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<mods:title id="DE4695991C2182C9C0B2B4934FF241AA">Tortonian teleost otoliths from northern Italy: taxonomic synthesis and stratigraphic significance</mods:title>
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<mods:namePart id="1B4D3ACC0D28669CE8214016B4D2556D">Brzobohatý, Rostislav</mods:namePart>
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<treatment id="61599612947FCE2F79B8A1FCFE589473" ID-DOI="http://doi.org/10.5281/zenodo.3848313" ID-GBIF-Taxon="164159942" ID-Zenodo-Dep="3848313" LSID="urn:lsid:plazi:treatment:61599612947FCE2F79B8A1FCFE589473" httpUri="http://treatment.plazi.org/id/61599612947FCE2F79B8A1FCFE589473" lastPageNumber="26" pageId="25" pageNumber="26">
<subSubSection id="A1EA748F947FCE2F79B8A1FCFB819697" box="[503,1085,1451,1478]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="E94F2704947FCE2F79B8A1FCFB819697" blockId="25.[503,1085,1451,1478]" box="[503,1085,1451,1478]" pageId="25" pageNumber="26">
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<taxonomicName id="2EF05C87947FCE2F79B8A1FCFB889697" ID-CoL="4PQL" ID-ENA="45204" authorityName="Linnaeus" authorityYear="1758" box="[503,1076,1451,1478]" class="Actinopterygii" family="Gobiidae" genus="Gobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="26" phylum="Chordata" rank="genus">
<emphasis id="DB84FB16947FCE2F79B8A1FCFD489697" bold="true" box="[503,756,1451,1478]" pageId="25" pageNumber="26">
<emphasis id="DB84FB16947FCE2F7A4AA1FCFDE79697" bold="true" box="[517,603,1451,1478]" italics="true" pageId="25" pageNumber="26">Gobius</emphasis>
<emphasis id="DB84FB16947FCE2F7A3DA1FCFD489697" bold="true" box="[626,756,1451,1478]" italics="true" pageId="25" pageNumber="26">bicornutus</emphasis>
</emphasis>
(
<bibRefCitation id="8D615AF5947FCE2F7B4BA1FCFB889697" author="Lin C. - H. &amp; Girone A. &amp; Nolf D." box="[772,1076,1451,1478]" pageId="25" pageNumber="26" pagination="249 - 261" refId="ref22326" refString="Lin C. - H., Girone A. &amp; Nolf D. 2015. Tortonian fish otoliths from turbiditic deposits in northern Italy: taxonomic and stratigraphic significance. Geobios 48: 249 - 261. https: // doi. org / 10.1016 / j. geobios. 2015.03.003" type="journal article" year="2015">Lin, Girone &amp; Nolf, 2015</bibRefCitation>
</taxonomicName>
)
</heading>
</paragraph>
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<subSubSection id="A1EA748F947FCE2F78F2A1BEFE589473" pageId="25" pageNumber="26" type="discussion">
<paragraph id="E94F2704947FCE2F78F2A1BEFE589473" blockId="25.[189,1399,1513,1826]" pageId="25" pageNumber="26">
This species was described, for the first time, from the turbiditic Tortonian deposits in the open generic nomenclature as “Gobiida”
<emphasis id="DB84FB16947FCE2F7A49A25AFDC49576" box="[518,632,1549,1575]" italics="true" pageId="25" pageNumber="26">bicornuta</emphasis>
by
<bibRefCitation id="8D615AF5947FCE2F7AE9A25AFCD29579" author="Lin C. - H. &amp; Girone A. &amp; Nolf D." box="[678,878,1549,1576]" pageId="25" pageNumber="26" pagination="249 - 261" refId="ref22326" refString="Lin C. - H., Girone A. &amp; Nolf D. 2015. Tortonian fish otoliths from turbiditic deposits in northern Italy: taxonomic and stratigraphic significance. Geobios 48: 249 - 261. https: // doi. org / 10.1016 / j. geobios. 2015.03.003" type="journal article" year="2015">
Lin
<emphasis id="DB84FB16947FCE2F7A97A259FCB09576" box="[728,780,1549,1575]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
. (2015)
</bibRefCitation>
. Following the nomenclature adopted in the present paper and based on diagnostic features, this species is here assigned to “
<emphasis id="DB84FB16947FCE2F7CCCA265FB23951D" box="[1155,1183,1586,1612]" italics="true" pageId="25" pageNumber="26">G.</emphasis>
<emphasis id="DB84FB16947FCE2F7CF9A266FA8F951A" box="[1206,1331,1585,1611]" italics="true" pageId="25" pageNumber="26">bicornutus</emphasis>
(
<bibRefCitation id="8D615AF5947FCE2F7D0AA266FE7D9521" author="Lin C. - H. &amp; Girone A. &amp; Nolf D." pageId="25" pageNumber="26" pagination="249 - 261" refId="ref22326" refString="Lin C. - H., Girone A. &amp; Nolf D. 2015. Tortonian fish otoliths from turbiditic deposits in northern Italy: taxonomic and stratigraphic significance. Geobios 48: 249 - 261. https: // doi. org / 10.1016 / j. geobios. 2015.03.003" type="journal article" year="2015">Lin, Girone &amp; Nolf, 2015</bibRefCitation>
). The otoliths of “
<emphasis id="DB84FB16947FCE2F7AEFA201FD009521" box="[672,700,1622,1648]" italics="true" pageId="25" pageNumber="26">G.</emphasis>
<emphasis id="DB84FB16947FCE2F7A9CA202FCEC953E" box="[723,848,1621,1647]" italics="true" pageId="25" pageNumber="26">bicornutus</emphasis>
are rare in Tortonian deposits. They are very thick and nearly square shaped, with large and horn-like expansions at the postero-dorsal corner. Their sulcus is elliptic without strongly delimitated margins (
<figureCitation id="71CB3B81947FCE2F7B0BA2CAFC0E95E6" box="[836,946,1693,1719]" captionStart="Fig" captionStartId="28.[189,232,1788,1814]" captionTargetBox="[189,1398,612,1746]" captionTargetId="figure@28.[189,1398,612,1746]" captionTargetPageId="28" captionText="Fig. 12. Tortonian fish otoliths from northern Italy. AC. “Gobius” bicornutus (Lin, Girone &amp; Nolf, 2015), SantAgata Fossili (IRSNB P 9836P 9838). D. Aphanopus carbo Lowe, 1839, Costa Vescovato (IRSNB P 9839). E. Nealotus tripes Johnson, 1865, Mondovi, Madonna della Neve (IRSNB P 9840). FG. Arnoglossus kokeni (Bassoli &amp; Schubert, 1906), Torrente Stirone (IRSNB P 9841P 9842). HI. Gobiidae indet., SantAgata Fossili (IRSNB P 9843P 9844). JK. Microchirus aff. variegatus (Donovan, 1808), Torrente Stirone (IRSNB P 98459846). 1 = ventral view; 2 = inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829329" httpUri="https://zenodo.org/record/3829329/files/figure.png" pageId="25" pageNumber="26">Fig. 12A</figureCitation>
). Some small specimens (
<figureCitation id="71CB3B81947FCE2F7CADA2CAFA8A95E6" box="[1250,1334,1693,1719]" captionStart="Fig" captionStartId="28.[189,232,1788,1814]" captionTargetBox="[189,1398,612,1746]" captionTargetId="figure@28.[189,1398,612,1746]" captionTargetPageId="28" captionText="Fig. 12. Tortonian fish otoliths from northern Italy. AC. “Gobius” bicornutus (Lin, Girone &amp; Nolf, 2015), SantAgata Fossili (IRSNB P 9836P 9838). D. Aphanopus carbo Lowe, 1839, Costa Vescovato (IRSNB P 9839). E. Nealotus tripes Johnson, 1865, Mondovi, Madonna della Neve (IRSNB P 9840). FG. Arnoglossus kokeni (Bassoli &amp; Schubert, 1906), Torrente Stirone (IRSNB P 9841P 9842). HI. Gobiidae indet., SantAgata Fossili (IRSNB P 9843P 9844). JK. Microchirus aff. variegatus (Donovan, 1808), Torrente Stirone (IRSNB P 98459846). 1 = ventral view; 2 = inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829329" httpUri="https://zenodo.org/record/3829329/files/figure.png" pageId="25" pageNumber="26">Fig. 12</figureCitation>
BC) of this species from SantAgata Fossili (
<bibRefCitation id="8D615AF5947FCE2F7AD4A296FC0E958A" author="Nolf D. &amp; Steurbaut E." box="[667,946,1729,1755]" pageId="25" pageNumber="26" pagination="143 - 197" refId="ref23235" refString="Nolf D. &amp; Steurbaut E. 1983. Revision des otolithes de teleosteens du Tortonien stratotypique et de Montegibbio (Miocene Superieur d'Italie septentrionale). Mededelingen van de Werkgroep voor Tertiaire en Kwartaire Geologie 20: 143 - 197." type="journal article" year="1983">Nolf &amp; Steurbaut 1983</bibRefCitation>
: fig. 1.1, point C) also show the hornshaped expansion at the postero-dorsal corner, a feature which can be therefore considered as consistent, regardless of otolith size.
</paragraph>
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