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<document id="560F6C436C3C6B12041B15CE03D7DA82" ID-CLB-Dataset="31292" ID-DOI="10.11646/zootaxa.4358.1.7" ID-GBIF-Dataset="67de317b-9544-4420-95d1-2577db7d9f57" ID-ISSN="1175-5326" ID-Zenodo-Dep="1067554" ID-ZooBank="84494502-9E85-49DA-8530-092AF8918D88" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511863712862" checkinUser="plazi" docAuthor="Gerringer, Mackenzie E., Linley, Thomas D., Jamieson, Alan J., Goetze, Erica &amp; Drazen, Jeffrey C." docDate="2017" docId="0391DE73FF98FF9198B7FA6F5F959253" docLanguage="en" docName="zootaxa.4358.1.7.pdf.imf" docOrigin="Zootaxa 4358 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudoliparis swirei Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen 2017, sp. nov." docType="treatment" docVersion="8" lastPageNumber="170" masterDocId="FFA8A60BFF9BFF989820FFA35C5E950B" masterDocTitle="Pseudoliparis swirei sp. nov.: A newly-discovered hadal snailfish (Scorpaeniformes: Liparidae) from the Mariana Trench" masterLastPageNumber="177" masterPageNumber="161" pageNumber="164" updateTime="1698505016645" updateUser="ExternalLinkService">
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<mods:title id="ECFD08BCED2F1C3AFA524973FF2F23EF">Pseudoliparis swirei sp. nov.: A newly-discovered hadal snailfish (Scorpaeniformes: Liparidae) from the Mariana Trench</mods:title>
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<mods:namePart id="0CB4914533A7DB6CC514B68AE05F2B9B">Gerringer, Mackenzie E.</mods:namePart>
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<mods:namePart id="418D2698BD3D1FF1FA683588F3E48E32">Linley, Thomas D.</mods:namePart>
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<mods:namePart id="5F888F28FA370DF6CE4FDC40DD29F7B6">Jamieson, Alan J.</mods:namePart>
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<mods:namePart id="D7CCBD62CD8A2D7C019B6DA0D9AB9BE4">Goetze, Erica</mods:namePart>
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<mods:namePart id="6BD3D495AD1A3F075DC08AB36AD66405">Drazen, Jeffrey C.</mods:namePart>
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<treatment id="0391DE73FF98FF9198B7FA6F5F959253" ID-DOI="http://doi.org/10.5281/zenodo.6016523" ID-GBIF-Taxon="137580722" ID-Zenodo-Dep="6016523" LSID="urn:lsid:plazi:treatment:0391DE73FF98FF9198B7FA6F5F959253" httpUri="http://treatment.plazi.org/id/0391DE73FF98FF9198B7FA6F5F959253" lastPageId="9" lastPageNumber="170" pageId="3" pageNumber="164">
<subSubSection id="C3223CEEFF98FF9B98B7FA6F5EBF90ED" box="[151,737,1484,1510]" pageId="3" pageNumber="164" type="nomenclature">
<paragraph id="8B876F65FF98FF9B98B7FA6F5EBF90ED" blockId="3.[151,737,1484,1543]" box="[151,737,1484,1510]" pageId="3" pageNumber="164">
<taxonomicName id="4C3814E6FF98FF9B98B7FA6F5E2290EE" ID-CoL="4NZRR" authority="Gerringer &amp; Linley" authorityName="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen" authorityYear="2017" box="[151,636,1484,1510]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="164" phylum="Chordata" rank="species" species="swirei" status="sp. nov.">
<emphasis id="B94CB377FF98FF9B98B7FA6F5DD590ED" bold="true" box="[151,395,1484,1510]" italics="true" pageId="3" pageNumber="164">Pseudoliparis swirei</emphasis>
<emphasis id="B94CB377FF98FF9B99B1FA6F5E2290EE" bold="true" box="[401,636,1484,1509]" pageId="3" pageNumber="164">Gerringer &amp; Linley</emphasis>
</taxonomicName>
<emphasis id="B94CB377FF98FF9B9AA3FA6F5EBF90ED" bold="true" box="[643,737,1484,1510]" pageId="3" pageNumber="164">
<taxonomicNameLabel id="A27F0E0CFF98FF9B9AA3FA6F5EBF90ED" box="[643,737,1484,1510]" pageId="3" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
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<subSubSection id="C3223CEEFF98FF9B98B7FA4D5D82930C" box="[151,476,1518,1543]" pageId="3" pageNumber="164" type="description">
<paragraph id="8B876F65FF98FF9B98B7FA4D5D82930C" blockId="3.[151,737,1484,1543]" box="[151,476,1518,1543]" pageId="3" pageNumber="164">
<figureCitation id="130373E0FF98FF9B98B7FA4D5D5C930C" box="[151,258,1518,1543]" captionStart="FIGURE 2" captionStartId="4.[151,250,1453,1475]" captionTargetBox="[474,1113,373,1432]" captionTargetId="figure@4.[474,1113,373,1432]" captionTargetPageId="4" captionText="FIGURE 2. A) In situ photograph of Pseudoliparis swirei sp. nov. at 6,198 m. B) a group at 7,485 m. C) Deck photograph of SIO 16-82/HADES 200049.D) Radiograph of SIO 16-86/HADES 200141. Image by Sandra Raredon. Scale indicator 5 cm." httpUri="https://zenodo.org/record/1067558/files/figure.png" pageId="3" pageNumber="164">Figures 2</figureCitation>
,
<figureCitation id="130373E0FF98FF9B992EFA4C5D40930C" box="[270,286,1519,1543]" captionStart="FIGURE 3" captionStartId="4.[151,250,1897,1919]" captionTargetBox="[151,1436,1555,1875]" captionTargetId="figure@4.[151,1436,1555,1875]" captionTargetPageId="4" captionText="FIGURE 3. Lateral view of Pseudoliparis swirei sp. nov.. Combined representation of holotype, paratypes, and freshly captured images of paratype USNM 438985/HADES 200133, juvenile, 151 mm.Drawings by Thomas D. Linley." httpUri="https://zenodo.org/record/1067560/files/figure.png" pageId="3" pageNumber="164">3</figureCitation>
,
<figureCitation id="130373E0FF98FF9B990AFA4C5D64930C" box="[298,314,1519,1543]" captionStart="FIGURE 4" captionStartId="5.[151,250,1449,1471]" captionTargetBox="[151,1436,199,1428]" captionTargetId="figure@5.[151,1436,193,1428]" captionTargetPageId="5" captionText="FIGURE 4. A) Ventral view drawing. B) Tooth pattern on maxillary jaw. Counted rows denoted on right side of image. C) Tooth structure of specimen USNM 438996/HADES 200024. D) Disk details of SIO 16-87/HADES 200025 stained with Alizarin Red S. E) Disk of USNM 438995/HADES 200085 stained with Cyanine Blue." httpUri="https://zenodo.org/record/1067562/files/figure.png" pageId="3" pageNumber="164">4</figureCitation>
,
<figureCitation id="130373E0FF98FF9B9966FA4D5D08930C" box="[326,342,1518,1543]" captionStart="FIGURE 5" captionStartId="6.[151,250,1945,1967]" captionTargetBox="[242,1275,843,1911]" captionTargetId="figure@6.[241,1319,842,1911]" captionTargetPageId="6" captionText="FIGURE 5. Pectoral girdle of USNM 438994/HADES 200027, female, 220 mm. Ventral drawing of USNM 438995/HADES 200085, female, 225 mm.Scapula, radials 14, coracoid shown. Drawingby Thomas D. Linley." httpUri="https://zenodo.org/record/1067564/files/figure.png" pageId="3" pageNumber="164">5</figureCitation>
;
<tableCitation id="C6BA5ADEFF98FF9B9943FA4D5D9C930C" box="[355,450,1518,1543]" captionStart="TABLE 2" captionStartId="7.[151,239,152,175]" captionText="TABLE 2. Measurements and counts of Pseudoliparis swirei sp. nov.. Measurements taken from preserved specimens, given in mm. Body depth measured from fresh photographs. Dorsal fin origin between vertebrae #, # are shown. Maximum and minimum values (mean and standard deviation) shown for measurements. Counts show range (median). Number of specimens examined for individual characters (n)." pageId="3" pageNumber="164">Tables 2</tableCitation>
,
<tableCitation id="C6BA5ADEFF98FF9B99EEFA4C5D82930C" box="[462,476,1519,1543]" captionStart="TABLE 3" captionStartId="8.[151,239,152,175]" captionText="TABLE 3. Ratios of Pseudoliparis swirei sp. nov.. Preserved and fresh measurements presented. Body depth taken from photographs. Predorsal fin length measured from radiograph." pageId="3" pageNumber="164">3</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3223CEEFF98FF9B98B7F9955F1293A3" pageId="3" pageNumber="164" type="reference_group">
<paragraph id="8B876F65FF98FF9B98B7F9955EB69347" blockId="3.[151,844,1590,1704]" box="[151,744,1590,1613]" pageId="3" pageNumber="164">
Mariana snailfish:
<bibRefCitation id="EFA91294FF98FF9B9974F9955E579347" author="Linley" box="[340,521,1590,1613]" pageId="3" pageNumber="164" refString="Linley, T. D., Gerringer, M. E., Yancey, P. H., Drazen, J. C., Weinstock, C. L. &amp; Jamieson, A. J. (2016) Fishes of the hadal zone including new species, in situ observations and depth records of Liparidae. Deep Sea Research Part I: Oceanographic Research Papers, 114, 99 - 110. https: // doi. org / 10.1016 / j. dsr. 2016.05.003" type="journal article" year="2016">
Linley
<emphasis id="B94CB377FF98FF9B99BDF99B5D8E9347" box="[413,464,1590,1613]" italics="true" pageId="3" pageNumber="164">et al.</emphasis>
2016
</bibRefCitation>
(page 105,
<figureCitation id="130373E0FF98FF9B9AA0F9955EBF9347" box="[640,737,1590,1612]" captionStart="FIGURE 4" captionStartId="5.[151,250,1449,1471]" captionTargetBox="[151,1436,199,1428]" captionTargetId="figure@5.[151,1436,193,1428]" captionTargetPageId="5" captionText="FIGURE 4. A) Ventral view drawing. B) Tooth pattern on maxillary jaw. Counted rows denoted on right side of image. C) Tooth structure of specimen USNM 438996/HADES 200024. D) Disk details of SIO 16-87/HADES 200025 stained with Alizarin Red S. E) Disk of USNM 438995/HADES 200085 stained with Cyanine Blue." httpUri="https://zenodo.org/record/1067562/files/figure.png" pageId="3" pageNumber="164">Figure 4a</figureCitation>
)
</paragraph>
<paragraph id="8B876F65FF98FF9B98B7F9F65EAE9360" blockId="3.[151,844,1590,1704]" box="[151,752,1621,1643]" pageId="3" pageNumber="164">
Mariana snailfish: Linley
<emphasis id="B94CB377FF98FF9B99BDF9F55D979360" box="[413,457,1621,1643]" italics="true" pageId="3" pageNumber="164">et al</emphasis>
. 2017 (page 42, Figure 6.43)
</paragraph>
<paragraph id="8B876F65FF98FF9B98B7F9D75F179381" blockId="3.[151,844,1590,1704]" box="[151,841,1652,1674]" pageId="3" pageNumber="164">
Mariana snailfish/Mariana liparid:
<bibRefCitation id="EFA91294FF98FF9B99D8F9D75E869381" author="Gerringer" box="[504,728,1652,1674]" pageId="3" pageNumber="164" refString="Gerringer, M. E., Popp, B. N., Linley, T. D., Jamieson, A. J. &amp; Drazen, J. C. (2017 a) Comparative feeding ecology of abyssal and hadal fishes through stomach content and amino acid isotope analysis. Deep-Sea Research Part I: Oceanographic Research Papers 121, 110 - 120. https: // doi. org / 10.1016 / j. dsr. 2017.01.003" type="journal article" year="2017" yearSuffix="a">
Gerringer
<emphasis id="B94CB377FF98FF9B9A41F9D65ECA9381" box="[609,660,1652,1674]" italics="true" pageId="3" pageNumber="164">et al.</emphasis>
2017a
</bibRefCitation>
(page 111)
</paragraph>
<paragraph id="8B876F65FF98FF9B98B7F9315F1293A3" blockId="3.[151,844,1590,1704]" box="[151,844,1682,1704]" pageId="3" pageNumber="164">
Mariana liparid/
<taxonomicName id="4C3814E6FF98FF9B991BF9315DC293A3" ID-CoL="624XH" box="[315,412,1682,1704]" class="Actinopterygii" family="Liparidae" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="164" phylum="Chordata" rank="family">Liparidae</taxonomicName>
sp. nov:
<bibRefCitation id="EFA91294FF98FF9B99D8F9315E8793A3" author="Gerringer" box="[504,729,1682,1704]" pageId="3" pageNumber="164" refString="Gerringer, M. E., Drazen, J. C. &amp; Yancey, P. H. (2017 b) Metabolic enzyme activities of abyssal and hadal fishes: pressure effects and a re-evaluation of depth-related changes. Deep-Sea Research Part I: Oceanographic Research Papers, 125, 135 - 146. https: // doi. org / 10.1016 / j. dsr. 2017.05.010" type="journal article" year="2017" yearSuffix="b">
Gerringer
<emphasis id="B94CB377FF98FF9B9A41F9305ECA93A3" box="[609,660,1682,1704]" italics="true" pageId="3" pageNumber="164">et al.</emphasis>
2017b
</bibRefCitation>
(page 137)
</paragraph>
</subSubSection>
<subSubSection id="C3223CEEFF98FF9C98B7F9765828944B" lastPageId="4" lastPageNumber="165" pageId="3" pageNumber="164" type="diagnosis">
<paragraph id="8B876F65FF98FF9B98B7F97659C392E2" blockId="3.[151,1437,1749,2026]" pageId="3" pageNumber="164">
<emphasis id="B94CB377FF98FF9B98B7F9765D4F93E5" bold="true" box="[151,273,1749,1774]" pageId="3" pageNumber="164">Diagnosis.</emphasis>
<bibRefCitation id="EFA91294FF98FF9B9938F9765E3693E5" author="Andriashev" box="[280,616,1749,1774]" pageId="3" pageNumber="164" refString="Andriashev, A. P. &amp; Pitruk, D. L. (1993) A review of the ultra-abyssal (hadal) genus Pseudoliparis (Scorpaeniformes, Liparidae) with a description of a new species from the Japan Trench. Voprosy ikhtiologii, 33, 325 - 330." type="journal article" year="1993">Andriashev and Pitruk (1993)</bibRefCitation>
define the genus
<taxonomicName id="4C3814E6FF98FF9B9B0FF9765F9593E6" baseAuthorityName="Andriashev" baseAuthorityYear="1955" box="[815,971,1749,1773]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="164" phylum="Chordata" rank="genus">
<emphasis id="B94CB377FF98FF9B9B0FF9765F9593E6" box="[815,971,1749,1773]" italics="true" pageId="3" pageNumber="164">Pseudoliparis</emphasis>
</taxonomicName>
as having a well-developed disk and one pair of nostrils and lacking pseudobranchia and pleural ribs, with four radials in the pectoral girdle, which has neither notches nor fenestrae. In this genus, the hypural plate is divided by a distal slit (
<bibRefCitation id="EFA91294FF98FF9B9CB2F8BE5C8D9251" author="Andriashev" pageId="3" pageNumber="164" refString="Andriashev, A. P. &amp; Pitruk, D. L. (1993) A review of the ultra-abyssal (hadal) genus Pseudoliparis (Scorpaeniformes, Liparidae) with a description of a new species from the Japan Trench. Voprosy ikhtiologii, 33, 325 - 330." type="journal article" year="1993">Andriashev and Pitruk, 1993</bibRefCitation>
). Like the other in this genus,
<taxonomicName id="4C3814E6FF98FF9B9A05F8E25F559252" authority="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen, 2017" authorityName="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen" authorityYear="2017" box="[549,779,1857,1881]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="164" phylum="Chordata" rank="species" species="swirei" status="sp. nov.">
<emphasis id="B94CB377FF98FF9B9A05F8E25F559252" box="[549,779,1857,1881]" italics="true" pageId="3" pageNumber="164">Pseudoliparis swirei</emphasis>
</taxonomicName>
<emphasis id="B94CB377FF98FF9B9B31F8E25F359251" bold="true" box="[785,875,1857,1882]" pageId="3" pageNumber="164">
<taxonomicNameLabel id="A27F0E0CFF98FF9B9B31F8E25F359251" box="[785,875,1857,1882]" pageId="3" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="130373E0FF98FF9B9B59F8E25FB89251" box="[889,998,1857,1882]" captionStart="FIGURE 2" captionStartId="4.[151,250,1453,1475]" captionTargetBox="[474,1113,373,1432]" captionTargetId="figure@4.[474,1113,373,1432]" captionTargetPageId="4" captionText="FIGURE 2. A) In situ photograph of Pseudoliparis swirei sp. nov. at 6,198 m. B) a group at 7,485 m. C) Deck photograph of SIO 16-82/HADES 200049.D) Radiograph of SIO 16-86/HADES 200141. Image by Sandra Raredon. Scale indicator 5 cm." httpUri="https://zenodo.org/record/1067558/files/figure.png" pageId="3" pageNumber="164">Figures 2</figureCitation>
,
<figureCitation id="130373E0FF98FF9B9BD2F8E1585E9251" box="[1010,1024,1858,1882]" captionStart="FIGURE 3" captionStartId="4.[151,250,1897,1919]" captionTargetBox="[151,1436,1555,1875]" captionTargetId="figure@4.[151,1436,1555,1875]" captionTargetPageId="4" captionText="FIGURE 3. Lateral view of Pseudoliparis swirei sp. nov.. Combined representation of holotype, paratypes, and freshly captured images of paratype USNM 438985/HADES 200133, juvenile, 151 mm.Drawings by Thomas D. Linley." httpUri="https://zenodo.org/record/1067560/files/figure.png" pageId="3" pageNumber="164">3</figureCitation>
) displays these characters, including a moderately well-developed disk, although this is easily damaged in collection.
<taxonomicName id="4C3814E6FF98FF9B9C05F8C659529276" authority="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen, 2017" authorityName="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen" authorityYear="2017" box="[1061,1292,1893,1917]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="164" phylum="Chordata" rank="species" species="swirei" status="sp. nov.">
<emphasis id="B94CB377FF98FF9B9C05F8C659529276" box="[1061,1292,1893,1917]" italics="true" pageId="3" pageNumber="164">Pseudoliparis swirei</emphasis>
</taxonomicName>
<emphasis id="B94CB377FF98FF9B9D34F8C659309275" bold="true" box="[1300,1390,1893,1918]" pageId="3" pageNumber="164">
<taxonomicNameLabel id="A27F0E0CFF98FF9B9D34F8C659309275" box="[1300,1390,1893,1918]" pageId="3" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from the two other known
<taxonomicName id="4C3814E6FF98FF9B9ABAF82A5F6892AA" baseAuthorityName="Andriashev" baseAuthorityYear="1955" box="[666,822,1929,1953]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="164" phylum="Chordata" rank="genus">
<emphasis id="B94CB377FF98FF9B9ABAF82A5F6892AA" box="[666,822,1929,1953]" italics="true" pageId="3" pageNumber="164">Pseudoliparis</emphasis>
</taxonomicName>
species with the following characters.
<taxonomicName id="4C3814E6FF98FF9B9D20F82A5C8592CE" authority="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen, 2017" authorityName="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen" authorityYear="2017" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="164" phylum="Chordata" rank="species" species="swirei" status="sp. nov.">
<emphasis id="B94CB377FF98FF9B9D20F82A5C8592CE" italics="true" pageId="3" pageNumber="164">Pseudoliparis swirei</emphasis>
</taxonomicName>
<emphasis id="B94CB377FF98FF9B98C1F80E5D6492CD" bold="true" box="[225,314,1965,1990]" pageId="3" pageNumber="164">
<taxonomicNameLabel id="A27F0E0CFF98FF9B98C1F80E5D6492CD" box="[225,314,1965,1990]" pageId="3" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C3814E6FF98FF9B99EDF80D5E1B92CE" baseAuthorityName="Andriashev &amp; Pitruk" baseAuthorityYear="1993" box="[461,581,1965,1989]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="164" phylum="Chordata" rank="species" species="belyaevi">
<emphasis id="B94CB377FF98FF9B99EDF80D5E1B92CE" box="[461,581,1965,1989]" italics="true" pageId="3" pageNumber="164">P. belyaevi</emphasis>
</taxonomicName>
in the presence of a distinct lower pectoral-fin lobe, similar to that seen in
<taxonomicName id="4C3814E6FF98FF9B9DA7F80D5E1A92E1" authority="(Andriashev, 1955)" authorityName="Andriashev" authorityYear="1955" baseAuthorityName="Andriashev" baseAuthorityYear="1955" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="164" phylum="Chordata" rank="species" species="amblystomopsis">
<emphasis id="B94CB377FF98FF9B9DA7F80D5D1492E2" italics="true" pageId="3" pageNumber="164">P. amblystomopsis</emphasis>
(Andriashev, 1955)
</taxonomicName>
.
<taxonomicName id="4C3814E6FF98FF9B9A7CF8725F1092E2" authorityName="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen" authorityYear="2017" box="[604,846,2001,2025]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="164" phylum="Chordata" rank="species" species="swirei">
<emphasis id="B94CB377FF98FF9B9A7CF8725F1092E2" box="[604,846,2001,2025]" italics="true" pageId="3" pageNumber="164">Pseudoliparis swirei</emphasis>
</taxonomicName>
has more dorsal-fin rays 55 (5158) than
<emphasis id="B94CB377FF98FF9B9DA7F87159C392E2" box="[1415,1437,2002,2025]" italics="true" pageId="3" pageNumber="164">P.</emphasis>
</paragraph>
<paragraph id="8B876F65FF9FFF9C98B7FF3B5828944B" blockId="4.[151,1436,151,320]" pageId="4" pageNumber="165">
<taxonomicName id="4C3814E6FF9FFF9C98B7FF3B5D1495BB" baseAuthorityName="Andriashev" baseAuthorityYear="1955" box="[151,330,152,176]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="4" pageNumber="165" phylum="Chordata" rank="species" species="amblystomopsis">
<emphasis id="B94CB377FF9FFF9C98B7FF3B5D1495BB" box="[151,330,152,176]" italics="true" pageId="4" pageNumber="165">amblystomopsis</emphasis>
</taxonomicName>
49 (4952), more anal-fin rays 48 (4349) compared to 43 (4245), and more vertebrae 61 (56 62), compared to 5557, although these ranges somewhat overlap. Head length is shorter in
<taxonomicName id="4C3814E6FF9FFF9C9CF2FF1E596F95DF" authority="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen, 2017" authorityName="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen" authorityYear="2017" box="[1234,1329,189,212]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="4" pageNumber="165" phylum="Chordata" rank="species" species="swirei" status="sp. nov.">
<emphasis id="B94CB377FF9FFF9C9CF2FF1E596F95DF" box="[1234,1329,189,212]" italics="true" pageId="4" pageNumber="165">P. swirei</emphasis>
</taxonomicName>
<emphasis id="B94CB377FF9FFF9C9D1FFF1F59C295DE" bold="true" box="[1343,1436,188,213]" pageId="4" pageNumber="165">
<taxonomicNameLabel id="A27F0E0CFF9FFF9C9D1FFF1F59C295DE" box="[1343,1436,188,213]" pageId="4" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(17.021.7 %SL) than
<taxonomicName id="4C3814E6FF9FFF9C99B4FF425E3C95F3" baseAuthorityName="Andriashev" baseAuthorityYear="1955" box="[404,610,224,248]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="4" pageNumber="165" phylum="Chordata" rank="species" species="amblystomopsis">
<emphasis id="B94CB377FF9FFF9C99B4FF425E3C95F3" box="[404,610,224,248]" italics="true" pageId="4" pageNumber="165">P. amblystomopsis</emphasis>
</taxonomicName>
(21.624.0 %SL). Comparisons were made according to ranges presented by
<bibRefCitation id="EFA91294FF9FFF9C989BFEA75DA09417" author="Andriashev" box="[187,510,260,285]" pageId="4" pageNumber="165" refString="Andriashev, A. P. &amp; Pitruk, D. L. (1993) A review of the ultra-abyssal (hadal) genus Pseudoliparis (Scorpaeniformes, Liparidae) with a description of a new species from the Japan Trench. Voprosy ikhtiologii, 33, 325 - 330." type="journal article" year="1993">Andriashev &amp; Pitruk (1993)</bibRefCitation>
.
<taxonomicName id="4C3814E6FF9FFF9C9A2AFEA75F549417" baseAuthorityName="Andriashev &amp; Pitruk" baseAuthorityYear="1993" box="[522,778,260,284]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="4" pageNumber="165" phylum="Chordata" rank="species" species="belyaevi">
<emphasis id="B94CB377FF9FFF9C9A2AFEA75F549417" box="[522,778,260,284]" italics="true" pageId="4" pageNumber="165">Pseudoliparis belyaevi</emphasis>
</taxonomicName>
is known only from the
<collectingCountry id="F32F2FF5FF9FFF9C9C06FEA658389417" box="[1062,1126,261,284]" name="Japan" pageId="4" pageNumber="165">Japan</collectingCountry>
Trench,
<taxonomicName id="4C3814E6FF9FFF9C9CECFEA659C29417" baseAuthorityName="Andriashev" baseAuthorityYear="1955" box="[1228,1436,260,284]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="4" pageNumber="165" phylum="Chordata" rank="species" species="amblystomopsis">
<emphasis id="B94CB377FF9FFF9C9CECFEA659C29417" box="[1228,1436,260,284]" italics="true" pageId="4" pageNumber="165">P. amblystomopsis</emphasis>
</taxonomicName>
from the
<collectingCountry id="F32F2FF5FF9FFF9C98DDFE8B5D639434" box="[253,317,296,319]" name="Japan" pageId="4" pageNumber="165">Japan</collectingCountry>
and Kurile-Kamchatka trenches,
<taxonomicName id="4C3814E6FF9FFF9C9A96FE8A5F489434" authorityName="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen" authorityYear="2017" box="[694,790,296,320]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="4" pageNumber="165" phylum="Chordata" rank="species" species="swirei">
<emphasis id="B94CB377FF9FFF9C9A96FE8A5F489434" box="[694,790,296,320]" italics="true" pageId="4" pageNumber="165">P. swirei</emphasis>
</taxonomicName>
only from the Mariana Trench.
</paragraph>
</subSubSection>
<caption id="DF473FEDFF9FFF9C98B7FA0E59DD90E9" httpUri="https://zenodo.org/record/1067558/files/figure.png" pageId="4" pageNumber="165" targetBox="[474,1113,373,1432]" targetPageId="4">
<paragraph id="8B876F65FF9FFF9C98B7FA0E59DD90E9" blockId="4.[151,1435,1453,1506]" pageId="4" pageNumber="165">
<emphasis id="B94CB377FF9FFF9C98B7FA0E5D4D90C9" bold="true" box="[151,275,1453,1475]" pageId="4" pageNumber="165">FIGURE 2.</emphasis>
A)
<emphasis id="B94CB377FF9FFF9C991CFA0D5D2290C8" box="[316,380,1454,1475]" italics="true" pageId="4" pageNumber="165">In situ</emphasis>
photograph of
<taxonomicName id="4C3814E6FF9FFF9C9A39FA0E5EB490C8" authority="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen, 2017" authorityName="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen" authorityYear="2017" box="[537,746,1453,1475]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="4" pageNumber="165" phylum="Chordata" rank="species" species="swirei" status="sp. nov.">
<emphasis id="B94CB377FF9FFF9C9A39FA0E5EB490C8" box="[537,746,1453,1475]" italics="true" pageId="4" pageNumber="165">Pseudoliparis swirei</emphasis>
</taxonomicName>
<emphasis id="B94CB377FF9FFF9C9AD0FA0E5F1F90C9" bold="true" box="[752,833,1453,1474]" pageId="4" pageNumber="165">
<taxonomicNameLabel id="A27F0E0CFF9FFF9C9AD0FA0E5F1F90C9" box="[752,833,1453,1474]" pageId="4" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
at 6,198 m. B) a group at 7,485 m. C) Deck photograph of SIO 16-82/HADES 200049. D) Radiograph of SIO 16-86/HADES 200141. Image by Sandra Raredon. Scale indicator 5 cm.
</paragraph>
</caption>
<caption id="DF473FEDFF9FFF9C98B7F8CA59489295" httpUri="https://zenodo.org/record/1067560/files/figure.png" pageId="4" pageNumber="165" targetBox="[151,1436,1555,1875]" targetPageId="4">
<paragraph id="8B876F65FF9FFF9C98B7F8CA59489295" blockId="4.[151,1435,1897,1950]" pageId="4" pageNumber="165">
<emphasis id="B94CB377FF9FFF9C98B7F8CA5D499275" bold="true" box="[151,279,1897,1919]" pageId="4" pageNumber="165">FIGURE 3.</emphasis>
Lateral view of
<taxonomicName id="4C3814E6FF9FFF9C99F2F8CA5EF99274" authority="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen, 2017" authorityName="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen" authorityYear="2017" box="[466,679,1897,1919]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="4" pageNumber="165" phylum="Chordata" rank="species" species="swirei" status="sp. nov.">
<emphasis id="B94CB377FF9FFF9C99F2F8CA5EF99274" box="[466,679,1897,1919]" italics="true" pageId="4" pageNumber="165">Pseudoliparis swirei</emphasis>
</taxonomicName>
<emphasis id="B94CB377FF9FFF9C9A92F8C95F569274" bold="true" box="[690,776,1898,1919]" pageId="4" pageNumber="165">
<taxonomicNameLabel id="A27F0E0CFF9FFF9C9A92F8C95F569274" box="[690,776,1898,1919]" pageId="4" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. Combined representation of holotype, paratypes, and freshly captured images of paratype USNM 438985/HADES 200133, juvenile, 151 mm. Drawings by Thomas D. Linley.
</paragraph>
</caption>
<caption id="DF473FEDFF9EFF9D98B7FA0A585290F7" httpUri="https://zenodo.org/record/1067562/files/figure.png" pageId="5" pageNumber="166" targetBox="[151,1436,199,1428]" targetPageId="5">
<paragraph id="8B876F65FF9EFF9D98B7FA0A585290F7" blockId="5.[151,1436,1449,1532]" pageId="5" pageNumber="166">
<emphasis id="B94CB377FF9EFF9D98B7FA0A5D4A90B5" bold="true" box="[151,276,1449,1471]" pageId="5" pageNumber="166">FIGURE 4.</emphasis>
A) Ventral view drawing. B) Tooth pattern on maxillary jaw. Counted rows denoted on right side of image. C) Tooth structure of specimen USNM 438996/HADES 200024. D) Disk details of SIO 16-87/HADES 200025 stained with Alizarin Red S. E) Disk of USNM 438995/HADES 200085 stained with Cyanine Blue.
</paragraph>
</caption>
<subSubSection id="C3223CEEFF9EFF9098E7F98A5F08935E" lastPageId="8" lastPageNumber="169" pageId="5" pageNumber="166" type="description">
<paragraph id="8B876F65FF9EFF9D98E7F98A5DF093D9" blockId="5.[151,1437,1577,2034]" pageId="5" pageNumber="166">
<emphasis id="B94CB377FF9EFF9D98E7F98A5D079349" bold="true" box="[199,345,1577,1602]" pageId="5" pageNumber="166">Description.</emphasis>
Vertebrae 61 (5662), dorsal-fin rays 55 (5158), anal-fin rays 48 (4349), caudal-fin rays 13 (1114), pectoral-fin rays 30 (2832), pectoral radials 4, pyloric caeca 7 (59). Ranges of measurements and counts are presented in
<tableCitation id="C6BA5ADEFF9EFF9D9972F9D25DF69382" box="[338,424,1649,1674]" captionStart="TABLE 2" captionStartId="7.[151,239,152,175]" captionText="TABLE 2. Measurements and counts of Pseudoliparis swirei sp. nov.. Measurements taken from preserved specimens, given in mm. Body depth measured from fresh photographs. Dorsal fin origin between vertebrae #, # are shown. Maximum and minimum values (mean and standard deviation) shown for measurements. Counts show range (median). Number of specimens examined for individual characters (n)." pageId="5" pageNumber="166">Table 2</tableCitation>
. Ratios are presented in
<tableCitation id="C6BA5ADEFF9EFF9D9A9FF9D25F4A9382" box="[703,788,1649,1674]" captionStart="TABLE 3" captionStartId="8.[151,239,152,175]" captionText="TABLE 3. Ratios of Pseudoliparis swirei sp. nov.. Preserved and fresh measurements presented. Body depth taken from photographs. Predorsal fin length measured from radiograph." pageId="5" pageNumber="166">Table 3</tableCitation>
. All individual measurements and counts are available in Supplementary
<tableCitation id="C6BA5ADEFF9EFF9D9970F9365DF393A5" box="[336,429,1685,1710]" captionStart="TABLE 1" captionStartId="1.[151,239,542,565]" captionText="TABLE 1. Collection information. Specimens collected on cruise FK141109 of the RV Falkor and SY1615 of the RV Shinyo-maru. Number of individual fish collected in each trap deployment (n) shown." pageId="5" pageNumber="166">Table 1</tableCitation>
. Micro-CT scans and a 3D rotatable model of the
<typeStatus id="5483D1C7FF9EFF9D9C1BF93658C393A5" box="[1083,1181,1685,1710]" pageId="5" pageNumber="166" type="holotype">holotype</typeStatus>
are available in the Supplementary Material.
</paragraph>
<paragraph id="8B876F65FF9EFF9E98E7F97E58D3944B" blockId="5.[151,1437,1577,2034]" lastBlockId="6.[151,1437,151,789]" lastPageId="6" lastPageNumber="167" pageId="5" pageNumber="166">
Head small, low, and wide, lateral profile anteriorly rounded and rising slowly to occiput, where the angle increases. Head depth about equal to body depth or lower, depending on abdominal fullness. Snout blunt, nostrils single, nares on horizontal with center of eye. Mouth broad, horizontal, subterminal, moderately large; upper jaw reaching to below middle of orbit, oral cleft reaching to below anterior edge of orbit. Teeth simple, sharp canines, innermost largest, arranged in approximately 9 (611 maxilla, 713 mandible) oblique, irregular rows of up to 20 (617 maxilla, 820 mandible) teeth each, forming a moderately wide band (24 teeth wide) in each jaw (
<figureCitation id="130373E0FF9EFF9D9D0DF83259D192A2" box="[1325,1423,1937,1962]" captionStart="FIGURE 4" captionStartId="5.[151,250,1449,1471]" captionTargetBox="[151,1436,199,1428]" captionTargetId="figure@5.[151,1436,193,1428]" captionTargetPageId="5" captionText="FIGURE 4. A) Ventral view drawing. B) Tooth pattern on maxillary jaw. Counted rows denoted on right side of image. C) Tooth structure of specimen USNM 438996/HADES 200024. D) Disk details of SIO 16-87/HADES 200025 stained with Alizarin Red S. E) Disk of USNM 438995/HADES 200085 stained with Cyanine Blue." httpUri="https://zenodo.org/record/1067562/files/figure.png" pageId="5" pageNumber="166">Figure 4</figureCitation>
). Larger individuals had more teeth per row and more rows of teeth. Maxilla with prominent symphyseal gap, slight gap present in mandible. Pharyngeal teeth well developed, long, sharp, strongly fixed on globular tooth plates. Eye very small, about 10% head length. Orbit large, its dorsal margin well below that of head. Gill opening small, located completely above pectoral fin, width 5% SL. Opercular flap fleshy, broadly triangular, opercle terminates in two small spines below the flesh. Cephalic pores small, easily damaged; few remaining. Eight widely separated infraorbital and mandibular pores, lacking raised rims. In fresh specimens, the infraorbital and mandibular pores appear sunken and anteriorly orientated within the gelatinous tissue of the head (
<figureCitation id="130373E0FF9DFF9E9C3DFE8458DE944B" box="[1053,1152,295,320]" captionStart="FIGURE 4" captionStartId="5.[151,250,1449,1471]" captionTargetBox="[151,1436,199,1428]" captionTargetId="figure@5.[151,1436,193,1428]" captionTargetPageId="5" captionText="FIGURE 4. A) Ventral view drawing. B) Tooth pattern on maxillary jaw. Counted rows denoted on right side of image. C) Tooth structure of specimen USNM 438996/HADES 200024. D) Disk details of SIO 16-87/HADES 200025 stained with Alizarin Red S. E) Disk of USNM 438995/HADES 200085 stained with Cyanine Blue." httpUri="https://zenodo.org/record/1067562/files/figure.png" pageId="6" pageNumber="167">Figure 4</figureCitation>
).
</paragraph>
<paragraph id="8B876F65FF9DFF9E98E7FEEF5F01961E" blockId="6.[151,1437,151,789]" pageId="6" pageNumber="167">
Pectoral fin divided into lobes by a moderately deep notch, rudimentary rays absent. Notch rays 5 (36), clearly more widely spaced than those of upper and lower lobes, more so in larger individuals. Upper and lower lobe rays closely spaced. Dorsal-most pectoral-fin ray on horizontal between level of upper jaw and lower margin of orbit. Symphysis of pectoral fins and anteriormost ray below rear of orbit. Upper lobe about 15.2% SL (12.6 19.8), lower about 7.8% SL (6.312.1). Upper lobe almost extending to anal-fin origin, lower lobe distinct, reaching well behind disk to below middle of upper lobe base. Pectoral radials four, fenestra absent; of four specimens examined (USNM 438996/HADES 200024, USNM 438994/HADES 200027, USNM 438997/HADES 200033, USNM 438989/HADES 200096); one (USNM 438996HADES 200024) had (1+1+1+1), and three (USNM 438994/HADES 200027, USNM 438997/HADES 200033, USNM 438989/HADES 200096) had (3+1) radials, generally round, notches and foramina absent (
<figureCitation id="130373E0FF9DFF9E9B2AFD2C5F3097A3" box="[778,878,655,680]" captionStart="FIGURE 5" captionStartId="6.[151,250,1945,1967]" captionTargetBox="[242,1275,843,1911]" captionTargetId="figure@6.[241,1319,842,1911]" captionTargetPageId="6" captionText="FIGURE 5. Pectoral girdle of USNM 438994/HADES 200027, female, 220 mm. Ventral drawing of USNM 438995/HADES 200085, female, 225 mm.Scapula, radials 14, coracoid shown. Drawingby Thomas D. Linley." httpUri="https://zenodo.org/record/1067564/files/figure.png" pageId="6" pageNumber="167">Figure 5</figureCitation>
). Radials gradually and irregularly decreasing in size from R1 (largest) to R4 (smallest). Distal radials present under the base of all rays, with the exception of the two most ventral rays (Supplementary
<figureCitation id="130373E0FF9DFF9E9A75FD745EEA97FB" box="[597,692,727,752]" captionStart="FIGURE 1" captionStartId="2.[151,250,1128,1150]" captionTargetBox="[166,1432,203,1100]" captionTargetId="figure@2.[151,1436,195,1104]" captionTargetPageId="2" captionText="FIGURE 1. Map of collection locations within the Mariana Trench. Bathymetry courtesy of JAMSTEC. Isobaths are added between 5,000 and 9,000 m at 1,000 m intervals." httpUri="https://zenodo.org/record/1067556/files/figure.png" pageId="6" pageNumber="167">Figure 1</figureCitation>
). Scapula double-headed, posterior head larger and broader than anterior head, coracoid with broad head and long slender helve.
</paragraph>
<caption id="DF473FEDFF9DFF9E98B7F83A583A92C5" httpUri="https://zenodo.org/record/1067564/files/figure.png" pageId="6" pageNumber="167" targetBox="[242,1275,843,1911]" targetPageId="6">
<paragraph id="8B876F65FF9DFF9E98B7F83A583A92C5" blockId="6.[151,1436,1945,1998]" pageId="6" pageNumber="167">
<emphasis id="B94CB377FF9DFF9E98B7F83A5D4D92A5" bold="true" box="[151,275,1945,1967]" pageId="6" pageNumber="167">FIGURE 5.</emphasis>
Pectoral girdle of USNM 438994/HADES 200027, female, 220 mm. Ventral drawing of USNM 438995/HADES 200085, female, 225 mm. Scapula, radials 14, coracoid shown. Drawing by Thomas D. Linley.
</paragraph>
</caption>
<caption id="DF473FEDFF9CFF9F98B7FF3B5F799404" pageId="7" pageNumber="168">
<paragraph id="8B876F65FF9CFF9F98B7FF3B5F799404" blockId="7.[151,1436,151,271]" pageId="7" pageNumber="168">
<emphasis id="B94CB377FF9CFF9F98B7FF3B5D5495A4" bold="true" box="[151,266,152,175]" pageId="7" pageNumber="168">TABLE 2.</emphasis>
Measurements and counts of
<taxonomicName id="4C3814E6FF9CFF9F9A6FFF345F7595A4" authority="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen, 2017" authorityName="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen" authorityYear="2017" box="[591,811,151,175]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="168" phylum="Chordata" rank="species" species="swirei" status="sp. nov.">
<emphasis id="B94CB377FF9CFF9F9A6FFF345F7595A4" box="[591,811,151,175]" italics="true" pageId="7" pageNumber="168">Pseudoliparis swirei</emphasis>
</taxonomicName>
<emphasis id="B94CB377FF9CFF9F9B12FF3B5FD795A4" bold="true" box="[818,905,152,175]" pageId="7" pageNumber="168">
<taxonomicNameLabel id="A27F0E0CFF9CFF9F9B12FF3B5FD795A4" box="[818,905,152,175]" pageId="7" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. Measurements taken from preserved specimens, given in mm. Body depth measured from fresh photographs. Dorsal fin origin between vertebrae #, # are shown. Maximum and minimum values (mean and standard deviation) shown for measurements. Counts show range (median). Number of specimens examined for individual characters (
<emphasis id="B94CB377FF9CFF9F9B2AFF5B5F499404" box="[778,791,248,271]" italics="true" pageId="7" pageNumber="168">n</emphasis>
).
</paragraph>
</caption>
<paragraph id="8B876F65FF9CFF9F9AF1FE8959309268" pageId="7" pageNumber="168">
<table id="F9389DC5FF9C006798BFFE8959309268" box="[159,1390,298,1891]" gridcols="4" gridrows="37" pageId="7" pageNumber="168">
<tr id="35086D27FF9C006798BFFE895930944B" box="[159,1390,298,320]" gridrow="0" pageId="7" pageNumber="168" rowspan-0="1">
<th id="76D9045BFF9C00679AF1FE895F71944B" box="[721,815,298,320]" gridcol="1" gridrow="0" pageId="7" pageNumber="168">Holotype</th>
<th id="76D9045BFF9C00679BB3FE8958D1944B" box="[915,1167,298,320]" gridcol="2" gridrow="0" pageId="7" pageNumber="168">Holotype and Paratypes</th>
<th id="76D9045BFF9C00679D75FE895930944B" box="[1365,1390,298,320]" gridcol="3" gridrow="0" pageId="7" pageNumber="168">n</th>
</tr>
<tr id="35086D27FF9C006798BFFEF659309461" box="[159,1390,341,362]" gridrow="1" pageId="7" pageNumber="168" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76D9045BFF9C006798BFFEF65DB49461" box="[159,490,341,362]" gridcol="0" gridrow="1" pageId="7" pageNumber="168">Measurements</th>
</tr>
<tr id="35086D27FF9C006798BFFEDE59309498" box="[159,1390,381,403]" gridrow="2" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFEDE5DB49498" box="[159,490,381,403]" gridcol="0" gridrow="2" pageId="7" pageNumber="168">Standard Length</th>
<td id="76D9045BFF9C00679AF1FEDE5F719498" box="[721,815,381,403]" gridcol="1" gridrow="2" pageId="7" pageNumber="168">104.0</td>
<td id="76D9045BFF9C00679BB3FEDE58D19498" box="[915,1167,381,403]" gridcol="2" gridrow="2" pageId="7" pageNumber="168">87.0226.0 (151.3±38.8)</td>
<td id="76D9045BFF9C00679D75FEDE59309498" box="[1365,1390,381,403]" gridcol="3" gridrow="2" pageId="7" pageNumber="168">24</td>
</tr>
<tr id="35086D27FF9C006798BFFE05593094B7" box="[159,1390,422,444]" gridrow="3" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFE055DB494B7" box="[159,490,422,444]" gridcol="0" gridrow="3" pageId="7" pageNumber="168">Total Length</th>
<td id="76D9045BFF9C00679AF1FE055F7194B7" box="[721,815,422,444]" gridcol="1" gridrow="3" pageId="7" pageNumber="168">112.0</td>
<td id="76D9045BFF9C00679BB3FE0558D194B7" box="[915,1167,422,444]" gridcol="2" gridrow="3" pageId="7" pageNumber="168">95.0237.0 (160.5±41.6)</td>
<td id="76D9045BFF9C00679D75FE05593094B7" box="[1365,1390,422,444]" gridcol="3" gridrow="3" pageId="7" pageNumber="168">21</td>
</tr>
<tr id="35086D27FF9C006798BFFE73593094ED" box="[159,1390,464,486]" gridrow="4" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFE735DB494ED" box="[159,490,464,486]" gridcol="0" gridrow="4" pageId="7" pageNumber="168">Head Length</th>
<td id="76D9045BFF9C00679AF1FE735F7194ED" box="[721,815,464,486]" gridcol="1" gridrow="4" pageId="7" pageNumber="168">18.9</td>
<td id="76D9045BFF9C00679BB3FE7358D194ED" box="[915,1167,464,486]" gridcol="2" gridrow="4" pageId="7" pageNumber="168">17.842.9 (31.2±8.0)</td>
<td id="76D9045BFF9C00679D75FE73593094ED" box="[1365,1390,464,486]" gridcol="3" gridrow="4" pageId="7" pageNumber="168">20</td>
</tr>
<tr id="35086D27FF9C006798BFFE5A59309704" box="[159,1390,505,527]" gridrow="5" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFE5A5DB49704" box="[159,490,505,527]" gridcol="0" gridrow="5" pageId="7" pageNumber="168">Head Depth at Occiput</th>
<td id="76D9045BFF9C00679AF1FE5A5F719704" box="[721,815,505,527]" gridcol="1" gridrow="5" pageId="7" pageNumber="168">~14</td>
<td id="76D9045BFF9C00679BB3FE5A58D19704" box="[915,1167,505,527]" gridcol="2" gridrow="5" pageId="7" pageNumber="168">20.532.7 (27.3±6.2)</td>
<td id="76D9045BFF9C00679D75FE5A59309704" box="[1365,1390,505,527]" gridcol="3" gridrow="5" pageId="7" pageNumber="168">3</td>
</tr>
<tr id="35086D27FF9C006798BFFD8159309733" box="[159,1390,546,568]" gridrow="6" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFD815DB49733" box="[159,490,546,568]" gridcol="0" gridrow="6" pageId="7" pageNumber="168">Head Width</th>
<td id="76D9045BFF9C00679AF1FD815F719733" box="[721,815,546,568]" gridcol="1" gridrow="6" pageId="7" pageNumber="168">13.1</td>
<td id="76D9045BFF9C00679BB3FD8158D19733" box="[915,1167,546,568]" gridcol="2" gridrow="6" pageId="7" pageNumber="168">2.641.4 (27.0±9.7)</td>
<td id="76D9045BFF9C00679D75FD8159309733" box="[1365,1390,546,568]" gridcol="3" gridrow="6" pageId="7" pageNumber="168">11</td>
</tr>
<tr id="35086D27FF9C006798BFFDEF59309769" box="[159,1390,588,610]" gridrow="7" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFDEF5DB49769" box="[159,490,588,610]" gridcol="0" gridrow="7" pageId="7" pageNumber="168">Snout Length</th>
<td id="76D9045BFF9C00679AF1FDEF5F719769" box="[721,815,588,610]" gridcol="1" gridrow="7" pageId="7" pageNumber="168">6.7</td>
<td id="76D9045BFF9C00679BB3FDEF58D19769" box="[915,1167,588,610]" gridcol="2" gridrow="7" pageId="7" pageNumber="168">5.615.2 (9.8±2.8)</td>
<td id="76D9045BFF9C00679D75FDEF59309769" box="[1365,1390,588,610]" gridcol="3" gridrow="7" pageId="7" pageNumber="168">26</td>
</tr>
<tr id="35086D27FF9C006798BFFDD659309780" box="[159,1390,629,651]" gridrow="8" pageId="7" pageNumber="168" rowspan-1="1">
<th id="76D9045BFF9C006798BFFDD65DB49780" box="[159,490,629,651]" gridcol="0" gridrow="8" pageId="7" pageNumber="168">Lower Lobe Distance</th>
<td id="76D9045BFF9C00679BB3FDD658D19780" box="[915,1167,629,651]" gridcol="2" gridrow="8" pageId="7" pageNumber="168">2.36.6 (4.7±1.4)</td>
<td id="76D9045BFF9C00679D75FDD659309780" box="[1365,1390,629,651]" gridcol="3" gridrow="8" pageId="7" pageNumber="168">16</td>
</tr>
<tr id="35086D27FF9C006798BFFD3D593097BF" box="[159,1390,670,692]" gridrow="9" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFD3D5DB497BF" box="[159,490,670,692]" gridcol="0" gridrow="9" pageId="7" pageNumber="168">Body Depth</th>
<td id="76D9045BFF9C00679AF1FD3D5F7197BF" box="[721,815,670,692]" gridcol="1" gridrow="9" pageId="7" pageNumber="168">18.0</td>
<td id="76D9045BFF9C00679BB3FD3D58D197BF" box="[915,1167,670,692]" gridcol="2" gridrow="9" pageId="7" pageNumber="168">18.062.0 (41.0±11.0)</td>
<td id="76D9045BFF9C00679D75FD3D593097BF" box="[1365,1390,670,692]" gridcol="3" gridrow="9" pageId="7" pageNumber="168">18</td>
</tr>
<tr id="35086D27FF9C006798BFFD6B593097D5" box="[159,1390,712,734]" gridrow="10" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFD6B5DB497D5" box="[159,490,712,734]" gridcol="0" gridrow="10" pageId="7" pageNumber="168">Orbit Width</th>
<td id="76D9045BFF9C00679AF1FD6B5F7197D5" box="[721,815,712,734]" gridcol="1" gridrow="10" pageId="7" pageNumber="168">3.5</td>
<td id="76D9045BFF9C00679BB3FD6B58D197D5" box="[915,1167,712,734]" gridcol="2" gridrow="10" pageId="7" pageNumber="168">2.86.1 (4.8±1.0)</td>
<td id="76D9045BFF9C00679D75FD6B593097D5" box="[1365,1390,712,734]" gridcol="3" gridrow="10" pageId="7" pageNumber="168">23</td>
</tr>
<tr id="35086D27FF9C006798BFFD525930960C" box="[159,1390,753,775]" gridrow="11" pageId="7" pageNumber="168" rowspan-1="1">
<th id="76D9045BFF9C006798BFFD525DB4960C" box="[159,490,753,775]" gridcol="0" gridrow="11" pageId="7" pageNumber="168">Disk Length</th>
<td id="76D9045BFF9C00679BB3FD5258D1960C" box="[915,1167,753,775]" gridcol="2" gridrow="11" pageId="7" pageNumber="168">2.99.0 (6.2±2.1)</td>
<td id="76D9045BFF9C00679D75FD525930960C" box="[1365,1390,753,775]" gridcol="3" gridrow="11" pageId="7" pageNumber="168">14</td>
</tr>
<tr id="35086D27FF9C006798BFFCB95930963B" box="[159,1390,794,816]" gridrow="12" pageId="7" pageNumber="168" rowspan-1="1">
<th id="76D9045BFF9C006798BFFCB95DB4963B" box="[159,490,794,816]" gridcol="0" gridrow="12" pageId="7" pageNumber="168">Gill Opening</th>
<td id="76D9045BFF9C00679BB3FCB958D1963B" box="[915,1167,794,816]" gridcol="2" gridrow="12" pageId="7" pageNumber="168">5.49.6 (8.1±2.4)</td>
<td id="76D9045BFF9C00679D75FCB95930963B" box="[1365,1390,794,816]" gridcol="3" gridrow="12" pageId="7" pageNumber="168">3</td>
</tr>
<tr id="35086D27FF9C006798BFFCE759309651" box="[159,1390,836,858]" gridrow="13" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFCE75DB49651" box="[159,490,836,858]" gridcol="0" gridrow="13" pageId="7" pageNumber="168">Upper Jaw Length</th>
<td id="76D9045BFF9C00679AF1FCE75F719651" box="[721,815,836,858]" gridcol="1" gridrow="13" pageId="7" pageNumber="168">~8.7</td>
<td id="76D9045BFF9C00679BB3FCE758D19651" box="[915,1167,836,858]" gridcol="2" gridrow="13" pageId="7" pageNumber="168">8.120.6 (13.8±3.3)</td>
<td id="76D9045BFF9C00679D75FCE759309651" box="[1365,1390,836,858]" gridcol="3" gridrow="13" pageId="7" pageNumber="168">31</td>
</tr>
<tr id="35086D27FF9C006798BFFCCE59309688" box="[159,1390,877,899]" gridrow="14" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFCCE5DB49688" box="[159,490,877,899]" gridcol="0" gridrow="14" pageId="7" pageNumber="168">Lower Jaw Length</th>
<td id="76D9045BFF9C00679AF1FCCE5F719688" box="[721,815,877,899]" gridcol="1" gridrow="14" pageId="7" pageNumber="168">8.2</td>
<td id="76D9045BFF9C00679BB3FCCE58D19688" box="[915,1167,877,899]" gridcol="2" gridrow="14" pageId="7" pageNumber="168">6.619.3 (12.7±3.1)</td>
<td id="76D9045BFF9C00679D75FCCE59309688" box="[1365,1390,877,899]" gridcol="3" gridrow="14" pageId="7" pageNumber="168">31</td>
</tr>
<tr id="35086D27FF9C006798BFFC35593096A7" box="[159,1390,918,940]" gridrow="15" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFC355DB496A7" box="[159,490,918,940]" gridcol="0" gridrow="15" pageId="7" pageNumber="168">Distance: Mandible to Disk</th>
<td id="76D9045BFF9C00679AF1FC355F7196A7" box="[721,815,918,940]" gridcol="1" gridrow="15" pageId="7" pageNumber="168">8.8</td>
<td id="76D9045BFF9C00679BB3FC3558D196A7" box="[915,1167,918,940]" gridcol="2" gridrow="15" pageId="7" pageNumber="168">8.822.3 (15.2±3.7)</td>
<td id="76D9045BFF9C00679D75FC35593096A7" box="[1365,1390,918,940]" gridcol="3" gridrow="15" pageId="7" pageNumber="168">25</td>
</tr>
<tr id="35086D27FF9C006798BFFC63593096DD" box="[159,1390,960,982]" gridrow="16" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFC635DB496DD" box="[159,490,960,982]" gridcol="0" gridrow="16" pageId="7" pageNumber="168">Distance: Snout to Anus</th>
<td id="76D9045BFF9C00679AF1FC635F7196DD" box="[721,815,960,982]" gridcol="1" gridrow="16" pageId="7" pageNumber="168">23.2</td>
<td id="76D9045BFF9C00679BB3FC6358D196DD" box="[915,1167,960,982]" gridcol="2" gridrow="16" pageId="7" pageNumber="168">23.256.7 (41.1±13.5)</td>
<td id="76D9045BFF9C00679D75FC63593096DD" box="[1365,1390,960,982]" gridcol="3" gridrow="16" pageId="7" pageNumber="168">6</td>
</tr>
<tr id="35086D27FF9C006798BFFC4A593096F4" box="[159,1390,1001,1023]" gridrow="17" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFC4A5DB496F4" box="[159,490,1001,1023]" gridcol="0" gridrow="17" pageId="7" pageNumber="168">Distance: Mandible to Anus</th>
<td id="76D9045BFF9C00679AF1FC4A5F7196F4" box="[721,815,1001,1023]" gridcol="1" gridrow="17" pageId="7" pageNumber="168">21.0</td>
<td id="76D9045BFF9C00679BB3FC4A58D196F4" box="[915,1167,1001,1023]" gridcol="2" gridrow="17" pageId="7" pageNumber="168">21.047.5 (35.3±12.0)</td>
<td id="76D9045BFF9C00679D75FC4A593096F4" box="[1365,1390,1001,1023]" gridcol="3" gridrow="17" pageId="7" pageNumber="168">5</td>
</tr>
<tr id="35086D27FF9C006798BFFBB159309123" box="[159,1390,1042,1064]" gridrow="18" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFBB15DB49123" box="[159,490,1042,1064]" gridcol="0" gridrow="18" pageId="7" pageNumber="168">Distance: Disk to Anus</th>
<td id="76D9045BFF9C00679AF1FBB15F719123" box="[721,815,1042,1064]" gridcol="1" gridrow="18" pageId="7" pageNumber="168">9.5</td>
<td id="76D9045BFF9C00679BB3FBB158D19123" box="[915,1167,1042,1064]" gridcol="2" gridrow="18" pageId="7" pageNumber="168">9.533.5 (18.7±8.9)</td>
<td id="76D9045BFF9C00679D75FBB159309123" box="[1365,1390,1042,1064]" gridcol="3" gridrow="18" pageId="7" pageNumber="168">5</td>
</tr>
<tr id="35086D27FF9C006798BFFB9F59309159" box="[159,1390,1084,1106]" gridrow="19" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFB9F5DB49159" box="[159,490,1084,1106]" gridcol="0" gridrow="19" pageId="7" pageNumber="168">Distance: Anus to Anal Fin</th>
<td id="76D9045BFF9C00679AF1FB9F5F719159" box="[721,815,1084,1106]" gridcol="1" gridrow="19" pageId="7" pageNumber="168">10.9</td>
<td id="76D9045BFF9C00679BB3FB9F58D19159" box="[915,1167,1084,1106]" gridcol="2" gridrow="19" pageId="7" pageNumber="168">10.925.9 (18.4±6.3)</td>
<td id="76D9045BFF9C00679D75FB9F59309159" box="[1365,1390,1084,1106]" gridcol="3" gridrow="19" pageId="7" pageNumber="168">7</td>
</tr>
<tr id="35086D27FF9C006798BFFBC659309170" box="[159,1390,1125,1147]" gridrow="20" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFBC65DB49170" box="[159,490,1125,1147]" gridcol="0" gridrow="20" pageId="7" pageNumber="168">Length Upper Pectoral Fin Lobe</th>
<td id="76D9045BFF9C00679AF1FBC65F719170" box="[721,815,1125,1147]" gridcol="1" gridrow="20" pageId="7" pageNumber="168">&gt;15.6</td>
<td id="76D9045BFF9C00679BB3FBC658D19170" box="[915,1167,1125,1147]" gridcol="2" gridrow="20" pageId="7" pageNumber="168">14.233.4 (22.8±4.5)</td>
<td id="76D9045BFF9C00679D75FBC659309170" box="[1365,1390,1125,1147]" gridcol="3" gridrow="20" pageId="7" pageNumber="168">28</td>
</tr>
<tr id="35086D27FF9C006798BFFB2D593091AF" box="[159,1390,1166,1188]" gridrow="21" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFB2D5DB491AF" box="[159,490,1166,1188]" gridcol="0" gridrow="21" pageId="7" pageNumber="168">Length Lower Pectoral Fin Lobe</th>
<td id="76D9045BFF9C00679AF1FB2D5F7191AF" box="[721,815,1166,1188]" gridcol="1" gridrow="21" pageId="7" pageNumber="168">8.9</td>
<td id="76D9045BFF9C00679BB3FB2D58D191AF" box="[915,1167,1166,1188]" gridcol="2" gridrow="21" pageId="7" pageNumber="168">7.322.9 (12.3±3.3)</td>
<td id="76D9045BFF9C00679D75FB2D593091AF" box="[1365,1390,1166,1188]" gridcol="3" gridrow="21" pageId="7" pageNumber="168">22</td>
</tr>
<tr id="35086D27FF9C006798BFFB42593091FC" box="[159,1390,1249,1271]" gridrow="22" pageId="7" pageNumber="168" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76D9045BFF9C006798BFFB425DB491FC" box="[159,490,1249,1271]" gridcol="0" gridrow="22" pageId="7" pageNumber="168">Counts</th>
</tr>
<tr id="35086D27FF9C006798BFFAA95930902B" box="[159,1390,1290,1312]" gridrow="23" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFAA95DB4902B" box="[159,490,1290,1312]" gridcol="0" gridrow="23" pageId="7" pageNumber="168">Total Vertebrae</th>
<td id="76D9045BFF9C00679AF1FAA95F71902B" box="[721,815,1290,1312]" gridcol="1" gridrow="23" pageId="7" pageNumber="168">58</td>
<td id="76D9045BFF9C00679BB3FAA958D1902B" box="[915,1167,1290,1312]" gridcol="2" gridrow="23" pageId="7" pageNumber="168">5662 (59)</td>
<td id="76D9045BFF9C00679D75FAA95930902B" box="[1365,1390,1290,1312]" gridcol="3" gridrow="23" pageId="7" pageNumber="168">19</td>
</tr>
<tr id="35086D27FF9C006798BFFA9759309041" box="[159,1390,1332,1354]" gridrow="24" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFA975DB49041" box="[159,490,1332,1354]" gridcol="0" gridrow="24" pageId="7" pageNumber="168">Abdominal Vertebrae</th>
<td id="76D9045BFF9C00679AF1FA975F719041" box="[721,815,1332,1354]" gridcol="1" gridrow="24" pageId="7" pageNumber="168">12</td>
<td id="76D9045BFF9C00679BB3FA9758D19041" box="[915,1167,1332,1354]" gridcol="2" gridrow="24" pageId="7" pageNumber="168">1114 (12)</td>
<td id="76D9045BFF9C00679D75FA9759309041" box="[1365,1390,1332,1354]" gridcol="3" gridrow="24" pageId="7" pageNumber="168">18</td>
</tr>
<tr id="35086D27FF9C006798BFFAFE59309078" box="[159,1390,1373,1395]" gridrow="25" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFAFE5DB49078" box="[159,490,1373,1395]" gridcol="0" gridrow="25" pageId="7" pageNumber="168">Caudal Vertebrae</th>
<td id="76D9045BFF9C00679AF1FAFE5F719078" box="[721,815,1373,1395]" gridcol="1" gridrow="25" pageId="7" pageNumber="168">46</td>
<td id="76D9045BFF9C00679BB3FAFE58D19078" box="[915,1167,1373,1395]" gridcol="2" gridrow="25" pageId="7" pageNumber="168">4449 (46)</td>
<td id="76D9045BFF9C00679D75FAFE59309078" box="[1365,1390,1373,1395]" gridcol="3" gridrow="25" pageId="7" pageNumber="168">17</td>
</tr>
<tr id="35086D27FF9C006798BFFA2559309097" box="[159,1390,1414,1436]" gridrow="26" pageId="7" pageNumber="168" rowspan-1="1">
<th id="76D9045BFF9C006798BFFA255DB49097" box="[159,490,1414,1436]" gridcol="0" gridrow="26" pageId="7" pageNumber="168">Dorsal Fin Origin</th>
<td id="76D9045BFF9C00679BB3FA2558D19097" box="[915,1167,1414,1436]" gridcol="2" gridrow="26" pageId="7" pageNumber="168">36 (4,5)</td>
<td id="76D9045BFF9C00679D75FA2559309097" box="[1365,1390,1414,1436]" gridcol="3" gridrow="26" pageId="7" pageNumber="168">12</td>
</tr>
<tr id="35086D27FF9C006798BFFA13593090E4" box="[159,1390,1456,1519]" gridrow="27" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFFA135DB490E4" box="[159,490,1456,1519]" gridcol="0" gridrow="27" pageId="7" pageNumber="168">Dorsal Fin Rays Anal Fin Rays</th>
<td id="76D9045BFF9C00679AF1FA135F7190E4" box="[721,815,1456,1519]" gridcol="1" gridrow="27" pageId="7" pageNumber="168">
~52
<emphasis id="B94CB377FF9CFF9F9AF1FA7B5E8390E4" bold="true" box="[721,733,1496,1519]" pageId="7" pageNumber="168">Ź</emphasis>
44
</td>
<td id="76D9045BFF9C00679BB3FA1358D190E4" box="[915,1167,1456,1519]" gridcol="2" gridrow="27" pageId="7" pageNumber="168">5158 (54) 4349 (47)</td>
<td id="76D9045BFF9C00679D75FA13593090E4" box="[1365,1390,1456,1519]" gridcol="3" gridrow="27" pageId="7" pageNumber="168">11 15</td>
</tr>
<tr id="35086D27FF9C006798BFF9A159309313" box="[159,1390,1538,1560]" gridrow="28" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFF9A15DB49313" box="[159,490,1538,1560]" gridcol="0" gridrow="28" pageId="7" pageNumber="168">Total Pectoral Rays</th>
<td id="76D9045BFF9C00679AF1F9A15F719313" box="[721,815,1538,1560]" gridcol="1" gridrow="28" pageId="7" pageNumber="168">30</td>
<td id="76D9045BFF9C00679BB3F9A158D19313" box="[915,1167,1538,1560]" gridcol="2" gridrow="28" pageId="7" pageNumber="168">2832 (30)</td>
<td id="76D9045BFF9C00679D75F9A159309313" box="[1365,1390,1538,1560]" gridcol="3" gridrow="28" pageId="7" pageNumber="168">24</td>
</tr>
<tr id="35086D27FF9C006798BFF98F59309349" box="[159,1390,1580,1602]" gridrow="29" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFF98F5DB49349" box="[159,490,1580,1602]" gridcol="0" gridrow="29" pageId="7" pageNumber="168">Pectoral Fin Rays (Upper Lobe)</th>
<td id="76D9045BFF9C00679AF1F98F5F719349" box="[721,815,1580,1602]" gridcol="1" gridrow="29" pageId="7" pageNumber="168">20</td>
<td id="76D9045BFF9C00679BB3F98F58D19349" box="[915,1167,1580,1602]" gridcol="2" gridrow="29" pageId="7" pageNumber="168">1823 (21)</td>
<td id="76D9045BFF9C00679D75F98F59309349" box="[1365,1390,1580,1602]" gridcol="3" gridrow="29" pageId="7" pageNumber="168">26</td>
</tr>
<tr id="35086D27FF9C006798BFF9F659309360" box="[159,1390,1621,1643]" gridrow="30" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFF9F65DB49360" box="[159,490,1621,1643]" gridcol="0" gridrow="30" pageId="7" pageNumber="168">Pectoral Rays (Notch)</th>
<td id="76D9045BFF9C00679AF1F9F65F719360" box="[721,815,1621,1643]" gridcol="1" gridrow="30" pageId="7" pageNumber="168">5</td>
<td id="76D9045BFF9C00679BB3F9F658D19360" box="[915,1167,1621,1643]" gridcol="2" gridrow="30" pageId="7" pageNumber="168">36 (5)</td>
<td id="76D9045BFF9C00679D75F9F659309360" box="[1365,1390,1621,1643]" gridcol="3" gridrow="30" pageId="7" pageNumber="168">27</td>
</tr>
<tr id="35086D27FF9C006798BFF9DD5930939F" box="[159,1390,1662,1684]" gridrow="31" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFF9DD5DB4939F" box="[159,490,1662,1684]" gridcol="0" gridrow="31" pageId="7" pageNumber="168">Pectoral Fin Rays (Lower Lobe)</th>
<td id="76D9045BFF9C00679AF1F9DD5F71939F" box="[721,815,1662,1684]" gridcol="1" gridrow="31" pageId="7" pageNumber="168">5</td>
<td id="76D9045BFF9C00679BB3F9DD58D1939F" box="[915,1167,1662,1684]" gridcol="2" gridrow="31" pageId="7" pageNumber="168">46 (5)</td>
<td id="76D9045BFF9C00679D75F9DD5930939F" box="[1365,1390,1662,1684]" gridcol="3" gridrow="31" pageId="7" pageNumber="168">26</td>
</tr>
<tr id="35086D27FF9C006798BFF90B593093B5" box="[159,1390,1704,1726]" gridrow="32" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFF90B5DB493B5" box="[159,490,1704,1726]" gridcol="0" gridrow="32" pageId="7" pageNumber="168">Total Caudal Fin Rays</th>
<td id="76D9045BFF9C00679AF1F90B5F7193B5" box="[721,815,1704,1726]" gridcol="1" gridrow="32" pageId="7" pageNumber="168">13</td>
<td id="76D9045BFF9C00679BB3F90B58D193B5" box="[915,1167,1704,1726]" gridcol="2" gridrow="32" pageId="7" pageNumber="168">1114 (13)</td>
<td id="76D9045BFF9C00679D75F90B593093B5" box="[1365,1390,1704,1726]" gridcol="3" gridrow="32" pageId="7" pageNumber="168">20</td>
</tr>
<tr id="35086D27FF9C006798BFF972593093EC" box="[159,1390,1745,1767]" gridrow="33" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFF9725DB493EC" box="[159,490,1745,1767]" gridcol="0" gridrow="33" pageId="7" pageNumber="168">Caudal Fin Rays (Upper)</th>
<td id="76D9045BFF9C00679AF1F9725F7193EC" box="[721,815,1745,1767]" gridcol="1" gridrow="33" pageId="7" pageNumber="168">5</td>
<td id="76D9045BFF9C00679BB3F97258D193EC" box="[915,1167,1745,1767]" gridcol="2" gridrow="33" pageId="7" pageNumber="168">46 (5)</td>
<td id="76D9045BFF9C00679D75F972593093EC" box="[1365,1390,1745,1767]" gridcol="3" gridrow="33" pageId="7" pageNumber="168">21</td>
</tr>
<tr id="35086D27FF9C006798BFF9595930921B" box="[159,1390,1786,1808]" gridrow="34" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFF9595DB4921B" box="[159,490,1786,1808]" gridcol="0" gridrow="34" pageId="7" pageNumber="168">Caudal Fin Rays (Lower)</th>
<td id="76D9045BFF9C00679AF1F9595F71921B" box="[721,815,1786,1808]" gridcol="1" gridrow="34" pageId="7" pageNumber="168">6</td>
<td id="76D9045BFF9C00679BB3F95958D1921B" box="[915,1167,1786,1808]" gridcol="2" gridrow="34" pageId="7" pageNumber="168">47 (6)</td>
<td id="76D9045BFF9C00679D75F9595930921B" box="[1365,1390,1786,1808]" gridcol="3" gridrow="34" pageId="7" pageNumber="168">21</td>
</tr>
<tr id="35086D27FF9C006798BFF88759309231" box="[159,1390,1828,1850]" gridrow="35" pageId="7" pageNumber="168">
<th id="76D9045BFF9C006798BFF8875DB49231" box="[159,490,1828,1850]" gridcol="0" gridrow="35" pageId="7" pageNumber="168">Caudal Fin Rays (Auxiliary)</th>
<td id="76D9045BFF9C00679AF1F8875F719231" box="[721,815,1828,1850]" gridcol="1" gridrow="35" pageId="7" pageNumber="168">2</td>
<td id="76D9045BFF9C00679BB3F88758D19231" box="[915,1167,1828,1850]" gridcol="2" gridrow="35" pageId="7" pageNumber="168">02 (1)</td>
<td id="76D9045BFF9C00679D75F88759309231" box="[1365,1390,1828,1850]" gridcol="3" gridrow="35" pageId="7" pageNumber="168">20</td>
</tr>
<tr id="35086D27FF9C006798BFF8EE59309268" box="[159,1390,1869,1891]" gridrow="36" pageId="7" pageNumber="168" rowspan-1="1">
<th id="76D9045BFF9C006798BFF8EE5DB49268" box="[159,490,1869,1891]" gridcol="0" gridrow="36" pageId="7" pageNumber="168">Pyloric Caeca</th>
<td id="76D9045BFF9C00679BB3F8EE58D19268" box="[915,1167,1869,1891]" gridcol="2" gridrow="36" pageId="7" pageNumber="168">59 (7)</td>
<td id="76D9045BFF9C00679D75F8EE59309268" box="[1365,1390,1869,1891]" gridcol="3" gridrow="36" pageId="7" pageNumber="168">17</td>
</tr>
</table>
</paragraph>
<caption id="DF473FEDFF93FF9098B7FF3B5F7D95C4" pageId="8" pageNumber="169">
<paragraph id="8B876F65FF93FF9098B7FF3B5F7D95C4" blockId="8.[151,1436,151,207]" pageId="8" pageNumber="169">
<emphasis id="B94CB377FF93FF9098B7FF3B5D5795A4" bold="true" box="[151,265,152,175]" pageId="8" pageNumber="169">TABLE 3.</emphasis>
Ratios of
<taxonomicName id="4C3814E6FF93FF909957FF345E0C95A4" authority="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen, 2017" authorityName="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen" authorityYear="2017" box="[375,594,151,175]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="8" pageNumber="169" phylum="Chordata" rank="species" species="swirei" status="sp. nov.">
<emphasis id="B94CB377FF93FF909957FF345E0C95A4" box="[375,594,151,175]" italics="true" pageId="8" pageNumber="169">Pseudoliparis swirei</emphasis>
</taxonomicName>
<emphasis id="B94CB377FF93FF909A79FF3B5EF095A4" bold="true" box="[601,686,152,175]" pageId="8" pageNumber="169">
<taxonomicNameLabel id="A27F0E0CFF93FF909A79FF3B5EF095A4" box="[601,686,152,175]" pageId="8" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. Preserved and fresh measurements presented. Body depth taken from photographs. Predorsal fin length measured from radiograph.
</paragraph>
</caption>
<paragraph id="8B876F65FF93FF909AC0FF4958B3940B" blockId="8.[499,1403,234,298]" box="[736,1261,234,256]" pageId="8" pageNumber="169">%SL %HL</paragraph>
<paragraph id="8B876F65FF93FF9099D3FEB75C8A917C" blockId="8.[499,1403,234,298]" lastBlockId="8.[155,1416,1121,1351]" pageId="8" pageNumber="169">
<typeStatus id="5483D1C7FF93FF9099D3FEB75E8A9421" box="[499,724,276,298]" pageId="8" pageNumber="169" type="holotype">Holotype Holotype</typeStatus>
and
<typeStatus id="5483D1C7FF93FF909B26FEB65F369421" box="[774,872,277,298]" pageId="8" pageNumber="169" type="paratype">Paratypes</typeStatus>
n
<typeStatus id="5483D1C7FF93FF909BE5FEB758FA9421" box="[965,1188,276,298]" pageId="8" pageNumber="169" type="holotype">Holotype Holotype</typeStatus>
and
<typeStatus id="5483D1C7FF93FF909CF6FEB659679421" box="[1238,1337,277,298]" pageId="8" pageNumber="169" type="paratype">Paratypes</typeStatus>
n Fresh
</paragraph>
<paragraph id="8B876F65FF93FF9098BBFEE85925912F" pageId="8" pageNumber="169">
<table id="F9389DC5FF93006798BBFEE859D6912F" box="[155,1416,331,1060]" gridcols="7" gridrows="18" pageId="8" pageNumber="169">
<tr id="35086D27FF93006798BBFEE859D6946A" box="[155,1416,331,353]" gridrow="0" pageId="8" pageNumber="169" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76D9045BFF93006798BBFEE85DE9946A" box="[155,439,331,353]" gridcol="0" gridrow="0" pageId="8" pageNumber="169">Preserved</th>
</tr>
<tr id="35086D27FF93006798BBFED759D69481" box="[155,1416,372,394]" gridrow="1" pageId="8" pageNumber="169" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76D9045BFF93006798BBFED75DE99481" box="[155,439,372,394]" gridcol="0" gridrow="1" pageId="8" pageNumber="169">Head Length</th>
<td id="76D9045BFF93006799D3FED75E7F9481" box="[499,545,372,394]" gridcol="1" gridrow="1" pageId="8" pageNumber="169">18.2</td>
<td id="76D9045BFF9300679A56FED75F129481" box="[630,844,372,394]" gridcol="2" gridrow="1" pageId="8" pageNumber="169">17.021.7 (19.8±1.1)</td>
<td id="76D9045BFF9300679BABFED75FFB9481" box="[907,933,372,394]" gridcol="3" gridrow="1" pageId="8" pageNumber="169">19</td>
</tr>
<tr id="35086D27FF93006798BBFE3D59D694BF" box="[155,1416,414,436]" gridrow="2" pageId="8" pageNumber="169">
<th id="76D9045BFF93006798BBFE3D5DE994BF" box="[155,439,414,436]" gridcol="0" gridrow="2" pageId="8" pageNumber="169">Head Width</th>
<td id="76D9045BFF93006799D3FE3D5E7F94BF" box="[499,545,414,436]" gridcol="1" gridrow="2" pageId="8" pageNumber="169">12.6</td>
<td id="76D9045BFF9300679A56FE3D5F1294BF" box="[630,844,414,436]" gridcol="2" gridrow="2" pageId="8" pageNumber="169">12.619.6 (15.8±2.5)</td>
<td id="76D9045BFF9300679BABFE3D5FFB94BF" box="[907,933,414,436]" gridcol="3" gridrow="2" pageId="8" pageNumber="169">8</td>
<td id="76D9045BFF9300679BE5FE3D5FA194BF" box="[965,1023,414,436]" gridcol="4" gridrow="2" pageId="8" pageNumber="169">69.3</td>
<td id="76D9045BFF9300679C66FE3D591094BF" box="[1094,1358,414,436]" gridcol="5" gridrow="2" pageId="8" pageNumber="169">68.2101.0 (81.0±11.5)</td>
<td id="76D9045BFF9300679D4EFE3D59D694BF" box="[1390,1416,414,436]" gridcol="6" gridrow="2" pageId="8" pageNumber="169">8</td>
</tr>
<tr id="35086D27FF93006798BBFE6459D694D6" box="[155,1416,455,477]" gridrow="3" pageId="8" pageNumber="169">
<th id="76D9045BFF93006798BBFE645DE994D6" box="[155,439,455,477]" gridcol="0" gridrow="3" pageId="8" pageNumber="169">Snout Length</th>
<td id="76D9045BFF93006799D3FE645E7F94D6" box="[499,545,455,477]" gridcol="1" gridrow="3" pageId="8" pageNumber="169">6.4</td>
<td id="76D9045BFF9300679A56FE645F1294D6" box="[630,844,455,477]" gridcol="2" gridrow="3" pageId="8" pageNumber="169">5.28.3 (6.5±0.7)</td>
<td id="76D9045BFF9300679BABFE645FFB94D6" box="[907,933,455,477]" gridcol="3" gridrow="3" pageId="8" pageNumber="169">21</td>
<td id="76D9045BFF9300679BE5FE645FA194D6" box="[965,1023,455,477]" gridcol="4" gridrow="3" pageId="8" pageNumber="169">35.4</td>
<td id="76D9045BFF9300679C66FE64591094D6" box="[1094,1358,455,477]" gridcol="5" gridrow="3" pageId="8" pageNumber="169">28.538.0 (32.9±2.6)</td>
<td id="76D9045BFF9300679D4EFE6459D694D6" box="[1390,1416,455,477]" gridcol="6" gridrow="3" pageId="8" pageNumber="169">18</td>
</tr>
<tr id="35086D27FF93006798BBFE5259D6970C" box="[155,1416,497,519]" gridrow="4" pageId="8" pageNumber="169">
<th id="76D9045BFF93006798BBFE525DE9970C" box="[155,439,497,519]" gridcol="0" gridrow="4" pageId="8" pageNumber="169">Orbit Width</th>
<td id="76D9045BFF93006799D3FE525E7F970C" box="[499,545,497,519]" gridcol="1" gridrow="4" pageId="8" pageNumber="169">3.4</td>
<td id="76D9045BFF9300679A56FE525F12970C" box="[630,844,497,519]" gridcol="2" gridrow="4" pageId="8" pageNumber="169">2.34.4 (3.2±0.6)</td>
<td id="76D9045BFF9300679BABFE525FFB970C" box="[907,933,497,519]" gridcol="3" gridrow="4" pageId="8" pageNumber="169">17</td>
<td id="76D9045BFF9300679BE5FE525FA1970C" box="[965,1023,497,519]" gridcol="4" gridrow="4" pageId="8" pageNumber="169">18.5</td>
<td id="76D9045BFF9300679C66FE525910970C" box="[1094,1358,497,519]" gridcol="5" gridrow="4" pageId="8" pageNumber="169">12.120.1 (16.0±2.3)</td>
<td id="76D9045BFF9300679D4EFE5259D6970C" box="[1390,1416,497,519]" gridcol="6" gridrow="4" pageId="8" pageNumber="169">16</td>
</tr>
<tr id="35086D27FF93006798BBFDB859D6973A" box="[155,1416,539,561]" gridrow="5" pageId="8" pageNumber="169">
<th id="76D9045BFF93006798BBFDB85DE9973A" box="[155,439,539,561]" gridcol="0" gridrow="5" pageId="8" pageNumber="169">Upper Jaw Length</th>
<td id="76D9045BFF93006799D3FDB85E7F973A" box="[499,545,539,561]" gridcol="1" gridrow="5" pageId="8" pageNumber="169">~8.4</td>
<td id="76D9045BFF9300679A56FDB85F12973A" box="[630,844,539,561]" gridcol="2" gridrow="5" pageId="8" pageNumber="169">7.410.5 (9.1±0.8)</td>
<td id="76D9045BFF9300679BABFDB85FFB973A" box="[907,933,539,561]" gridcol="3" gridrow="5" pageId="8" pageNumber="169">20</td>
<td id="76D9045BFF9300679BE5FDB85FA1973A" box="[965,1023,539,561]" gridcol="4" gridrow="5" pageId="8" pageNumber="169">~46.0</td>
<td id="76D9045BFF9300679C66FDB85910973A" box="[1094,1358,539,561]" gridcol="5" gridrow="5" pageId="8" pageNumber="169">38.352.2 (45.9±3.3)</td>
<td id="76D9045BFF9300679D4EFDB859D6973A" box="[1390,1416,539,561]" gridcol="6" gridrow="5" pageId="8" pageNumber="169">17</td>
</tr>
<tr id="35086D27FF93006798BBFDE759D69751" box="[155,1416,580,602]" gridrow="6" pageId="8" pageNumber="169">
<th id="76D9045BFF93006798BBFDE75DE99751" box="[155,439,580,602]" gridcol="0" gridrow="6" pageId="8" pageNumber="169">Lower Jaw Length</th>
<td id="76D9045BFF93006799D3FDE75E7F9751" box="[499,545,580,602]" gridcol="1" gridrow="6" pageId="8" pageNumber="169">7.9</td>
<td id="76D9045BFF9300679A56FDE75F129751" box="[630,844,580,602]" gridcol="2" gridrow="6" pageId="8" pageNumber="169">6.510.6 (8.3±1.0)</td>
<td id="76D9045BFF9300679BABFDE75FFB9751" box="[907,933,580,602]" gridcol="3" gridrow="6" pageId="8" pageNumber="169">20</td>
<td id="76D9045BFF9300679BE5FDE75FA19751" box="[965,1023,580,602]" gridcol="4" gridrow="6" pageId="8" pageNumber="169">43.4</td>
<td id="76D9045BFF9300679C66FDE759109751" box="[1094,1358,580,602]" gridcol="5" gridrow="6" pageId="8" pageNumber="169">33.753.2 (42.2±4.7)</td>
<td id="76D9045BFF9300679D4EFDE759D69751" box="[1390,1416,580,602]" gridcol="6" gridrow="6" pageId="8" pageNumber="169">17</td>
</tr>
<tr id="35086D27FF93006798BBFDCD59D6978F" box="[155,1416,622,644]" gridrow="7" pageId="8" pageNumber="169" rowspan-1="1" rowspan-4="1">
<th id="76D9045BFF93006798BBFDCD5DE9978F" box="[155,439,622,644]" gridcol="0" gridrow="7" pageId="8" pageNumber="169">Upper Pectoral Fin Length</th>
<td id="76D9045BFF9300679A56FDCD5F12978F" box="[630,844,622,644]" gridcol="2" gridrow="7" pageId="8" pageNumber="169">12.619.8 (15.2±2.2)</td>
<td id="76D9045BFF9300679BABFDCD5FFB978F" box="[907,933,622,644]" gridcol="3" gridrow="7" pageId="8" pageNumber="169">21</td>
<td id="76D9045BFF9300679C66FDCD5910978F" box="[1094,1358,622,644]" gridcol="5" gridrow="7" pageId="8" pageNumber="169">60.295.4 (75.4±11.3)</td>
<td id="76D9045BFF9300679D4EFDCD59D6978F" box="[1390,1416,622,644]" gridcol="6" gridrow="7" pageId="8" pageNumber="169">16</td>
</tr>
<tr id="35086D27FF93006798BBFD3459D697A6" box="[155,1416,663,685]" gridrow="8" pageId="8" pageNumber="169">
<th id="76D9045BFF93006798BBFD345DE997A6" box="[155,439,663,685]" gridcol="0" gridrow="8" pageId="8" pageNumber="169">Lower Pectoral Fin Length</th>
<td id="76D9045BFF93006799D3FD345E7F97A6" box="[499,545,663,685]" gridcol="1" gridrow="8" pageId="8" pageNumber="169">8.6</td>
<td id="76D9045BFF9300679A56FD345F1297A6" box="[630,844,663,685]" gridcol="2" gridrow="8" pageId="8" pageNumber="169">6.312.1 (7.8±1.4)</td>
<td id="76D9045BFF9300679BABFD345FFB97A6" box="[907,933,663,685]" gridcol="3" gridrow="8" pageId="8" pageNumber="169">16</td>
<td id="76D9045BFF9300679BE5FD345FA197A6" box="[965,1023,663,685]" gridcol="4" gridrow="8" pageId="8" pageNumber="169">47.1</td>
<td id="76D9045BFF9300679C66FD34591097A6" box="[1094,1358,663,685]" gridcol="5" gridrow="8" pageId="8" pageNumber="169">30.061.6 (39.0±7.7)</td>
<td id="76D9045BFF9300679D4EFD3459D697A6" box="[1390,1416,663,685]" gridcol="6" gridrow="8" pageId="8" pageNumber="169">14</td>
</tr>
<tr id="35086D27FF93006798BBFD6259D697DC" box="[155,1416,705,727]" gridrow="9" pageId="8" pageNumber="169" rowspan-1="1" rowspan-4="1">
<th id="76D9045BFF93006798BBFD625DE997DC" box="[155,439,705,727]" gridcol="0" gridrow="9" pageId="8" pageNumber="169">Gill Opening</th>
<td id="76D9045BFF9300679A56FD625F1297DC" box="[630,844,705,727]" gridcol="2" gridrow="9" pageId="8" pageNumber="169">4.45.3 (5.0±0.5)</td>
<td id="76D9045BFF9300679BABFD625FFB97DC" box="[907,933,705,727]" gridcol="3" gridrow="9" pageId="8" pageNumber="169">3</td>
<td id="76D9045BFF9300679C66FD62591097DC" box="[1094,1358,705,727]" gridcol="5" gridrow="9" pageId="8" pageNumber="169">22.326.5 (24.9±2.3)</td>
<td id="76D9045BFF9300679D4EFD6259D697DC" box="[1390,1416,705,727]" gridcol="6" gridrow="9" pageId="8" pageNumber="169">3</td>
</tr>
<tr id="35086D27FF93006798BBFD4859D6960A" box="[155,1416,747,769]" gridrow="10" pageId="8" pageNumber="169" rowspan-1="1" rowspan-4="1">
<th id="76D9045BFF93006798BBFD485DE9960A" box="[155,439,747,769]" gridcol="0" gridrow="10" pageId="8" pageNumber="169">Disk Length</th>
<td id="76D9045BFF9300679A56FD485F12960A" box="[630,844,747,769]" gridcol="2" gridrow="10" pageId="8" pageNumber="169">2.45.6 (3.9±0.9)</td>
<td id="76D9045BFF9300679BABFD485FFB960A" box="[907,933,747,769]" gridcol="3" gridrow="10" pageId="8" pageNumber="169">13</td>
<td id="76D9045BFF9300679C66FD485910960A" box="[1094,1358,747,769]" gridcol="5" gridrow="10" pageId="8" pageNumber="169">12.125.9 (20.0±4.0)</td>
<td id="76D9045BFF9300679D4EFD4859D6960A" box="[1390,1416,747,769]" gridcol="6" gridrow="10" pageId="8" pageNumber="169">12</td>
</tr>
<tr id="35086D27FF93006798BBFCB759D69621" box="[155,1416,788,810]" gridrow="11" pageId="8" pageNumber="169">
<th id="76D9045BFF93006798BBFCB75DE99621" box="[155,439,788,810]" gridcol="0" gridrow="11" pageId="8" pageNumber="169">Distance: Disk to Anus</th>
<td id="76D9045BFF93006799D3FCB75E7F9621" box="[499,545,788,810]" gridcol="1" gridrow="11" pageId="8" pageNumber="169">9.1</td>
<td id="76D9045BFF9300679A56FCB75F129621" box="[630,844,788,810]" gridcol="2" gridrow="11" pageId="8" pageNumber="169">8.615.6 (10.8±3.2)</td>
<td id="76D9045BFF9300679BABFCB75FFB9621" box="[907,933,788,810]" gridcol="3" gridrow="11" pageId="8" pageNumber="169">4</td>
<td id="76D9045BFF9300679BE5FCB75FA19621" box="[965,1023,788,810]" gridcol="4" gridrow="11" pageId="8" pageNumber="169">50.3</td>
<td id="76D9045BFF9300679C66FCB759109621" box="[1094,1358,788,810]" gridcol="5" gridrow="11" pageId="8" pageNumber="169">43.183.8 (59.0±21.7)</td>
<td id="76D9045BFF9300679D4EFCB759D69621" box="[1390,1416,788,810]" gridcol="6" gridrow="11" pageId="8" pageNumber="169">3</td>
</tr>
<tr id="35086D27FF93006798BBFC9D59D6965F" box="[155,1416,830,852]" gridrow="12" pageId="8" pageNumber="169">
<th id="76D9045BFF93006798BBFC9D5DE9965F" box="[155,439,830,852]" gridcol="0" gridrow="12" pageId="8" pageNumber="169">Distance: Mandible to Disk</th>
<td id="76D9045BFF93006799D3FC9D5E7F965F" box="[499,545,830,852]" gridcol="1" gridrow="12" pageId="8" pageNumber="169">8.5</td>
<td id="76D9045BFF9300679A56FC9D5F12965F" box="[630,844,830,852]" gridcol="2" gridrow="12" pageId="8" pageNumber="169">7.713.7 (10.0±1.4)</td>
<td id="76D9045BFF9300679BABFC9D5FFB965F" box="[907,933,830,852]" gridcol="3" gridrow="12" pageId="8" pageNumber="169">20</td>
<td id="76D9045BFF9300679BE5FC9D5FA1965F" box="[965,1023,830,852]" gridcol="4" gridrow="12" pageId="8" pageNumber="169">46.6</td>
<td id="76D9045BFF9300679C66FC9D5910965F" box="[1094,1358,830,852]" gridcol="5" gridrow="12" pageId="8" pageNumber="169">39.859.8 (49.7±5.4)</td>
<td id="76D9045BFF9300679D4EFC9D59D6965F" box="[1390,1416,830,852]" gridcol="6" gridrow="12" pageId="8" pageNumber="169">17</td>
</tr>
<tr id="35086D27FF93006798BBFCC459D69676" box="[155,1416,871,893]" gridrow="13" pageId="8" pageNumber="169" rowspan-1="1" rowspan-4="1">
<th id="76D9045BFF93006798BBFCC45DE99676" box="[155,439,871,893]" gridcol="0" gridrow="13" pageId="8" pageNumber="169">Lower Lobe Distance</th>
<td id="76D9045BFF9300679A56FCC45F129676" box="[630,844,871,893]" gridcol="2" gridrow="13" pageId="8" pageNumber="169">1.84.7 (3.1±0.9)</td>
<td id="76D9045BFF9300679BABFCC45FFB9676" box="[907,933,871,893]" gridcol="3" gridrow="13" pageId="8" pageNumber="169">12</td>
<td id="76D9045BFF9300679C66FCC459109676" box="[1094,1358,871,893]" gridcol="5" gridrow="13" pageId="8" pageNumber="169">8.323.0 (15.2±4.7)</td>
<td id="76D9045BFF9300679D4EFCC459D69676" box="[1390,1416,871,893]" gridcol="6" gridrow="13" pageId="8" pageNumber="169">10</td>
</tr>
<tr id="35086D27FF93006798BBFC3259D696AC" box="[155,1416,913,935]" gridrow="14" pageId="8" pageNumber="169">
<th id="76D9045BFF93006798BBFC325DE996AC" box="[155,439,913,935]" gridcol="0" gridrow="14" pageId="8" pageNumber="169">Predorsal Fin Length</th>
<td id="76D9045BFF93006799D3FC325E7F96AC" box="[499,545,913,935]" gridcol="1" gridrow="14" pageId="8" pageNumber="169">32.0</td>
<td id="76D9045BFF9300679A56FC325F1296AC" box="[630,844,913,935]" gridcol="2" gridrow="14" pageId="8" pageNumber="169">22.732.6 (27.8±3.1)</td>
<td id="76D9045BFF9300679BABFC325FFB96AC" box="[907,933,913,935]" gridcol="3" gridrow="14" pageId="8" pageNumber="169">13</td>
<td id="76D9045BFF9300679BE5FC325FA196AC" box="[965,1023,913,935]" gridcol="4" gridrow="14" pageId="8" pageNumber="169">176.4</td>
<td id="76D9045BFF9300679C66FC32591096AC" box="[1094,1358,913,935]" gridcol="5" gridrow="14" pageId="8" pageNumber="169">118.1176.4 (142.2±18.2)</td>
<td id="76D9045BFF9300679D4EFC3259D696AC" box="[1390,1416,913,935]" gridcol="6" gridrow="14" pageId="8" pageNumber="169">13</td>
</tr>
<tr id="35086D27FF93006798BBFC1859D696DA" box="[155,1416,955,977]" gridrow="15" pageId="8" pageNumber="169">
<th id="76D9045BFF93006798BBFC185DE996DA" box="[155,439,955,977]" gridcol="0" gridrow="15" pageId="8" pageNumber="169">Distance: Snout to Anus</th>
<td id="76D9045BFF93006799D3FC185E7F96DA" box="[499,545,955,977]" gridcol="1" gridrow="15" pageId="8" pageNumber="169">22.3</td>
<td id="76D9045BFF9300679A56FC185F1296DA" box="[630,844,955,977]" gridcol="2" gridrow="15" pageId="8" pageNumber="169">22.329.3 (26.0±3.0)</td>
<td id="76D9045BFF9300679BABFC185FFB96DA" box="[907,933,955,977]" gridcol="3" gridrow="15" pageId="8" pageNumber="169">6</td>
<td id="76D9045BFF9300679BE5FC185FA196DA" box="[965,1023,955,977]" gridcol="4" gridrow="15" pageId="8" pageNumber="169">122.8</td>
<td id="76D9045BFF9300679C66FC18591096DA" box="[1094,1358,955,977]" gridcol="5" gridrow="15" pageId="8" pageNumber="169">114.6143.3 (130.0±11.3)</td>
<td id="76D9045BFF9300679D4EFC1859D696DA" box="[1390,1416,955,977]" gridcol="6" gridrow="15" pageId="8" pageNumber="169">5</td>
</tr>
<tr id="35086D27FF93006798BBFC4759D696F1" box="[155,1416,996,1018]" gridrow="16" pageId="8" pageNumber="169">
<th id="76D9045BFF93006798BBFC475DE996F1" box="[155,439,996,1018]" gridcol="0" gridrow="16" pageId="8" pageNumber="169">Distance: Mandible to Anus</th>
<td id="76D9045BFF93006799D3FC475E7F96F1" box="[499,545,996,1018]" gridcol="1" gridrow="16" pageId="8" pageNumber="169">20.2</td>
<td id="76D9045BFF9300679A56FC475F1296F1" box="[630,844,996,1018]" gridcol="2" gridrow="16" pageId="8" pageNumber="169">20.227.6 (24.3±3.4)</td>
<td id="76D9045BFF9300679BABFC475FFB96F1" box="[907,933,996,1018]" gridcol="3" gridrow="16" pageId="8" pageNumber="169">4</td>
<td id="76D9045BFF9300679BE5FC475FA196F1" box="[965,1023,996,1018]" gridcol="4" gridrow="16" pageId="8" pageNumber="169">111.1</td>
<td id="76D9045BFF9300679C66FC47591096F1" box="[1094,1358,996,1018]" gridcol="5" gridrow="16" pageId="8" pageNumber="169">111.1134.8 (120.1±12.8)</td>
<td id="76D9045BFF9300679D4EFC4759D696F1" box="[1390,1416,996,1018]" gridcol="6" gridrow="16" pageId="8" pageNumber="169">3</td>
</tr>
<tr id="35086D27FF93006798BBFBAD59D6912F" box="[155,1416,1038,1060]" gridrow="17" pageId="8" pageNumber="169">
<th id="76D9045BFF93006798BBFBAD5DE9912F" box="[155,439,1038,1060]" gridcol="0" gridrow="17" pageId="8" pageNumber="169">Distance: Anus to Anal Fin</th>
<td id="76D9045BFF93006799D3FBAD5E7F912F" box="[499,545,1038,1060]" gridcol="1" gridrow="17" pageId="8" pageNumber="169">10.5</td>
<td id="76D9045BFF9300679A56FBAD5F12912F" box="[630,844,1038,1060]" gridcol="2" gridrow="17" pageId="8" pageNumber="169">8.715.9 (12.5±2.9)</td>
<td id="76D9045BFF9300679BABFBAD5FFB912F" box="[907,933,1038,1060]" gridcol="3" gridrow="17" pageId="8" pageNumber="169">5</td>
<td id="76D9045BFF9300679BE5FBAD5FA1912F" box="[965,1023,1038,1060]" gridcol="4" gridrow="17" pageId="8" pageNumber="169">57.7</td>
<td id="76D9045BFF9300679C66FBAD5910912F" box="[1094,1358,1038,1060]" gridcol="5" gridrow="17" pageId="8" pageNumber="169">46.566.6 (59.1±9.3)</td>
<td id="76D9045BFF9300679D4EFBAD59D6912F" box="[1390,1416,1038,1060]" gridcol="6" gridrow="17" pageId="8" pageNumber="169">4</td>
</tr>
</table>
</paragraph>
<paragraph id="8B876F65FF93FF9098BBFB295FFB91AB" blockId="8.[155,1416,1121,1351]" box="[155,933,1162,1184]" pageId="8" pageNumber="169">Head Length 20.6 14.521.9 (18.7±1.8) 33</paragraph>
<paragraph id="8B876F65FF93FF9098BBFB175F08935E" blockId="8.[155,1416,1121,1351]" lastBlockId="8.[151,1437,1415,2017]" pageId="8" pageNumber="169">
Snout Length 7.2 4.39.3 (6.4±1.0) 33 35.0 26.345.5 (33.8±5.3) 37 Eye Width 2.1 0.92.9 (1.8±0.5) 33 10.0 5.315.4 (9.9±2.4) 37 Body Depth 18.6 18.631.2 (24.5±3.1) 18 90.0 90.0156.8 (128.7±17.9) 18 Preanal Fin Length 37.1 36.549.5 (42.0±3.5) 33 180.0 180.0295.8 (228.0±26.1) 37 Disk present, oval, longer than wide, below cheek and gill cavity between pectoral fin notches; well behind pectoral symphysis. Bones fully developed but weakly calcified; all elements present. Disk and pectoral girdle supported by a pair of clearly visible and strongly developed muscles extending anteriorly to pectoral symphysis, probably infracarinalis anterior (D.L. Stein,
<emphasis id="B94CB377FF93FF909AB5FA575FF39307" box="[661,941,1524,1548]" italics="true" pageId="8" pageNumber="169">personal communication</emphasis>
). Disk structure supporting a thin layer of tissue, often damaged or missing entirely; disk margin only slightly thicker than more central tissue. In cross section, disk rays clearly flattened as if to support disk margin.
</paragraph>
</subSubSection>
<subSubSection id="C3223CEEFF93FF9198E7F9FC581C9333" lastPageId="9" lastPageNumber="170" pageId="8" pageNumber="169" type="discussion">
<paragraph id="8B876F65FF93FF9198E7F9FC5FE89416" blockId="8.[151,1437,1415,2017]" lastBlockId="9.[151,1437,151,1880]" lastPageId="9" lastPageNumber="170" pageId="8" pageNumber="169">
Body depth dependent on reproductive state and fullness of stomach, usually much deeper than head depth, but shallow above vertebral column. Abdominal cavity long; peritoneum and body cavity extending to about 40% standard length. Total vertebrae 61 (5662); anterior 13 (1114), caudal 46 (4449). In the 19 individuals in good enough condition to be radiographed, a double ray is present at or near anal fin first ray, usually between the second and third haemal spines of the caudal vertebrae. Pre-dorsal length about 27% SL (18.632.6), dorsal fin origin between fourth and fifth vertebrae (origin after vertebrae 35). Pre-anal-fin length about 42% SL (36.349.5)*. Anus far posterior to disk, roughly 2/3 of distance from disk to anal fin origin. Pyloric caeca usually 7 (59), located left ventrally in body cavity; thick, digitate, usually separated into two distinct size classes, most commonly 4 short and 3 long, longest about 8.7% SL (5.911.7), shortest 3.7% SL (2.45.3). Longer caeca generally grouped together. Hypural with obvious suture; caudal fin most commonly of 13 (1114) rays, ventral one or two often rudimentary. Skin thin, transparent; subdermal extracellular matrix (SECM;
<bibRefCitation id="EFA91294FF93FF909BD1F86A588C92EB" author="Eastman" box="[1009,1234,1992,2016]" pageId="8" pageNumber="169" refString="Eastman, J., Hikida, R. &amp; Devries, A. (1994) Buoyancy studies and microscopy of skin and subdermal extracellular matrix of the antarctic snailfish, Paraliparis devriesi. Journal of Morphology, 220, 85 - 101. https: // doi. org / 10.1002 / jmor. 1052200108" type="journal article" year="1994">
Eastman
<emphasis id="B94CB377FF93FF909C79F86A58CC92EB" box="[1113,1170,1992,2016]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
1994
</bibRefCitation>
) thick below skin and between muscle bands. Total and standard lengths were approximately 10% shorter after preservation. The subdermal extracellular matrix is also lost after capture and preservation, resulting in changes to shape and proportion (Gerringer
<emphasis id="B94CB377FF92FF9199B5FF425D8E95F3" box="[405,464,224,248]" italics="true" pageId="9" pageNumber="170">et al.</emphasis>
unpublished data). With increased use of visual
<emphasis id="B94CB377FF92FF919C21FF43581695FC" box="[1025,1096,224,247]" italics="true" pageId="9" pageNumber="170">in situ</emphasis>
techniques, reporting of both fresh and preserved specimen features will become increasingly useful.
</paragraph>
<paragraph id="8B876F65FF92FF9198E7FE8458B194D2" blockId="9.[151,1437,151,1880]" pageId="9" pageNumber="170">
The 37 individuals used for description varied in size from
<quantity id="4CC0C280FF92FF919BB1FE8458439434" box="[913,1053,295,320]" metricMagnitude="-1" metricUnit="m" metricValue="1.6199999999999999" metricValueMax="2.35" metricValueMin="0.89" pageId="9" pageNumber="170" unit="mm" value="162.0" valueMax="235.0" valueMin="89.0">89235 mm</quantity>
SL, apparently covering a wide developmental range for the species. Some characters correlated significantly with ontogeny, explaining much of the variation in ratios. Both the upper and lower pectoral-fin lobe lengths as a percentage of SL decreased significantly with increasing SL (upper: F1,25=11.88, p&lt;0.01, R2=0.322; lower F1,23=5.05, p&lt;0.05, R2=0.180). Proportional orbit width decreased with increasing standard length (F1,18=26.25, p&lt;0.01, R2=0.593).
</paragraph>
<paragraph id="8B876F65FF92FF9198E7FE4B5E06979B" blockId="9.[151,1437,151,1880]" pageId="9" pageNumber="170">In life, body pinkish-white, skin and peritoneum transparent; internal organs (liver, stomach, pyloric caeca) and muscles of trunk clearly visible through skin and thin abdominal wall. Anterior bundles of epaxial muscle thick, becoming less densely packed posteriorly. Some larger specimens with dusky skin on head. Pyloric caeca orange; most individuals entirely lacked both internal and external pigmentation. In alcohol, except for those with dusky heads, specimens uniformly pale.</paragraph>
<paragraph id="8B876F65FF92FF9198E7FD3858E19030" blockId="9.[151,1437,151,1880]" pageId="9" pageNumber="170">
Phylogenetic inference supports placement within the genus
<taxonomicName id="4C3814E6FF92FF919BB7FD38586D97B8" baseAuthorityName="Andriashev" baseAuthorityYear="1955" box="[919,1075,667,691]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="genus">
<emphasis id="B94CB377FF92FF919BB7FD38586D97B8" box="[919,1075,667,691]" italics="true" pageId="9" pageNumber="170">Pseudoliparis</emphasis>
</taxonomicName>
, with
<taxonomicName id="4C3814E6FF92FF919CA5FD3F58B797B8" authority="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen, 2017" authorityName="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen" authorityYear="2017" box="[1157,1257,668,691]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="species" species="swirei" status="sp. nov.">
<emphasis id="B94CB377FF92FF919CA5FD3F58B797B8" box="[1157,1257,668,691]" italics="true" pageId="9" pageNumber="170">P. swirei</emphasis>
</taxonomicName>
<emphasis id="B94CB377FF92FF919CD6FD38590897BF" bold="true" box="[1270,1366,667,692]" pageId="9" pageNumber="170">
<taxonomicNameLabel id="A27F0E0CFF92FF919CD6FD38590897BF" box="[1270,1366,667,692]" pageId="9" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
more closely related to
<taxonomicName id="4C3814E6FF92FF91997CFD635D8D97DC" baseAuthorityName="Andriashev &amp; Pitruk" baseAuthorityYear="1993" box="[348,467,703,727]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="species" species="belyaevi">
<emphasis id="B94CB377FF92FF91997CFD635D8D97DC" box="[348,467,703,727]" italics="true" pageId="9" pageNumber="170">P. belyaevi</emphasis>
</taxonomicName>
of the
<collectingCountry id="F32F2FF5FF92FF919A01FD635E3F97DC" box="[545,609,704,727]" name="Japan" pageId="9" pageNumber="170">Japan</collectingCountry>
Trench than to the Kermadec Trench liparids (
<taxonomicName id="4C3814E6FF92FF919C49FD1C59CA97DC" baseAuthorityName="Nielsen" baseAuthorityYear="1964" box="[1129,1428,703,727]" class="Actinopterygii" family="Liparidae" genus="Notoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="species" species="kermadecensis">
<emphasis id="B94CB377FF92FF919C49FD1C59CA97DC" box="[1129,1428,703,727]" italics="true" pageId="9" pageNumber="170">Notoliparis kermadecensis</emphasis>
</taxonomicName>
;
<taxonomicName id="4C3814E6FF92FF9198B7FD475D4A97F0" baseAuthorityName="Stein" baseAuthorityYear="2016" box="[151,276,740,763]" class="Actinopterygii" family="Liparidae" genus="Notoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="species" species="stewarti">
<emphasis id="B94CB377FF92FF9198B7FD475D4A97F0" box="[151,276,740,763]" italics="true" pageId="9" pageNumber="170">N. stewarti</emphasis>
</taxonomicName>
). Phylogenetic relationships of
<taxonomicName id="4C3814E6FF92FF919AA1FD475E8197F0" authority="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen, 2017" authorityName="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen" authorityYear="2017" box="[641,735,740,763]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="species" species="swirei" status="sp. nov.">
<emphasis id="B94CB377FF92FF919AA1FD475E8197F0" box="[641,735,740,763]" italics="true" pageId="9" pageNumber="170">P. swirei</emphasis>
</taxonomicName>
<emphasis id="B94CB377FF92FF919ACBFD405F1997F7" bold="true" box="[747,839,739,764]" pageId="9" pageNumber="170">
<taxonomicNameLabel id="A27F0E0CFF92FF919ACBFD405F1997F7" box="[747,839,739,764]" pageId="9" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and closely-related species based on 16S, COI, and Cyt-
<emphasis id="B94CB377FF92FF9198E9FCA45C899614" box="[201,215,775,799]" italics="true" pageId="9" pageNumber="170">b</emphasis>
are presented in
<figureCitation id="130373E0FF92FF9199B4FCA45DAA962B" box="[404,500,775,800]" captionStart="FIGURE 6" captionStartId="10.[151,250,1073,1095]" captionTargetBox="[352,1239,212,1026]" captionTargetId="figure@10.[340,1247,193,1052]" captionTargetPageId="10" captionText="FIGURE 6. Phylogenetic relationships of Pseudoliparis swirei sp. nov. to closely related hadal liparids, P. belyaevi from the Japan Trench and Notoliparis kermadecensis and Notoliparis stewarti from the Kermadec Trench. Maximum likelihood (ML) trees based on A) cytochrome c oxidase subunit I (1399 bp), B) cytochrome b (1007 bp), and C) 16S rRNA (1472 bp). ML bootstrap support (&gt;70%) and Bayesianposterior probability (&gt;0.70) valuesareshown as ML/BI.Bold values indicate thenode that supports placement of P. swirei sp. nov. within the genus Pseudoliparis. Aptocyclus ventricosus (GenBank accession NC008129.1) was used to root each tree." httpUri="https://zenodo.org/record/1067566/files/figure.png" pageId="9" pageNumber="170">Figure 6</figureCitation>
, with estimates of evolutionary divergence among species reported in
<tableCitation id="C6BA5ADEFF92FF919D25FCA459059614" box="[1285,1371,775,800]" captionStart="TABLE 4" captionStartId="12.[119,198,512,533]" captionText="TABLE 4. Evolutionary đivergence between Pseudoliparis swirei sp. nov. anđ close relatives. Average genetic đistance among species calculateđ for (A) cytochrome c oxiđase subunit I (644 bp), (B) cytochrome b (744 bp), anđ (C) 16S rRNA (699 bp), baseđ on the Tamura-Nei substitution mođel with gamma-đistributeđ rate variation across sites. Sequences from P.swirei sp.nov., N.kermadecensis,N.stewarti đerive fromthis stuđy, P.belyaevi are unpublisheđ đata fromT. Satoh (National Museum of Nature anđ Science, Japan), sequences from C. rastrinus, C. cypselurus, A. ventricosus were from NCBI (GenBank Accession Numbers: AB565515.1, AB565517.1, AB565629.1, FJ164433.1," pageId="9" pageNumber="170">Table 4</tableCitation>
. Both the 16S and Cyt-
<emphasis id="B94CB377FF92FF919978FC885D389648" box="[344,358,811,835]" italics="true" pageId="9" pageNumber="170">b</emphasis>
trees support placement of
<taxonomicName id="4C3814E6FF92FF919A84FC8F5F5B9648" authority="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen, 2017" authorityName="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen" authorityYear="2017" box="[676,773,812,835]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="species" species="swirei" status="sp. nov.">
<emphasis id="B94CB377FF92FF919A84FC8F5F5B9648" box="[676,773,812,835]" italics="true" pageId="9" pageNumber="170">P. swirei</emphasis>
</taxonomicName>
<emphasis id="B94CB377FF92FF919B2DFC885F36964F" bold="true" box="[781,872,811,836]" pageId="9" pageNumber="170">
<taxonomicNameLabel id="A27F0E0CFF92FF919B2DFC885F36964F" box="[781,872,811,836]" pageId="9" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
as most closely related to
<taxonomicName id="4C3814E6FF92FF919CBAFC8F594A9648" baseAuthorityName="Andriashev &amp; Pitruk" baseAuthorityYear="1993" box="[1178,1300,811,835]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="species" species="belyaevi">
<emphasis id="B94CB377FF92FF919CBAFC8F594A9648" box="[1178,1300,811,835]" italics="true" pageId="9" pageNumber="170">P. belyaevi</emphasis>
</taxonomicName>
(&gt;74% ML bootstrap support,&gt;0.96 Bayesian posterior probability), with highly concordant topology. Genetic distances between the two species of
<taxonomicName id="4C3814E6FF92FF9199ECFCD05E369680" baseAuthorityName="Andriashev" baseAuthorityYear="1955" box="[460,616,883,907]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="genus">
<emphasis id="B94CB377FF92FF9199ECFCD05E369680" box="[460,616,883,907]" italics="true" pageId="9" pageNumber="170">Pseudoliparis</emphasis>
</taxonomicName>
are 0.6% at Cyt-
<emphasis id="B94CB377FF92FF919B07FCD05F6B9680" box="[807,821,883,907]" italics="true" pageId="9" pageNumber="170">b</emphasis>
and 0.7% at 16S rRNA (
<tableCitation id="C6BA5ADEFF92FF919C6EFCD058FB9680" box="[1102,1189,883,908]" captionStart="TABLE 4" captionStartId="12.[119,198,512,533]" captionText="TABLE 4. Evolutionary đivergence between Pseudoliparis swirei sp. nov. anđ close relatives. Average genetic đistance among species calculateđ for (A) cytochrome c oxiđase subunit I (644 bp), (B) cytochrome b (744 bp), anđ (C) 16S rRNA (699 bp), baseđ on the Tamura-Nei substitution mođel with gamma-đistributeđ rate variation across sites. Sequences from P.swirei sp.nov., N.kermadecensis,N.stewarti đerive fromthis stuđy, P.belyaevi are unpublisheđ đata fromT. Satoh (National Museum of Nature anđ Science, Japan), sequences from C. rastrinus, C. cypselurus, A. ventricosus were from NCBI (GenBank Accession Numbers: AB565515.1, AB565517.1, AB565629.1, FJ164433.1," pageId="9" pageNumber="170">Table 4</tableCitation>
), with distances of 1 1.4% (Cyt-
<emphasis id="B94CB377FF92FF919934FC345D7C96A4" box="[276,290,919,943]" italics="true" pageId="9" pageNumber="170">b</emphasis>
) and 0.81% (16S rRNA) to the
<taxonomicName id="4C3814E6FF92FF919ABEFC345F4096A4" baseAuthorityName="Andriashev" baseAuthorityYear="1975" box="[670,798,919,943]" class="Actinopterygii" family="Liparidae" genus="Notoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="genus">
<emphasis id="B94CB377FF92FF919ABEFC345F4096A4" box="[670,798,919,943]" italics="true" pageId="9" pageNumber="170">Notoliparis</emphasis>
</taxonomicName>
species. COI lacked sufficient polymorphism to resolve the relevant nodes, with low bootstrap support observed for the placement of
<taxonomicName id="4C3814E6FF92FF919C0FFC1F58D196D8" authority="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen, 2017" authorityName="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen" authorityYear="2017" box="[1071,1167,956,979]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="species" species="swirei" status="sp. nov.">
<emphasis id="B94CB377FF92FF919C0FFC1F58D196D8" box="[1071,1167,956,979]" italics="true" pageId="9" pageNumber="170">P. swirei</emphasis>
</taxonomicName>
<emphasis id="B94CB377FF92FF919CBEFC1858A296DF" bold="true" box="[1182,1276,955,980]" pageId="9" pageNumber="170">
<taxonomicNameLabel id="A27F0E0CFF92FF919CBEFC1858A296DF" box="[1182,1276,955,980]" pageId="9" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
relative to P.
<taxonomicName id="4C3814E6FF92FF9198B7FC7C5CAA96FC" baseAuthorityName="Andriashev &amp; Pitruk" baseAuthorityYear="1993" box="[151,244,991,1015]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="species" species="belyaevi">
<emphasis id="B94CB377FF92FF9198B7FC7C5CAA96FC" box="[151,244,991,1015]" italics="true" pageId="9" pageNumber="170">belyaevi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3814E6FF92FF91990AFC435DB196FC" baseAuthorityName="Nielsen" baseAuthorityYear="1964" box="[298,495,991,1015]" class="Actinopterygii" family="Liparidae" genus="Notoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="species" species="kermadecensis">
<emphasis id="B94CB377FF92FF91990AFC435DB196FC" box="[298,495,991,1015]" italics="true" pageId="9" pageNumber="170">N. kermadecensis</emphasis>
</taxonomicName>
(
<figureCitation id="130373E0FF92FF9199DEFC7C5E0196F3" box="[510,607,991,1016]" captionStart="FIGURE 6" captionStartId="10.[151,250,1073,1095]" captionTargetBox="[352,1239,212,1026]" captionTargetId="figure@10.[340,1247,193,1052]" captionTargetPageId="10" captionText="FIGURE 6. Phylogenetic relationships of Pseudoliparis swirei sp. nov. to closely related hadal liparids, P. belyaevi from the Japan Trench and Notoliparis kermadecensis and Notoliparis stewarti from the Kermadec Trench. Maximum likelihood (ML) trees based on A) cytochrome c oxidase subunit I (1399 bp), B) cytochrome b (1007 bp), and C) 16S rRNA (1472 bp). ML bootstrap support (&gt;70%) and Bayesianposterior probability (&gt;0.70) valuesareshown as ML/BI.Bold values indicate thenode that supports placement of P. swirei sp. nov. within the genus Pseudoliparis. Aptocyclus ventricosus (GenBank accession NC008129.1) was used to root each tree." httpUri="https://zenodo.org/record/1067566/files/figure.png" pageId="9" pageNumber="170">Figure 6</figureCitation>
). Genetic distances among species also were the lowest at COI, at &lt;1% or three nucleotide substitutions observed between the sequences of
<taxonomicName id="4C3814E6FF92FF919B5DFBA058479110" baseAuthorityName="Andriashev" baseAuthorityYear="1955" box="[893,1049,1027,1051]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="genus">
<emphasis id="B94CB377FF92FF919B5DFBA058479110" box="[893,1049,1027,1051]" italics="true" pageId="9" pageNumber="170">Pseudoliparis</emphasis>
</taxonomicName>
. Phylogenetic analyses conducted using the more closely-related
<taxonomicName id="4C3814E6FF92FF9199CFFB8B5E279134" box="[495,633,1064,1087]" class="Actinopterygii" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="genus">
<emphasis id="B94CB377FF92FF9199CFFB8B5E279134" box="[495,633,1064,1087]" italics="true" pageId="9" pageNumber="170">Careproctus</emphasis>
</taxonomicName>
outgroup taxa, but with a shorter alignment (696-bp 16S rRNA, 744-bp Cyt-
<emphasis id="B94CB377FF92FF9198E9FBE85C899168" box="[201,215,1099,1123]" italics="true" pageId="9" pageNumber="170">b</emphasis>
, 645-bp COI) found congruent results to the trees presented in
<figureCitation id="130373E0FF92FF919BE1FBE8587D916F" box="[961,1059,1099,1124]" captionStart="FIGURE 6" captionStartId="10.[151,250,1073,1095]" captionTargetBox="[352,1239,212,1026]" captionTargetId="figure@10.[340,1247,193,1052]" captionTargetPageId="10" captionText="FIGURE 6. Phylogenetic relationships of Pseudoliparis swirei sp. nov. to closely related hadal liparids, P. belyaevi from the Japan Trench and Notoliparis kermadecensis and Notoliparis stewarti from the Kermadec Trench. Maximum likelihood (ML) trees based on A) cytochrome c oxidase subunit I (1399 bp), B) cytochrome b (1007 bp), and C) 16S rRNA (1472 bp). ML bootstrap support (&gt;70%) and Bayesianposterior probability (&gt;0.70) valuesareshown as ML/BI.Bold values indicate thenode that supports placement of P. swirei sp. nov. within the genus Pseudoliparis. Aptocyclus ventricosus (GenBank accession NC008129.1) was used to root each tree." httpUri="https://zenodo.org/record/1067566/files/figure.png" pageId="9" pageNumber="170">Figure 6</figureCitation>
(
<taxonomicName id="4C3814E6FF92FF919C17FBE858EC9168" box="[1079,1202,1099,1123]" class="Actinopterygii" family="Cyclopteridae" genus="Aptocyclus" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="genus">
<emphasis id="B94CB377FF92FF919C17FBE858EC9168" box="[1079,1202,1099,1123]" italics="true" pageId="9" pageNumber="170">Aptocyclus</emphasis>
</taxonomicName>
outgroup), but with lower bootstrap support for most nodes (Supplementary
<figureCitation id="130373E0FF92FF919B30FBCC5F2F918C" box="[784,881,1135,1160]" captionStart="FIGURE 2" captionStartId="4.[151,250,1453,1475]" captionTargetBox="[474,1113,373,1432]" captionTargetId="figure@4.[474,1113,373,1432]" captionTargetPageId="4" captionText="FIGURE 2. A) In situ photograph of Pseudoliparis swirei sp. nov. at 6,198 m. B) a group at 7,485 m. C) Deck photograph of SIO 16-82/HADES 200049.D) Radiograph of SIO 16-86/HADES 200141. Image by Sandra Raredon. Scale indicator 5 cm." httpUri="https://zenodo.org/record/1067558/files/figure.png" pageId="9" pageNumber="170">Figure 2</figureCitation>
). The 16S rRNA phylogeny supported placement of the new species into
<taxonomicName id="4C3814E6FF92FF91999DFB305E0791A0" baseAuthorityName="Andriashev" baseAuthorityYear="1955" box="[445,601,1171,1195]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="genus">
<emphasis id="B94CB377FF92FF91999DFB305E0791A0" box="[445,601,1171,1195]" italics="true" pageId="9" pageNumber="170">Pseudoliparis</emphasis>
</taxonomicName>
(67% ML bootstrap support/0.75 Bayesian posterior probability), and moderate support was observed at Cyt-
<emphasis id="B94CB377FF92FF919A71FB145E0191C4" box="[593,607,1207,1231]" italics="true" pageId="9" pageNumber="170">b</emphasis>
(0.82 Bayesian posterior probability, ML unresolved). There was still no significant resolution of generic placement in the COI phylogeny with the
<taxonomicName id="4C3814E6FF92FF919C16FB78589E91F9" box="[1078,1216,1243,1266]" class="Actinopterygii" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="genus">
<emphasis id="B94CB377FF92FF919C16FB78589E91F9" box="[1078,1216,1243,1266]" italics="true" pageId="9" pageNumber="170">Careproctus</emphasis>
</taxonomicName>
outgroup species. Sequences from
<taxonomicName id="4C3814E6FF92FF919979FB5C5E1D901C" authority="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen, 2017" authorityName="Gerringer &amp; Linley &amp; Jamieson &amp; Goetze &amp; Drazen" authorityYear="2017" box="[345,579,1279,1303]" class="Actinopterygii" family="Liparidae" genus="Pseudoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="species" species="swirei" status="sp. nov.">
<emphasis id="B94CB377FF92FF919979FB5C5E1D901C" box="[345,579,1279,1303]" italics="true" pageId="9" pageNumber="170">Pseudoliparis swirei</emphasis>
</taxonomicName>
<emphasis id="B94CB377FF92FF919A6DFB5C5EF49013" bold="true" box="[589,682,1279,1304]" pageId="9" pageNumber="170">
<taxonomicNameLabel id="A27F0E0CFF92FF919A6DFB5C5EF49013" box="[589,682,1279,1304]" pageId="9" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B94CB377FF92FF919A9CFB5C58BF901C" box="[700,1249,1279,1303]" italics="true" pageId="9" pageNumber="170">
<taxonomicName id="4C3814E6FF92FF919A9CFB5C5FB1901C" baseAuthorityName="Nielsen" baseAuthorityYear="1964" box="[700,1007,1279,1303]" class="Actinopterygii" family="Liparidae" genus="Notoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="species" species="kermadecensis">Notoliparis kermadecensis</taxonomicName>
,
<taxonomicName id="4C3814E6FF92FF919BDDFB5C58BF901C" baseAuthorityName="Stein" baseAuthorityYear="2016" box="[1021,1249,1279,1303]" class="Actinopterygii" family="Liparidae" genus="Notoliparis" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="170" phylum="Chordata" rank="species" species="stewarti">Notoliparis stewarti</taxonomicName>
</emphasis>
(this study) are available under GenBank accession numbers
<accessionNumber id="946BF286FF92FF919AB4FA805F4E9037" box="[660,784,1315,1340]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY659176" pageId="9" pageNumber="170">KY659176</accessionNumber>
<accessionNumber id="946BF286FF92FF919B3FFA805FC59037" box="[799,923,1315,1340]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY659204" pageId="9" pageNumber="170">KY659204</accessionNumber>
(Supplementary
<tableCitation id="C6BA5ADEFF92FF919C7CFA8058EF9030" box="[1116,1201,1315,1340]" captionStart="TABLE 2" captionStartId="7.[151,239,152,175]" captionText="TABLE 2. Measurements and counts of Pseudoliparis swirei sp. nov.. Measurements taken from preserved specimens, given in mm. Body depth measured from fresh photographs. Dorsal fin origin between vertebrae #, # are shown. Maximum and minimum values (mean and standard deviation) shown for measurements. Counts show range (median). Number of specimens examined for individual characters (n)." pageId="9" pageNumber="170">Table 2</tableCitation>
).
</paragraph>
<paragraph id="8B876F65FF92FF9198E7FAE4581C9333" blockId="9.[151,1437,151,1880]" pageId="9" pageNumber="170">
<emphasis id="B94CB377FF92FF9198E7FAE45D2C906B" bold="true" box="[199,370,1351,1376]" pageId="9" pageNumber="170">Reproduction.</emphasis>
<typeStatus id="5483D1C7FF92FF919959FAE45DBF906B" box="[377,481,1351,1376]" pageId="9" pageNumber="170" type="holotype">Holotype</typeStatus>
is immature. Ripe females had eggs up to
<quantity id="4CC0C280FF92FF919B9CFAE4584F9054" box="[956,1041,1351,1375]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="9" pageNumber="170" unit="mm" value="9.4">9.4 mm</quantity>
diameter, among the largest teleost eggs recorded (
<bibRefCitation id="EFA91294FF92FF919966FAC85E37908F" author="Tyler" box="[326,617,1387,1412]" pageId="9" pageNumber="170" refString="Tyler, C. R. &amp; Sumpter, J. P. (1996) Oocyte growth and development in teleosts. Reviews in Fish Biology and Fisheries, 6 (3), 287 - 318. https: // doi. org / 10.1007 / BF 00122584" type="journal article" year="1996">Tyler and Sumpter, 1996</bibRefCitation>
),
<quantity id="4CC0C280FF92FF919A5EFAC85E899088" box="[638,727,1387,1411]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="170" unit="mm" value="0.4">0.4 mm</quantity>
smaller than the largest record (
<bibRefCitation id="EFA91294FF92FF919C6DFAC859049088" author="Matallanas" box="[1101,1370,1387,1412]" pageId="9" pageNumber="170" refString="Matallanas, J., Rucabado, J., Lloris, D. &amp; Olivar, M. P. (1990) Early stages of development and reproductive biology of the South-American eelpout Austrolycus depressiceps Regan, 1913 (Teleostei: Zoarcidae). Scientia Marina, 54 (3), 257 - 261." type="journal article" year="1990">
Matallanas
<emphasis id="B94CB377FF92FF919CF4FACF59509088" box="[1236,1294,1387,1411]" italics="true" pageId="9" pageNumber="170">et al.</emphasis>
, 1990
</bibRefCitation>
). The eggs were unsorted within gonad, with the largest eggs free and interspersed within a matrix of smaller eggs. No developmental structures were visible within even the largest eggs. Two distinct size classes of eggs present with up to 23 large eggs (&gt;
<quantity id="4CC0C280FF92FF9199AFFA745D8F90E4" box="[399,465,1495,1520]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="170" unit="mm" value="5.0">5 mm</quantity>
) and up to 851 small eggs of less than half the diameter of the larger size class. There were rarely intermediate stages (
<figureCitation id="130373E0FF92FF919A24FA585E389318" box="[516,614,1531,1556]" captionStart="FIGURE 7" captionStartId="10.[151,250,1944,1966]" captionTargetBox="[180,1395,1327,1907]" captionTargetId="figure@10.[165,1409,1298,1920]" captionTargetPageId="10" captionText="FIGURE 7. Egg sizes and frequencies from four individuals. Binned into 0.5 mm increments.Small peaks show 22, 23, 7, 14 large eggs for specimens USNM 438994/HADES 200027, SIO 16-81/HADES 200039, SIO 16-87/HADES 200025, USNM 438995/HADES 200085." httpUri="https://zenodo.org/record/1067568/files/figure.png" pageId="9" pageNumber="170">Figure 7</figureCitation>
). Individuals with only small eggs had maximum egg sizes ranging from
<quantity id="4CC0C280FF92FF9198B7F9BC5D69933C" box="[151,311,1567,1591]" metricMagnitude="-3" metricUnit="m" metricValue="1.0499999999999998" metricValueMax="1.4" metricValueMin="0.7" pageId="9" pageNumber="170" unit="mm" value="1.0499999999999998" valueMax="1.4" valueMin="0.7">0.7 to 1.4 mm</quantity>
. Genital papilla visible in freshly collected males, oriented anteriorly.
</paragraph>
</subSubSection>
<subSubSection id="C3223CEEFF92FF9198E7F9E05C8893AF" pageId="9" pageNumber="170" type="distribution">
<paragraph id="8B876F65FF92FF9198E7F9E05C8893AF" blockId="9.[151,1437,151,1880]" pageId="9" pageNumber="170">
<emphasis id="B94CB377FF92FF9198E7F9E05D3E9357" bold="true" box="[199,352,1603,1628]" pageId="9" pageNumber="170">Distribution.</emphasis>
Known only from the Mariana Trench at capture depths from
<quantity id="4CC0C280FF92FF919C01F9E058929350" box="[1057,1228,1603,1628]" metricMagnitude="3" metricUnit="m" metricValue="7.432" metricValueMax="7.965999999999999" metricValueMin="6.897999999999999" pageId="9" pageNumber="170" unit="m" value="7432.0" valueMax="7966.0" valueMin="6898.0">6,8987,966 m</quantity>
, individuals likely this species were recognized in video at depths
<quantity id="4CC0C280FF92FF919A87F9C45F0E9374" box="[679,848,1639,1664]" metricMagnitude="3" metricUnit="m" metricValue="7.147999999999999" metricValueMax="8.097999999999999" metricValueMin="6.1979999999999995" pageId="9" pageNumber="170" unit="m" value="7148.0" valueMax="8098.0" valueMin="6198.0">6,1988,098 m</quantity>
(
<bibRefCitation id="EFA91294FF92FF919B7EF9C45879938B" author="Linley" box="[862,1063,1639,1664]" pageId="9" pageNumber="170" refString="Linley, T. D., Gerringer, M. E., Yancey, P. H., Drazen, J. C., Weinstock, C. L. &amp; Jamieson, A. J. (2016) Fishes of the hadal zone including new species, in situ observations and depth records of Liparidae. Deep Sea Research Part I: Oceanographic Research Papers, 114, 99 - 110. https: // doi. org / 10.1016 / j. dsr. 2016.05.003" type="journal article" year="2016">
Linley
<emphasis id="B94CB377FF92FF919B8FF9CB5FB99374" box="[943,999,1639,1663]" italics="true" pageId="9" pageNumber="170">et al.</emphasis>
2016
</bibRefCitation>
; Jamieson &amp; Linley, unpublished data).
</paragraph>
</subSubSection>
<subSubSection id="C3223CEEFF92FF9198E7F90C5F959253" pageId="9" pageNumber="170" type="etymology">
<paragraph id="8B876F65FF92FF9198E7F90C5F959253" blockId="9.[151,1437,151,1880]" pageId="9" pageNumber="170">
<emphasis id="B94CB377FF92FF9198E7F90C5D1093C3" bold="true" box="[199,334,1711,1736]" pageId="9" pageNumber="170">Etymology.</emphasis>
The Mariana Trench famously houses the oceans deepest point, at Challenger Deep, named for the
<emphasis id="B94CB377FF92FF9198E0F9705D2193E0" box="[192,383,1747,1771]" italics="true" pageId="9" pageNumber="170">HMS Challenger</emphasis>
expedition which discovered the trench in 1875. Their deepest sounding of
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, then the greatest known ocean depth, was christened Swire Deep after Herbert Swire, the ships First Navigating Sublieutenant (
<bibRefCitation id="EFA91294FF92FF919964F8B85DB29238" author="Corfield" box="[324,492,1819,1844]" pageId="9" pageNumber="170" refString="Corfield, R. (2003) The Silent Landscape the Scientific Voyage of HMS Challenger. Washington, D. C.: Joseph Henry Press, Washington, D. C., 285 pp." type="book" year="2003">Corfield 2003</bibRefCitation>
). We name this fish in his honor, in acknowledgment and gratitude of the crew members that have supported oceanographic research throughout history.
</paragraph>
</subSubSection>
</treatment>
</document>