509 lines
57 KiB
XML
509 lines
57 KiB
XML
<document id="F524AA4F9C80D30BFC9F757FA3B51522" ID-CLB-Dataset="24056" ID-DOI="10.1080/00222933.2019.1709669" ID-GBIF-Dataset="175ef2e7-06f6-4404-a6be-224dac1a06ec" ID-ISSN="1464-5262" ID-Zenodo-Dep="3666538" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581606429779" checkinUser="valdenar" docAuthor="Freitag, Hendrik, Molls, Christian, Bouma, Aglaia M., Garces, Jhoana M., Rossato, Marzia, Cosentino, Emanuela & Delledonne, Massimo" docDate="2019" docId="052D87A158741300FEB9AA7AFDE2FD60" docLanguage="en" docName="J_NAT_HIST.53.41–42.2593–2620.pdf.imf" docOrigin="Journal of Natural History 53 (41)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Grouvellinus luciaensis Freitag, Molls and Bouma 2019, sp. nov" docType="treatment" docVersion="13" lastPageNumber="2612" masterDocId="F914FFD958641314FF8EA961FFF3FFB2" masterDocTitle="Additional new species of Grouvellinus Champion 1923 (Insectaı Coleopteraı Elmidae) discovered by citizen scientists and DNA barcoded in the field applying a novel MinIONbased workflow" masterLastPageNumber="2620" masterPageNumber="2593" pageNumber="2608" updateTime="1698782781787" updateUser="ExternalLinkService">
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<mods:title id="C0AA567B4EC820A7853AE6D580D13C95">Additional new species of Grouvellinus Champion 1923 (Insectaı Coleopteraı Elmidae) discovered by citizen scientists and DNA barcoded in the field applying a novel MinIONbased workflow</mods:title>
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<mods:namePart id="7F7AD0C0EA68310095FE31A0A5E67DF0">Freitag, Hendrik</mods:namePart>
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<mods:namePart id="266EC6D4055CC908E15431A846818EA7">Molls, Christian</mods:namePart>
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<mods:namePart id="719F90E2488088210C882FA10C279055">Bouma, Aglaia M.</mods:namePart>
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<mods:roleTerm id="B453ABBE09FD00F731E15F3724956444">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="171A545BDAF5477CCAEDE60B2FDDD66E">Garces, Jhoana M.</mods:namePart>
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<mods:roleTerm id="3CEB30C507ADA40ECD15D2E826503FE7">Author</mods:roleTerm>
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<mods:namePart id="B7BB6CC0AF8F4C44C2596523F42CD880">Rossato, Marzia</mods:namePart>
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<mods:namePart id="4500A578CF3C32B5349DD7DAEC336D77">Cosentino, Emanuela</mods:namePart>
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<mods:roleTerm id="469306B310339D81EF595F88EBCCFE7F">Author</mods:roleTerm>
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<mods:namePart id="5BEBC4A1206E1247EE01A9C28FABCFDE">Delledonne, Massimo</mods:namePart>
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<mods:typeOfResource id="70B7C4D1E9B8D3B2AFB0C7675FB88C0D">text</mods:typeOfResource>
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<mods:title id="14D8E6E12B865ACDF8BB8E069BF2EB20">Journal of Natural History</mods:title>
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<mods:part id="B8C8B922E31A5F8FABB830CE88F3BBFD">
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<mods:date id="F802B1950167097C65E49C20044DE58D">2019</mods:date>
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<mods:number id="C8562C3A5C3EEA94956CE753C21971CB">2020-01-24</mods:number>
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<mods:number id="F58E56993966E6BDA01B55F9B3DA5E8A">53</mods:number>
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<mods:number id="490635DB4012DD8AF89810D321E73CD2">41</mods:number>
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<mods:start id="C8F2C4E50218D910DB19929A80799BFD">2593</mods:start>
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<mods:classification id="126CF4F341AC955E4D23081B11EB73CF">journal article</mods:classification>
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<mods:identifier id="338937A6AE0A6437DEBA353BD139947E" type="CLB-Dataset">24056</mods:identifier>
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<mods:identifier id="874B578561151CDFDA6D29D6DE308DBC" type="DOI">10.1080/00222933.2019.1709669</mods:identifier>
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<treatment id="052D87A158741300FEB9AA7AFDE2FD60" ID-DOI="http://doi.org/10.5281/zenodo.3671826" ID-GBIF-Taxon="162136040" ID-Zenodo-Dep="3671826" LSID="urn:lsid:plazi:treatment:052D87A158741300FEB9AA7AFDE2FD60" httpUri="http://treatment.plazi.org/id/052D87A158741300FEB9AA7AFDE2FD60" lastPageId="20" lastPageNumber="2612" pageId="16" pageNumber="2608">
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<subSubSection id="C59E653C58741304FEB9AA7AFCC7FCE4" pageId="16" pageNumber="2608" type="nomenclature">
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<paragraph id="8D3B36B758741304FEB9AA7AFC1EFC86" blockId="16.[311,1005,795,854]" box="[311,1005,795,820]" pageId="16" pageNumber="2608">
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<heading id="D67381DB58741304FEB9AA7AFC1AFC86" box="[311,1001,795,820]" pageId="16" pageNumber="2608">
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||
<taxonomicName id="4A844D3458741304FEB9AA7AFC74FC86" authority="Freitag, Molls and Bouma" authorityName="Freitag, Molls and Bouma" authorityYear="2019" box="[311,903,795,820]" class="Insecta" family="Elmidae" genus="Grouvellinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="2608" phylum="Arthropoda" rank="species" species="luciaensis" status="sp. nov.">
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<emphasis id="BFF0EAA558741304FEB9AA7AFDA3FC86" bold="true" box="[311,592,795,820]" italics="true" pageId="16" pageNumber="2608">Grouvellinus luciaensis</emphasis>
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Freitag, Molls and Bouma
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</taxonomicName>
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||
,
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<taxonomicNameLabel id="A4C357DE58741304FC1CAA7AFC1AFC86" box="[914,1001,795,820]" pageId="16" pageNumber="2608" rank="species">sp. nov</taxonomicNameLabel>
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</heading>
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||
.
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</paragraph>
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||
<paragraph id="8D3B36B758741304FE7EAA5CFCC7FCE4" blockId="16.[311,1005,795,854]" box="[496,820,829,854]" pageId="16" pageNumber="2608">
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<heading id="D67381DB58741304FE7EAA5CFCC7FCE4" box="[496,820,829,854]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="2608" reason="5">
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(
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||
<figureCitation id="15BF2A3258741304FE76AA5CFD90FCE4" box="[504,611,829,854]" captionStart="Figures 6–9" captionStartId="7.[160,235,1337,1360]" captionTargetBox="[163,1153,148,1289]" captionTargetId="figure@7.[163,1153,142,1306]" captionTargetPageId="7" captionText="Figures 6–9. Habitus of the Grouvellinus species collected from the Tawan Hills Park. 6: G. luciaensis, sp. nov. (paratype male from Lucia River); 7: G. borneensis, sp. nov. (paratype male from Tawau River environment); 8: G. sp.1 (female from Tawau River environment); 9: G. sp.2 (female from Lucia River). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3666544" httpUri="https://zenodo.org/record/3666544/files/figure.png" pageId="16" pageNumber="2608">Figures 6</figureCitation>
|
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,
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<figureCitation id="15BF2A3258741304FDFFAA5CFD7DFCE4" box="[625,654,829,854]" captionStart="Figure 14" captionStartId="18.[160,225,1168,1191]" captionTargetBox="[163,1153,142,1137]" captionTargetId="figure@18.[163,1153,142,1137]" captionTargetPageId="18" captionText="Figure 14. Grouvellinus luciaensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum; (b) anterioventral aspect with head and prosternum; (c) ventral aspect with prosternum and mesoventrite, (d) posterioventral aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666554" httpUri="https://zenodo.org/record/3666554/files/figure.png" pageId="16" pageNumber="2608">14</figureCitation>
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(a
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<emphasis id="BFF0EAA558741304FD2DAA5CFD42FCE4" box="[675,689,829,854]" italics="true" pageId="16" pageNumber="2608">–</emphasis>
|
||
d), 15(a
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<emphasis id="BFF0EAA558741304FC87AA5CFCE4FCE4" box="[777,791,829,854]" italics="true" pageId="16" pageNumber="2608">–</emphasis>
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||
c))
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</heading>
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C59E653C58741304FF2EAAFBFD54FC67" pageId="16" pageNumber="2608" type="distribution">
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||
<paragraph id="8D3B36B758741304FF2EAAFBFD54FC67" blockId="16.[160,1155,922,981]" pageId="16" pageNumber="2608">
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||
<emphasis id="BFF0EAA558741304FF2EAAFBFEB0FC01" bold="true" box="[160,323,922,947]" italics="true" pageId="16" pageNumber="2608">
|
||
<typeStatus id="523F881558741304FF2EAAFBFF2BFC01" box="[160,216,922,947]" pageId="16" pageNumber="2608">Type</typeStatus>
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||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="F593762758741304FEDCAAFAFE47FC01" box="[338,436,923,947]" name="Malaysia" pageId="16" pageNumber="2608">Malaysia</collectingCountry>
|
||
,
|
||
<collectingRegion id="4F40F85558741304FE4BAAFAFDF9FC01" box="[453,522,923,947]" country="Malaysia" name="Sabah" pageId="16" pageNumber="2608">Sabah</collectingRegion>
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||
(on Borneo Island), Tawau Hills Park, Lucia River,
|
||
<geoCoordinate id="E8B0507058741304FBE3AAFAFF08FC67" degrees="4" direction="north" minutes="28" orientation="latitude" pageId="16" pageNumber="2608" precision="15" seconds="34" value="4.476111">
|
||
4° 28
|
||
<emphasis id="BFF0EAA558741304FF35AADCFF32FC67" bold="true" box="[187,193,957,981]" italics="true" pageId="16" pageNumber="2608">ʹ</emphasis>
|
||
34
|
||
<emphasis id="BFF0EAA558741304FF53AADCFF1AFC67" bold="true" box="[221,233,957,981]" italics="true" pageId="16" pageNumber="2608">ʹʹ</emphasis>
|
||
N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E8B0507058741304FE89AADCFE63FC67" box="[263,400,957,981]" degrees="117" direction="east" minutes="55" orientation="longitude" pageId="16" pageNumber="2608" precision="15" seconds="38" value="117.92722">
|
||
117°55
|
||
<emphasis id="BFF0EAA558741304FEDAAADCFEA9FC67" bold="true" box="[340,346,957,981]" italics="true" pageId="16" pageNumber="2608">ʹ</emphasis>
|
||
38
|
||
<emphasis id="BFF0EAA558741304FEFBAADCFE72FC67" bold="true" box="[373,385,957,981]" italics="true" pageId="16" pageNumber="2608">ʹʹ</emphasis>
|
||
E
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||
</geoCoordinate>
|
||
,
|
||
<quantity id="4A7C9B5258741304FE15AADCFE12FC67" box="[411,481,957,981]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="16" pageNumber="2608" unit="m" value="750.0">750 m</quantity>
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||
a.s.l. (
|
||
<figureCitation id="15BF2A3258741304FDA8AADCFD8DFC67" box="[550,638,957,981]" captionStart="Figure 1" captionStartId="3.[160,225,1413,1436]" captionTargetBox="[163,1153,142,1381]" captionTargetId="figure@3.[163,1153,142,1381]" captionTargetPageId="3" captionText="Figure 1. (a) Co-authors and assistants of the Taxon Expedition to Tawau Hills Park, Sabah, Malaysian Borneo, performing microhabitat-specific sampling; (b) Co-authors during beetle collection at a blacklight trap installed in the vicinity of the Tawau River; (c) The first author (second from the right) instructing Taxon Expedition participants on the identification of riffle beetles; (d) the transportable molecular field laboratory at the Tawau Park Station; (e) Type locality of Grouvellinus borneensis, sp. nov., G. cruxniger, sp. nov., G. nigerquadratus, sp. nov., and collection site of G. sp. 1; (f) Type locality of Grouvellinus luciaensis, sp. nov., and collection site of G. andrekuipersi, G. nigerquadratus, sp. nov. and G. sp. 2." figureDoi="http://doi.org/10.5281/zenodo.3666540" httpUri="https://zenodo.org/record/3666540/files/figure.png" pageId="16" pageNumber="2608">Figure 1</figureCitation>
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||
(f)).
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C59E653C58741304FF2EAD69FBD4FAA1" pageId="16" pageNumber="2608" type="materials_examined">
|
||
<paragraph id="8D3B36B758741304FF2EAD69FBD4FAA1" blockId="16.[160,1157,1032,1299]" pageId="16" pageNumber="2608">
|
||
<emphasis id="BFF0EAA558741304FF2EAD69FEA2FB93" bold="true" box="[160,337,1032,1057]" italics="true" pageId="16" pageNumber="2608">Type material.</emphasis>
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||
<materialsCitation id="3DEC3CEA58741304FEEFAD68FB73FBD4" ID-GBIF-Occurrence="2571115426" collectorName="Taxon Expedition" country="Malaysia" elevation="750" latitude="4.476111" location="March" longLatPrecision="21" longitude="117.92722" municipality="Lucia River" pageId="16" pageNumber="2608" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
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||
<emphasis id="BFF0EAA558741304FEEFAD68FE3CFB93" bold="true" box="[353,463,1033,1057]" pageId="16" pageNumber="2608">
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<typeStatus id="523F881558741304FEEFAD68FE3CFB93" box="[353,463,1033,1057]" pageId="16" pageNumber="2608" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
3 (
|
||
<specimenCount id="9B82FD3E58741304FD8BAD68FDD5FB93" box="[517,550,1033,1057]" pageId="16" pageNumber="2608" type="generic">SP</specimenCount>
|
||
):
|
||
<emphasis id="BFF0EAA558741304FDB3AD69FDB0FB92" box="[573,579,1032,1056]" italics="true" pageId="16" pageNumber="2608">‘</emphasis>
|
||
<collectingCountry id="F593762758741304FDCCAD68FD49FB93" box="[578,698,1033,1057]" name="Malaysia" pageId="16" pageNumber="2608">MALAYSIA</collectingCountry>
|
||
:
|
||
<collectingRegion id="4F40F85558741304FD45AD68FCE0FB93" box="[715,787,1033,1057]" country="Malaysia" name="Sabah" pageId="16" pageNumber="2608">Sabah</collectingRegion>
|
||
:
|
||
<collectingMunicipality id="6D5FACCD58741304FCADAD68FC56FB93" box="[803,933,1033,1057]" pageId="16" pageNumber="2608">Lucia River</collectingMunicipality>
|
||
; rock surface, run; primary forest; ca.
|
||
<geoCoordinate id="E8B0507058741304FEFBAD4AFE14FBF1" box="[373,487,1067,1091]" degrees="4" direction="north" minutes="28" orientation="latitude" pageId="16" pageNumber="2608" precision="15" seconds="34" value="4.476111">
|
||
4°28
|
||
<emphasis id="BFF0EAA558741304FE29AD4AFE5EFBF1" bold="true" box="[423,429,1067,1091]" italics="true" pageId="16" pageNumber="2608">ʹ</emphasis>
|
||
34
|
||
<emphasis id="BFF0EAA558741304FE46AD4AFE27FBF1" bold="true" box="[456,468,1067,1091]" italics="true" pageId="16" pageNumber="2608">ʹʹ</emphasis>
|
||
N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E8B0507058741304FE7AAD4AFD8DFBF1" box="[500,638,1067,1091]" degrees="117" direction="east" minutes="55" orientation="longitude" pageId="16" pageNumber="2608" precision="15" seconds="38" value="117.92722">
|
||
117°55
|
||
<emphasis id="BFF0EAA558741304FDCFAD4AFDB4FBF1" bold="true" box="[577,583,1067,1091]" italics="true" pageId="16" pageNumber="2608">ʹ</emphasis>
|
||
38
|
||
<emphasis id="BFF0EAA558741304FDECAD4AFD9DFBF1" bold="true" box="[610,622,1067,1091]" italics="true" pageId="16" pageNumber="2608">ʹʹ</emphasis>
|
||
E
|
||
</geoCoordinate>
|
||
, ca.
|
||
<quantity id="4A7C9B5258741304FD29AD4AFD03FBF1" box="[679,752,1067,1091]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="16" pageNumber="2608" unit="m" value="750.0">
|
||
<elevation id="06A9D18458741304FD29AD4AFD03FBF1" box="[679,752,1067,1091]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="16" pageNumber="2608" unit="m" value="750.0">750 m</elevation>
|
||
</quantity>
|
||
a.s.l., leg.
|
||
<collectorName id="2071536158741304FCE9AD4AFBDFFBF1" box="[871,1068,1067,1091]" pageId="16" pageNumber="2608">Taxon Expedition</collectorName>
|
||
participants 10.
|
||
<location id="885B606C58741304FE8DAD2FFEB8FBD4" LSID="urn:lsid:plazi:treatment:052D87A158741300FEB9AA7AFDE2FD60:885B606C58741304FE8DAD2FFEB8FBD4" box="[259,331,1102,1126]" country="Malaysia" latitude="4.476111" longLatPrecision="21" longitude="117.92722" municipality="Lucia River" name="March" pageId="16" pageNumber="2608" stateProvince="Sabah">March</location>
|
||
.2018 (
|
||
<quantity id="4A7C9B5258741304FE19AD2FFE46FBD4" box="[407,437,1102,1126]" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" pageId="16" pageNumber="2608" unit="g" value="6.0">6g</quantity>
|
||
)M
|
||
<emphasis id="BFF0EAA558741304FE5CAD2FFE2BFBD4" box="[466,472,1102,1126]" italics="true" pageId="16" pageNumber="2608">’</emphasis>
|
||
, terminal parts of abdomen incl. aedeagus glued separately
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3DEC3CEA58741304FF2EAD10FCD5FB7C" ID-GBIF-Occurrence="2571115425" collectorName="Taxon Expedition" country="Malaysia" elevation="750" latitude="4.476111" location="March" longLatPrecision="21" longitude="117.92722" municipality="Lucia River" pageId="16" pageNumber="2608" specimenCount="13" specimenCount-female="11" stateProvince="Sabah" typeStatus="paratype">
|
||
<emphasis id="BFF0EAA558741304FF2EAD10FEE4FB3B" bold="true" box="[160,279,1137,1161]" pageId="16" pageNumber="2608">
|
||
<typeStatus id="523F881558741304FF2EAD10FEE4FB3B" box="[160,279,1137,1161]" pageId="16" pageNumber="2608" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9B82FD3E58741304FEA8AD10FEBBFB3A" box="[294,328,1136,1161]" pageId="16" pageNumber="2608" type="female">3♀</specimenCount>
|
||
(
|
||
<specimenCount id="9B82FD3E58741304FED8AD10FE86FB3B" box="[342,373,1137,1161]" pageId="16" pageNumber="2608" type="generic">SP</specimenCount>
|
||
) same data as holotype; 13 [H12],
|
||
<specimenCount id="9B82FD3E58741304FC83AD10FCDCFB3A" box="[781,815,1136,1161]" pageId="16" pageNumber="2608" type="female">2♀</specimenCount>
|
||
(
|
||
<specimenCount id="9B82FD3E58741304FCB0AD10FCAEFB3B" box="[830,861,1137,1161]" pageId="16" pageNumber="2608" type="generic">SP</specimenCount>
|
||
)
|
||
<emphasis id="BFF0EAA558741304FCE2AD11FC81FB3A" box="[876,882,1136,1160]" italics="true" pageId="16" pageNumber="2608">‘</emphasis>
|
||
<collectingCountry id="F593762758741304FCFCAD10FC19FB3B" box="[882,1002,1137,1161]" name="Malaysia" pageId="16" pageNumber="2608">MALAYSIA</collectingCountry>
|
||
:
|
||
<collectingRegion id="4F40F85558741304FC7BAD10FBCEFB3B" box="[1013,1085,1137,1161]" country="Malaysia" name="Sabah" pageId="16" pageNumber="2608">Sabah</collectingRegion>
|
||
:
|
||
<collectingMunicipality id="6D5FACCD58741304FBC7AD10FF29FB19" pageId="16" pageNumber="2608">Lucia River</collectingMunicipality>
|
||
; submerged wood, run; primary forest; ca.
|
||
<geoCoordinate id="E8B0507058741304FD44ADF2FCCFFB19" box="[714,828,1171,1195]" degrees="4" direction="north" minutes="28" orientation="latitude" pageId="16" pageNumber="2608" precision="15" seconds="34" value="4.476111">
|
||
4°28
|
||
<emphasis id="BFF0EAA558741304FD72ADF2FCF1FB19" bold="true" box="[764,770,1171,1195]" italics="true" pageId="16" pageNumber="2608">ʹ</emphasis>
|
||
34
|
||
<emphasis id="BFF0EAA558741304FC93ADF2FCDAFB19" bold="true" box="[797,809,1171,1195]" italics="true" pageId="16" pageNumber="2608">ʹʹ</emphasis>
|
||
N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E8B0507058741304FCC7ADF2FC20FB19" box="[841,979,1171,1195]" degrees="117" direction="east" minutes="55" orientation="longitude" pageId="16" pageNumber="2608" precision="15" seconds="38" value="117.92722">
|
||
117°55
|
||
<emphasis id="BFF0EAA558741304FC18ADF2FC6FFB19" bold="true" box="[918,924,1171,1195]" italics="true" pageId="16" pageNumber="2608">ʹ</emphasis>
|
||
38
|
||
<emphasis id="BFF0EAA558741304FC39ADF2FC30FB19" bold="true" box="[951,963,1171,1195]" italics="true" pageId="16" pageNumber="2608">ʹʹ</emphasis>
|
||
E
|
||
</geoCoordinate>
|
||
, ca.
|
||
<quantity id="4A7C9B5258741304FC72ADF2FBB5FB19" box="[1020,1094,1171,1195]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="16" pageNumber="2608" unit="m" value="750.0">
|
||
<elevation id="06A9D18458741304FC72ADF2FBB5FB19" box="[1020,1094,1171,1195]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="16" pageNumber="2608" unit="m" value="750.0">750 m</elevation>
|
||
</quantity>
|
||
a.s.l., leg.
|
||
<collectorName id="2071536158741304FF5AADD7FE6FFB7C" box="[212,412,1206,1230]" pageId="16" pageNumber="2608">Taxon Expedition</collectorName>
|
||
participants 10.
|
||
<location id="885B606C58741304FDDBADD7FD6EFB7C" LSID="urn:lsid:plazi:treatment:052D87A158741300FEB9AA7AFDE2FD60:885B606C58741304FDDBADD7FD6EFB7C" box="[597,669,1206,1230]" country="Malaysia" latitude="4.476111" longLatPrecision="21" longitude="117.92722" municipality="Lucia River" name="March" pageId="16" pageNumber="2608" stateProvince="Sabah">March</location>
|
||
.2018 (
|
||
<specimenCount id="9B82FD3E58741304FD63ADD7FCF7FB7C" box="[749,772,1206,1230]" pageId="16" pageNumber="2608" type="female">6f</specimenCount>
|
||
)M
|
||
<emphasis id="BFF0EAA558741304FCAFADD7FCD5FB7C" box="[801,806,1206,1230]" italics="true" pageId="16" pageNumber="2608">’</emphasis>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3DEC3CEA58741304FCB9ADD7FBD4FAA1" ID-GBIF-Occurrence="2571115427" collectorName="Taxon Expedition" country="Malaysia" elevation="750" latitude="4.476111" location="March" longLatPrecision="21" longitude="117.92722" municipality="Lucia River" pageId="16" pageNumber="2608" specimenCount="2" specimenCount-female="1" stateProvince="Sabah">
|
||
3
|
||
<specimenCount id="9B82FD3E58741304FCD9ADD7FC8AFB7C" box="[855,889,1206,1230]" pageId="16" pageNumber="2608" type="female">1♀</specimenCount>
|
||
[H22] (
|
||
<specimenCount id="9B82FD3E58741304FC5CADD7FC02FB7C" box="[978,1009,1206,1230]" pageId="16" pageNumber="2608" type="generic">SP</specimenCount>
|
||
)
|
||
<emphasis id="BFF0EAA558741304FB8DADD7FBFAFB7C" box="[1027,1033,1206,1230]" italics="true" pageId="16" pageNumber="2608">‘</emphasis>
|
||
<collectingCountry id="F593762758741304FB87ADD7FB72FB7C" box="[1033,1153,1206,1230]" name="Malaysia" pageId="16" pageNumber="2608">MALAYSIA</collectingCountry>
|
||
:
|
||
<collectingRegion id="4F40F85558741304FF2EADB8FF1BFB43" box="[160,232,1241,1265]" country="Malaysia" name="Sabah" pageId="16" pageNumber="2608">Sabah</collectingRegion>
|
||
:
|
||
<collectingMunicipality id="6D5FACCD58741304FF77ADB8FE8FFB43" box="[249,380,1241,1265]" pageId="16" pageNumber="2608">Lucia River</collectingMunicipality>
|
||
; submerged root packs, run; primary forest; ca.
|
||
<geoCoordinate id="E8B0507058741304FC40ADB8FBCCFB43" box="[974,1087,1240,1265]" degrees="4" direction="north" minutes="28" orientation="latitude" pageId="16" pageNumber="2608" precision="15" seconds="34" value="4.476111">
|
||
4°28
|
||
<emphasis id="BFF0EAA558741304FC71ADB9FBF6FB42" bold="true" box="[1023,1029,1240,1264]" italics="true" pageId="16" pageNumber="2608">ʹ</emphasis>
|
||
34
|
||
<emphasis id="BFF0EAA558741304FBAEADB9FBDFFB42" bold="true" box="[1056,1068,1240,1264]" italics="true" pageId="16" pageNumber="2608">ʹʹ</emphasis>
|
||
N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E8B0507058741304FBDCADB8FF0BFAA1" degrees="117" direction="east" minutes="55" orientation="longitude" pageId="16" pageNumber="2608" precision="15" seconds="38" value="117.92722">
|
||
117° 55
|
||
<emphasis id="BFF0EAA558741304FF35AD9AFF32FAA1" bold="true" box="[187,193,1275,1299]" italics="true" pageId="16" pageNumber="2608">ʹ</emphasis>
|
||
38
|
||
<emphasis id="BFF0EAA558741304FF53AD9AFF1AFAA1" bold="true" box="[221,233,1275,1299]" italics="true" pageId="16" pageNumber="2608">ʹʹ</emphasis>
|
||
E
|
||
</geoCoordinate>
|
||
, ca.
|
||
<quantity id="4A7C9B5258741304FEAEAD9AFE94FAA1" box="[288,359,1275,1299]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="16" pageNumber="2608" unit="m" value="750.0">
|
||
<elevation id="06A9D18458741304FEAEAD9AFE94FAA1" box="[288,359,1275,1299]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="16" pageNumber="2608" unit="m" value="750.0">750 m</elevation>
|
||
</quantity>
|
||
a.s.l., leg.
|
||
<collectorName id="2071536158741304FE54AD9AFD6EFAA1" box="[474,669,1275,1299]" pageId="16" pageNumber="2608">Taxon Expedition</collectorName>
|
||
participants 10.
|
||
<location id="885B606C58741304FCC3AD9AFC66FAA1" LSID="urn:lsid:plazi:treatment:052D87A158741300FEB9AA7AFDE2FD60:885B606C58741304FCC3AD9AFC66FAA1" box="[845,917,1275,1299]" country="Malaysia" latitude="4.476111" longLatPrecision="21" longitude="117.92722" municipality="Lucia River" name="March" pageId="16" pageNumber="2608" stateProvince="Sabah">March</location>
|
||
.2018 (6h)M
|
||
<emphasis id="BFF0EAA558741304FB95AD9AFBD2FAA1" box="[1051,1057,1275,1299]" italics="true" pageId="16" pageNumber="2608">’</emphasis>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C59E653C58741304FF2EAC27FF11FA16" pageId="16" pageNumber="2608" type="etymology">
|
||
<paragraph id="8D3B36B758741304FF2EAC27FEEEFAED" blockId="16.[160,1155,1350,1444]" box="[160,285,1350,1375]" pageId="16" pageNumber="2608">
|
||
<emphasis id="BFF0EAA558741304FF2EAC27FEEEFAED" bold="true" box="[160,285,1350,1375]" italics="true" pageId="16" pageNumber="2608">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758741304FF2EAC08FF11FA16" blockId="16.[160,1155,1350,1444]" pageId="16" pageNumber="2608">
|
||
The species is named in reference to its
|
||
<typeStatus id="523F881558741304FDD5AC08FD7EFA33" box="[603,653,1385,1409]" pageId="16" pageNumber="2608">type</typeStatus>
|
||
locality, Mt. Lucia and Lucia River originating there.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C59E653C58741300FF2EACB7FE72FDF5" lastPageId="20" lastPageNumber="2612" pageId="16" pageNumber="2608" type="description">
|
||
<paragraph id="8D3B36B758741304FF2EACB7FED5FA5D" blockId="16.[160,1155,1494,1727]" box="[160,294,1494,1519]" pageId="16" pageNumber="2608">
|
||
<emphasis id="BFF0EAA558741304FF2EACB7FED5FA5D" bold="true" box="[160,294,1494,1519]" italics="true" pageId="16" pageNumber="2608">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758741304FF2EAC9BFBADF987" blockId="16.[160,1155,1494,1727]" pageId="16" pageNumber="2608">
|
||
Body elongate obovate, males
|
||
<quantity id="4A7C9B5258741304FD84AC9BFD5DF9A0" box="[522,686,1530,1554]" metricMagnitude="-3" metricUnit="m" metricValue="1.935" metricValueMax="1.96" metricValueMin="1.91" pageId="16" pageNumber="2608" unit="mm" value="1.935" valueMax="1.96" valueMin="1.91">
|
||
1.91
|
||
<emphasis id="BFF0EAA558741304FDB6AC9BFDB6F9A0" box="[568,581,1530,1554]" italics="true" pageId="16" pageNumber="2608">–</emphasis>
|
||
1.96 mm
|
||
</quantity>
|
||
, females
|
||
<quantity id="4A7C9B5258741304FC93AC9BFC4DF9A0" box="[797,958,1530,1554]" metricMagnitude="-3" metricUnit="m" metricValue="2.005" metricValueMax="2.04" metricValueMin="1.97" pageId="16" pageNumber="2608" unit="mm" value="2.005" valueMax="2.04" valueMin="1.97">
|
||
1.97
|
||
<emphasis id="BFF0EAA558741304FCC5AC9BFCABF9A0" box="[843,856,1530,1554]" italics="true" pageId="16" pageNumber="2608">–</emphasis>
|
||
2.04 mm
|
||
</quantity>
|
||
long (CL); males
|
||
<quantity id="4A7C9B5258741304FF2EAF7CFEB2F987" box="[160,321,1564,1589]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="9.1" metricValueMin="8.9" pageId="16" pageNumber="2608" unit="mm" value="0.9" valueMax="0.91" valueMin="0.89">
|
||
0.89
|
||
<emphasis id="BFF0EAA558741304FF41AF7DFF2FF986" box="[207,220,1564,1588]" italics="true" pageId="16" pageNumber="2608">–</emphasis>
|
||
0.91 mm
|
||
</quantity>
|
||
, females ca.
|
||
<quantity id="4A7C9B5258741304FE41AF7CFDC2F987" box="[463,561,1565,1589]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" pageId="16" pageNumber="2608" unit="mm" value="0.96">0.96 mm</quantity>
|
||
wide (EW), ca. 2.1 times as long as wide (CL/EW).
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758741304FF4EAF5EFEDBF90D" blockId="16.[160,1155,1494,1727]" pageId="16" pageNumber="2608">
|
||
Dorsal colouration (
|
||
<figureCitation id="15BF2A3258741304FE2EAF5EFE0DF9E5" box="[416,510,1599,1623]" captionStart="Figures 6–9" captionStartId="7.[160,235,1337,1360]" captionTargetBox="[163,1153,148,1289]" captionTargetId="figure@7.[163,1153,142,1306]" captionTargetPageId="7" captionText="Figures 6–9. Habitus of the Grouvellinus species collected from the Tawan Hills Park. 6: G. luciaensis, sp. nov. (paratype male from Lucia River); 7: G. borneensis, sp. nov. (paratype male from Tawau River environment); 8: G. sp.1 (female from Tawau River environment); 9: G. sp.2 (female from Lucia River). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3666544" httpUri="https://zenodo.org/record/3666544/files/figure.png" pageId="16" pageNumber="2608">Figure 6</figureCitation>
|
||
) almost entirely dark brown to black; legs paler, at least distal half of femora and claws golden brown; maxillary and labial palps dark golden brown; pubescence shiny yellowish, rather inconspicuous. Ventral side (
|
||
<figureCitation id="15BF2A3258741304FC58AFE4FBB0F92F" box="[982,1091,1669,1693]" captionStart="Figure 14" captionStartId="18.[160,225,1168,1191]" captionTargetBox="[163,1153,142,1137]" captionTargetId="figure@18.[163,1153,142,1137]" captionTargetPageId="18" captionText="Figure 14. Grouvellinus luciaensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum; (b) anterioventral aspect with head and prosternum; (c) ventral aspect with prosternum and mesoventrite, (d) posterioventral aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666554" httpUri="https://zenodo.org/record/3666554/files/figure.png" pageId="16" pageNumber="2608">Figure 14</figureCitation>
|
||
(b
|
||
<emphasis id="BFF0EAA558741304FBD6AFE5FB96F92E" box="[1112,1125,1668,1692]" italics="true" pageId="16" pageNumber="2608">–</emphasis>
|
||
d)) dark brown.
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758751305FF4EA9EFFC01FEE6" blockId="17.[160,1156,142,1726]" pageId="17" pageNumber="2609">
|
||
Head (
|
||
<figureCitation id="15BF2A3258751305FE84A9EFFE80FF14" box="[266,371,142,166]" captionStart="Figure 14" captionStartId="18.[160,225,1168,1191]" captionTargetBox="[163,1153,142,1137]" captionTargetId="figure@18.[163,1153,142,1137]" captionTargetPageId="18" captionText="Figure 14. Grouvellinus luciaensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum; (b) anterioventral aspect with head and prosternum; (c) ventral aspect with prosternum and mesoventrite, (d) posterioventral aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666554" httpUri="https://zenodo.org/record/3666554/files/figure.png" pageId="17" pageNumber="2609">Figure 14</figureCitation>
|
||
(a))
|
||
<quantity id="4A7C9B5258751305FE11A9EFFDCEFF14" box="[415,573,142,166]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" metricValueMax="4.3" metricValueMin="3.9" pageId="17" pageNumber="2609" unit="mm" value="0.41" valueMax="0.43" valueMin="0.39">
|
||
0.39
|
||
<emphasis id="BFF0EAA558751305FE43A9EFFE29FF14" box="[461,474,142,166]" italics="true" pageId="17" pageNumber="2609">–</emphasis>
|
||
0.43 mm
|
||
</quantity>
|
||
wide (HW); ID
|
||
<quantity id="4A7C9B5258751305FD68A9EFFC7BFF14" box="[742,904,142,166]" metricMagnitude="-4" metricUnit="m" metricValue="2.15" metricValueMax="2.4" metricValueMin="1.9" pageId="17" pageNumber="2609" unit="mm" value="0.215" valueMax="0.24" valueMin="0.19">
|
||
0.19
|
||
<emphasis id="BFF0EAA558751305FC9BA9EFFCD1FF14" box="[789,802,142,166]" italics="true" pageId="17" pageNumber="2609">–</emphasis>
|
||
0.24 mm
|
||
</quantity>
|
||
; disc of frons, clypeus, and anterior and lateral areas of labrum sparsely pubescent; clypeus with two pairs of conspicuous sublateroanterior setae and entire frontal and lateral margins of labrum densely pubescent; punctures small; intervals glabrous; except for rugulouse frontal portion bordering eyes. Frontoclypeal suture straight, distinct. Eyes moderately large (eye diameter smaller than ID), slightly protruding. Antennae genus-typical.
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758751305FF4EA83FFBF4FD46" blockId="17.[160,1156,142,1726]" pageId="17" pageNumber="2609">
|
||
Pronotum (
|
||
<figureCitation id="15BF2A3258751305FEB0A83FFE55FEC4" box="[318,422,350,374]" captionStart="Figure 14" captionStartId="18.[160,225,1168,1191]" captionTargetBox="[163,1153,142,1137]" captionTargetId="figure@18.[163,1153,142,1137]" captionTargetPageId="18" captionText="Figure 14. Grouvellinus luciaensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum; (b) anterioventral aspect with head and prosternum; (c) ventral aspect with prosternum and mesoventrite, (d) posterioventral aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666554" httpUri="https://zenodo.org/record/3666554/files/figure.png" pageId="17" pageNumber="2609">Figure 14</figureCitation>
|
||
(a)) in males:
|
||
<quantity id="4A7C9B5258751305FDB6A83FFD24FEC4" box="[568,727,350,374]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" metricValueMax="5.4" metricValueMin="5.2" pageId="17" pageNumber="2609" unit="mm" value="0.53" valueMax="0.54" valueMin="0.52">
|
||
0.52
|
||
<emphasis id="BFF0EAA558751305FDE8A83FFD80FEC4" box="[614,627,350,374]" italics="true" pageId="17" pageNumber="2609">–</emphasis>
|
||
0.54 mm
|
||
</quantity>
|
||
, in females:
|
||
<quantity id="4A7C9B5258751305FCD3A83FFC09FEC4" box="[861,1018,350,374]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="5.699999999999999" metricValueMin="5.3" pageId="17" pageNumber="2609" unit="mm" value="0.55" valueMax="0.57" valueMin="0.53">
|
||
0.53
|
||
<emphasis id="BFF0EAA558751305FC02A83FFC6AFEC4" box="[908,921,350,374]" italics="true" pageId="17" pageNumber="2609">–</emphasis>
|
||
0.57 mm
|
||
</quantity>
|
||
long (PL); in males:
|
||
<quantity id="4A7C9B5258751305FF7CA8E0FE64FE2B" box="[242,407,385,409]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" metricValueMax="6.7" metricValueMin="6.5" pageId="17" pageNumber="2609" unit="mm" value="0.66" valueMax="0.67" valueMin="0.65">
|
||
0.65
|
||
<emphasis id="BFF0EAA558751305FEAFA8E0FEDDFE2B" box="[289,302,385,409]" italics="true" pageId="17" pageNumber="2609">–</emphasis>
|
||
0.67 mm
|
||
</quantity>
|
||
, in females:
|
||
<quantity id="4A7C9B5258751305FDA2A8E0FD3DFE2B" box="[556,718,385,409]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" metricValueMax="7.199999999999999" metricValueMin="6.6000000000000005" pageId="17" pageNumber="2609" unit="mm" value="0.69" valueMax="0.72" valueMin="0.66">
|
||
0.66
|
||
<emphasis id="BFF0EAA558751305FDD5A8E0FD9BFE2B" box="[603,616,385,409]" italics="true" pageId="17" pageNumber="2609">–</emphasis>
|
||
0.72 mm
|
||
</quantity>
|
||
wide (PW), wider than long (PL/PW), widest posterior 0
|
||
<emphasis id="BFF0EAA558751305FEF6A8C2FE76FE09" box="[376,389,419,443]" italics="true" pageId="17" pageNumber="2609">–</emphasis>
|
||
0.3, distinctly narrower than elytra, anteriorly attenuate; anterior margin slightly convex; pronotal disc moderately vaulted; entire pronotum moderately sparsely punctate; punctures small and shallowly impressed; median carina absent; with a pair of small and moderately deep posterior-median pits (anterior of the scutellum); sublateral carinae distinct and short; oblique impression moderately deep, narrow and glabrous, extending approx. anterior 0.35
|
||
<emphasis id="BFF0EAA558751305FDF4AB30FD74FDDB" box="[634,647,593,617]" italics="true" pageId="17" pageNumber="2609">–</emphasis>
|
||
0.65; laterobasal impression shallow, rugose; setae moderately short; interstices glabrous and flat. Hypomeron densely punctate, punctures large. Prosternum (
|
||
<figureCitation id="15BF2A3258751305FE7EABF7FDA8FD1C" box="[496,603,662,686]" captionStart="Figure 14" captionStartId="18.[160,225,1168,1191]" captionTargetBox="[163,1153,142,1137]" captionTargetId="figure@18.[163,1153,142,1137]" captionTargetPageId="18" captionText="Figure 14. Grouvellinus luciaensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum; (b) anterioventral aspect with head and prosternum; (c) ventral aspect with prosternum and mesoventrite, (d) posterioventral aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666554" httpUri="https://zenodo.org/record/3666554/files/figure.png" pageId="17" pageNumber="2609">Figure 14</figureCitation>
|
||
(b)) short; lateral portions with pubescence (plastron); median portion including process medially broadly impressed, slightly rugulose; prosternal process trapezoidal, slightly longer than wide, medially impressed.
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758751305FF4EAB9FFBECFB26" blockId="17.[160,1156,142,1726]" pageId="17" pageNumber="2609">
|
||
Scutellum (
|
||
<figureCitation id="15BF2A3258751305FEB2AB9FFE56FCA4" box="[316,421,766,790]" captionStart="Figure 14" captionStartId="18.[160,225,1168,1191]" captionTargetBox="[163,1153,142,1137]" captionTargetId="figure@18.[163,1153,142,1137]" captionTargetPageId="18" captionText="Figure 14. Grouvellinus luciaensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum; (b) anterioventral aspect with head and prosternum; (c) ventral aspect with prosternum and mesoventrite, (d) posterioventral aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666554" httpUri="https://zenodo.org/record/3666554/files/figure.png" pageId="17" pageNumber="2609">Figure 14</figureCitation>
|
||
(a)) subtriangular, flat, glabrous. Elytra (
|
||
<figureCitation id="15BF2A3258751305FCD9AB9FFC41FCA4" box="[855,946,766,790]" captionStart="Figures 6–9" captionStartId="7.[160,235,1337,1360]" captionTargetBox="[163,1153,148,1289]" captionTargetId="figure@7.[163,1153,142,1306]" captionTargetPageId="7" captionText="Figures 6–9. Habitus of the Grouvellinus species collected from the Tawan Hills Park. 6: G. luciaensis, sp. nov. (paratype male from Lucia River); 7: G. borneensis, sp. nov. (paratype male from Tawau River environment); 8: G. sp.1 (female from Tawau River environment); 9: G. sp.2 (female from Lucia River). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3666544" httpUri="https://zenodo.org/record/3666544/files/figure.png" pageId="17" pageNumber="2609">Figure 6</figureCitation>
|
||
) elongate, subparallel anterior 0.15
|
||
<emphasis id="BFF0EAA558751305FEEDAA40FE83FC8B" box="[355,368,801,825]" italics="true" pageId="17" pageNumber="2609">–</emphasis>
|
||
0.60, moderately convex dorsally, in males:
|
||
<quantity id="4A7C9B5258751305FCD0AA40FC0DFC8B" box="[862,1022,801,825]" metricMagnitude="-3" metricUnit="m" metricValue="1.4249999999999998" metricValueMax="1.44" metricValueMin="1.41" pageId="17" pageNumber="2609" unit="mm" value="1.4249999999999998" valueMax="1.44" valueMin="1.41">
|
||
1.41
|
||
<emphasis id="BFF0EAA558751305FC05AA40FC6BFC8B" box="[907,920,801,825]" italics="true" pageId="17" pageNumber="2609">–</emphasis>
|
||
1.44 mm
|
||
</quantity>
|
||
, in females:
|
||
<quantity id="4A7C9B5258751305FF2EAA25FEC9FCEE" box="[160,314,835,860]" metricMagnitude="-3" metricUnit="m" metricValue="1.46" metricValueMax="1.48" metricValueMin="1.44" pageId="17" pageNumber="2609" unit="mm" value="1.46" valueMax="1.48" valueMin="1.44">
|
||
1.44
|
||
<emphasis id="BFF0EAA558751305FF43AA22FF29FCE9" box="[205,218,835,859]" italics="true" pageId="17" pageNumber="2609">–</emphasis>
|
||
1.48 mm
|
||
</quantity>
|
||
long (EL), 1.5
|
||
<emphasis id="BFF0EAA558751305FE47AA22FE25FCE9" box="[457,470,835,859]" italics="true" pageId="17" pageNumber="2609">–</emphasis>
|
||
1.6 times as long as wide (EL/EW), widest in anterior half; apices narrow, distinctly separately rounded; with eight longitudinal, distinctly impressed rows of punctures (striae); stria 8 reaching only up to 0.4 posterior; punctures somewhat regularly arranged, distinctly larger (diameter larger than interstices) and very deeply impressed in anterior and lateral portions, increasingly shallower and smaller apicad; interstices glabrous to finely rugulouse; median elytral portion, especially along interval 1 with additional very small setiferous secondary punctures; pubescence inconspicuous, consisting of very small adpressed setae and moderately large erected setae; interval 8 evenly carinate, but not serrate; interval 7 very slightly carinate; interval 5 (and intervals
|
||
<quantity id="4A7C9B5258751305FCDFAD38FC67FBC3" box="[849,916,1113,1137]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="10.16" metricValueMin="5.08" pageId="17" pageNumber="2609" unit="in" value="3.0" valueMax="4.0" valueMin="2.0">
|
||
2
|
||
<emphasis id="BFF0EAA558751305FCD0AD38FC98FBC3" box="[862,875,1113,1137]" italics="true" pageId="17" pageNumber="2609">–</emphasis>
|
||
4 in
|
||
</quantity>
|
||
lesser extent) broadly elevated in subanterior portion; lateral elytral margin carinate with fine serration.
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758751305FF4EADFFFBC9FAAC" blockId="17.[160,1156,142,1726]" pageId="17" pageNumber="2609">
|
||
Mesoventrite (
|
||
<figureCitation id="15BF2A3258751305FEEAADFFFE22FB04" box="[356,465,1182,1206]" captionStart="Figure 14" captionStartId="18.[160,225,1168,1191]" captionTargetBox="[163,1153,142,1137]" captionTargetId="figure@18.[163,1153,142,1137]" captionTargetPageId="18" captionText="Figure 14. Grouvellinus luciaensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum; (b) anterioventral aspect with head and prosternum; (c) ventral aspect with prosternum and mesoventrite, (d) posterioventral aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666554" httpUri="https://zenodo.org/record/3666554/files/figure.png" pageId="17" pageNumber="2609">Figure 14</figureCitation>
|
||
(c)) with two pairs of shallow grooves behind procoxae and medially, respectively. Metaventrite (
|
||
<figureCitation id="15BF2A3258751305FDC1ADA0FD33FB6B" box="[591,704,1217,1241]" captionStart="Figure 14" captionStartId="18.[160,225,1168,1191]" captionTargetBox="[163,1153,142,1137]" captionTargetId="figure@18.[163,1153,142,1137]" captionTargetPageId="18" captionText="Figure 14. Grouvellinus luciaensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum; (b) anterioventral aspect with head and prosternum; (c) ventral aspect with prosternum and mesoventrite, (d) posterioventral aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666554" httpUri="https://zenodo.org/record/3666554/files/figure.png" pageId="17" pageNumber="2609">Figure 14</figureCitation>
|
||
(d)) moderately densely punctate, less densely at disc; longitudinal impression along entire median suture; lateral portions rather flat with inconspicuous plastron and a row of large punctures behind mesocoxae.
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758751305FF4EAC48FCDEFA7E" blockId="17.[160,1156,142,1726]" pageId="17" pageNumber="2609">
|
||
Ventrite 1 (
|
||
<figureCitation id="15BF2A3258751305FEB5AC48FE55FAF3" box="[315,422,1321,1345]" captionStart="Figure 14" captionStartId="18.[160,225,1168,1191]" captionTargetBox="[163,1153,142,1137]" captionTargetId="figure@18.[163,1153,142,1137]" captionTargetPageId="18" captionText="Figure 14. Grouvellinus luciaensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum; (b) anterioventral aspect with head and prosternum; (c) ventral aspect with prosternum and mesoventrite, (d) posterioventral aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666554" httpUri="https://zenodo.org/record/3666554/files/figure.png" pageId="17" pageNumber="2609">Figure 14</figureCitation>
|
||
(d)) with pair of longitudinal carinae bordering disc; disc glabrous, sparsely punctate; lateral portions inconspicously pubescent (plastron); broad lateral portions of ventrites 1
|
||
<emphasis id="BFF0EAA558751305FE28AC0FFE40FA34" box="[422,435,1390,1414]" italics="true" pageId="17" pageNumber="2609">–</emphasis>
|
||
4 and almost entire ventrite 5 (
|
||
<figureCitation id="15BF2A3258751305FCA7AC0FFC6BFA34" box="[809,920,1390,1414]" captionStart="Figure 14" captionStartId="18.[160,225,1168,1191]" captionTargetBox="[163,1153,142,1137]" captionTargetId="figure@18.[163,1153,142,1137]" captionTargetPageId="18" captionText="Figure 14. Grouvellinus luciaensis, sp. nov. (paratypes): (a) anteriodorsal aspect with pronotum; (b) anterioventral aspect with head and prosternum; (c) ventral aspect with prosternum and mesoventrite, (d) posterioventral aspect with metaventrite and ventrites 1–5." figureDoi="http://doi.org/10.5281/zenodo.3666554" httpUri="https://zenodo.org/record/3666554/files/figure.png" pageId="17" pageNumber="2609">Figure 14</figureCitation>
|
||
(d)) densely covered with rather inconspicuous micro-plastron; ventrite 5 densely covered with setiferous tubercles and with apical fringe of moderately long setae.
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758751305FF4EACB7FF13F90C" blockId="17.[160,1156,142,1726]" pageId="17" pageNumber="2609">
|
||
Legs (
|
||
<figureCitation id="15BF2A3258751305FF70ACB7FEA7FA5C" box="[254,340,1494,1518]" captionStart="Figures 6–9" captionStartId="7.[160,235,1337,1360]" captionTargetBox="[163,1153,148,1289]" captionTargetId="figure@7.[163,1153,142,1306]" captionTargetPageId="7" captionText="Figures 6–9. Habitus of the Grouvellinus species collected from the Tawan Hills Park. 6: G. luciaensis, sp. nov. (paratype male from Lucia River); 7: G. borneensis, sp. nov. (paratype male from Tawau River environment); 8: G. sp.1 (female from Tawau River environment); 9: G. sp.2 (female from Lucia River). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3666544" httpUri="https://zenodo.org/record/3666544/files/figure.png" pageId="17" pageNumber="2609">Figure 6</figureCitation>
|
||
) approximately as long as body; hindleg longest; tibia longer than tarsus and femur in all legs; proximal portion of mesofemur with loose fringe of moderately long trichoid setae at anterior-dorsal ridge (in both sexes); distal area of all tibiae with more or less densely covered with short trichoid setae; hind tibia additionally with loose longitudinal fringe of robust spine-like setae; coxae, trochanter, and particularly inner and posterior faces of femora and tibiae covered with conspicuous plastron setae. Legs not conspicuously varying between sexes.
|
||
</paragraph>
|
||
<caption id="D9FB663F58761306FF2EADF1FCB6FB50" ID-DOI="http://doi.org/10.5281/zenodo.3666554" ID-Zenodo-Dep="3666554" httpUri="https://zenodo.org/record/3666554/files/figure.png" pageId="18" pageNumber="2610" startId="18.[160,225,1168,1191]" targetBox="[163,1153,142,1137]" targetPageId="18">
|
||
<paragraph id="8D3B36B758761306FF2EADF1FCB6FB50" blockId="18.[160,1156,1168,1250]" pageId="18" pageNumber="2610">
|
||
<emphasis id="BFF0EAA558761306FF2EADF1FEFAFB15" bold="true" box="[160,265,1168,1191]" pageId="18" pageNumber="2610">Figure 14.</emphasis>
|
||
<taxonomicName id="4A844D3458761306FE9FADF1FE19FB15" authority="Freitag & Molls & Bouma & Garces & Rossato & Cosentino & Delledonne, 2019" authorityName="Freitag & Molls & Bouma & Garces & Rossato & Cosentino & Delledonne" authorityYear="2019" box="[273,490,1168,1191]" class="Insecta" family="Elmidae" genus="Grouvellinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="2610" phylum="Arthropoda" rank="species" species="luciaensis" status="sp. nov.">
|
||
<emphasis id="BFF0EAA558761306FE9FADF1FE19FB15" box="[273,490,1168,1191]" italics="true" pageId="18" pageNumber="2610">Grouvellinus luciaensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A4C357DE58761306FE76ADF1FDBFFB15" box="[504,588,1168,1191]" pageId="18" pageNumber="2610" rank="species">
|
||
<emphasis id="BFF0EAA558761306FE76ADF1FDB4FB15" bold="true" box="[504,583,1168,1191]" pageId="18" pageNumber="2610">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(paratypes): (a) anteriodorsal aspect with pronotum; (b) anterioventral aspect with head and prosternum; (c) ventral aspect with prosternum and mesoventrite, (d) posterioventral aspect with metaventrite and ventrites 1
|
||
<emphasis id="BFF0EAA558761306FCA8ADAAFCC0FB50" box="[806,819,1227,1250]" italics="true" pageId="18" pageNumber="2610">–</emphasis>
|
||
5.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8D3B36B758761306FF4EAC4BFE51F9C8" blockId="18.[160,1156,1322,1727]" pageId="18" pageNumber="2610">
|
||
Aedeagus (
|
||
<figureCitation id="15BF2A3258761306FECDAC4BFE47FAF0" box="[323,436,1322,1346]" captionStart="Figure 15" captionStartId="19.[160,225,1527,1550]" captionTargetBox="[163,1153,142,1496]" captionTargetId="figure@19.[163,1153,142,1496]" captionTargetPageId="19" captionText="Figure 15. Grouvellinus luciaensis, sp. nov. (paratypes): (a) aedeagus, lateral; (b) aedeagus, ventral; (c) ovipositor (half), ventral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3666556" httpUri="https://zenodo.org/record/3666556/files/figure.png" pageId="18" pageNumber="2610">Figure 15</figureCitation>
|
||
(a,b)) ca. 635 μm long, ca. 190 μm wide. Base reaching basal 0.38 of total aedeagus length. Median lobe ca. four times as long as wide, moderately overreaching parameres, abruptly tapering in apical third towards the slender conical, rounded apex. Ventral sac apically inflated overreaching median lobe and parameres, internally densely stippled and with dense submedian fringes of moderately long, thin spines; apically the spines get longer and reach the apical opening. Parameres apically moderately conical (in both, ventral and lateral view), usually with more than 50 trichoid setae in apical two-third, most of them at the outer ventral face; most apical two setae longest and inserted at the apex. Male sternite IX as in previous new species.
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758761307FF4EAFE4FCA7F90B" blockId="18.[160,1156,1322,1727]" lastBlockId="19.[160,1155,1627,1721]" lastPageId="19" lastPageNumber="2611" pageId="18" pageNumber="2610">
|
||
Ovipositor (
|
||
<figureCitation id="15BF2A3258761306FECBAFE4FE41F92F" box="[325,434,1669,1693]" captionStart="Figure 15" captionStartId="19.[160,225,1527,1550]" captionTargetBox="[163,1153,142,1496]" captionTargetId="figure@19.[163,1153,142,1496]" captionTargetPageId="19" captionText="Figure 15. Grouvellinus luciaensis, sp. nov. (paratypes): (a) aedeagus, lateral; (b) aedeagus, ventral; (c) ovipositor (half), ventral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3666556" httpUri="https://zenodo.org/record/3666556/files/figure.png" pageId="18" pageNumber="2610">Figure 15</figureCitation>
|
||
(c)) total length ca. 790 μm; stylus ca. 40 μm long, moderately bent outwards with three short sensilla; coxite ca. 375 μm long with scattered extremely short, acute setae and four hook-like bent sensilla, apically moderately broadened at outer margin, valvifer ca. 430 μm long, caudal portion slightly sclerotised and with very few scattered, extremely short, acute setae; fibula almost straight.
|
||
</paragraph>
|
||
<caption id="D9FB663F58771307FF2EAC96FD92F99E" ID-DOI="http://doi.org/10.5281/zenodo.3666556" ID-Zenodo-Dep="3666556" httpUri="https://zenodo.org/record/3666556/files/figure.png" pageId="19" pageNumber="2611" startId="19.[160,225,1527,1550]" targetBox="[163,1153,142,1496]" targetPageId="19">
|
||
<paragraph id="8D3B36B758771307FF2EAC96FD92F99E" blockId="19.[160,1155,1526,1580]" pageId="19" pageNumber="2611">
|
||
<emphasis id="BFF0EAA558771307FF2EAC96FEF5F9BC" bold="true" box="[160,262,1527,1550]" pageId="19" pageNumber="2611">Figure 15.</emphasis>
|
||
<taxonomicName id="4A844D3458771307FE83AC97FE10F9BF" authority="Freitag & Molls & Bouma & Garces & Rossato & Cosentino & Delledonne, 2019" authorityName="Freitag & Molls & Bouma & Garces & Rossato & Cosentino & Delledonne" authorityYear="2019" box="[269,483,1526,1549]" class="Insecta" family="Elmidae" genus="Grouvellinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="2611" phylum="Arthropoda" rank="species" species="luciaensis" status="sp. nov.">
|
||
<emphasis id="BFF0EAA558771307FE83AC97FE10F9BF" box="[269,483,1526,1549]" italics="true" pageId="19" pageNumber="2611">Grouvellinus luciaensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A4C357DE58771307FE60AC96FDB3F9BC" box="[494,576,1527,1550]" pageId="19" pageNumber="2611" rank="species">
|
||
<emphasis id="BFF0EAA558771307FE60AC96FDC8F9BC" bold="true" box="[494,571,1527,1550]" pageId="19" pageNumber="2611">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(paratypes): (a) aedeagus, lateral; (b) aedeagus, ventral; (c) ovipositor (half), ventral. Scale bar = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8D3B36B758701300FF2EA9EFFE77FF15" blockId="20.[160,995,142,202]" box="[160,388,142,167]" pageId="20" pageNumber="2612">
|
||
<emphasis id="BFF0EAA558701300FF2EA9EFFE77FF15" bold="true" box="[160,388,142,167]" italics="true" pageId="20" pageNumber="2612">Sexual dimorphism</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758701300FF2EA9D3FC10FF78" blockId="20.[160,995,142,202]" box="[160,995,178,202]" pageId="20" pageNumber="2612">Females with separately rounded and slightly more pointed elytral apices.</paragraph>
|
||
<paragraph id="8D3B36B758701300FF2EA997FE6EFEBD" blockId="20.[160,1155,246,583]" box="[160,413,246,271]" pageId="20" pageNumber="2612">
|
||
<emphasis id="BFF0EAA558701300FF2EA997FE6EFEBD" bold="true" box="[160,413,246,271]" italics="true" pageId="20" pageNumber="2612">Differential diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758701300FF2EA878FBB2FDB0" blockId="20.[160,1155,246,583]" pageId="20" pageNumber="2612">
|
||
<taxonomicName id="4A844D3458701300FF2EA878FE66FE80" authority="Freitag & Molls & Bouma & Garces & Rossato & Cosentino & Delledonne, 2019" authorityName="Freitag & Molls & Bouma & Garces & Rossato & Cosentino & Delledonne" authorityYear="2019" box="[160,405,281,306]" class="Insecta" family="Elmidae" genus="Grouvellinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="2612" phylum="Arthropoda" rank="species" species="luciaensis" status="sp. nov.">
|
||
<emphasis id="BFF0EAA558701300FF2EA878FE66FE80" box="[160,405,281,306]" italics="true" pageId="20" pageNumber="2612">Grouvellinus luciaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A4C357DE58701300FE12A87BFE01FE80" box="[412,498,282,306]" pageId="20" pageNumber="2612" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be distinguished from all known Bornean
|
||
<taxonomicName id="4A844D3458701300FC73A878FB70FE80" authorityName="Champion" authorityYear="1923" box="[1021,1155,281,306]" class="Insecta" family="Elmidae" genus="Grouvellinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="2612" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BFF0EAA558701300FC73A878FB70FE80" box="[1021,1155,281,306]" italics="true" pageId="20" pageNumber="2612">Grouvellinus</emphasis>
|
||
</taxonomicName>
|
||
spp. by its moderately large size (CL) of
|
||
<quantity id="4A7C9B5258701300FDEDA85DFCF2FEE6" box="[611,769,316,340]" metricMagnitude="-3" metricUnit="m" metricValue="1.975" metricValueMax="2.04" metricValueMin="1.91" pageId="20" pageNumber="2612" unit="mm" value="1.975" valueMax="2.04" valueMin="1.91">
|
||
1.91
|
||
<emphasis id="BFF0EAA558701300FD1FA85DFD6DFEE6" box="[657,670,316,340]" italics="true" pageId="20" pageNumber="2612">–</emphasis>
|
||
2.04 mm
|
||
</quantity>
|
||
(vs. ≤
|
||
<quantity id="4A7C9B5258701300FCCAA85DFC6FFEE6" box="[836,924,316,340]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="20" pageNumber="2612" unit="mm" value="1.9">1.9 mm</quantity>
|
||
, or ≥
|
||
<quantity id="4A7C9B5258701300FC5BA85DFBDFFEE6" box="[981,1068,316,340]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="20" pageNumber="2612" unit="mm" value="2.3">2.3 mm</quantity>
|
||
), somewhat straight conical elytral posterior portion, very deeply impressed elytral striae in anterior and lateral portions, and a non-serrate 8th elytral interval. Furthermore, it is recognisable by the slight and uniform reddish-brownish tinge of the body, comparably pale femora, the aedeagal median lobe that is abruptly tapering in apical third and the comparably broad parameres with more than 50 trichoid setae in apical two-third.
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758701300FF4EAB6DFE72FDF5" blockId="20.[160,1155,246,583]" pageId="20" pageNumber="2612">
|
||
The species varies by about 11
|
||
<emphasis id="BFF0EAA558701300FD95AB6DFDDBFD96" box="[539,552,524,548]" italics="true" pageId="20" pageNumber="2612">–</emphasis>
|
||
17% genetic distance (654 bp CO1 barcode) from the Bornean congeners.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C59E653C58701300FF2EAB15FDE2FD60" pageId="20" pageNumber="2612" type="distribution">
|
||
<paragraph id="8D3B36B758701300FF2EAB15FEDFFD3F" blockId="20.[160,1156,628,722]" box="[160,300,628,653]" pageId="20" pageNumber="2612">
|
||
<emphasis id="BFF0EAA558701300FF2EAB15FEDFFD3F" bold="true" box="[160,300,628,653]" italics="true" pageId="20" pageNumber="2612">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8D3B36B758701300FF2EABF6FDE2FD60" blockId="20.[160,1156,628,722]" pageId="20" pageNumber="2612">
|
||
This species is only known from the
|
||
<typeStatus id="523F881558701300FDC3ABF6FD8CFD1D" box="[589,639,663,687]" pageId="20" pageNumber="2612">type</typeStatus>
|
||
locality, upper Lucia River, Tawau Hills Park, Sabah, Borneo Island (
|
||
<figureCitation id="15BF2A3258701300FE15ABDBFDF7FD60" box="[411,516,698,722]" captionStart="Figure 18" captionStartId="25.[160,225,1675,1698]" captionTargetBox="[163,1153,977,1643]" captionTargetId="figure@25.[163,1153,977,1643]" captionTargetPageId="25" captionText="Figure 18. Map of Sabah and Brunei (Borneo) with collecting site of Grouvellinus species with new species record in red fonts." figureDoi="http://doi.org/10.5281/zenodo.3666562" httpUri="https://zenodo.org/record/3666562/files/figure.png" pageId="20" pageNumber="2612">Figure 18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |