360FF2DD3E9FE9A880F" bold="true" box="[199,298,762,788]" pageId="18" pageNumber="169">Habitat.</emphasis>
Specimens were collected from moss, mat plants, and litter samples.
</paragraph>
<paragraph id="8D4F36F48B207360FF2DD20DFD1F889A" blockId="18.[151,1437,150,897]" pageId="18" pageNumber="169">
<emphasis id="BF84EAE68B207360FF2DD20DFEC08823" bold="true" box="[199,368,798,824]" pageId="18" pageNumber="169">Relationships.</emphasis>
The structure of the male genitalia of
<taxonomicName id="4AF04D778B207360FCF2D20CFC308822" authorityName="Sokolov" authorityYear="2023" box="[792,896,799,825]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="hobbit">
<emphasis id="BF84EAE68B207360FCF2D20CFC308822" box="[792,896,799,825]" italics="true" pageId="18" pageNumber="169">N. hobbit</emphasis>
</taxonomicName>
suggests its relatedness to the other species with trisetose parameres, a well-developed rC-sclerite with small V-contour in the internal sac of median lobe, such as
<taxonomicName id="4AF04D778B207360FA69D250FE91889B" authorityName="Sokolov" authorityYear="2023" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="canaanensis">
<emphasis id="BF84EAE68B207360FA69D250FE91889B" italics="true" pageId="18" pageNumber="169">N. canaanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4AF04D778B207360FEB2D274FE54889B" authorityName="Sokolov" authorityYear="2023" box="[344,484,871,896]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="ovipennis">
<emphasis id="BF84EAE68B207360FEB2D274FE54889B" box="[344,484,871,896]" italics="true" pageId="18" pageNumber="169">N. ovipennis</emphasis>
</taxonomicName>
, described below.
</paragraph>
</subSubSection>
</treatment>
</document>