114 lines
11 KiB
XML
114 lines
11 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/fr.25.83781" ID-Pensoft-Pub="2193-0074-1-99" ID-Pensoft-UUID="2F5D6AE2EEB55A17ACF1623B06B4EA8D" ID-ZooBank="7A16698D4F1848D29D9651A6E0CC15AC" ModsDocID="2193-0074-25-1-99" checkinTime="1667366289781" checkinUser="pensoft" docAuthor="Vasilyan, Davit, Cernansky, Andrej, Szyndlar, Zbigniew & Moers, Thomas" docDate="2022" docId="582E10CB531754898341C6ED8194E1C4" docLanguage="en" docName="FossilRecord 25(1): 99-145" docOrigin="Fossil Record 25 (1)" docPubDate="2022-05-10" docSource="http://dx.doi.org/10.3897/fr.25.83781" docTitle="N. ” indet." docType="treatment" docVersion="1" id="2F5D6AE2EEB55A17ACF1623B06B4EA8D" lastPageNumber="99" masterDocId="2F5D6AE2EEB55A17ACF1623B06B4EA8D" masterDocTitle="Amphibian and reptilian fauna from the early Miocene of Echzell, Germany" masterLastPageNumber="145" masterPageNumber="99" pageNumber="99" updateTime="1667366289781" updateUser="pensoft">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>Amphibian and reptilian fauna from the early Miocene of Echzell, Germany</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Vasilyan, Davit</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8712-0678</mods:nameIdentifier>
|
|
<mods:affiliation>JURASSICA Museum, Route de Fontenais 21. 2900 Porrentruy, Switzerland & Department of Geosciences, University of Fribourg, Chemin du musee 6, 1700, Fribourg, Switzerland</mods:affiliation>
|
|
<mods:nameIdentifier type="email">davit.vasilyan@jurassica.ch</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Cernansky, Andrej</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8920-2503</mods:nameIdentifier>
|
|
<mods:affiliation>Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynska dolina, Ilkovicova 6, 842 15 Bratislava, Slovakia</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Szyndlar, Zbigniew</mods:namePart>
|
|
<mods:affiliation>Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Slawkowska 17, 31 - 016 Krakow, Poland</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Moers, Thomas</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2268-5824</mods:nameIdentifier>
|
|
<mods:affiliation>Department of Palaeobiology, Swedish Museum of Natural History, P. O. Box 50007, 104 05 Stockholm, Sweden & Bolin Centre for Climate Research, Stockholm University, Stockholm, Sweden</mods:affiliation>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>Fossil Record</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2022</mods:date>
|
|
<mods:detail type="pubDate">
|
|
<mods:number>2022-05-10</mods:number>
|
|
</mods:detail>
|
|
<mods:detail type="volume">
|
|
<mods:number>25</mods:number>
|
|
</mods:detail>
|
|
<mods:detail type="issue">
|
|
<mods:number>1</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>99</mods:start>
|
|
<mods:end>145</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location>
|
|
<mods:url>http://dx.doi.org/10.3897/fr.25.83781</mods:url>
|
|
</mods:location>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/fr.25.83781</mods:identifier>
|
|
<mods:identifier type="Pensoft-Pub">2193-0074-1-99</mods:identifier>
|
|
<mods:identifier type="ZooBank">7A16698D4F1848D29D9651A6E0CC15AC</mods:identifier>
|
|
<mods:identifier type="Pensoft-UUID">2F5D6AE2EEB55A17ACF1623B06B4EA8D</mods:identifier>
|
|
</mods:mods>
|
|
<treatment LSID="urn:lsid:plazi:treatment:582E10CB531754898341C6ED8194E1C4" httpUri="http://treatment.plazi.org/id/582E10CB531754898341C6ED8194E1C4" lastPageNumber="99" pageId="0" pageNumber="99">
|
|
<subSubSection pageId="0" pageNumber="99" type="nomenclature">
|
|
<paragraph pageId="0" pageNumber="99">
|
|
"
|
|
<taxonomicName LSID="582E10CB-5317-5489-8341-C6ED8194E1C4" authority="” indet." authorityName="” indet." family="Colubridae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Squamata" pageId="0" pageNumber="99" rank="subFamily" subFamily="Colubrinae">Colubrinae " indet.</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="99" type="description">
|
|
<paragraph pageId="0" pageNumber="99">
|
|
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Snakes from the Echzell. A-E. Bavarioboa cf. hermi (HLMD-Ez 2148), middle trunk vertebra; F-J. " Colubrinae " indet. (HLMD-Ez 2159), middle trunk vertebra; K, L. Naja cf. romani (HLMD-Ez 2151), middle trunk vertebra. All vertebrae in (A, F, K) lateral, (B, G, L) dorsal, (C, H, M) ventral, (D, I, N) anterior and (E, J, O) posterior views." figureDoi="10.3897/fr.25.83781.figure18" httpUri="https://binary.pensoft.net/fig/682238" pageId="0" pageNumber="99">Fig. 18F-J</figureCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="99" type="materials_examined">
|
|
<paragraph pageId="0" pageNumber="99">Material.</paragraph>
|
|
<paragraph pageId="0" pageNumber="99">Seventy trunk vertebrae HLMD-Ez 2159 and HLMD-Ez 2159a.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="99" type="description">
|
|
<paragraph pageId="0" pageNumber="99">Description.</paragraph>
|
|
<paragraph pageId="0" pageNumber="99">
|
|
Vertebrae: All vertebrae classified here as "
|
|
<taxonomicName authorityName="Oppel" authorityYear="1811" lsidName="" pageId="0" pageNumber="99" rank="subFamily" subFamily="Colubrinae">Colubrinae</taxonomicName>
|
|
" are preserved in more or less fragmentary state; most are badly preserved. The largest vertebra, coming from the middle trunk portion of the column HLMD-Ez 2159 (Fig.
|
|
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Snakes from the Echzell. A-E. Bavarioboa cf. hermi (HLMD-Ez 2148), middle trunk vertebra; F-J. " Colubrinae " indet. (HLMD-Ez 2159), middle trunk vertebra; K, L. Naja cf. romani (HLMD-Ez 2151), middle trunk vertebra. All vertebrae in (A, F, K) lateral, (B, G, L) dorsal, (C, H, M) ventral, (D, I, N) anterior and (E, J, O) posterior views." figureDoi="10.3897/fr.25.83781.figure18" httpUri="https://binary.pensoft.net/fig/682238" pageId="0" pageNumber="99">18F-J</figureCitation>
|
|
), belonged to a relatively small-sized snake (centrum length measures 4.5 mm, centrum width - 3.6, centrum length / centrum width ratio equals 1.25). It lacks both prezygapophyseal processes and right postzygapophysis. The centrum of most vertebrae is subtriangular in ventral view and slightly longer than wide in the largest vertebrae. In a number of smaller vertebrae, interpreted as posterior trunk vertebrae, the centrum is more elongated. The subcentral ridges are well developed. The haemal keel is distinct and slightly widening before reaching the condyle base. The neural arch is moderately vaulted, not accompanied by any epizygapophyseal spine. The neural spine (partly preserved in few vertebrae only) is twice longer than high approximately. The zygosphenal roof (preserved in few vertebrae) is roughly straight or consisting of three indistinct lobes. The prezygapophyseal articular facets are oval (Fig.
|
|
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Snakes from the Echzell. A-E. Bavarioboa cf. hermi (HLMD-Ez 2148), middle trunk vertebra; F-J. " Colubrinae " indet. (HLMD-Ez 2159), middle trunk vertebra; K, L. Naja cf. romani (HLMD-Ez 2151), middle trunk vertebra. All vertebrae in (A, F, K) lateral, (B, G, L) dorsal, (C, H, M) ventral, (D, I, N) anterior and (E, J, O) posterior views." figureDoi="10.3897/fr.25.83781.figure18" httpUri="https://binary.pensoft.net/fig/682238" pageId="0" pageNumber="99">18G</figureCitation>
|
|
), the postzygapophyseal facets are usually subsquare in shape (Fig.
|
|
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Snakes from the Echzell. A-E. Bavarioboa cf. hermi (HLMD-Ez 2148), middle trunk vertebra; F-J. " Colubrinae " indet. (HLMD-Ez 2159), middle trunk vertebra; K, L. Naja cf. romani (HLMD-Ez 2151), middle trunk vertebra. All vertebrae in (A, F, K) lateral, (B, G, L) dorsal, (C, H, M) ventral, (D, I, N) anterior and (E, J, O) posterior views." figureDoi="10.3897/fr.25.83781.figure18" httpUri="https://binary.pensoft.net/fig/682238" pageId="0" pageNumber="99">18H</figureCitation>
|
|
). The prezygapophyseal processes are damaged in all vertebrae. Remnants of the processes preserved in one (?posterior) vertebra indicate that the apophyses may have been slender and relatively long (as long or almost as long as the prezygophyseal articular facets). The paradiapophyses (usually eroded) are moderately developed, with the dia- and parapophyseal portions of roughly equal length (Fig.
|
|
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Snakes from the Echzell. A-E. Bavarioboa cf. hermi (HLMD-Ez 2148), middle trunk vertebra; F-J. " Colubrinae " indet. (HLMD-Ez 2159), middle trunk vertebra; K, L. Naja cf. romani (HLMD-Ez 2151), middle trunk vertebra. All vertebrae in (A, F, K) lateral, (B, G, L) dorsal, (C, H, M) ventral, (D, I, N) anterior and (E, J, O) posterior views." figureDoi="10.3897/fr.25.83781.figure18" httpUri="https://binary.pensoft.net/fig/682238" pageId="0" pageNumber="99">18I</figureCitation>
|
|
). The cotyle and condyle are slightly depressed dorso-ventrally. The lateral, subcentral and paracotylar foramina are small but distinct.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="99" type="remarks">
|
|
<paragraph pageId="0" pageNumber="99">Remarks.</paragraph>
|
|
<paragraph pageId="0" pageNumber="99">
|
|
Owing to the bad preservation state of the aforementioned remains, we refrain from identifying the fossil snake to the genus level. The remains resemble roughly those of a number of small
|
|
<normalizedToken originalValue="“colubrines”">"colubrines"</normalizedToken>
|
|
(i.e., "
|
|
<taxonomicName authorityName="Oppell" authorityYear="1811" class="Reptilia" family="Colubridae" kingdom="Animalia" lsidName="" order="Squamata" pageId="0" pageNumber="99" phylum="Chordata" rank="family">Colubridae</taxonomicName>
|
|
" devoid of hypapophyses throughout the column) reported from several early Miocene sites (
|
|
<bibRefCitation DOI="https://doi.org/10.2113/gssgfbull.183.6.661" author="Szyndlar, Z" journalOrPublisher="Bulletin de la Societe Geologique de France" pageId="0" pageNumber="99" pagination="661 - 681" refId="B124" refString="Szyndlar, Z, 2012. Early Oligocene to Pliocene Colubridae of Europe: a review. Bulletin de la Societe Geologique de France 183: 661 - 681, DOI: https://doi.org/10.2113/gssgfbull.183.6.661" title="Early Oligocene to Pliocene Colubridae of Europe: a review." url="https://doi.org/10.2113/gssgfbull.183.6.661" volume="183" year="2012">Szyndlar 2012</bibRefCitation>
|
|
). We cannot exclude the possibility that the fossils belonged to more than one taxon, but it cannot be proved.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |