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<document id="6E8CFAE2FCC760FFD77AD221D5296F70" ID-DOI="10.5281/zenodo.281437" ID-GBIF-Dataset="d2ba1c60-b8e6-4623-8231-d98150162598" ID-ISSN="1175-5326" ID-Zenodo-Dep="281437" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460488764929" checkinUser="plazi" docAuthor="Geiger, Daniel L. &amp; Marshall, Bruce A." docDate="2012" docId="D077A52CA330FFFAFF18FF0161AB18CF" docLanguage="en" docName="zt03344p033.pdf" docOrigin="Zootaxa 3344" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Anatoma megascutula Geiger &amp; Marshall, 2012, new species" docType="treatment" docVersion="7" lastPageNumber="26" masterDocId="2C4EDD54A327FFE3FF8FFF9661441F76" masterDocTitle="New species of Scissurellidae, Anatomidae, and Larocheidae (Mollusca: Gastropoda: Vetigastropoda) from New Zealand and beyond" masterLastPageNumber="33" masterPageNumber="1" pageNumber="24" updateTime="1698309732748" updateUser="plazi">
<mods:mods id="CE38FAA6715C8C75AA49B81394E879DB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="BC4CEBFCF2ADFD717D64A1EBDEFF25DE">New species of Scissurellidae, Anatomidae, and Larocheidae (Mollusca: Gastropoda: Vetigastropoda) from New Zealand and beyond</mods:title>
</mods:titleInfo>
<mods:name id="1DECAEF29B736B79CD49578EFC51562C" type="personal">
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<mods:namePart id="A80B275D4A048A018E66EB9BC506CB30">Geiger, Daniel L.</mods:namePart>
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<mods:name id="A2CF2BCBC31ACF6D62C28C037033A1DC" type="personal">
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<mods:namePart id="D2E72F0E7ADC5E40686F082B1E791C2D">Marshall, Bruce A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="469658E09F1F156AEEB8985FCDFB8312">text</mods:typeOfResource>
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<mods:title id="A970D5E4AD8B67635ED8F17C0F7F4842">Zootaxa</mods:title>
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<mods:part id="BA8CCD90BDE98BF11929ABFD24773D9F">
<mods:date id="C75021BC9E95D43CE75864349CDAF931">2012</mods:date>
<mods:detail id="1ED5E6A6082088CA63845E693EC46DA3" type="volume">
<mods:number id="673F3A52CABA4CFD237B597CE826EC25">3344</mods:number>
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<mods:start id="4AAAAF3FB21B15287BB9CEE02B75B61E">1</mods:start>
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<mods:classification id="4AD1B63C43C202271E288517C96B5315">journal article</mods:classification>
<mods:identifier id="F57AB635E185D990D29DED14F7BD0E38" type="DOI">10.5281/zenodo.281437</mods:identifier>
<mods:identifier id="21CBD51380BC6AF2E0FDD22C46A3CF05" type="GBIF-Dataset">d2ba1c60-b8e6-4623-8231-d98150162598</mods:identifier>
<mods:identifier id="696B8F606C8F9117C42EA7D9B6378772" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BE5E1FCA58238F9E09228825973F8119" type="Zenodo-Dep">281437</mods:identifier>
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<treatment id="D077A52CA330FFFAFF18FF0161AB18CF" ID-DOI="http://doi.org/10.5281/zenodo.6166692" ID-GBIF-Taxon="119519060" ID-Zenodo-Dep="6166692" LSID="urn:lsid:zoobank.org:act:A749E8F1-B2AB-4022-A387-873D58EEB368" httpUri="http://treatment.plazi.org/id/D077A52CA330FFFAFF18FF0161AB18CF" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
<subSubSection id="10C447B1A330FFF4FF18FF01600E1FA4" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="5861143AA330FFF4FF18FF0163791FC7" blockId="23.[151,573,151,210]" box="[151,573,151,177]" pageId="23" pageNumber="24">
<heading id="0329A356A330FFF4FF18FF0163791FC7" bold="true" box="[151,573,151,177]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<emphasis id="6AAAC828A330FFF4FF18FF0163791FC7" bold="true" box="[151,573,151,177]" pageId="23" pageNumber="24">
<taxonomicName id="9FDE6FB9A330FFF4FF18FF0160E21FC7" ID-CoL="DJCH" box="[151,422,151,177]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="megascutula" status="sp. nov.">
<emphasis id="6AAAC828A330FFF4FF18FF0160E21FC7" bold="true" box="[151,422,151,177]" italics="true" pageId="23" pageNumber="24">Anatoma megascutula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71997553A330FFF4FE22FF0163791FC7" box="[429,573,151,177]" pageId="23" pageNumber="24" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="5861143AA330FFF4FF18FF2C600E1FA4" blockId="23.[151,573,151,210]" box="[151,330,186,210]" pageId="23" pageNumber="24">
(
<figureCitation id="C0E508BFA330FFF4FF11FF2C60071FA4" box="[158,323,186,210]" captionStart-0="FIGURE 18" captionStart-1="FIGURE 19" captionStart-2="FIGURE 20" captionStartId-0="23.[151,250,1486,1509]" captionStartId-1="24.[151,250,1555,1578]" captionStartId-2="25.[151,250,709,732]" captionTargetBox-0="[151,1436,541,1472]" captionTargetBox-1="[151,1435,197,1534]" captionTargetBox-2="[151,1436,199,693]" captionTargetId-0="figure@23.[151,1436,541,1473]" captionTargetId-1="figure@24.[151,1436,197,1535]" captionTargetId-2="figure@25.[151,1436,199,693]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionTargetPageId-2="25" captionText-0="FIGURE 18. Anatoma megascutula new species. Holotype. 800 – 850 m, east of Rapa, Austral Islands, French Polynesia, 27.618 ˚ S, 144.257 ˚ W (MNHN 24954). Scale bar shell = 1 mm. Scale bar protoconch = 100 µm." captionText-1="FIGURE 19. Anatoma megascutula new species. Paratypes. A. Norfolk Ridge, New Caledonia, 820 – 1220 m (MNHN 24959). B. Tongatapu, Tonga, 456 – 510 m (MNHN 24955). C. Fiji, 820 – 863 m (MNHN 24956). Scale bars shell = 1 mm. Scale bars protoconch = 100 µm." captionText-2="FIGURE 20. Anatoma megascutula new species. Radula. Porthos Bank, Norfolk Ridge, New Caledonia, 680 – 980 m (MNHN). A. Central field. B. Marginal teeth. Scale bars = 100 µm." httpUri-0="https://zenodo.org/record/281453/files/figure.png" httpUri-1="https://zenodo.org/record/281454/files/figure.png" httpUri-2="https://zenodo.org/record/281455/files/figure.png" pageId="23" pageNumber="24">Figures 18–20</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="10C447B1A330FFF4FF18FE9163061ED9" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="5861143AA330FFF4FF18FE9165AD1E11" blockId="23.[151,1436,262,503]" pageId="23" pageNumber="24">
<emphasis id="6AAAC828A330FFF4FF18FE91600E1E69" bold="true" box="[151,330,262,287]" pageId="23" pageNumber="24">
<typeStatus id="8765AA98A330FFF4FF18FE9161941E69" box="[151,208,263,287]" pageId="23" pageNumber="24">Type</typeStatus>
material.
</emphasis>
<typeStatus id="8765AA98A330FFF4FED4FE9160871E69" box="[347,451,263,287]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
(
<collectionCode id="3ECF8CFFA330FFF4FE56FE9163751E69" LSID="urn:lsid:biocol.org:col:34988" box="[473,561,263,287]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="24">MNHN</collectionCode>
24954:
<figureCitation id="C0E508BFA330FFF4FD12FE9163AB1E69" box="[669,751,263,287]" captionStart="FIGURE 18" captionStartId="23.[151,250,1486,1509]" captionTargetBox="[151,1436,541,1472]" captionTargetId="figure@23.[151,1436,541,1473]" captionTargetPageId="23" captionText="FIGURE 18. Anatoma megascutula new species. Holotype. 800 – 850 m, east of Rapa, Austral Islands, French Polynesia, 27.618 ˚ S, 144.257 ˚ W (MNHN 24954). Scale bar shell = 1 mm. Scale bar protoconch = 100 µm." httpUri="https://zenodo.org/record/281453/files/figure.png" pageId="23" pageNumber="24">Fig. 18</figureCitation>
). 5.03 ×
<quantity id="9F26B9DFA330FFF4FCE5FE91629C1E69" box="[874,984,263,287]" metricMagnitude="-3" metricUnit="m" metricValue="3.36" pageId="23" pageNumber="24" unit="mm" value="3.36">3.36 mm</quantity>
.
<typeStatus id="8765AA98A330FFF4FC64FE9E65181E56" box="[1003,1116,264,288]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="20C954AAA330FFF4FBE1FE9165831E69" box="[1134,1223,263,287]" name="Norfolk Island" pageId="23" pageNumber="24">Norfolk</collectingCountry>
Ridge, 24.867˚S, 168.183˚E,
<quantity id="9F26B9DFA330FFF4FEAEFEBD60EA1E35" box="[289,430,299,323]" metricMagnitude="3" metricUnit="m" metricValue="1.02" metricValueMax="1.22" metricValueMin="0.82" pageId="23" pageNumber="24" unit="m" value="1020.0" valueMax="1220.0" valueMin="820.0">820–1220 m</quantity>
(
<collectionCode id="3ECF8CFFA330FFF4FE4BFEBD63581E35" LSID="urn:lsid:biocol.org:col:34988" box="[452,540,299,323]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="24">MNHN</collectionCode>
24959, 1:
<figureCitation id="C0E508BFA330FFF4FD27FEBD63BD1E35" box="[680,761,299,323]" captionStart="FIGURE 19" captionStartId="24.[151,250,1555,1578]" captionTargetBox="[151,1435,197,1534]" captionTargetId="figure@24.[151,1436,197,1535]" captionTargetPageId="24" captionText="FIGURE 19. Anatoma megascutula new species. Paratypes. A. Norfolk Ridge, New Caledonia, 820 – 1220 m (MNHN 24959). B. Tongatapu, Tonga, 456 – 510 m (MNHN 24955). C. Fiji, 820 – 863 m (MNHN 24956). Scale bars shell = 1 mm. Scale bars protoconch = 100 µm." httpUri="https://zenodo.org/record/281454/files/figure.png" pageId="23" pageNumber="24">Fig. 19</figureCitation>
A); Tongatapu,
<collectingCountry id="20C954AAA330FFF4FC39FEBA62BA1E32" box="[950,1022,300,324]" name="Tonga" pageId="23" pageNumber="24">Tonga</collectingCountry>
, 21.083˚S, 175.367˚W,
<quantity id="9F26B9DFA330FFF4FA92FEBD64D81E35" box="[1309,1436,299,323]" metricMagnitude="2" metricUnit="m" metricValue="4.83" metricValueMax="5.1" metricValueMin="4.56" pageId="23" pageNumber="24" unit="m" value="483.0" valueMax="510.0" valueMin="456.0">456–510 m</quantity>
(
<collectionCode id="3ECF8CFFA330FFF4FF11FED961B21E11" LSID="urn:lsid:biocol.org:col:34988" box="[158,246,335,359]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="24">MNHN</collectionCode>
24955, 1:
<figureCitation id="C0E508BFA330FFF4FEE2FED960FB1E11" box="[365,447,335,359]" captionStart="FIGURE 19" captionStartId="24.[151,250,1555,1578]" captionTargetBox="[151,1435,197,1534]" captionTargetId="figure@24.[151,1436,197,1535]" captionTargetPageId="24" captionText="FIGURE 19. Anatoma megascutula new species. Paratypes. A. Norfolk Ridge, New Caledonia, 820 – 1220 m (MNHN 24959). B. Tongatapu, Tonga, 456 – 510 m (MNHN 24955). C. Fiji, 820 – 863 m (MNHN 24956). Scale bars shell = 1 mm. Scale bars protoconch = 100 µm." httpUri="https://zenodo.org/record/281454/files/figure.png" pageId="23" pageNumber="24">Fig. 19</figureCitation>
B);
<collectingCountry id="20C954AAA330FFF4FE67FED963561E11" box="[488,530,335,359]" name="Fiji" pageId="23" pageNumber="24">Fiji</collectingCountry>
, 17.300˚S, 179.550˚W,
<quantity id="9F26B9DFA330FFF4FC94FED962DD1E11" box="[795,921,335,359]" metricMagnitude="2" metricUnit="m" metricValue="8.415" metricValueMax="8.629999999999999" metricValueMin="8.2" pageId="23" pageNumber="24" unit="m" value="841.5" valueMax="863.0" valueMin="820.0">820–863 m</quantity>
(
<collectionCode id="3ECF8CFFA330FFF4FC27FED965441E11" LSID="urn:lsid:biocol.org:col:34988" box="[936,1024,335,359]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="24">MNHN</collectionCode>
24956, 1:
<figureCitation id="C0E508BFA330FFF4FBF8FED965831E11" box="[1143,1223,335,359]" captionStart="FIGURE 19" captionStartId="24.[151,250,1555,1578]" captionTargetBox="[151,1435,197,1534]" captionTargetId="figure@24.[151,1436,197,1535]" captionTargetPageId="24" captionText="FIGURE 19. Anatoma megascutula new species. Paratypes. A. Norfolk Ridge, New Caledonia, 820 – 1220 m (MNHN 24959). B. Tongatapu, Tonga, 456 – 510 m (MNHN 24955). C. Fiji, 820 – 863 m (MNHN 24956). Scale bars shell = 1 mm. Scale bars protoconch = 100 µm." httpUri="https://zenodo.org/record/281454/files/figure.png" pageId="23" pageNumber="24">Fig. 19</figureCitation>
C).
</paragraph>
<paragraph id="5861143AA330FFF4FF48FEE263061ED9" blockId="23.[151,1436,262,503]" pageId="23" pageNumber="24">
<emphasis id="6AAAC828A330FFF4FF48FEE260221EFA" bold="true" box="[199,358,371,396]" pageId="23" pageNumber="24">
<typeStatus id="8765AA98A330FFF4FF48FEE260461EFA" box="[199,258,372,396]" pageId="23" pageNumber="24">Type</typeStatus>
locality.
</emphasis>
E of Rapa, Austral Islands,
<collectingCountry id="20C954AAA330FFF4FD26FEE5622B1EFD" box="[681,879,371,395]" name="French Polynesia" pageId="23" pageNumber="24">French Polynesia</collectingCountry>
, 27.618˚S, 144.257˚W,
<quantity id="9F26B9DFA330FFF4FBF4FEE565B81EFD" box="[1147,1276,371,395]" metricMagnitude="2" metricUnit="m" metricValue="8.25" metricValueMax="8.5" metricValueMin="8.0" pageId="23" pageNumber="24" unit="m" value="825.0" valueMax="850.0" valueMin="800.0">800–850 m</quantity>
,
<date id="2C6032FAA330FFF4FA86FEE264DC1EFD" box="[1289,1432,371,395]" pageId="23" pageNumber="24" value="2002-11-07">7 Nov. 2002</date>
, N.
<collectingCountry id="20C954AAA330FFF4FF3CFE01618E1ED9" box="[179,202,407,431]" name="Iceland" pageId="23" pageNumber="24">O</collectingCountry>
. Alis (BENTHAUS stn CP1891).
</paragraph>
</subSubSection>
<subSubSection id="10C447B1A330FFF4FF48FE2A63E31E81" pageId="23" pageNumber="24" type="etymology">
<paragraph id="5861143AA330FFF4FF48FE2A63E31E81" blockId="23.[151,1436,262,503]" pageId="23" pageNumber="24">
<emphasis id="6AAAC828A330FFF4FF48FE2A600A1EA2" bold="true" box="[199,334,444,468]" pageId="23" pageNumber="24">Etymology.</emphasis>
Mega-, Greek prefix for large; -scutula, Latin for saucer, shallow bowl, referring to the large size of this flat-shelled species. Noun in apposition.
</paragraph>
</subSubSection>
<caption id="0CA144B2A330FFF4FF18FA5865201975" httpUri="https://zenodo.org/record/281453/files/figure.png" pageId="23" pageNumber="24" targetBox="[151,1436,541,1472]" targetPageId="23">
<paragraph id="5861143AA330FFF4FF18FA5865201975" blockId="23.[151,1436,1486,1539]" pageId="23" pageNumber="24">
<emphasis id="6AAAC828A330FFF4FF18FA5860671A95" bold="true" box="[151,291,1486,1509]" pageId="23" pageNumber="24">FIGURE 18.</emphasis>
<taxonomicName id="9FDE6FB9A330FFF4FEA1FA5963501A92" box="[302,532,1486,1508]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="megascutula" status="sp. nov.">
<emphasis id="6AAAC828A330FFF4FEA1FA5963501A92" box="[302,532,1486,1508]" italics="true" pageId="23" pageNumber="24">Anatoma megascutula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71997553A330FFF4FD91FA5963DB1A92" box="[542,671,1486,1508]" pageId="23" pageNumber="24" rank="species">new species</taxonomicNameLabel>
. Holotype. 800–850 m, east of Rapa, Austral Islands, French Polynesia, 27.618˚S, 144.257˚W (MNHN 24954). Scale bar shell = 1 mm. Scale bar protoconch = 100 µm.
</paragraph>
</caption>
<subSubSection id="10C447B1A330FFFAFF48F9BE60E31BF4" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24" type="description">
<paragraph id="5861143AA330FFF4FF48F9BE60391887" blockId="23.[151,1437,1576,2033]" pageId="23" pageNumber="24">
<emphasis id="6AAAC828A330FFF4FF48F9BE601D1937" bold="true" box="[199,345,1576,1601]" pageId="23" pageNumber="24">Description.</emphasis>
Shell large (to
<quantity id="9F26B9DFA330FFF4FD87F9BF63251937" box="[520,609,1577,1601]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="23" pageNumber="24" unit="mm" value="6.8">6.8 mm</quantity>
.
<figureCitation id="C0E508BFA330FFF4FDE2F9BF63A91937" box="[621,749,1577,1601]" captionStart-0="FIGURE 18" captionStart-1="FIGURE 19" captionStartId-0="23.[151,250,1486,1509]" captionStartId-1="24.[151,250,1555,1578]" captionTargetBox-0="[151,1436,541,1472]" captionTargetBox-1="[151,1435,197,1534]" captionTargetId-0="figure@23.[151,1436,541,1473]" captionTargetId-1="figure@24.[151,1436,197,1535]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionText-0="FIGURE 18. Anatoma megascutula new species. Holotype. 800 – 850 m, east of Rapa, Austral Islands, French Polynesia, 27.618 ˚ S, 144.257 ˚ W (MNHN 24954). Scale bar shell = 1 mm. Scale bar protoconch = 100 µm." captionText-1="FIGURE 19. Anatoma megascutula new species. Paratypes. A. Norfolk Ridge, New Caledonia, 820 – 1220 m (MNHN 24959). B. Tongatapu, Tonga, 456 – 510 m (MNHN 24955). C. Fiji, 820 – 863 m (MNHN 24956). Scale bars shell = 1 mm. Scale bars protoconch = 100 µm." httpUri-0="https://zenodo.org/record/281453/files/figure.png" httpUri-1="https://zenodo.org/record/281454/files/figure.png" pageId="23" pageNumber="24">Figs 18–19</figureCitation>
), trochiform depressed, discoidal. Protoconch of 0.75 whorl, flocculent sculpture, apertural varix not connected to embryonic cap, apertural margin shallow sinusoid. Teleoconch I of 0.75–0.875 whorl, approximately 20–30 axial cords, no spirals. Teleoconch II of 3.3 whorls, suture lightly impressed, no sutsel except for last quarter whorl with slight descent showing 1 spiral thread. Shoulder broadly convex, many barely perceptible, low axial cords becoming somewhat more noticeable with growth; first low spiral thread after 0.125–0.8 whorl, increasing in strength to become low to distinct spiral cords, approximately a dozen after first teleoconch I whorl, approximately 25 at apertural margin, forming indistinct thickenings at intersection with axials. Base without constriction below selenizone, continuously sloping with umbilicus; axial sculpture as on shoulder if usually somewhat stronger; approximately 22 spirals, distinct threads from selenizone onwards, changing at mid-base into steps. Umbilicus variable; open, bordered by carina, walls straight, smooth; additionally with distinct funiculus; closed by parietal callus [see Remarks]. Aperture oblong D-shaped. Selenizone at periphery, keels moderately strong, moderately elevated (usually low due to erosion); lunules distinct; slit open, margins converging.
</paragraph>
<caption id="0CA144B2A33FFFFBFF18F985603D1911" httpUri="https://zenodo.org/record/281454/files/figure.png" pageId="24" pageNumber="25" targetBox="[151,1435,197,1534]" targetPageId="24">
<paragraph id="5861143AA33FFFFBFF18F985603D1911" blockId="24.[151,1437,1555,1639]" pageId="24" pageNumber="25">
<emphasis id="6AAAC828A33FFFFBFF18F985605B195F" bold="true" box="[151,287,1555,1578]" pageId="24" pageNumber="25">FIGURE 19.</emphasis>
<taxonomicName id="9FDE6FB9A33FFFFBFEA9F983634C195C" box="[294,520,1556,1578]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="megascutula" status="sp. nov.">
<emphasis id="6AAAC828A33FFFFBFEA9F983634C195C" box="[294,520,1556,1578]" italics="true" pageId="24" pageNumber="25">Anatoma megascutula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71997553A33FFFFBFD81F98363CF195C" box="[526,651,1556,1578]" pageId="24" pageNumber="25" rank="species">new species</taxonomicNameLabel>
. Paratypes. A. Norfolk Ridge, New Caledonia, 820–1220 m (MNHN 24959). B. Tongatapu, Tonga, 456–510 m (MNHN 24955). C. Fiji, 820–863 m (MNHN 24956). Scale bars shell = 1 mm. Scale bars protoconch = 100 µm.
</paragraph>
</caption>
<paragraph id="5861143AA33FFFFBFF48F92E652C18DE" blockId="24.[151,1437,1720,2033]" pageId="24" pageNumber="25">
Radula (
<figureCitation id="C0E508BFA33FFFFBFEA7F92E603819A6" box="[296,380,1720,1744]" captionStart="FIGURE 20" captionStartId="25.[151,250,709,732]" captionTargetBox="[151,1436,199,693]" captionTargetId="figure@25.[151,1436,199,693]" captionTargetPageId="25" captionText="FIGURE 20. Anatoma megascutula new species. Radula. Porthos Bank, Norfolk Ridge, New Caledonia, 680 – 980 m (MNHN). A. Central field. B. Marginal teeth. Scale bars = 100 µm." httpUri="https://zenodo.org/record/281455/files/figure.png" pageId="24" pageNumber="25">Fig. 20</figureCitation>
). Rachidian tooth trapezoid, apical cusp largest, 4–5 on each side in continuous arc. Lateral tooth 1 broadened, apical cusp slightly larger than 7–8 distal ones on cutting edge. Lateral teeth 3–4 similar, apical cusp largest, 3–5 along outer margin of cutting edge. Lateral tooth 4 reduced, apical cusp largest, three small ones on each side. Lateral tooth 5 enlarged by moderate broadening, apical cusp largest, about 12 small cusps along inner edge, approximately 8 not reaching apical cusp along outer edge. Inner marginal teeth with triangular tip, apical cusp largest, approximately 8 minor cusps on each side. Outer marginal teeth with indistinct separation of shaft and tip, tip with multiple fine bristles. Radular interlock of central field moderate.
</paragraph>
<paragraph id="5861143AA33FFFFBFF48F82263FD1886" blockId="24.[151,1437,1720,2033]" pageId="24" pageNumber="25">
Operculum corneous, thin, round, multispiral, with central nucleus, covering approximately 70% of aperture.
<emphasis id="6AAAC828A33FFFFBFF48F84E60241887" bold="true" box="[199,352,2008,2033]" pageId="24" pageNumber="25">Distribution.</emphasis>
Western Pacific,
<quantity id="9F26B9DFA33FFFFBFDAAF84E63F11886" box="[549,693,2008,2032]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.65" metricValueMin="0.45" pageId="24" pageNumber="25" unit="m" value="1050.0" valueMax="1650.0" valueMin="450.0">450–1650 m</quantity>
.
</paragraph>
<caption id="0CA144B2A33EFFFAFF18FD5363931D8C" httpUri="https://zenodo.org/record/281455/files/figure.png" pageId="25" pageNumber="26" targetBox="[151,1436,199,693]" targetPageId="25">
<paragraph id="5861143AA33EFFFAFF18FD5363931D8C" blockId="25.[151,1437,709,762]" pageId="25" pageNumber="26">
<emphasis id="6AAAC828A33EFFFAFF18FD53605B1DAC" bold="true" box="[151,287,709,732]" pageId="25" pageNumber="26">FIGURE 20.</emphasis>
<taxonomicName id="9FDE6FB9A33EFFFAFEABFD5063411DAD" box="[292,517,709,731]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="megascutula" status="sp. nov.">
<emphasis id="6AAAC828A33EFFFAFEABFD5063411DAD" box="[292,517,709,731]" italics="true" pageId="25" pageNumber="26">Anatoma megascutula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71997553A33EFFFAFD84FD5063CC1DAD" box="[523,648,709,731]" pageId="25" pageNumber="26" rank="species">new species</taxonomicNameLabel>
. Radula. Porthos Bank, Norfolk Ridge, New Caledonia, 680–980 m (MNHN). A. Central field. B. Marginal teeth. Scale bars = 100 µm.
</paragraph>
</caption>
<paragraph id="5861143AA33EFFFAFF48FCB060E31BF4" blockId="25.[151,1437,806,1977]" pageId="25" pageNumber="26">
<emphasis id="6AAAC828A33EFFFAFF48FCB060E61C49" bold="true" box="[199,418,806,831]" pageId="25" pageNumber="26">Specimen records.</emphasis>
<collectingCountry id="20C954AAA33EFFFAFE25FCB063541C48" box="[426,528,806,830]" name="Australia" pageId="25" pageNumber="26">Australia</collectingCountry>
. N of
<collectingCountry id="20C954AAA33EFFFAFDD4FCB062461C48" box="[603,770,806,830]" name="Norfolk Island" pageId="25" pageNumber="26">Norfolk Island</collectingCountry>
,
<collectingCountry id="20C954AAA33EFFFAFC81FCB062231C48" box="[782,871,806,830]" name="Norfolk Island" pageId="25" pageNumber="26">Norfolk</collectingCountry>
Ridge, 26.432˚S, 167.181˚E,
<quantity id="9F26B9DFA33EFFFAFB36FCB0647C1C48" box="[1209,1336,806,830]" metricMagnitude="2" metricUnit="m" metricValue="7.62" metricValueMax="7.74" metricValueMin="7.5" pageId="25" pageNumber="26" unit="m" value="762.0" valueMax="774.0" valueMin="750.0">750–774 m</quantity>
(NMNZ M.301394, 7). Coral Sea. 21.317˚S, 157.967˚E,
<quantity id="9F26B9DFA33EFFFAFD48FCDC624A1C14" box="[711,782,842,866]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="25" pageNumber="26" unit="m" value="970.0">970 m</quantity>
(MNHN, 1).
<collectingCountry id="20C954AAA33EFFFAFC3DFCDD65261C14" box="[946,1122,842,866]" name="New Caledonia" pageId="25" pageNumber="26">New Caledonia</collectingCountry>
. Grand Passage, 18.808˚S, 163.190˚E, 683–
<quantity id="9F26B9DFA33EFFFAFED2FCF860E01CF0" box="[349,420,878,902]" metricMagnitude="2" metricUnit="m" metricValue="6.69" pageId="25" pageNumber="26" unit="m" value="669.0">669 m</quantity>
(MNHN, 2). 18.900˚S, 163.317˚E,
<quantity id="9F26B9DFA33EFFFAFCE4FCF862F61CF0" box="[875,946,878,902]" metricMagnitude="2" metricUnit="m" metricValue="5.25" pageId="25" pageNumber="26" unit="m" value="525.0">525 m</quantity>
(MNHN, 1); Loyalty Basin, 20.983˚S, 166.983˚E,
<quantity id="9F26B9DFA33EFFFAFE92FC0460FC1CDC" box="[285,440,914,938]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" metricValueMax="1.64" metricValueMin="1.6" pageId="25" pageNumber="26" unit="m" value="1620.0" valueMax="1640.0" valueMin="1600.0">1600–1640 m</quantity>
(MNHN, 3); 23.050˚S, 167.533˚E,
<quantity id="9F26B9DFA33EFFFAFCD7FC0462931CDC" box="[856,983,914,938]" metricMagnitude="2" metricUnit="m" metricValue="8.275" metricValueMax="8.3" metricValueMin="8.25" pageId="25" pageNumber="26" unit="m" value="827.5" valueMax="830.0" valueMin="825.0">825–830 m</quantity>
(MNHN, 1); Loyalty Ridge, 24.767˚S, 170.117˚E,
<quantity id="9F26B9DFA33EFFFAFE96FC2060DC1CB8" box="[281,408,950,974]" metricMagnitude="2" metricUnit="m" metricValue="8.525" metricValueMax="8.55" metricValueMin="8.5" pageId="25" pageNumber="26" unit="m" value="852.5" valueMax="855.0" valueMin="850.0">850–855 m</quantity>
(MNHN, 1);
<collectingCountry id="20C954AAA33EFFFAFDBEFC2063CE1CB8" box="[561,650,950,974]" name="Norfolk Island" pageId="25" pageNumber="26">Norfolk</collectingCountry>
Ridge, 24.867˚S, 168.183˚E,
<quantity id="9F26B9DFA33EFFFAFC57FC2065201CB8" box="[984,1124,950,974]" metricMagnitude="3" metricUnit="m" metricValue="1.02" metricValueMax="1.22" metricValueMin="0.82" pageId="25" pageNumber="26" unit="m" value="1020.0" valueMax="1220.0" valueMin="820.0">820–1220 m</quantity>
(MNHN, 1); Porthos Bank,
<collectingCountry id="20C954AAA33EFFFAFF18FC4C61B41C84" box="[151,240,986,1010]" name="Norfolk Island" pageId="25" pageNumber="26">Norfolk</collectingCountry>
Ridge, 25.333˚S, 168.950˚E,
<quantity id="9F26B9DFA33EFFFAFDCCFC4C63861C84" box="[579,706,986,1010]" metricMagnitude="2" metricUnit="m" metricValue="8.3" metricValueMax="9.8" metricValueMin="6.8" pageId="25" pageNumber="26" unit="m" value="830.0" valueMax="980.0" valueMin="680.0">680–980 m</quantity>
(
<collectingCountry id="20C954AAA33EFFFAFD5EFC4C62131C84" box="[721,855,986,1010]" name="Norfolk Island" pageId="25" pageNumber="26">NORFOLK</collectingCountry>
2 DW 2068, MNHN, 1).
<collectingCountry id="20C954AAA33EFFFAFBF0FC4D64531C84" box="[1151,1303,986,1010]" name="New Zealand" pageId="25" pageNumber="26">New Zealand</collectingCountry>
. Graveyard Seamount Complex, Chatham Rise, 42.766˚S, 179.989˚W,
<quantity id="9F26B9DFA33EFFFAFCC1FC6862891B60" box="[846,973,1022,1046]" metricMagnitude="2" metricUnit="m" metricValue="8.16" metricValueMax="8.75" metricValueMin="7.57" pageId="25" pageNumber="26" unit="m" value="816.0" valueMax="875.0" valueMin="757.0">757–875 m</quantity>
(NMNZ M.174849, 1).
<collectingCountry id="20C954AAA33EFFFAFB7AFC6864581B60" box="[1269,1308,1022,1046]" name="Fiji" pageId="25" pageNumber="26">Fiji</collectingCountry>
. 17.300˚S, 179.550˚W,
<quantity id="9F26B9DFA33EFFFAFEADFBB460E51B4C" box="[290,417,1058,1082]" metricMagnitude="2" metricUnit="m" metricValue="8.415" metricValueMax="8.629999999999999" metricValueMin="8.2" pageId="25" pageNumber="26" unit="m" value="841.5" valueMax="863.0" valueMin="820.0">820–863 m</quantity>
(MNHN, 1).
<collectingCountry id="20C954AAA33EFFFAFDB3FBB563C51B4D" box="[572,641,1059,1083]" name="Tonga" pageId="25" pageNumber="26">Tonga</collectingCountry>
. 20.633˚S, 178.050˚W,
<quantity id="9F26B9DFA33EFFFAFC1FFBB4654B1B4C" box="[912,1039,1058,1082]" metricMagnitude="2" metricUnit="m" metricValue="4.77" metricValueMax="4.779999999999999" metricValueMin="4.76" pageId="25" pageNumber="26" unit="m" value="477.0" valueMax="478.0" valueMin="476.0">476–478 m</quantity>
(MNHN, 1). Tongatapu, 21.083˚S, 175.367˚W,
<quantity id="9F26B9DFA33EFFFAFEA5FBD060EC1B28" box="[298,424,1094,1118]" metricMagnitude="2" metricUnit="m" metricValue="4.83" metricValueMax="5.1" metricValueMin="4.56" pageId="25" pageNumber="26" unit="m" value="483.0" valueMax="510.0" valueMin="456.0">456–510 m</quantity>
(MNHN, 1).
<collectingCountry id="20C954AAA33EFFFAFDD3FBD062621B28" box="[604,806,1094,1118]" name="French Polynesia" pageId="25" pageNumber="26">French Polynesia</collectingCountry>
. E of Rapa, Austral Islands, 27.618˚S, 144.257˚W,
<quantity id="9F26B9DFA33EFFFAFF18FBFC60511BF4" box="[151,277,1130,1154]" metricMagnitude="2" metricUnit="m" metricValue="8.25" metricValueMax="8.5" metricValueMin="8.0" pageId="25" pageNumber="26" unit="m" value="825.0" valueMax="850.0" valueMin="800.0">800–850 m</quantity>
(MNHN, 1).
</paragraph>
</subSubSection>
<subSubSection id="10C447B1A33EFFFAFF48FB1961AB18CF" pageId="25" pageNumber="26" type="discussion">
<paragraph id="5861143AA33EFFFAFF48FB19604119E9" blockId="25.[151,1437,806,1977]" pageId="25" pageNumber="26">
<emphasis id="6AAAC828A33EFFFAFF48FB19607F1BD1" bold="true" box="[199,315,1167,1191]" pageId="25" pageNumber="26">Remarks.</emphasis>
<taxonomicName id="9FDE6FB9A33EFFFAFECDFB1863A71BD0" authority="A. Adams, 1862" authorityName="A. Adams" authorityYear="1862" box="[322,739,1166,1190]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="japonica">
<emphasis id="6AAAC828A33EFFFAFECDFB1863541BD0" box="[322,528,1166,1190]" italics="true" pageId="25" pageNumber="26">Anatoma japonica</emphasis>
(A. Adams, 1862)
</taxonomicName>
from the Indian Ocean and Indo-Pacific has a wider sutsel on whorl 2 (0.5–1× vs. 0× width of selenizone), has stronger axial than spiral sculpture, and has a narrower selenizone.
<taxonomicName id="9FDE6FB9A33EFFFAFF18FB4060241B98" box="[151,352,1238,1262]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="maxima">
<emphasis id="6AAAC828A33EFFFAFF18FB4060241B98" box="[151,352,1238,1262]" italics="true" pageId="25" pageNumber="26">Anatoma maxima</emphasis>
</taxonomicName>
from the Indo-Malayan Archipelago has coarser sculpture (14 vs. 25 spirals on shoulder at apertural margin; distinct vs. indistinct axials) in which axials and spirals form points at intersections, a less rapid expansion of the whorls (1.4 vs. 1.8), and a wider sutsel at the apertural margin showing 2–3 spiral threads.
<taxonomicName id="9FDE6FB9A33EFFFAFF18FAD560161A2D" box="[151,338,1347,1371]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="rainesi">
<emphasis id="6AAAC828A33EFFFAFF18FAD560161A2D" box="[151,338,1347,1371]" italics="true" pageId="25" pageNumber="26">Anatoma rainesi</emphasis>
</taxonomicName>
from the Eastern and Western Pacific has a sutsel on the early whorls, has strong axial cords on teleoconch I, and the selenizone is closer to the periphery of the shell.
<taxonomicName id="9FDE6FB9A33EFFFAFC53FAF1641E1A09" authority="Geiger, 2008" authorityName="Geiger" authorityYear="2008" box="[988,1370,1383,1407]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="rapaensis">
<emphasis id="6AAAC828A33EFFFAFC53FAF165FE1A09" box="[988,1210,1383,1407]" italics="true" pageId="25" pageNumber="26">Anatoma rapaensis</emphasis>
Geiger, 2008
</taxonomicName>
from
<collectingCountry id="20C954AAA33EFFFAFF18FA1C601D1AD4" box="[151,345,1418,1442]" name="French Polynesia" pageId="25" pageNumber="26">French Polynesia</collectingCountry>
shares the overall discoidal shape with no sutsel, but the protoconch lacks an apertural varix, the teleoconch I is shorter (0.2 vs.&gt;0.75 whorl, the axial sculpture is more pronounced forming sharp points at the intersection with the spirals.
<taxonomicName id="9FDE6FB9A33EFFFAFE57FA4562071A9C" authority="Mestayer, 1916" authorityName="Mestayer" authorityYear="1916" box="[472,835,1490,1515]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="regia">
<emphasis id="6AAAC828A33EFFFAFE57FA4563391A9D" box="[472,637,1491,1515]" italics="true" pageId="25" pageNumber="26">Anatoma regia</emphasis>
(Mestayer, 1916)
</taxonomicName>
from
<collectingCountry id="20C954AAA33EFFFAFC08FA4565581A9C" box="[903,1052,1490,1514]" name="New Zealand" pageId="25" pageNumber="26">New Zealand</collectingCountry>
has a narrower but still noticeable sutsel on early whorls, has a narrower selenizone, and its axial sculpture is stronger than the spiral sculpture.
<taxonomicName id="9FDE6FB9A33EFFFAFF18F98D63911944" authority="Golikov &amp; Sirenko, 1980" authorityName="Golikov &amp; Sirenko" authorityYear="1980" box="[151,725,1562,1587]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="disciformis">
<emphasis id="6AAAC828A33EFFFAFF18F98D60C21945" box="[151,390,1563,1587]" italics="true" pageId="25" pageNumber="26">Anatoma disciformis</emphasis>
(Golikov &amp; Sirenko, 1980)
</taxonomicName>
from the boreal north Pacific has an overall taller shell, a protoconch with strong flocculent sculpture, and much stronger axial cords.
<taxonomicName id="9FDE6FB9A33EFFFAFC7BF9A9641A1921" authority="Jeffreys, 1877" authorityName="Jeffreys" authorityYear="1877" box="[1012,1374,1599,1623]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="tenuis">
<emphasis id="6AAAC828A33EFFFAFC7BF9A965E01921" box="[1012,1188,1599,1623]" italics="true" pageId="25" pageNumber="26">Anatoma tenuis</emphasis>
(Jeffreys, 1877)
</taxonomicName>
from the temperate north Atlantic has a shorter teleoconch I (0.4 vs. 0.75 whorl), and has stronger axial than spiral sculpture.
</paragraph>
<paragraph id="5861143AA33EFFFAFF48F93C61AB1824" blockId="25.[151,1437,806,1977]" pageId="25" pageNumber="26">The number of axials on teleoconch I is difficult to count due to erosion of the early parts of the whorl. The condition of the umbilicus is quite variable in the limited number of specimens available. The more open condition is found in smaller specimens, while the callus closing the umbilicus is encountered in the largest specimens, which suggest an ontogenetic change. Other conchological characters all suggest that the specimens belong to a single species.</paragraph>
<paragraph id="5861143AA33EFFFAFF48F8C961AB18CF" blockId="25.[151,1437,806,1977]" pageId="25" pageNumber="26">
The radula of
<taxonomicName id="9FDE6FB9A33EFFFAFEE1F8C9635B1801" box="[366,543,1887,1911]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="megascutula">
<emphasis id="6AAAC828A33EFFFAFEE1F8C9635B1801" box="[366,543,1887,1911]" italics="true" pageId="25" pageNumber="26">A. megascutula</emphasis>
</taxonomicName>
shows an unusual arrangement in the central field. Lateral teeth 1–3 are not similar to one another, lateral tooth 1 is much broadened, and lateral tooth 4 is not as reduced as in most other species.
</paragraph>
</subSubSection>
</treatment>
</document>
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