treatments-xml/data/05/59/87/055987E28B207361FF7DD2C1FC0689B2.xml
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<document id="3CCB9559C80858BFD5DF2191C73B577F" ID-DOI="10.11646/zootaxa.5375.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10184224" ID-ZooBank="F3D0E008-556C-4FAD-BF51-4F1A714325DA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700642529072" checkinUser="plazi" docAuthor="Sokolov, Igor M." docDate="2023" docId="055987E28B207361FF7DD2C1FC0689B2" docLanguage="en" docName="zootaxa.5375.2.1.pdf" docOrigin="Zootaxa 5375 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5375.2.1/52321" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nesamblyops karamea Sokolov 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="170" masterDocId="F960FF9A8B327372FFEAD113FFB08B1B" masterDocTitle="Fourteen new species of the genus Nesamblyops Jeannel (Coleoptera: Carabidae: Anillini) from the South Island of New Zealand with redescription of the genus and description of a new subtribe" masterLastPageNumber="192" masterPageNumber="151" pageNumber="169" updateTime="1700643183691" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4647F1D874829394B8F31D0F409F645F">Fourteen new species of the genus Nesamblyops Jeannel (Coleoptera: Carabidae: Anillini) from the South Island of New Zealand with redescription of the genus and description of a new subtribe</mods:title>
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<mods:namePart id="3BB54693C312DB9380BC4B1BB991BBFF">Sokolov, Igor M.</mods:namePart>
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<mods:date id="00F610EA7D9914E3127F6595458137FF">2023</mods:date>
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<mods:number id="3A80ED6893CC6E29B83A167BAC7BB53A">2023-11-22</mods:number>
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<mods:identifier id="38F34AC24B64DFC535D854F8EE5B1245" type="DOI">10.11646/zootaxa.5375.2.1</mods:identifier>
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<treatment id="055987E28B207361FF7DD2C1FC0689B2" LSID="urn:lsid:plazi:treatment:055987E28B207361FF7DD2C1FC0689B2" httpUri="http://treatment.plazi.org/id/055987E28B207361FF7DD2C1FC0689B2" lastPageId="19" lastPageNumber="170" pageId="18" pageNumber="169">
<subSubSection id="C5EA657F8B207360FF7DD2C1FDA488F6" box="[151,532,978,1005]" pageId="18" pageNumber="169" type="nomenclature">
<paragraph id="8D4F36F48B207360FF7DD2C1FDA488F6" blockId="18.[151,534,978,1041]" box="[151,532,978,1005]" pageId="18" pageNumber="169">
<heading id="D60781988B207360FF7DD2C1FDA488F6" bold="true" box="[151,532,978,1005]" fontSize="11" level="1" pageId="18" pageNumber="169" reason="1">
<emphasis id="BF84EAE68B207360FF7DD2C1FDA488F6" bold="true" box="[151,532,978,1005]" pageId="18" pageNumber="169">
<taxonomicName id="4AF04D778B207360FF7DD2C1FE1888F6" authority="Sokolov, 2023" authorityName="Sokolov" authorityYear="2023" box="[151,424,978,1005]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="karamea" status="sp. nov.">
<emphasis id="BF84EAE68B207360FF7DD2C1FE1888F6" bold="true" box="[151,424,978,1005]" italics="true" pageId="18" pageNumber="169">Nesamblyops karamea</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A4B7579D8B207360FE5DD2C0FDA488F6" box="[439,532,979,1005]" pageId="18" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B207360FF7DD2E4FDA68F0A" box="[151,534,1015,1041]" pageId="18" pageNumber="169" type="description">
<paragraph id="8D4F36F48B207360FF7DD2E4FDA68F0A" blockId="18.[151,534,978,1041]" box="[151,534,1015,1041]" pageId="18" pageNumber="169">
<figureCitation id="15CB2A718B207360FF7DD2E4FEAD8F0A" box="[151,285,1015,1041]" captionStart="FIGURE 9" captionStartId="15.[151,250,1649,1673]" captionTargetBox="[155,1436,181,1621]" captionTargetId="figure-15@15.[151,1436,181,1625]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect.A—N. canaanensis (Canaan, Nelson, SO), B—N. disjunctus (Fletchers Creek, Buller, SO), C—N. hobbit (9mi. N Karamea, Nelson, SO), D—N. karamea (Karamea River Gorge, Nelson, SO), E—N. kuscheli (Flora Hut, MountArthur, Nelson, SO), F—N. magnificus (Allison Reserve,Akatore, Dunedin, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184245" httpUri="https://zenodo.org/record/10184245/files/figure.png" pageId="18" pageNumber="169">Figures. 9D</figureCitation>
,
<figureCitation id="15CB2A718B207360FEC3D2E4FEEC8F0A" box="[297,348,1015,1041]" captionStart="FIGURE 12" captionStartId="26.[151,250,1215,1239]" captionTargetBox="[157,1430,181,1192]" captionTargetId="figure-32@26.[157,1430,181,1192]" captionTargetPageId="26" captionText="FIGURE 12. Digital images of pronota of New Zealand Nesamblyops species, dorsal aspect. A—N. canaanensis (Canaan, Nelson, SO), B—N. disjunctus (Fletchers Creek, Buller, SO), C—N. hobbit (9mi. N Karamea, Nelson, SO), D—N. karamea (Karamea River Gorge, Nelson, SO), E—N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO), F—N. magnificus (Allison Reserve, Akatore, Dunedin, SO), G—N. montanus (Lake Sylvester, Kahurangi National Park, Nelson, SO), H—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), I—N. ovipennis (Mount Arthur, Nelson, SO). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10184251" httpUri="https://zenodo.org/record/10184251/files/figure.png" pageId="18" pageNumber="169">12D</figureCitation>
,
<figureCitation id="15CB2A718B207360FE82D2E4FE1E8F0A" box="[360,430,1015,1041]" captionStart="FIGURE 14" captionStartId="30.[151,250,1365,1389]" captionTargetBox="[157,1433,187,1336]" captionTargetId="figure-32@30.[151,1436,181,1341]" captionTargetPageId="30" captionText="FIGURE 14. Line drawings of male genitalia of New Zealand Nesamblyops species. N. canaanensis (Canaan, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. disjunctus (Fletchers Creek, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. hobbit (Mount Domett, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. karamea (Karamea River Gorge, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. magnificus (Allison Reserve, Akatore, Dunedin, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Legend: rCs—reversed C-contour; Vs—V-contour. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184255" httpUri="https://zenodo.org/record/10184255/files/figure.png" pageId="18" pageNumber="169">14JL</figureCitation>
,
<figureCitation id="15CB2A718B207360FE56D2E4FE5E8F0A" box="[444,494,1015,1041]" captionStart="FIGURE 17" captionStartId="34.[151,250,1191,1215]" captionTargetBox="[163,1421,187,1168]" captionTargetId="figure-32@34.[151,1436,181,1168]" captionTargetPageId="34" captionText="FIGURE 17. Line drawings of ring sclerite of New Zealand Nesamblyops species, male genitalia, dorsal aspect. A—N. canaanensis (Canaan, Nelson, SO), B—N. disjunctus (Fletchers Creek, Buller, SO), C—N. hobbit (Mount Domett, Nelson, SO), D—N. karamea (Karamea River Gorge, Nelson, SO), E—N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO), F—N. montanus (Relax Shelter, Mount Robert Ridge, Nelson, SO), G—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), H—N. ovipennis (Mount Arthur, Nelson, SO), I—N. rotundicollis (Onekaka, Nelson, SO), J—N. subcaecus (Margarets Tarn Lake, Arthurs Pass, Mid Canterbury, SO), K—N. viator (Secretary Island, Grono Bay, Fjordland, SO), L—N. victoriae (Capleston, Victoria Range, Buller, SO). Scale = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10184261" httpUri="https://zenodo.org/record/10184261/files/figure.png" pageId="18" pageNumber="169">17D</figureCitation>
,
<figureCitation id="15CB2A718B207360FE10D2E4FDA68F0A" box="[506,534,1015,1041]" captionStart="FIGURE 19" captionStartId="36.[151,250,1381,1405]" captionTargetBox="[151,1436,181,1357]" captionTargetId="figure-32@36.[151,1436,181,1357]" captionTargetPageId="36" captionText="FIGURE 19. Distributional records for the Nesamblyops species and images of male median lobe, obtained from specimens, collected in the line-connected with images localities. Green circles—N. canaanensis; black circles—N. hobbit; blue circles— N. ovipennis; yellow circles—N. karamea; red circles—N. kuscheli. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10184267" httpUri="https://zenodo.org/record/10184267/files/figure.png" pageId="18" pageNumber="169">19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B207360FF7DD52CFB508FDE" pageId="18" pageNumber="169" type="materials_examined">
<paragraph id="8D4F36F48B207360FF7DD52CFB508FDE" blockId="18.[151,1437,1086,2013]" pageId="18" pageNumber="169">
<emphasis id="BF84EAE68B207360FF7DD52CFEF68F43" bold="true" box="[151,326,1086,1113]" pageId="18" pageNumber="169">Type material.</emphasis>
<materialsCitation id="3D983CA98B207360FEA5D52CFEDE8F66" collectionCode="NZAC, DSIR" collectorName="Karamea River Gorge. F. Alack" country="New Zealand" county="No" location="Karamea River" municipality="River Gorge." pageId="18" pageNumber="169" specimenCount="1" typeStatus="holotype">
<emphasis id="BF84EAE68B207360FEA5D52CFE548F42" bold="true" box="[335,484,1087,1113]" italics="true" pageId="18" pageNumber="169">
<typeStatus id="524B88568B207360FEA5D52CFE548F42" box="[335,484,1087,1113]" pageId="18" pageNumber="169" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
, male, in
<collectionCode id="EBE1AE318B207360FDBDD52CFD168F42" box="[599,678,1087,1113]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="18" pageNumber="169" type="Other institutional type">NZAC</collectionCode>
, labeled: \
<collectorName id="200553228B207360FCCED52CFB348F42" box="[804,1156,1087,1113]" pageId="18" pageNumber="169">
<location id="882F602F8B207360FCCED52CFC608F42" LSID="urn:lsid:plazi:treatment:055987E28B207361FF7DD2C1FC0689B2:882F602F8B207360FCCED52CFC608F42" box="[804,976,1087,1113]" country="New Zealand" county="No" municipality="River Gorge." name="Karamea River" pageId="18" pageNumber="169">
Karamea
<collectingMunicipality id="6D2BAC8E8B207360FC78D52CFC608F42" box="[914,976,1087,1113]" pageId="18" pageNumber="169">River</collectingMunicipality>
</location>
Gorge. F.Alack
</collectorName>
24.vi.67 \ litter 67/216 \
<collectionCode id="EBE1AE318B207360FF7DD570FF678F66" box="[151,215,1123,1149]" country="New Zealand" httpUri="http://grbio.org/cool/6qmb-xjj9" name="Department of Scientific and Industrial Research" pageId="18" pageNumber="169">DSIR</collectionCode>
\
<collectingCounty id="642E4E788B207360FF01D570FEBD8F66" box="[235,269,1123,1149]" pageId="18" pageNumber="169">No</collectingCounty>
\
<collectingCountry id="F5E776648B207360FECBD570FEF68F66" box="[289,326,1123,1149]" name="New Zealand" pageId="18" pageNumber="169">NZ</collectingCountry>
PB
</materialsCitation>
\.
<emphasis id="BF84EAE68B207360FE63D570FD948F66" bold="true" box="[393,548,1123,1149]" italics="true" pageId="18" pageNumber="169">
<typeStatus id="524B88568B207360FE63D570FD948F66" box="[393,548,1123,1149]" pageId="18" pageNumber="169" type="paratype">PARATYPES</typeStatus>
</emphasis>
(
<specimenCount id="9BF6FD7D8B207360FDD9D570FD0F8F66" box="[563,703,1123,1149]" count="9" pageId="18" pageNumber="169" type="generic">9 specimens</specimenCount>
, dissected 2 exx.),
<specimenCount id="9BF6FD7D8B207360FC65D570FC698F66" box="[911,985,1123,1149]" count="1" pageId="18" pageNumber="169" type="male">1 male</specimenCount>
and
<specimenCount id="9BF6FD7D8B207360FBE4D570FBC98F66" box="[1038,1145,1123,1149]" count="2" pageId="18" pageNumber="169" type="female">2 females</specimenCount>
labeled same as
<typeStatus id="524B88568B207360FAD8D570FA288F66" box="[1330,1432,1123,1149]" pageId="18" pageNumber="169" type="holotype">holotype</typeStatus>
;
<materialsCitation id="3D983CA98B207360FF7DD594FB898FBA" box="[151,1081,1159,1185]" collectionCode="DSIR" collectorName="Karamea Val. F. Alack" country="New Zealand" location="No" pageId="18" pageNumber="169" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9BF6FD7D8B207360FF7DD594FF5C8FBA" box="[151,236,1159,1185]" count="2" pageId="18" pageNumber="169" type="male">2 males</specimenCount>
labeled: \
<collectorName id="200553228B207360FEB4D594FDE18FBA" box="[350,593,1159,1185]" pageId="18" pageNumber="169">Karamea Val. F.Alack</collectorName>
21.vi.67 \ litter 67/218 \
<collectionCode id="EBE1AE318B207360FC89D594FC138FBA" box="[867,931,1159,1185]" country="New Zealand" httpUri="http://grbio.org/cool/6qmb-xjj9" name="Department of Scientific and Industrial Research" pageId="18" pageNumber="169">DSIR</collectionCode>
\
<location id="882F602F8B207360FC5CD594FC688FBA" LSID="urn:lsid:plazi:treatment:055987E28B207361FF7DD2C1FC0689B2:882F602F8B207360FC5CD594FC688FBA" box="[950,984,1159,1185]" country="New Zealand" name="No" pageId="18" pageNumber="169">No</location>
\
<collectingCountry id="F5E776648B207360FC06D594FBA18FBA" box="[1004,1041,1159,1185]" name="New Zealand" pageId="18" pageNumber="169">NZ</collectingCountry>
PB
</materialsCitation>
\;
<specimenCount id="9BF6FD7D8B207360FBBED594FB2E8FBA" box="[1108,1182,1159,1185]" count="1" pageId="18" pageNumber="169" type="male">1 male</specimenCount>
and
<specimenCount id="9BF6FD7D8B207360FB39D594FA8E8FBA" box="[1235,1342,1159,1185]" count="3" pageId="18" pageNumber="169" type="female">3 females</specimenCount>
labeled: \
<specimenCount id="9BF6FD7D8B207360FF4CD5B8FF618FDE" box="[166,209,1195,1221]" count="6" pageId="18" pageNumber="169" type="generic">
<quantity id="4A089B118B207360FF4CD5B8FF7D8FDE" box="[166,205,1195,1221]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="169" unit="m" value="6.0">6m</quantity>
<materialsCitation id="3D983CA98B207360FF27D5B8FF618FDE" box="[205,209,1195,1221]" collectionCode="R, DSIR" collectorName="Little Wanganui &amp; F. Alack" country="New Zealand" location="West" municipality="No" pageId="18" pageNumber="169" specimenCount="1" stateProvince="Nelson" typeStatus="paratype">.</materialsCitation>
</specimenCount>
up
<collectorName id="200553228B207360FF11D5B8FE1F8FDE" box="[251,431,1195,1221]" pageId="18" pageNumber="169">Little Wanganui</collectorName>
<collectionCode id="EBE1AE318B207360FE5CD5B8FE798FDE" box="[438,457,1195,1221]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="18" pageNumber="169">R</collectionCode>
.
<location id="882F602F8B207360FE3CD5B8FDBD8FDE" LSID="urn:lsid:plazi:treatment:055987E28B207361FF7DD2C1FC0689B2:882F602F8B207360FE3CD5B8FDBD8FDE" box="[470,525,1195,1221]" country="New Zealand" municipality="No" name="West" pageId="18" pageNumber="169" stateProvince="Nelson">West</location>
<collectingRegion id="4F34F8168B207360FDFED5B8FDD78FDE" box="[532,615,1195,1221]" country="New Zealand" name="Nelson" pageId="18" pageNumber="169">Nelson</collectingRegion>
. \
<collectorName id="200553228B207360FD6AD5B8FD678FDE" box="[640,727,1195,1221]" pageId="18" pageNumber="169">F.Alack</collectorName>
24.vi.67 \ litter 67/211 \
<collectionCode id="EBE1AE318B207360FC04D5B8FB9E8FDE" box="[1006,1070,1195,1221]" country="New Zealand" httpUri="http://grbio.org/cool/6qmb-xjj9" name="Department of Scientific and Industrial Research" pageId="18" pageNumber="169">DSIR</collectionCode>
\
<collectingMunicipality id="6D2BAC8E8B207360FBAED5B8FBD68FDE" box="[1092,1126,1195,1221]" pageId="18" pageNumber="169">No</collectingMunicipality>
\
<collectingCountry id="F5E776648B207360FB96D5B8FB118FDE" box="[1148,1185,1195,1221]" name="New Zealand" pageId="18" pageNumber="169">NZ</collectingCountry>
PB \.
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B207360FF2DD5DCFD568E16" pageId="18" pageNumber="169" type="etymology">
<paragraph id="8D4F36F48B207360FF2DD5DCFD568E16" blockId="18.[151,1437,1086,2013]" pageId="18" pageNumber="169">
<emphasis id="BF84EAE68B207360FF2DD5DCFEFE8FF2" bold="true" box="[199,334,1231,1257]" pageId="18" pageNumber="169">Etymology.</emphasis>
The specific epithet is a noun in apposition in the nominative case and refers to the name of the river gorge from which the new species is described.
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B207360FF2DD404FE488EDA" pageId="18" pageNumber="169" type="materials_examined">
<paragraph id="8D4F36F48B207360FF2DD404FF7D8E4E" blockId="18.[151,1437,1086,2013]" pageId="18" pageNumber="169">
<materialsCitation id="3D983CA98B207360FF2DD404FF798E4E" country="New Zealand" county="South Island" location="Karamea River Gorge area" municipality="Kahurangi National Park" pageId="18" pageNumber="169" specimenCount="1" stateProvince="West Coast" typeStatus="holotype">
<emphasis id="BF84EAE68B207360FF2DD404FED88E2B" bold="true" box="[199,360,1302,1329]" pageId="18" pageNumber="169">
<typeStatus id="524B88568B207360FF2DD404FEB28E2A" box="[199,258,1303,1329]" pageId="18" pageNumber="169">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F5E776648B207360FE9BD404FDBC8E2A" box="[369,524,1303,1329]" name="New Zealand" pageId="18" pageNumber="169">New Zealand</collectingCountry>
,
<collectingCounty id="642E4E788B207360FDF3D404FD1B8E2A" box="[537,683,1303,1329]" pageId="18" pageNumber="169">South Island</collectingCounty>
,
<collectingRegion id="4F34F8168B207360FD52D404FC8C8E2A" box="[696,828,1303,1329]" country="New Zealand" name="West Coast" pageId="18" pageNumber="169">West Coast</collectingRegion>
,
<collectingMunicipality id="6D2BAC8E8B207360FCA3D404FBD88E2A" box="[841,1128,1303,1329]" pageId="18" pageNumber="169">Kahurangi National Park</collectingMunicipality>
, the
<location id="882F602F8B207360FB4BD404FF798E4E" LSID="urn:lsid:plazi:treatment:055987E28B207361FF7DD2C1FC0689B2:882F602F8B207360FB4BD404FF798E4E" country="New Zealand" county="South Island" municipality="Kahurangi National Park" name="Karamea River Gorge area" pageId="18" pageNumber="169" stateProvince="West Coast">Karamea River Gorge area</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8D4F36F48B207360FF2DD44DFE488EDA" blockId="18.[151,1437,1086,2013]" pageId="18" pageNumber="169">
<emphasis id="BF84EAE68B207360FF2DD44DFEED8E63" bold="true" box="[199,349,1374,1400]" pageId="18" pageNumber="169">Recognition.</emphasis>
Adults of this species (
<figureCitation id="15CB2A718B207360FD69D44CFD528E62" box="[643,738,1375,1401]" captionStart="FIGURE 9" captionStartId="15.[151,250,1649,1673]" captionTargetBox="[155,1436,181,1621]" captionTargetId="figure-15@15.[151,1436,181,1625]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect.A—N. canaanensis (Canaan, Nelson, SO), B—N. disjunctus (Fletchers Creek, Buller, SO), C—N. hobbit (9mi. N Karamea, Nelson, SO), D—N. karamea (Karamea River Gorge, Nelson, SO), E—N. kuscheli (Flora Hut, MountArthur, Nelson, SO), F—N. magnificus (Allison Reserve,Akatore, Dunedin, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184245" httpUri="https://zenodo.org/record/10184245/files/figure.png" pageId="18" pageNumber="169">Fig. 9D</figureCitation>
) are practically indistinguishable from the adults of many
<taxonomicName id="4AF04D778B207360FF7DD490FE388E86" box="[151,392,1411,1437]" pageId="18" pageNumber="169">
<emphasis id="BF84EAE68B207360FF7DD490FE9C8E86" box="[151,300,1411,1437]" italics="true" pageId="18" pageNumber="169">Nesamblyops</emphasis>
species
</taxonomicName>
based on external characters (e.g.,
<figureCitation id="15CB2A718B207360FCCED490FC318E86" box="[804,897,1411,1437]" captionStart="FIGURE 9" captionStartId="15.[151,250,1649,1673]" captionTargetBox="[155,1436,181,1621]" captionTargetId="figure-15@15.[151,1436,181,1625]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect.A—N. canaanensis (Canaan, Nelson, SO), B—N. disjunctus (Fletchers Creek, Buller, SO), C—N. hobbit (9mi. N Karamea, Nelson, SO), D—N. karamea (Karamea River Gorge, Nelson, SO), E—N. kuscheli (Flora Hut, MountArthur, Nelson, SO), F—N. magnificus (Allison Reserve,Akatore, Dunedin, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184245" httpUri="https://zenodo.org/record/10184245/files/figure.png" pageId="18" pageNumber="169">Figs 9C</figureCitation>
,
<figureCitation id="15CB2A718B207360FC7BD490FC718E86" box="[913,961,1411,1437]" captionStart="FIGURE 10" captionStartId="21.[151,250,1533,1557]" captionTargetBox="[155,1431,188,1505]" captionTargetId="figure-15@21.[154,1434,181,1509]" captionTargetPageId="21" captionText="FIGURE 10. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect. A—N. montanus (Lake Sylvester, Kahurangi National Park, Nelson, SO), B—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), C—N. ovipennis (Mount Arthur, Nelson, SO), D—N. rotundicollis (Onekaka, Nelson, SO), E—N. solitarius (Italian Creek, Capleston, Buller, SO), F—N. subcaecus (Sharp) (Greymouth, Buller, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184247" httpUri="https://zenodo.org/record/10184247/files/figure.png" pageId="18" pageNumber="169">10C</figureCitation>
) and are distinguished from them by the structures of the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C5EA657F8B207361FF2DD4D9FC0689B2" lastPageId="19" lastPageNumber="170" pageId="18" pageNumber="169" type="description">
<paragraph id="8D4F36F48B207360FF2DD4D9FABE8EFE" blockId="18.[151,1437,1086,2013]" box="[199,1294,1482,1509]" pageId="18" pageNumber="169">
<emphasis id="BF84EAE68B207360FF2DD4D9FEE98EFF" bold="true" box="[199,345,1482,1508]" pageId="18" pageNumber="169">Description.</emphasis>
Of moderate size for genus (SBL range
<quantity id="4A089B118B207360FCF4D4D8FC738EFE" box="[798,963,1483,1509]" metricMagnitude="-3" metricUnit="m" metricValue="1.79" metricValueMax="1.91" metricValueMin="1.67" pageId="18" pageNumber="169" unit="mm" value="1.79" valueMax="1.91" valueMin="1.67">1.671.91 mm</quantity>
, mean 1.79±
<quantity id="4A089B118B207360FBBAD4D8FB768EFE" box="[1104,1222,1483,1509]" metricMagnitude="-5" metricUnit="m" metricValue="7.6" pageId="18" pageNumber="169" unit="mm" value="0.076">0.076 mm</quantity>
, n=9).
</paragraph>
<paragraph id="8D4F36F48B207360FF2DD4FCFDAB8D4A" blockId="18.[151,1437,1086,2013]" pageId="18" pageNumber="169">
<emphasis id="BF84EAE68B207360FF2DD4FCFE978D12" box="[199,295,1519,1545]" italics="true" pageId="18" pageNumber="169">Habitus.</emphasis>
Body form (
<figureCitation id="15CB2A718B207360FE44D4FCFDB38D12" box="[430,515,1519,1545]" captionStart="FIGURE 9" captionStartId="15.[151,250,1649,1673]" captionTargetBox="[155,1436,181,1621]" captionTargetId="figure-15@15.[151,1436,181,1625]" captionTargetPageId="15" captionText="FIGURE 9. Digital images of habitus of New Zealand Nesamblyops species, dorsal aspect.A—N. canaanensis (Canaan, Nelson, SO), B—N. disjunctus (Fletchers Creek, Buller, SO), C—N. hobbit (9mi. N Karamea, Nelson, SO), D—N. karamea (Karamea River Gorge, Nelson, SO), E—N. kuscheli (Flora Hut, MountArthur, Nelson, SO), F—N. magnificus (Allison Reserve,Akatore, Dunedin, SO). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10184245" httpUri="https://zenodo.org/record/10184245/files/figure.png" pageId="18" pageNumber="169">Fig. 9D</figureCitation>
) markedly convex, elongate ovoid, general proportions wide (WE/SBL0.40±0.010), head narrow relative to pronotum (WH/WPm 0.68±0.015), proportions of pronotum in comparison to elytra average for genus (WPm/WE 0.75±0.013).
</paragraph>
<paragraph id="8D4F36F48B207360FF2DD748FCFD8D6E" blockId="18.[151,1437,1086,2013]" box="[199,845,1627,1653]" pageId="18" pageNumber="169">
<emphasis id="BF84EAE68B207360FF2DD748FEBD8D6E" box="[199,269,1627,1653]" italics="true" pageId="18" pageNumber="169">Color.</emphasis>
Body color brunneorufous, appendages testaceous.
</paragraph>
<paragraph id="8D4F36F48B207360FF2DD76CFA3B8C1E" blockId="18.[151,1437,1086,2013]" pageId="18" pageNumber="169">
<emphasis id="BF84EAE68B207360FF2DD76CFEF18D82" box="[199,321,1663,1689]" italics="true" pageId="18" pageNumber="169">Prothorax.</emphasis>
Pronotum (
<figureCitation id="15CB2A718B207360FE2FD76CFD9C8D82" box="[453,556,1663,1689]" captionStart="FIGURE 12" captionStartId="26.[151,250,1215,1239]" captionTargetBox="[157,1430,181,1192]" captionTargetId="figure-32@26.[157,1430,181,1192]" captionTargetPageId="26" captionText="FIGURE 12. Digital images of pronota of New Zealand Nesamblyops species, dorsal aspect. A—N. canaanensis (Canaan, Nelson, SO), B—N. disjunctus (Fletchers Creek, Buller, SO), C—N. hobbit (9mi. N Karamea, Nelson, SO), D—N. karamea (Karamea River Gorge, Nelson, SO), E—N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO), F—N. magnificus (Allison Reserve, Akatore, Dunedin, SO), G—N. montanus (Lake Sylvester, Kahurangi National Park, Nelson, SO), H—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), I—N. ovipennis (Mount Arthur, Nelson, SO). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10184251" httpUri="https://zenodo.org/record/10184251/files/figure.png" pageId="18" pageNumber="169">Fig. 12D</figureCitation>
) moderately long in comparison to elytra (LP/LE 0.42±0.010) and moderately transverse (WPm/LP 1.22±0.024), with lateral margins arcuately constricted posteriorly (WPm/WPp 1.31±0.049). Anterior angles indistinct, posterior angles obtuse (107119°), rounded. Width between posterior angles greater than between anterior angles (WPa/WPp 0.89±0.027). Basal margin almost rectilinear, slightly convex at middle.
</paragraph>
<paragraph id="8D4F36F48B207360FF2DD61CFD578C6A" blockId="18.[151,1437,1086,2013]" pageId="18" pageNumber="169">
<emphasis id="BF84EAE68B207360FF2DD61CFEA58C32" box="[199,277,1807,1833]" italics="true" pageId="18" pageNumber="169">Elytra.</emphasis>
Ovoid, narrowly depressed along suture, comparatively long (LE/SBL 0.59±0.007) and moderately wide (WE/LE 0.68±0.020). Humeri completely rounded. Lateral margins slightly divergent at basal half, subparallel at middle and evenly rounded to apex in apical third.
</paragraph>
<paragraph id="8D4F36F48B207361FF2DD668FE8D8A5A" blockId="18.[151,1437,1086,2013]" lastBlockId="19.[151,1437,151,681]" lastPageId="19" lastPageNumber="170" pageId="18" pageNumber="169">
<emphasis id="BF84EAE68B207360FF2DD668FEC58C8E" box="[199,373,1915,1941]" italics="true" pageId="18" pageNumber="169">Male genitalia.</emphasis>
Median lobe of aedeagus (
<figureCitation id="15CB2A718B207360FD46D668FCBF8C8E" box="[684,783,1915,1941]" captionStart="FIGURE 14" captionStartId="30.[151,250,1365,1389]" captionTargetBox="[157,1433,187,1336]" captionTargetId="figure-32@30.[151,1436,181,1341]" captionTargetPageId="30" captionText="FIGURE 14. Line drawings of male genitalia of New Zealand Nesamblyops species. N. canaanensis (Canaan, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. disjunctus (Fletchers Creek, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. hobbit (Mount Domett, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. karamea (Karamea River Gorge, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. magnificus (Allison Reserve, Akatore, Dunedin, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Legend: rCs—reversed C-contour; Vs—V-contour. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184255" httpUri="https://zenodo.org/record/10184255/files/figure.png" pageId="18" pageNumber="169">Fig. 14L</figureCitation>
) moderately arcuate and moderately twisted. Shaft short, convex dorsally, tapering in apical half. Apex moderately curved dorsally with rounded tip. Apical orifice long, occupies almost half of the shaft length. Ventral margin of median lobe straight. Walls of shaft without poriferous canals. Dorsal copulatory sclerites with well-developed rC-sclerite and with moderately long V-contour (
<figureCitation id="15CB2A718B217361FAC1D184FA3C8BAA" box="[1323,1420,151,177]" captionStart="FIGURE 14" captionStartId="30.[151,250,1365,1389]" captionTargetBox="[157,1433,187,1336]" captionTargetId="figure-32@30.[151,1436,181,1341]" captionTargetPageId="30" captionText="FIGURE 14. Line drawings of male genitalia of New Zealand Nesamblyops species. N. canaanensis (Canaan, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. disjunctus (Fletchers Creek, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. hobbit (Mount Domett, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. karamea (Karamea River Gorge, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. magnificus (Allison Reserve, Akatore, Dunedin, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Legend: rCs—reversed C-contour; Vs—V-contour. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184255" httpUri="https://zenodo.org/record/10184255/files/figure.png" pageId="19" pageNumber="170">Fig. 14L</figureCitation>
). Additionally, internal sac possesses curved narrowly sclerotized fold apico-ventrally from rC-sclerite. Left paramere (
<figureCitation id="15CB2A718B217361FF75D1CCFF4D8BE2" box="[159,253,223,249]" captionStart="FIGURE 14" captionStartId="30.[151,250,1365,1389]" captionTargetBox="[157,1433,187,1336]" captionTargetId="figure-32@30.[151,1436,181,1341]" captionTargetPageId="30" captionText="FIGURE 14. Line drawings of male genitalia of New Zealand Nesamblyops species. N. canaanensis (Canaan, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. disjunctus (Fletchers Creek, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. hobbit (Mount Domett, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. karamea (Karamea River Gorge, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. magnificus (Allison Reserve, Akatore, Dunedin, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Legend: rCs—reversed C-contour; Vs—V-contour. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184255" httpUri="https://zenodo.org/record/10184255/files/figure.png" pageId="19" pageNumber="170">Fig. 14J</figureCitation>
) comparatively narrow, with slightly tapering apex, bearing three long setae. Right paramere (
<figureCitation id="15CB2A718B217361FAC7D1CCFA238BE2" box="[1325,1427,223,249]" captionStart="FIGURE 14" captionStartId="30.[151,250,1365,1389]" captionTargetBox="[157,1433,187,1336]" captionTargetId="figure-32@30.[151,1436,181,1341]" captionTargetPageId="30" captionText="FIGURE 14. Line drawings of male genitalia of New Zealand Nesamblyops species. N. canaanensis (Canaan, Nelson, SO): A—left paramere, left lateral aspect, B—right paramere, right lateral aspect, C—median lobe, right lateral aspect. N. disjunctus (Fletchers Creek, Buller, SO): D—left paramere, left lateral aspect, E—right paramere, right lateral aspect, F—median lobe, right lateral aspect. N. hobbit (Mount Domett, Nelson, SO): G—left paramere, left lateral aspect, H—right paramere, right lateral aspect, I—median lobe, right lateral aspect. N. karamea (Karamea River Gorge, Nelson, SO): J—left paramere, left lateral aspect, K—right paramere, right lateral aspect, L—median lobe, right lateral aspect. N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO): M—left paramere, left lateral aspect, N—right paramere, right lateral aspect, O—median lobe, right lateral aspect. N. magnificus (Allison Reserve, Akatore, Dunedin, SO): P—left paramere, left lateral aspect, Q—right paramere, right lateral aspect, R—median lobe, right lateral aspect. Legend: rCs—reversed C-contour; Vs—V-contour. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10184255" httpUri="https://zenodo.org/record/10184255/files/figure.png" pageId="19" pageNumber="170">Fig. 14K</figureCitation>
) of moderate length, bearing three long setae, which are slightly shorter than the length of paramere. Ring sclerite as in
<figureCitation id="15CB2A718B217361FF38D034FE898A5A" box="[210,313,295,321]" captionStart="FIGURE 17" captionStartId="34.[151,250,1191,1215]" captionTargetBox="[163,1421,187,1168]" captionTargetId="figure-32@34.[151,1436,181,1168]" captionTargetPageId="34" captionText="FIGURE 17. Line drawings of ring sclerite of New Zealand Nesamblyops species, male genitalia, dorsal aspect. A—N. canaanensis (Canaan, Nelson, SO), B—N. disjunctus (Fletchers Creek, Buller, SO), C—N. hobbit (Mount Domett, Nelson, SO), D—N. karamea (Karamea River Gorge, Nelson, SO), E—N. kuscheli (Flora Hut, Mount Arthur, Nelson, SO), F—N. montanus (Relax Shelter, Mount Robert Ridge, Nelson, SO), G—N. moorei (Kangaroo Creek, Mawhera Forest, Buller, SO), H—N. ovipennis (Mount Arthur, Nelson, SO), I—N. rotundicollis (Onekaka, Nelson, SO), J—N. subcaecus (Margarets Tarn Lake, Arthurs Pass, Mid Canterbury, SO), K—N. viator (Secretary Island, Grono Bay, Fjordland, SO), L—N. victoriae (Capleston, Victoria Range, Buller, SO). Scale = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10184261" httpUri="https://zenodo.org/record/10184261/files/figure.png" pageId="19" pageNumber="170">Fig. 17D</figureCitation>
.
</paragraph>
<paragraph id="8D4F36F48B217361FF2DD058FD298A7E" blockId="19.[151,1437,151,681]" box="[199,665,331,357]" pageId="19" pageNumber="170">
<emphasis id="BF84EAE68B217361FF2DD058FE5D8A7E" box="[199,493,331,357]" italics="true" pageId="19" pageNumber="170">Female internal genitalia.</emphasis>
Not examined.
</paragraph>
<paragraph id="8D4F36F48B217361FF2DD07DFC758AB6" blockId="19.[151,1437,151,681]" pageId="19" pageNumber="170">
<emphasis id="BF84EAE68B217361FF2DD07DFDB58A93" bold="true" box="[199,517,366,392]" pageId="19" pageNumber="170">Geographical distribution.</emphasis>
The range of the species occupies the northern part of the
<collectingRegion id="4F34F8168B217361FB7ED07CFAA38A92" box="[1172,1299,367,393]" country="New Zealand" name="West Coast" pageId="19" pageNumber="170">West Coast</collectingRegion>
presumably between the Karamea and the Wanganui Rivers (
<figureCitation id="15CB2A718B217361FD53D080FCBB8AB6" box="[697,779,403,429]" captionStart="FIGURE 19" captionStartId="36.[151,250,1381,1405]" captionTargetBox="[151,1436,181,1357]" captionTargetId="figure-32@36.[151,1436,181,1357]" captionTargetPageId="36" captionText="FIGURE 19. Distributional records for the Nesamblyops species and images of male median lobe, obtained from specimens, collected in the line-connected with images localities. Green circles—N. canaanensis; black circles—N. hobbit; blue circles— N. ovipennis; yellow circles—N. karamea; red circles—N. kuscheli. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10184267" httpUri="https://zenodo.org/record/10184267/files/figure.png" pageId="19" pageNumber="170">Fig. 19</figureCitation>
, yellow circles).
</paragraph>
<paragraph id="8D4F36F48B217361FF2DD0A5FC868ACA" blockId="19.[151,1437,151,681]" box="[199,822,438,465]" pageId="19" pageNumber="170">
<emphasis id="BF84EAE68B217361FF2DD0A5FE9A8ACB" bold="true" box="[199,298,438,464]" pageId="19" pageNumber="170">Habitat.</emphasis>
Specimens were collected from litter samples.
</paragraph>
<paragraph id="8D4F36F48B217361FF2DD0C9FC0689B2" blockId="19.[151,1437,151,681]" pageId="19" pageNumber="170">
<emphasis id="BF84EAE68B217361FF2DD0C9FEC08AEF" bold="true" box="[199,368,474,500]" pageId="19" pageNumber="170">Relationships.</emphasis>
The structure of the male genitalia of
<taxonomicName id="4AF04D778B217361FCCDD0C8FC1B8AEE" authorityName="Sokolov" authorityYear="2023" box="[807,939,475,501]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="karamea">
<emphasis id="BF84EAE68B217361FCCDD0C8FC1B8AEE" box="[807,939,475,501]" italics="true" pageId="19" pageNumber="170">N. karamea</emphasis>
</taxonomicName>
suggests its relatedness to the other species with well-developed rC-sclerite and V-contour in the internal sac of the median lobe (cf. with the sclerites of
<taxonomicName id="4AF04D778B217361FA69D0ECFF408926" authorityName="Sokolov" authorityYear="2023" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="BF84EAE68B217361FA69D0ECFF408926" italics="true" pageId="19" pageNumber="170">N. carltoni</emphasis>
</taxonomicName>
,
<figureCitation id="15CB2A718B217361FF14D330FEDC8926" box="[254,364,546,573]" captionStart="FIGURE 5" captionStartId="9.[151,250,1796,1820]" captionTargetBox="[185,1401,181,1771]" captionTargetId="figure-15@9.[185,1401,181,1772]" captionTargetPageId="9" captionText="FIGURE 5. Digital illustrations of structural features of pterothorax of Anillini, ventral aspect: AD—subtribe Nesamblyopina; A—Nesamblyops oreobius (Broun); B—Nesamblyops subcaecus (Sharp); C—Nesamblyops brouni Sokolov; D—Nesamblyops viator n.sp.; E—Anillinus langdoni Sokolov and Carlton; F—Anillinus Cherokee Sokolov and Carlton; G—Geocharidius longinoi Sokolov and Kavanaugh; H—Geocharidius disjunctus Sokolov and Kavanaugh; I—Zapotecanillus longinoi Sokolov; J—Zeanillus pellucidus Sokolov; K—Zeanillus pallidus (Broun); L—Zeanillus nunni Sokolov; M—Pelodiaetodes insularis Sokolov; N—Pelodiaetodes nunni Sokolov; O—Pelodiaetodes moorei Sokolov; P—Pelodiaetus sulcatipennis Jeannel.Legend: mtes—metendosternite. Scales intentionally omitted." figureDoi="http://doi.org/10.5281/zenodo.10184235" httpUri="https://zenodo.org/record/10184235/files/figure.png" pageId="19" pageNumber="170">Fig. 5F, p</figureCitation>
. 190,
<bibRefCitation id="E9614B058B217361FE5AD330FDE18926" author="Sokolov, I. M." box="[432,593,547,573]" pageId="19" pageNumber="170" pagination="179 - 201" refId="ref27417" refString="Sokolov, I. M. (2023) Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa, 5230 (2), 179 - 201. https: // doi. org / 10.11646 / zootaxa. 5230.2.3" type="journal article" year="2023">Sokolov 2023</bibRefCitation>
). However, some details in the internal sac such as a presence of additional curved sclerotized fold apico-ventrally point out on isolated position of
<taxonomicName id="4AF04D778B217361FC39D354FBE8897A" authorityName="Sokolov" authorityYear="2023" box="[979,1112,583,609]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="karamea">
<emphasis id="BF84EAE68B217361FC39D354FBE8897A" box="[979,1112,583,609]" italics="true" pageId="19" pageNumber="170">N. karamea</emphasis>
</taxonomicName>
within the genus. Taking in account the trisetose parameres and a shape of median lobe this species may have a remote relatedness to the group of species, which includes
<taxonomicName id="4AF04D778B217361FE2ED39CFDDF89B3" authorityName="Sokolov" authorityYear="2023" box="[452,623,655,680]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="canaanensis">
<emphasis id="BF84EAE68B217361FE2ED39CFDDF89B3" box="[452,623,655,680]" italics="true" pageId="19" pageNumber="170">N. canaanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4AF04D778B217361FD97D39CFD5689B2" authorityName="Sokolov" authorityYear="2023" box="[637,742,655,681]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="hobbit">
<emphasis id="BF84EAE68B217361FD97D39CFD5689B2" box="[637,742,655,681]" italics="true" pageId="19" pageNumber="170">N. hobbit</emphasis>
</taxonomicName>
, and
<taxonomicName id="4AF04D778B217361FCC9D39CFC1F89B3" authorityName="Sokolov" authorityYear="2023" box="[803,943,655,680]" class="Insecta" family="Carabidae" genus="Nesamblyops" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="ovipennis">
<emphasis id="BF84EAE68B217361FCC9D39CFC1F89B3" box="[803,943,655,680]" italics="true" pageId="19" pageNumber="170">N. ovipennis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>