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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.284.3983" ID-PMC="PMC3677377" ID-Pensoft-Pub="1313-2970-284-1" ID-Pensoft-UUID="192EFF970E2BFFDF3161FFE53C4BFFAA" ID-PubMed="23794843" ID-Zenodo-Dep="578323" ID-ZooBank="33B15A74746D4A82A865EA1E7E55A9BB" ModsDocID="1313-2970-284-1" checkinTime="1451247469560" checkinUser="pensoft" docAuthor="Shi, Hongliang, Zhou, Hongzhang &amp; Liang, Hongbin" docDate="2013" docId="99C7B97C83EE583DA1BC8D97402DE8B2" docLanguage="en" docName="ZooKeys 284: 1-129" docOrigin="ZooKeys 284" docPubDate="2013-04-04" docSource="http://dx.doi.org/10.3897/zookeys.284.3983" docTitle="Diamella arrowi Shi &amp; Zhou &amp; Liang 2013, comb. n." docType="treatment" docVersion="3" id="192EFF970E2BFFDF3161FFE53C4BFFAA" lastPageNumber="46" masterDocId="192EFF970E2BFFDF3161FFE53C4BFFAA" masterDocTitle="Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera" masterLastPageNumber="129" masterPageNumber="1" pageNumber="45" updateTime="1668155586682" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shi, Hongliang</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; Graduate School of Chinese Academy of Sciences, Beijing 100039, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhou, Hongzhang</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liang, Hongbin</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
<mods:nameIdentifier type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-04-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>284</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>129</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.284.3983</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.284.3983</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-284-1</mods:identifier>
<mods:identifier type="ZooBank">33B15A74746D4A82A865EA1E7E55A9BB</mods:identifier>
<mods:identifier type="Pensoft-UUID">192EFF970E2BFFDF3161FFE53C4BFFAA</mods:identifier>
<mods:identifier type="Zenodo-Dep">578323</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182235615" LSID="urn:lsid:plazi:treatment:99C7B97C83EE583DA1BC8D97402DE8B2" httpUri="http://treatment.plazi.org/id/99C7B97C83EE583DA1BC8D97402DE8B2" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<subSubSection pageId="44" pageNumber="45" type="nomenclature">
<paragraph pageId="44" pageNumber="45">
<taxonomicName LSID="99C7B97C-83EE-583D-A1BC-8D97402DE8B2" authority="(Jedlicka, 1935 a)" authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Jedlicka" baseAuthorityYear="1935" class="Insecta" family="Carabidae" genus="Diamella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Diamella arrowi" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="arrowi" status="comb. n.">
Diamella arrowi (
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, 1935a)
</taxonomicName>
<taxonomicNameLabel pageId="44" pageNumber="45">comb. n.</taxonomicNameLabel>
<figureCitation captionStart="Figures 4348" captionStartId="F8" captionText="Figures 43 - 48. Type materials, scale bars = 2.0 mm: 43 Holotype of Anchista pilosa sp. n. 44 Holotype of Metallanchista laticollis sp. n. 45 Holotype of Diamella kaszabi Jedlicka [= Diamella kaszabi (Jedlicka)] 46 Lectotype of Calleida cupreomicans Oberthuer [= Diamella cupreomicans (Oberthuer)] 47 Lectotype of Allocota aerata Bates [= Diamella cupreomicans (Oberthuer)] 48 Holotype of Lachnoderma arrowi Jedlicka [= Diamella arrowi (Jedlicka)]." httpUri="https://binary.pensoft.net/fig/29681" pageId="44" pageNumber="45">Habitus: Fig. 48</figureCitation>
<figureCitation captionStart="Figure 78" captionStartId="F23" captionText="Figure 78. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Diamella arrowi (Jedlicka), holotype." httpUri="https://binary.pensoft.net/fig/29696" pageId="44" pageNumber="45">male genitalia: Figs 78</figureCitation>
<figureCitation captionStart="Figures 95110" captionStartId="F40" captionText="Figures 95 - 110. Left and right parameres of male genitalia, scale bars = 0.2 mm: 95 Paraphaea binotata (Dejean), a specimen from Guangxi 96 Paraphaea minor sp. n., holotype 97 Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir 98 Anchista pilosa sp. n., holotype 99 Metallanchista laticollis sp. n., holotype 100 Physodera amplicollis van de Poll, a specimen from Taiwan 101 Diamella cupreomicans (Oberthuer), a specimen from Yunnan 102 Diamella arrowi (Jedlicka), holotype 103 Allocota aurata (Bates), a specimen from Tonkin 104 Allocota aurata (Bates), a specimen from Shaanxi 105 Allocota bicolor sp. n., holotype 106 Lachnoderma asperum Bates, a specimen from Taiwan 107 Dasiosoma ivorense Basilewsky, a paratype from Cote d'Ivoire 108 Dasiosoma quadraticolle sp. n., holotype 109 Orionella kathmanduensis (Kirschenhofer), a specimen from Nepal 110 Endynomena pradieri (Fairmaire), a specimen from Fujian." httpUri="https://binary.pensoft.net/fig/29716" pageId="44" pageNumber="45">102</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="44" pageNumber="45" type="reference_group">
<paragraph pageId="44" pageNumber="45">
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Jedlicka" baseAuthorityYear="1935" class="Insecta" family="Carabidae" genus="Diamella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="arrowi" status="comb. n.">Diamella arrowi</taxonomicName>
<bibRefCitation author="Jedlicka, A" journalOrPublisher="Sbornik Entomologickeho Oddeleni pri Zoologichych Sbirkach Narodniho Musea v Praze" pageId="87" pageNumber="88" refId="B54" refString="Jedlicka, A, 1935a. Neue Carabiden aus Ostasien. (10. Teil.). Praha: A. Jedlicka, 20 pp." title="Neue Carabiden aus Ostasien. (10. Teil.). Praha: A. Jedlicka, 20 pp." year="1935 a">
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
1935a
</bibRefCitation>
: 15 (original:
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Insecta" family="Carabidae" genus="Lachnoderma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus" status="comb. n.">
<emphasis italics="true" pageId="44" pageNumber="45">Lachnoderma</emphasis>
</taxonomicName>
; type locality: Philippines; holotype deposited in NHML);
<bibRefCitation author="Jedlicka, A" journalOrPublisher="Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden" pageId="87" pageNumber="88" pagination="269 - 579" refId="B58" refString="Jedlicka, A, 1963. Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden 28: 269 - 579" title="Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae)." volume="28" year="1963">
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
1963
</bibRefCitation>
: 305 (
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus" status="comb. n.">
<emphasis italics="true" pageId="44" pageNumber="45">Allocota</emphasis>
</taxonomicName>
);
<bibRefCitation author="Kirschenhofer, E" journalOrPublisher="Linzer Biologische Beitraege" pageId="87" pageNumber="88" pagination="757 - 799" refId="B62" refString="Kirschenhofer, E, 1996. Weitere neue und wenig bekannte Carabidae aus Sued- und Suedostasien (Coleoptera: Carabidae: Lebiinae: Colliurinae, Callistinae, Hexagoninae, Brachininae, Panagaeinae). II. Teil. Linzer Biologische Beitraege 28 (2): 757 - 799" title="Weitere neue und wenig bekannte Carabidae aus Sued- und Suedostasien (Coleoptera: Carabidae: Lebiinae: Colliurinae, Callistinae, Hexagoninae, Brachininae, Panagaeinae). II. Teil." volume="28" year="1996">Kirschenhofer 1996</bibRefCitation>
: 764 (
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus" status="comb. n.">
<emphasis italics="true" pageId="44" pageNumber="45">Allocota</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="type examined">
<paragraph pageId="44" pageNumber="45">Type examined.</paragraph>
<paragraph lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<emphasis bold="true" pageId="44" pageNumber="45">Holotype</emphasis>
of
<taxonomicName authorityName="Jedlicka" authorityYear="1935" class="Insecta" family="Carabidae" genus="Lachnoderma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="arrowi">
<emphasis italics="true" pageId="44" pageNumber="45">Lachnoderma arrowi</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, by original designation (NHML): male, body length = 7.3 mm, board mounted, &quot;Philippine Is. / Coll.
<pageBreakToken pageId="45" pageNumber="46" start="start">Bottcher</pageBreakToken>
. / B. M. 1929-201&quot;; &quot;
<emphasis italics="true" pageId="45" pageNumber="46">96</emphasis>
&quot;;
<normalizedToken originalValue="“Type”">&quot;Type&quot;</normalizedToken>
[round label with red ring]; &quot;
<emphasis italics="true" pageId="45" pageNumber="46">Lachnoderma</emphasis>
/
<emphasis italics="true" pageId="45" pageNumber="46">Arrowi / Jedl. sp. n</emphasis>
. / det. ING.
<normalizedToken originalValue="JEDLIČKA”">JEDLICKA&quot;</normalizedToken>
[pink label] [Figs 48, 78, 102].
</paragraph>
</subSubSection>
<subSubSection pageId="45" pageNumber="46" type="notes on type">
<paragraph pageId="45" pageNumber="46">Notes on type.</paragraph>
<paragraph pageId="45" pageNumber="46">
The original literature clearly indicated that this specimen (
<figureCitation captionStart="Figures 4348" captionStartId="F8" captionText="Figures 43 - 48. Type materials, scale bars = 2.0 mm: 43 Holotype of Anchista pilosa sp. n. 44 Holotype of Metallanchista laticollis sp. n. 45 Holotype of Diamella kaszabi Jedlicka [= Diamella kaszabi (Jedlicka)] 46 Lectotype of Calleida cupreomicans Oberthuer [= Diamella cupreomicans (Oberthuer)] 47 Lectotype of Allocota aerata Bates [= Diamella cupreomicans (Oberthuer)] 48 Holotype of Lachnoderma arrowi Jedlicka [= Diamella arrowi (Jedlicka)]." httpUri="https://binary.pensoft.net/fig/29681" pageId="45" pageNumber="46">Fig. 48</figureCitation>
) is the holotype. But precise locality was not provided.
</paragraph>
</subSubSection>
<subSubSection pageId="45" pageNumber="46" type="diagnosis">
<paragraph pageId="45" pageNumber="46">Diagnosis.</paragraph>
<paragraph pageId="45" pageNumber="46">
Vertex moderately tumid, not forming horn-like humps at posterior supraorbital seta insertions; pronotum front angles not narrowed, lateral margins completely rounded in middle; elytra dark blue, strongly metallic; elytra with striae very shallow, with accessory setae only on odd intervals. Male genitalia of this species resemble
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Oberthuer" baseAuthorityYear="1883" class="Insecta" family="Carabidae" genus="Diamella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="cupreomicans">
<emphasis italics="true" pageId="45" pageNumber="46">Diamella cupreomicans</emphasis>
</taxonomicName>
, but: (1) apical lamella very short and wide, apex rounded; (2) in lateral view, median lobe of aedeagus slender, ventral margin less expanded near base.
</paragraph>
</subSubSection>
<subSubSection pageId="45" pageNumber="46" type="description">
<paragraph pageId="45" pageNumber="46">Description.</paragraph>
<paragraph pageId="45" pageNumber="46">
Body length 7.3 mm; head and pronotum reddish brown, with luster but not metallic, lateral explanate areas of pronotum paler; elytra metallic blue with disc somewhat reddish brown; mouthparts and antennae reddish brown, apices of terminal labial palpomeres yellow; legs brown, tarsomeres slightly paler; ventral side brownish.
<emphasis bold="true" pageId="45" pageNumber="46">Head</emphasis>
without microsculpture or punctures, sparsely pubescent on vertex; tempora slightly longer than half length of eyes; vertex moderately tumid, posterior supraorbital setae distant from eyes, seta insertions slightly humped, but not horn-like; labrum slightly widened to apex, apical margin deeply emarginate.
<emphasis bold="true" pageId="45" pageNumber="46">Pronotum</emphasis>
slightly wider than head, widest slightly before middle; ratio PW/PL 1.45; pronotal base briefly but distinctly lobed; front angles wide; lateral margins strongly expanded in middle, completely rounded, slightly sinuate before hind angles; hind angles subrectangular, hardly projected; disc slightly convex, microsculpture indistinct; lateral explanate areas wide and even; basal foveae shallow; disc nearly impunctate, with a few punctures along median line and in basal foveae; disc with long setae except area along median line, some setae distributed to middle of lateral explanate areas; lateral margins with long accessory setae through their full lengths, sparser after middle; median line distinct, nearly reaching basal and apical margins; disc not rugose.
<emphasis bold="true" pageId="45" pageNumber="46">Elytra</emphasis>
wider than pronotum, shoulders not narrowed, hind wings well developed; elytral lateral borders narrow, not reflexed up; striae very shallow, finely punctate; scutellary intervals with three large pores (basal pores); intervals flat, without microsculpture, odd intervals with secondary pores, more abundant on 7th interval; primary pores indistinct among secondary pores, 3rd and 5th intervals with some larger pores distinguishable; setae not present along striae; umbilical series of 9th interval composed of 15 pores; disc without distinct depression; lateral margins slightly depressed at basal third; 7th and 8th intervals slightly tumid near apex; elytral lateral margins with dense and fine setae; epipleura pubescent.
<emphasis bold="true" pageId="45" pageNumber="46">Ventral side</emphasis>
. Prosternum with long pubescence, proepisterna glabrous; mesosternum nearly glabrous; abdomen hairy.
<emphasis bold="true" pageId="45" pageNumber="46">Male genitalia</emphasis>
with median lobe of aedeagus straight and flat, gently expanded to apex; dorsal side slightly pubescent subapically; left lateral margin slightly curved in middle; apical orifice opened apically; apical lamella placed on right side, short and rounded, distinctly wider than length in dorsal view; internal sac with main flagellum thick and slightly corkscrewed, distinctly grooved ventrally, nearly reaching apical orifice; trumpet-form expansion strongly expanded, nearly half length of main flagellum (
<figureCitation captionStart="Figure 78" captionStartId="F23" captionText="Figure 78. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Diamella arrowi (Jedlicka), holotype." httpUri="https://binary.pensoft.net/fig/29696" pageId="45" pageNumber="46">Fig. 78</figureCitation>
).
<emphasis bold="true" pageId="45" pageNumber="46">Female genitalia</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="45" pageNumber="46" type="distribution">
<paragraph pageId="45" pageNumber="46">Distribution</paragraph>
<paragraph pageId="45" pageNumber="46">
(
<figureCitation captionStart="Map 5" captionStartId="F49" captionText="Map 5. Known distribution of genus Diamella: ● Diamella cupreomicans (Oberthuer) ◆ Diamella kaszabi (Jedlicka) ★ Diamella arrowi (Jedlicka)." httpUri="https://binary.pensoft.net/fig/29727" pageId="45" pageNumber="46">Map 5</figureCitation>
). Only known from the type locality, the Philippines.
</paragraph>
</subSubSection>
</treatment>
</document>