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<document id="ACF6468066B4D501553AC387F0E29459" ID-CLB-Dataset="22102" ID-DOI="10.5852/ejt.2017.322" ID-GBIF-Dataset="2922dbf4-12ec-49d4-ac36-565434419f51" ID-ISSN="2118-9773" ID-Zenodo-Dep="3829305" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589562461714" checkinUser="carolina" docAuthor="Lin, Chien-Hsiang, Brzobohatý, Rostislav, Nolf, Dirk &amp; Girone, Angela" docDate="2017" docId="61599612946CCE3D7A16A0FEFBA69034" docLanguage="en" docName="ejt-2017-322.pdf.imf" docOrigin="European Journal of Taxonomy 322" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Neobythites auriculatus Lin &amp; Brzobohatý &amp; Nolf &amp; Girone 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="12" masterDocId="9D60EE6A9466CE36784FA457FFBC9351" masterDocTitle="Tortonian teleost otoliths from northern Italy: taxonomic synthesis and stratigraphic significance" masterLastPageNumber="44" masterPageNumber="1" pageNumber="11" updateTime="1698839868927" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DDE4FF0B94F5775D8CF1E616343E57C5">Tortonian teleost otoliths from northern Italy: taxonomic synthesis and stratigraphic significance</mods:title>
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<mods:namePart id="34D2A4265C888068546FE35493BF8DCB">Lin, Chien-Hsiang</mods:namePart>
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<mods:namePart id="850D31F17744148DF131A21F911BAA72">Brzobohatý, Rostislav</mods:namePart>
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<mods:affiliation id="3560ED751FEF041B7AD56060FE392A65">Email: rosta @ sci. muni. cz &amp; urn: lsid: zoobank. org: author: A 354 D 828 - 3 D 07 - 46 F 5 - 9 D 27 - 4 F 9 D 7 B 208 F 12</mods:affiliation>
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<mods:namePart id="7A1BA123204A59A71835A30BC4D18A47">Nolf, Dirk</mods:namePart>
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<mods:namePart id="F74271F762FCDDC66E0C969AE308C2DD">Girone, Angela</mods:namePart>
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<mods:affiliation id="029B9B0C99F343DB9458C12A4D7F57DD">Email: angela. girone @ uniba. it &amp; urn: lsid: zoobank. org: author: B 4212 D 22 - 1 D 1 B- 48 E 9 - 8 ABB-C 1 D 1 E 05573 EF</mods:affiliation>
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<treatment id="61599612946CCE3D7A16A0FEFBA69034" ID-DOI="http://doi.org/10.5281/zenodo.3848335" ID-GBIF-Taxon="164159952" ID-Zenodo-Dep="3848335" LSID="urn:lsid:plazi:treatment:61599612946CCE3D7A16A0FEFBA69034" httpUri="http://treatment.plazi.org/id/61599612946CCE3D7A16A0FEFBA69034" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
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<paragraph id="E94F2704946CCE3C7A16A0FEFC679795" blockId="10.[380,1208,1193,1294]" box="[601,987,1193,1220]" pageId="10" pageNumber="11">
<heading id="B2079068946CCE3C7A16A0FEFC679795" box="[601,987,1193,1220]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="11" reason="2">
<taxonomicName id="2EF05C87946CCE3C7A16A0FEFCC59795" authority="Lin &amp; Brzobohatý &amp; Nolf &amp; Girone, 2017" authorityName="Lin &amp; Brzobohatý &amp; Nolf &amp; Girone" authorityYear="2017" box="[601,889,1193,1220]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="auriculatus" status="sp. nov.">
<emphasis id="DB84FB16946CCE3C7A16A0FEFCC59795" bold="true" box="[601,889,1193,1220]" italics="true" pageId="10" pageNumber="11">Neobythites auriculatus</emphasis>
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<taxonomicNameLabel id="C0B7466D946CCE3C7BCFA0FDFC679795" box="[896,987,1194,1220]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
<uuid id="9D561DD1946CCE3C7AD9A098FB0497BB" box="[662,1208,1231,1258]" pageId="10" pageNumber="11">06D09029-80AE-461E-9135-01594A825560</uuid>
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<paragraph id="E94F2704946CCE3C7AA1A0A3FCFA965F" blockId="10.[380,1208,1193,1294]" box="[750,838,1268,1294]" pageId="10" pageNumber="11">
<figureCitation id="71CB3B81946CCE3C7AA1A0A3FCFA965F" box="[750,838,1268,1294]" captionStart="Fig" captionStartId="24.[189,232,1614,1640]" captionTargetBox="[234,1353,265,1594]" captionTargetId="figure@24.[234,1353,265,1594]" captionTargetPageId="24" captionText="Fig. 9. Tortonian fish otoliths from northern Italy. A. Echiodon heinzelini Huyghebaert &amp; Nolf, 1979, Torrente Stirone(IRSNB P 9787).B. Hoplobrotula aff.armata(Temminck&amp; Schlegel, 1846),SantAlosio (IRSNB P 9788). C. Neobythites auriculatus sp. nov., SantAlosio (IRSNB P 9688 (holotype). DE. Carapus acus (Brünnich, 1768); D. Torrente Stirone, E. SantAgata Fossili (IRSNB P 9789P 9790). FG. Bythitidae indet., Torrente Stirone (IRSNB P 9791P 9792). H. Grammonus bassolii (Nolf, 1980), Torrente Stirone (IRSNB P 9793). I. Chaunax lobatus (Bassoli, 1906), Montegibbio (IRSNB P 9794). JL. Scopelogadus sp.; J. Alba, Tanaro (5 m), K. Stazzano, L. Alba, Tanaro (50 m) (IRSNB P 9795P 9797). M. Mugilidae indet., SantAlosio (IRSNB P 9798). N. Phycis musicki Cohen &amp; Lavenberg, 1984, Torrente Stirone (IRSNB P 9799). O. Melamphaes sp., Alba, Tanaro (50 m) (IRSNB P 9802). P. Micromesistius planatus (Bassoli &amp; Schubert, 1906), Montegibbio (IRSNB P 9803). QR. “Scorpaena” zibinica (Bassoli, 1909), Torrente Stirone (IRSNB P 9800P 9801). 1 = ventral view; 2 = inner view; 3 = anterior view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829323" httpUri="https://zenodo.org/record/3829323/files/figure.png" pageId="10" pageNumber="11">Fig. 9C</figureCitation>
</paragraph>
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<subSubSection id="A1EA748F946CCE3C78F2A16FFBA8969F" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="E94F2704946CCE3C78F2A16FFE8A9603" blockId="10.[189,310,1336,1362]" box="[189,310,1336,1362]" pageId="10" pageNumber="11">
<heading id="B2079068946CCE3C78F2A16FFE8A9603" bold="true" box="[189,310,1336,1362]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="DB84FB16946CCE3C78F2A16FFE8A9603" bold="true" box="[189,310,1336,1362]" pageId="10" pageNumber="11">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="E94F2704946CCE3C78F2A13EFBA8969F" blockId="10.[189,1399,1385,1486]" pageId="10" pageNumber="11">The species is characterised by very elongate otoliths with moderate thickness. The highest part of the otolith is in the middle. Both the dorsal and ventral rims are curved. The huge, wide sulcus bears a large and oblong ostial colliculum and a much shorter, oval caudal colliculum.</paragraph>
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<subSubSection id="A1EA748F946CCE3C78F2A1AFFAAA9539" pageId="10" pageNumber="11" type="etymology">
<paragraph id="E94F2704946CCE3C78F2A1AFFEF89543" blockId="10.[189,324,1528,1554]" box="[189,324,1528,1554]" pageId="10" pageNumber="11">
<heading id="B2079068946CCE3C78F2A1AFFEF89543" bold="true" box="[189,324,1528,1554]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="DB84FB16946CCE3C78F2A1AFFEF89543" bold="true" box="[189,324,1528,1554]" pageId="10" pageNumber="11">Etymology</emphasis>
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<paragraph id="E94F2704946CCE3C78F2A27FFAAA9539" blockId="10.[189,1399,1576,1640]" pageId="10" pageNumber="11">
The species epithet is derived from the Latin
<taxonomicName id="2EF05C87946CCE3C7A9BA27FFCE69513" authorityName="Lin &amp; Brzobohatý &amp; Nolf &amp; Girone" authorityYear="2017" box="[724,858,1576,1602]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="auriculatus">
<emphasis id="DB84FB16946CCE3C7A9BA27FFCE69513" box="[724,858,1576,1602]" italics="true" pageId="10" pageNumber="11">auriculatus</emphasis>
</taxonomicName>
,
<emphasis id="DB84FB16946CCE3C7B27A27EFCCB9513" box="[872,887,1577,1602]" italics="true" pageId="10" pageNumber="11">a</emphasis>
,
<emphasis id="DB84FB16946CCE3C7BCAA27EFC159513" box="[901,937,1577,1602]" italics="true" pageId="10" pageNumber="11">um</emphasis>
: having ears; alludes to the two obtuse little spines at both the anterior and posterior ends of the dorsal rim, suggesting two small ears.
</paragraph>
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<subSubSection id="A1EA748F946CCE3C78F2A2C5FBA39453" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="E94F2704946CCE3C78F2A2C5FE5095FA" blockId="10.[189,492,1681,1708]" box="[189,492,1681,1708]" pageId="10" pageNumber="11">
<heading id="B2079068946CCE3C78F2A2C5FE5095FA" bold="true" box="[189,492,1681,1708]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="DB84FB16946CCE3C78F2A2C5FE5095FA" bold="true" box="[189,492,1681,1708]" pageId="10" pageNumber="11">Type material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="59982D59946CCE3C78A5A295FBA39453" ID-GBIF-Occurrence="2620279301" collectionCode="IRSNB" country="Italy" location="ITALY" pageId="10" pageNumber="11" specimenCode="IRSNB P 9688" specimenCount="1" typeStatus="holotype">
<paragraph id="E94F2704946CCE3C78A5A295FEE0958D" blockId="10.[189,1055,1730,1794]" box="[234,348,1730,1756]" pageId="10" pageNumber="11">
<emphasis id="DB84FB16946CCE3C78A5A295FEE0958D" bold="true" box="[234,348,1730,1756]" pageId="10" pageNumber="11">
<typeStatus id="364B99A6946CCE3C78A5A295FEE0958D" box="[234,348,1730,1756]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="E94F2704946CCE3C78F2A2BFFBA39453" blockId="10.[189,1055,1730,1794]" box="[189,1055,1768,1794]" pageId="10" pageNumber="11">
<collectingCountry id="91E76794946CCE3C78F2A2BFFEAE9453" box="[189,274,1768,1794]" name="Italy" pageId="10" pageNumber="11">ITALY</collectingCountry>
: a left otolith, the only known specimen (
<specimenCode id="B9568F7F946CCE3C7AB9A2BFFC169453" box="[758,938,1768,1794]" pageId="10" pageNumber="11">IRSNB P 9688</specimenCode>
,
<figureCitation id="71CB3B81946CCE3C7BF9A2BFFBAD9453" box="[950,1041,1768,1794]" captionStart="Fig" captionStartId="24.[189,232,1614,1640]" captionTargetBox="[234,1353,265,1594]" captionTargetId="figure@24.[234,1353,265,1594]" captionTargetPageId="24" captionText="Fig. 9. Tortonian fish otoliths from northern Italy. A. Echiodon heinzelini Huyghebaert &amp; Nolf, 1979, Torrente Stirone(IRSNB P 9787).B. Hoplobrotula aff.armata(Temminck&amp; Schlegel, 1846),SantAlosio (IRSNB P 9788). C. Neobythites auriculatus sp. nov., SantAlosio (IRSNB P 9688 (holotype). DE. Carapus acus (Brünnich, 1768); D. Torrente Stirone, E. SantAgata Fossili (IRSNB P 9789P 9790). FG. Bythitidae indet., Torrente Stirone (IRSNB P 9791P 9792). H. Grammonus bassolii (Nolf, 1980), Torrente Stirone (IRSNB P 9793). I. Chaunax lobatus (Bassoli, 1906), Montegibbio (IRSNB P 9794). JL. Scopelogadus sp.; J. Alba, Tanaro (5 m), K. Stazzano, L. Alba, Tanaro (50 m) (IRSNB P 9795P 9797). M. Mugilidae indet., SantAlosio (IRSNB P 9798). N. Phycis musicki Cohen &amp; Lavenberg, 1984, Torrente Stirone (IRSNB P 9799). O. Melamphaes sp., Alba, Tanaro (50 m) (IRSNB P 9802). P. Micromesistius planatus (Bassoli &amp; Schubert, 1906), Montegibbio (IRSNB P 9803). QR. “Scorpaena” zibinica (Bassoli, 1909), Torrente Stirone (IRSNB P 9800P 9801). 1 = ventral view; 2 = inner view; 3 = anterior view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829323" httpUri="https://zenodo.org/record/3829323/files/figure.png" pageId="10" pageNumber="11">Fig. 9C</figureCitation>
).
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="A1EA748F946CCE3C78F2A37BFCDA9427" pageId="10" pageNumber="11" type="distribution">
<paragraph id="E94F2704946CCE3C78F2A37BFE479414" blockId="10.[189,507,1835,1862]" box="[189,507,1835,1862]" pageId="10" pageNumber="11">
<heading id="B2079068946CCE3C78F2A37BFE479414" bold="true" box="[189,507,1835,1862]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="DB84FB16946CCE3C78F2A37BFE479414" bold="true" box="[189,507,1835,1862]" pageId="10" pageNumber="11">Type locality and horizon</emphasis>
</heading>
</paragraph>
<paragraph id="E94F2704946CCE3C78F2A30BFCDA9427" blockId="10.[189,870,1884,1911]" box="[189,870,1884,1911]" pageId="10" pageNumber="11">
<collectingCountry id="91E76794946CCE3C78F2A30BFEAE9427" box="[189,274,1884,1910]" name="Italy" pageId="10" pageNumber="11">ITALY</collectingCountry>
: Tortonian Clay at Sant Alosio, surface collection.
</paragraph>
</subSubSection>
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<heading id="B2079068946CCE3C78F2A3F7FDAC94EB" bold="true" box="[189,528,1952,1978]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="DB84FB16946CCE3C78F2A3F7FDAC94EB" bold="true" box="[189,528,1952,1978]" pageId="10" pageNumber="11">
Dimensions of the
<typeStatus id="364B99A6946CCE3C79EBA3F7FDAC94EB" box="[420,528,1952,1978]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="E94F2704946CCE3C78F2A386FC1C94BA" blockId="10.[189,928,2000,2027]" box="[189,928,2000,2027]" pageId="10" pageNumber="11">
Length =
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, height =
<quantity id="2E088AE1946CCE3C7A52A386FD2494BA" box="[541,664,2001,2027]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="10" pageNumber="11" unit="mm" value="10.3">10.30 mm</quantity>
, thickness =
<quantity id="2E088AE1946CCE3C7B7FA386FC2794BA" box="[816,923,2001,2027]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="10" pageNumber="11" unit="mm" value="3.6">3.60 mm</quantity>
.
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<paragraph id="E94F2704946DCE3D78F2A55CFEF29274" blockId="11.[189,334,267,293]" box="[189,334,267,293]" pageId="11" pageNumber="12">
<heading id="B2079068946DCE3D78F2A55CFEF29274" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="DB84FB16946DCE3D78F2A55CFEF29274" bold="true" box="[189,334,267,293]" pageId="11" pageNumber="12">Description</emphasis>
</heading>
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<paragraph id="E94F2704946DCE3D78F2A56CFE1F919D" blockId="11.[189,1399,315,716]" pageId="11" pageNumber="12">
The
<typeStatus id="364B99A6946DCE3D78BEA56CFEE49204" box="[241,344,315,341]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
is a very large elongate elliptic otolith. It is moderately thick, with a convex inner face and concave outer face, but on the outer face, a swollen structure is observed in the middle of the ventral portion (
<figureCitation id="71CB3B81946DCE3D796FA5D1FEC292F1" box="[288,382,390,416]" captionStart="Fig" captionStartId="24.[189,232,1614,1640]" captionTargetBox="[234,1353,265,1594]" captionTargetId="figure@24.[234,1353,265,1594]" captionTargetPageId="24" captionText="Fig. 9. Tortonian fish otoliths from northern Italy. A. Echiodon heinzelini Huyghebaert &amp; Nolf, 1979, Torrente Stirone(IRSNB P 9787).B. Hoplobrotula aff.armata(Temminck&amp; Schlegel, 1846),SantAlosio (IRSNB P 9788). C. Neobythites auriculatus sp. nov., SantAlosio (IRSNB P 9688 (holotype). DE. Carapus acus (Brünnich, 1768); D. Torrente Stirone, E. SantAgata Fossili (IRSNB P 9789P 9790). FG. Bythitidae indet., Torrente Stirone (IRSNB P 9791P 9792). H. Grammonus bassolii (Nolf, 1980), Torrente Stirone (IRSNB P 9793). I. Chaunax lobatus (Bassoli, 1906), Montegibbio (IRSNB P 9794). JL. Scopelogadus sp.; J. Alba, Tanaro (5 m), K. Stazzano, L. Alba, Tanaro (50 m) (IRSNB P 9795P 9797). M. Mugilidae indet., SantAlosio (IRSNB P 9798). N. Phycis musicki Cohen &amp; Lavenberg, 1984, Torrente Stirone (IRSNB P 9799). O. Melamphaes sp., Alba, Tanaro (50 m) (IRSNB P 9802). P. Micromesistius planatus (Bassoli &amp; Schubert, 1906), Montegibbio (IRSNB P 9803). QR. “Scorpaena” zibinica (Bassoli, 1909), Torrente Stirone (IRSNB P 9800P 9801). 1 = ventral view; 2 = inner view; 3 = anterior view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829323" httpUri="https://zenodo.org/record/3829323/files/figure.png" pageId="11" pageNumber="12">Fig. 9C</figureCitation>
<figureCitation id="71CB3B81946DCE3D7931A5D1FE3592F1" box="[382,393,390,416]" captionStart="Fig" captionStartId="2.[189,232,1997,2023]" captionTargetBox="[189,1398,1032,1956]" captionTargetId="figure@2.[189,1398,1032,1956]" captionTargetPageId="2" captionText="Fig. 1. Map showing the sampling sites." figureDoi="http://doi.org/10.5281/zenodo.3829307" httpUri="https://zenodo.org/record/3829307/files/figure.png" pageId="11" pageNumber="12">1</figureCitation>
). The anterior rim is short and rounded; the posterior rim, which is more or less straight at the extended end, bears two lobes. The dorsal and ventral rims are gently curved, so that the otoliths are widest in the middle. The margin of the ventral rim is smooth. There are two small protrusions at the anterior and posterior angles of the dorsal rim. The sulcus is wide, occupies about one-third of the inner face, and is completely filled by collicula. The cristae are well marked. The collicula are both well developed, but are not completely separated at the collum. Only a large constriction of the crista inferior delimits the ostial and caudal portion. The surface of collicula is irregular. The caudal colliculum is short and oval; the ostial colliculum is about four times as long as the caudal one. The ventral area is wider than the dorsal one.
</paragraph>
</subSubSection>
<subSubSection id="A1EA748F946DCE3D78F2A6A2FBA69034" pageId="11" pageNumber="12" type="discussion">
<paragraph id="E94F2704946DCE3D78F2A6A2FE93905E" blockId="11.[189,303,757,783]" box="[189,303,757,783]" pageId="11" pageNumber="12">
<heading id="B2079068946DCE3D78F2A6A2FE93905E" bold="true" box="[189,303,757,783]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="DB84FB16946DCE3D78F2A6A2FE93905E" bold="true" box="[189,303,757,783]" pageId="11" pageNumber="12">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="E94F2704946DCE3D78F2A772FBA69034" blockId="11.[189,1398,805,869]" pageId="11" pageNumber="12">
This huge otolith apparently belongs to an adult (old) individual of an extinct species and it shows most similarity to those of the Recent genus
<taxonomicName id="2EF05C87946DCE3D7AC7A71CFBAA9034" authority="Goode &amp; Bean, 1886" authorityName="Goode &amp; Bean" authorityYear="1886" box="[648,1046,843,869]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="DB84FB16946DCE3D7AC7A71CFCAF9034" box="[648,787,843,869]" italics="true" pageId="11" pageNumber="12">Neobythites</emphasis>
Goode &amp; Bean, 1886
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>