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<mods:title id="4A9DC042A1B0A1F67D72148675D9CD3B">Tortonian teleost otoliths from northern Italy: taxonomic synthesis and stratigraphic significance</mods:title>
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<mods:namePart id="9CCECC50FEAC8C7FEE6F8637B98F7071">Lin, Chien-Hsiang</mods:namePart>
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<mods:namePart id="966AA7700A8630F11591EC345EE8751C">Brzobohatý, Rostislav</mods:namePart>
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<treatment id="615996129468CE3879A2A222FC8894BA" ID-DOI="http://doi.org/10.5281/zenodo.3848317" ID-GBIF-Taxon="164159972" ID-Zenodo-Dep="3848317" LSID="urn:lsid:plazi:treatment:615996129468CE3879A2A222FC8894BA" httpUri="http://treatment.plazi.org/id/615996129468CE3879A2A222FC8894BA" lastPageNumber="15" pageId="14" pageNumber="15">
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<heading id="B20790689468CE3879A2A222FBFB95DE" box="[493,1095,1653,1680]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="15" reason="2">
<taxonomicName id="2EF05C879468CE3879A2A222FBFB95DE" authority="Nolf &amp; Brzobohaty, 1994" authorityName="Nolf &amp; Brzobohaty" authorityYear="1994" box="[493,1095,1653,1680]" class="Actinopterygii" family="Alepocephalidae" genus="Xenodermichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Osmeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="senesi">
<emphasis id="DB84FB169468CE3879A2A222FCAD95DE" bold="true" box="[493,785,1653,1680]" italics="true" pageId="14" pageNumber="15">Xenodermichthys senesi</emphasis>
<bibRefCitation id="8D615AF59468CE387B56A222FBFB95DE" author="Nolf D. &amp; Brzobohaty R." box="[793,1095,1653,1679]" pageId="14" pageNumber="15" pagination="225 - 252" refId="ref22679" refString="Nolf D. &amp; Brzobohaty R. 1994. Fish otoliths from the Late Oligocene (Eger and Kiscell Formations) in the Eger area (northeastern Hungary). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Sciences de la Terre 64: 225 - 252." type="journal article" year="1994">
Nolf &amp;
<bibRefCitation id="8D615AF59468CE387B3BA222FBFB95DE" author="Brzobohaty R." box="[884,1095,1653,1679]" pageId="14" pageNumber="15" pagination="67 - 93" refId="ref21136" refString="Brzobohaty R. 1994. Die Fischotolithen des Badenien von Gainfarn, Niederosterreich (Mittelmiozan, Wiener Becken). Annalen des Naturhistorischen Museums Wien 96 A: 67 - 93." type="journal article" year="1994">Brzobohatý, 1994</bibRefCitation>
</bibRefCitation>
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1EA748F9468CE3878F2A2E0FC8894BA" pageId="14" pageNumber="15" type="discussion">
<paragraph id="E94F27049468CE3878F2A2E0FC8894BA" blockId="14.[189,1399,1718,2027]" pageId="14" pageNumber="15">
A large specimen (
<figureCitation id="71CB3B819468CE3879D8A2E1FE529580" box="[407,494,1718,1745]" captionStart="Fig" captionStartId="13.[189,232,1720,1746]" captionTargetBox="[189,1398,265,1701]" captionTargetId="figure@13.[189,1398,265,1701]" captionTargetPageId="13" captionText="Fig. 2. Tortonian fish otoliths from northern Italy. A. Pterothrissus umbonatus (Koken, 1884), Montegibbio (IRSNB P 9855). B. Paraconger sp., Torrente Stirone (IRSNB P 9856). C. Xenomystax sp., Torrente Stirone (IRSNB P 9857). D. Conger conger (Linnaeus, 1758), Torrente Stirone (IRSNB P 9858). E. Argentina sp., Torrente Stirone (IRSNB P 9859). F. Rhynchoconger pantanellii (Bassoli &amp; Schubert, 1906), Torrente Stirone (IRSNB P 9860). GI. Bathycongrus nagymarosyi (Nolf &amp; Brzobohatý, 1994), Torrente Stirone (IRSNB P 9699P 9701). JK. Gnathophis mystax (Delaroche, 1809), Torrente Stirone (IRSNB P 9702P 9703). LM. Nansenia sp., Mondovi, Madonna della Neve (IRSNB P 9704P 9705). NP. Xenodermichthys senesi Nolf &amp; Brzobohatý, 1994, Mondovi, Madonna della Neve (IRSNB P 9706P 9708). 1 = ventral view; 2 = inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829309" httpUri="https://zenodo.org/record/3829309/files/figure.png" pageId="14" pageNumber="15">Fig. 2P</figureCitation>
) in our material is somewhat higher in its overall shape and the antero-dorsal angle is more pronounced than those of the similar-sized Chattian
<typeStatus id="364B99A69468CE387B98A28DFBB695A5" box="[983,1034,1754,1780]" pageId="14" pageNumber="15">type</typeStatus>
specimens from the
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Basin, southwest
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(
<bibRefCitation id="8D615AF59468CE3879ACA2AAFCB49446" author="Nolf D. &amp; Brzobohaty R." box="[483,776,1789,1815]" pageId="14" pageNumber="15" pagination="225 - 252" refId="ref22679" refString="Nolf D. &amp; Brzobohaty R. 1994. Fish otoliths from the Late Oligocene (Eger and Kiscell Formations) in the Eger area (northeastern Hungary). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Sciences de la Terre 64: 225 - 252." type="journal article" year="1994">
Nolf &amp;
<bibRefCitation id="8D615AF59468CE387A74A2AAFCB49446" author="Brzobohaty R." box="[571,776,1789,1815]" pageId="14" pageNumber="15" pagination="67 - 93" refId="ref21136" refString="Brzobohaty R. 1994. Die Fischotolithen des Badenien von Gainfarn, Niederosterreich (Mittelmiozan, Wiener Becken). Annalen des Naturhistorischen Museums Wien 96 A: 67 - 93." type="journal article" year="1994">Brzobohatý 1994</bibRefCitation>
</bibRefCitation>
: pl. 2, figs 5, 11); the smaller specimens in our material (
<figureCitation id="71CB3B819468CE387889A377FEB5946B" box="[198,265,1824,1850]" captionStart="Fig" captionStartId="13.[189,232,1720,1746]" captionTargetBox="[189,1398,265,1701]" captionTargetId="figure@13.[189,1398,265,1701]" captionTargetPageId="13" captionText="Fig. 2. Tortonian fish otoliths from northern Italy. A. Pterothrissus umbonatus (Koken, 1884), Montegibbio (IRSNB P 9855). B. Paraconger sp., Torrente Stirone (IRSNB P 9856). C. Xenomystax sp., Torrente Stirone (IRSNB P 9857). D. Conger conger (Linnaeus, 1758), Torrente Stirone (IRSNB P 9858). E. Argentina sp., Torrente Stirone (IRSNB P 9859). F. Rhynchoconger pantanellii (Bassoli &amp; Schubert, 1906), Torrente Stirone (IRSNB P 9860). GI. Bathycongrus nagymarosyi (Nolf &amp; Brzobohatý, 1994), Torrente Stirone (IRSNB P 9699P 9701). JK. Gnathophis mystax (Delaroche, 1809), Torrente Stirone (IRSNB P 9702P 9703). LM. Nansenia sp., Mondovi, Madonna della Neve (IRSNB P 9704P 9705). NP. Xenodermichthys senesi Nolf &amp; Brzobohatý, 1994, Mondovi, Madonna della Neve (IRSNB P 9706P 9708). 1 = ventral view; 2 = inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829309" httpUri="https://zenodo.org/record/3829309/files/figure.png" pageId="14" pageNumber="15">Fig. 2</figureCitation>
NO) are characterised by thick otoliths and a convex outer face, and are reasonably similar to the
<typeStatus id="364B99A69468CE3878F2A313FF4C940F" box="[189,240,1860,1886]" pageId="14" pageNumber="15">type</typeStatus>
series from
<collectingRegion id="2B34E9E69468CE3879CEA314FE49940C" box="[385,501,1859,1885]" country="France" name="Aquitaine" pageId="14" pageNumber="15">Aquitaine</collectingRegion>
(
<bibRefCitation id="8D615AF59468CE387A4AA314FC8F940C" author="Nolf D. &amp; Brzobohaty R." box="[517,819,1859,1885]" pageId="14" pageNumber="15" pagination="225 - 252" refId="ref22679" refString="Nolf D. &amp; Brzobohaty R. 1994. Fish otoliths from the Late Oligocene (Eger and Kiscell Formations) in the Eger area (northeastern Hungary). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Sciences de la Terre 64: 225 - 252." type="journal article" year="1994">
Nolf &amp;
<bibRefCitation id="8D615AF59468CE387A2CA314FC8F940C" author="Brzobohaty R." box="[611,819,1859,1885]" pageId="14" pageNumber="15" pagination="67 - 93" refId="ref21136" refString="Brzobohaty R. 1994. Die Fischotolithen des Badenien von Gainfarn, Niederosterreich (Mittelmiozan, Wiener Becken). Annalen des Naturhistorischen Museums Wien 96 A: 67 - 93." type="journal article" year="1994">Brzobohatý 1994</bibRefCitation>
</bibRefCitation>
: pl. 2, figs 610). Because modern alepocephalid otoliths commonly show a great intraspecific variability (e.g.,
<bibRefCitation id="8D615AF59468CE387BDDA331FB0794D0" author="Nolf D. &amp; Brzobohaty R." box="[914,1211,1894,1921]" pageId="14" pageNumber="15" pagination="225 - 252" refId="ref22679" refString="Nolf D. &amp; Brzobohaty R. 1994. Fish otoliths from the Late Oligocene (Eger and Kiscell Formations) in the Eger area (northeastern Hungary). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Sciences de la Terre 64: 225 - 252." type="journal article" year="1994">
Nolf &amp;
<bibRefCitation id="8D615AF59468CE387BA2A331FB0794D0" author="Brzobohaty R." box="[1005,1211,1894,1921]" pageId="14" pageNumber="15" pagination="67 - 93" refId="ref21136" refString="Brzobohaty R. 1994. Die Fischotolithen des Badenien von Gainfarn, Niederosterreich (Mittelmiozan, Wiener Becken). Annalen des Naturhistorischen Museums Wien 96 A: 67 - 93." type="journal article" year="1994">Brzobohatý 1994</bibRefCitation>
</bibRefCitation>
: pl. 2, figs 14), the specimens in question are all assigned to
<taxonomicName id="2EF05C879468CE387A9DA3DDFC8794F2" authorityName="Nolf &amp; Brzobohaty" authorityYear="1994" box="[722,827,1930,1956]" class="Actinopterygii" family="Alepocephalidae" genus="Xenodermichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Osmeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="senesi">
<emphasis id="DB84FB169468CE387A9DA3DDFD5894F2" box="[722,740,1930,1955]" italics="true" pageId="14" pageNumber="15">X</emphasis>
.
<emphasis id="DB84FB169468CE387ABCA3DDFC8794F2" box="[755,827,1930,1955]" italics="true" pageId="14" pageNumber="15">senesi</emphasis>
</taxonomicName>
. The studied material, including large and small specimens, allows us to refer the otoliths figured by
<bibRefCitation id="8D615AF59468CE387B69A3FAFC5E9499" author="Lin C. - H. &amp; Girone A. &amp; Nolf D." box="[806,994,1965,1992]" pageId="14" pageNumber="15" pagination="249 - 261" refId="ref22326" refString="Lin C. - H., Girone A. &amp; Nolf D. 2015. Tortonian fish otoliths from turbiditic deposits in northern Italy: taxonomic and stratigraphic significance. Geobios 48: 249 - 261. https: // doi. org / 10.1016 / j. geobios. 2015.03.003" type="journal article" year="2015">
Lin
<emphasis id="DB84FB169468CE387B19A3F9FC369496" box="[854,906,1965,1991]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. (2015
</bibRefCitation>
: fig. 2(5), as
<taxonomicName id="2EF05C879468CE387C33A3FAFAF49496" authorityName="Gunther" authorityYear="1878" box="[1148,1352,1965,1991]" class="Actinopterygii" family="Alepocephalidae" genus="Xenodermichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Osmeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="DB84FB169468CE387C33A3FAFAF49496" box="[1148,1352,1965,1991]" italics="true" pageId="14" pageNumber="15">Xenodermichthys</emphasis>
</taxonomicName>
aff.
<emphasis id="DB84FB169468CE3878F2A386FF4194BB" box="[189,253,2001,2026]" italics="true" pageId="14" pageNumber="15">copei</emphasis>
(Gill, 1884)) to juvenile specimens of
<taxonomicName id="2EF05C879468CE387A8AA386FC9194BB" authorityName="Nolf &amp; Brzobohaty" authorityYear="1994" box="[709,813,2001,2027]" class="Actinopterygii" family="Alepocephalidae" genus="Xenodermichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Osmeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="senesi">
<emphasis id="DB84FB169468CE387A8AA386FD6B94BB" box="[709,727,2001,2026]" italics="true" pageId="14" pageNumber="15">X</emphasis>
.
<emphasis id="DB84FB169468CE387AAAA386FC9194BB" box="[741,813,2001,2026]" italics="true" pageId="14" pageNumber="15">senesi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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