treatments-xml/data/E8/51/1E/E8511E53FFC5EF656A8AFC74FDABF8D4.xml
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<document id="690F94FDBF6927757B60CF5818F55101" ID-DOI="10.5281/zenodo.6391684" ID-ISSN="1942-1354" ID-Zenodo-Dep="6391684" ID-ZooBank="CBD29188-143B-44DF-BE21-1654D50D8621" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648501406612" checkinUser="felipe" docAuthor="Phillips, Robert A." docDate="2022" docId="E8511E53FFC5EF656A8AFC74FDABF8D4" docLanguage="en" docName="InsectaMundi.2022.0907.1-214.pdf" docOrigin="Insecta Mundi 2022 (907)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Leptoconops (Holoconops) reesi Clastrier and Wirth" docType="treatment" docVersion="3" lastPageNumber="24" masterDocId="1468662BFFDCEF7C6A4AFFC3FFCCFFE4" masterDocTitle="Culicoides Latreille and Leptoconops Skuse biting midges of the southwestern United States with emphasis on the Canyonlands of southeastern Utah (Diptera: Ceratopogonidae)" masterLastPageNumber="214" masterPageNumber="1" pageNumber="24" updateTime="1699308763882" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="9E73D38BD4F99AEDD50DE5BA3EFE7CB4">Culicoides Latreille and Leptoconops Skuse biting midges of the southwestern United States with emphasis on the Canyonlands of southeastern Utah (Diptera: Ceratopogonidae)</mods:title>
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<mods:name id="8B2A291BEB13FB58A80E3524BD0B53F3" type="personal">
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<mods:namePart id="A02ABD5042F0A2612CC39ED3D1D463A3">Phillips, Robert A.</mods:namePart>
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<mods:date id="DA3FDB2B02D6ABE71F2FA45673463227">2022</mods:date>
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<treatment id="E8511E53FFC5EF656A8AFC74FDABF8D4" ID-DOI="http://doi.org/10.5281/zenodo.6391738" ID-Zenodo-Dep="6391738" LSID="urn:lsid:plazi:treatment:E8511E53FFC5EF656A8AFC74FDABF8D4" httpUri="http://treatment.plazi.org/id/E8511E53FFC5EF656A8AFC74FDABF8D4" lastPageNumber="24" pageId="25" pageNumber="24">
<subSubSection id="28E2FCCEFFC5EF656A8AFC74FC9DFC30" box="[192,849,951,980]" pageId="25" pageNumber="24" type="nomenclature">
<paragraph id="6047AF45FFC5EF656A8AFC74FC9DFC30" blockId="25.[192,849,951,980]" box="[192,849,951,980]" pageId="25" pageNumber="24">
<heading id="3B0F1829FFC5EF656A8AFC74FC9DFC30" box="[192,849,951,980]" fontSize="12" level="2" pageId="25" pageNumber="24" reason="2">
<taxonomicName id="A7F8D4C6FFC5EF656A8AFC74FC9DFC30" ID-CoL="8CHV6" authority="Clastrier and Wirth" authorityName="Clastrier and Wirth" box="[192,849,951,980]" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="24" phylum="Arthropoda" rank="species" species="reesi" subGenus="Holoconops">
<emphasis id="528C7357FFC5EF656A8AFC74FC9DFC30" bold="true" box="[192,849,951,980]" pageId="25" pageNumber="24">
<emphasis id="528C7357FFC5EF656A8AFC74FE91FC37" bold="true" box="[192,349,951,979]" italics="true" pageId="25" pageNumber="24">Leptoconops</emphasis>
(
<emphasis id="528C7357FFC5EF656B24FC74FDCCFC30" bold="true" box="[366,512,951,980]" italics="true" pageId="25" pageNumber="24">Holoconops</emphasis>
)
<emphasis id="528C7357FFC5EF65685BFC74FD87FC37" bold="true" box="[529,587,951,979]" italics="true" pageId="25" pageNumber="24">reesi</emphasis>
Clastrier and Wirth
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="28E2FCCEFFC5EF656AAAFC21FB8EFB75" pageId="25" pageNumber="24" type="reference_group">
<paragraph id="6047AF45FFC5EF656AAAFC21FE5CFBF3" blockId="25.[224,1442,993,1169]" pageId="25" pageNumber="24">
<treatmentCitationGroup id="40E8886BFFC5EF656AAAFC21FE5CFBF3" pageId="25" pageNumber="24">
<treatmentCitation id="E1598954FFC5EF656AAAFC21FC9BFC1D" author="Clastrier J &amp; Wirth WW" box="[224,855,993,1017]" page="42" pageId="25" pageNumber="24" year="1978">
<taxonomicName id="A7F8D4C6FFC5EF656AAAFC21FC9BFC1D" ID-CoL="8CHV6" authority="Clastrier and Wirth, 1978: 42" authorityName="Clastrier and Wirth" authorityPageNumber="42" authorityYear="1978" box="[224,855,993,1017]" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="24" phylum="Arthropoda" rank="species" species="reesi" subGenus="Holoconops">
<emphasis id="528C7357FFC5EF656AAAFC21FE91FC1D" box="[224,349,994,1017]" italics="true" pageId="25" pageNumber="24">Leptoconops</emphasis>
(
<emphasis id="528C7357FFC5EF656B26FC22FE2DFC1D" box="[364,481,993,1017]" italics="true" pageId="25" pageNumber="24">Holoconops</emphasis>
)
<emphasis id="528C7357FFC5EF656BBBFC21FDD3FC1D" box="[497,543,994,1017]" italics="true" pageId="25" pageNumber="24">reesi</emphasis>
<bibRefCitation id="0469D2B4FFC5EF65686CFC22FC9BFC1D" author="Clastrier J &amp; Wirth WW" box="[550,855,993,1017]" pageId="25" pageNumber="24" pagination="1 - 58" refId="ref128879" refString="Clastrier J, Wirth WW. 1978. The Leptoconops kerteszi complex in North America (Diptera: Ceratopogonidae). United States Department of Agriculture Technical Bulletin 1573: 1 - 58." type="journal article" year="1978">Clastrier and Wirth, 1978: 42</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(key; female, male; fig. female antenna, palpus, spermathecae;
<collectingRegion id="A23C61A7FFC5EF656B04FC3CFE49FBF3" box="[334,389,1023,1047]" country="United States of America" name="Utah" pageId="25" pageNumber="24">Utah</collectingRegion>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="6047AF45FFC5EF656AAAFBDCFC2CFBD2" blockId="25.[224,1442,993,1169]" box="[224,992,1054,1078]" pageId="25" pageNumber="24">
<taxonomicName id="A7F8D4C6FFC5EF656AAAFBDCFE7DFBD2" ID-CoL="8CHTL" authorityName="Kieffer" authorityYear="1908" box="[224,433,1054,1078]" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="24" phylum="Arthropoda" rank="species" species="kerteszi">
<emphasis id="528C7357FFC5EF656AAAFBDCFE7DFBD2" box="[224,433,1054,1078]" italics="true" pageId="25" pageNumber="24">Leptoconops kerteszi</emphasis>
</taxonomicName>
, misidentified:
<bibRefCitation id="0469D2B4FFC5EF65681AFBDDFCEAFBD2" author="Rees DM &amp; Smith JV" box="[592,806,1054,1078]" pageId="25" pageNumber="24" pagination="9 - 15" refId="ref138691" refString="Rees DM, Smith JV. 1950. Effective control methods used on biting gnats in Utah during 1949 (Diptera: Ceratopogonidae). Mosquito News 10: 9 - 15." type="journal article" year="1950">
Rees and
<collectingRegion id="A23C61A7FFC5EF6568F8FBDDFD3CFBD2" box="[690,752,1054,1078]" country="Bermuda" name="Smith's" pageId="25" pageNumber="24">Smith</collectingRegion>
1950
</bibRefCitation>
(in part; biology).
</paragraph>
<paragraph id="6047AF45FFC5EF656AAAFBFEFB8EFB75" blockId="25.[224,1442,993,1169]" pageId="25" pageNumber="24">
<taxonomicName id="A7F8D4C6FFC5EF656AAAFBFEFDF3FBB0" ID-CoL="8CHTL" authorityName="Kieffer" authorityYear="1908" box="[224,575,1084,1108]" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="24" phylum="Arthropoda" rank="species" species="kerteszi" subGenus="Holoconops">
<emphasis id="528C7357FFC5EF656AAAFBFEFE91FBB0" box="[224,349,1085,1108]" italics="true" pageId="25" pageNumber="24">Leptoconops</emphasis>
(
<emphasis id="528C7357FFC5EF656B27FBFFFE2EFBB0" box="[365,482,1084,1108]" italics="true" pageId="25" pageNumber="24">Holoconops</emphasis>
)
<emphasis id="528C7357FFC5EF656BBBFBFFFDF3FBB0" box="[497,575,1084,1108]" italics="true" pageId="25" pageNumber="24">kerteszi</emphasis>
</taxonomicName>
, misidentified:
<bibRefCitation id="0469D2B4FFC5EF6568ABFBFFFC58FBB0" author="Wirth WW" box="[737,916,1084,1108]" pageId="25" pageNumber="24" pagination="95 - 266" refId="ref142682" refString="Wirth WW. 1952 a. The Heleidae of California. University of California Publications in Entomology 9: 95 - 266." type="journal article" year="1952">Wirth 1952a: 113</bibRefCitation>
(in part; key; female; male genitalia; biology).
<bibRefCitation id="0469D2B4FFC5EF656F30FBFEFEAEFB97" author="Fox I." pageId="25" pageNumber="24" pagination="214 - 285" refId="ref130377" refString="Fox I. 1955. A catalogue of the bloodsucking midges of the Americas (Culicoides, Leptoconops and Lasiohelea) with keys to the subgenera and Nearctic species, a geographic index, and bibliography. Journal of Agriculture of University of Puerto Rico 39: 214 - 285." type="journal article" year="1955">Fox 1955: 263</bibRefCitation>
(key; taxonomy).
<bibRefCitation id="0469D2B4FFC5EF656851FB98FCF4FB97" author="Wirth WW &amp; Atchley WR" box="[539,824,1115,1139]" pageId="25" pageNumber="24" pagination="1 - 57" refId="ref142906" refString="Wirth WW, Atchley WR. 1973. A review of the North American Leptoconops (Diptera: Ceratopogonidae). Graduate Studies Texas Tech University 5: 1 - 57." type="journal article" year="1973">Wirth and Atchley 1973: 45</bibRefCitation>
(in part; key; female, male; fig. female wing, head, genitalia, spermathecae, hind tibial comb, fore tarsomeres 1 and 2, male genitalia; biology).
</paragraph>
</subSubSection>
<subSubSection id="28E2FCCEFFC5EF656A8AFB62FE0BF976" pageId="25" pageNumber="24" type="diagnosis">
<paragraph id="6047AF45FFC5EF656A8AFB62FE0BF976" blockId="25.[191,1442,1185,1682]" pageId="25" pageNumber="24">
<emphasis id="528C7357FFC5EF656A8AFB62FEF3FB5F" bold="true" box="[192,319,1185,1211]" pageId="25" pageNumber="24">Diagnosis.</emphasis>
(
<tableCitation id="2D7A9AFEFFC5EF656B07FB62FE63FB5F" box="[333,431,1185,1211]" captionStart="Table 13" captionStartId="150.[192,255,781,809]" captionText="Table 13.Quantitative characters of Leptoconops. Data in () are variations. Measurements and ratios are expressed as means. T9alp = male tergite9 apicolateral processes. f13/f12 = length of flagellomere13 divided by the length of flagellomere12. Wing, proboscis, teeth, setae, and spine data are compiled from Clastrier and Wirth (1978) for the Kerteszi group species and Wirth and Atchley (1973) for the others. Species marked (r) were not examined." pageId="25" pageNumber="24">Table 13</tableCitation>
) Body, including head capsule, light yellowish brown, femora and basal portion of tibiae brown, apical portion yellowish, all tarsomeres 1, usually 2, yellowish; median pair of distal clypeal setae ≥0.8 as far apart from each other as from corresponding lateral setae (as in
<figureCitation id="F8C3B3C0FFC5EF6569ADFB27FB85FB1A" box="[999,1097,1252,1278]" captionStart="Figures 915. 9" captionStartId="164.[192,279,1765,1791]" captionTargetBox="[196,1431,251,1723]" captionTargetId="figure-103@164.[192,1440,251,1730]" captionTargetPageId="164" captionText="Figures 915. 9) Leptoconops americanus female clypeus, setae (s). 10) L. knowltoni female clypeus. 11) L. knowltoni female palpal segment 3. 12) L. carteri female palpal segment 3 (Yolo County, CA). 13) L. werneri female tarsomeres 45 and claw (paratype, Imperial County, CA [UCRC]). 14) L. werneri male genitalia, ventral view, tergite 9 apicolateral processes (T9alp), meso-posterior lobe setae (s) (Riverside County, CA [UCRC]). 15) L. belkini male genitalia, ventral view, tergite 9 sclerotized lobe (L) (Riverside County, CA [UCRC])." figureDoi="http://doi.org/10.5281/zenodo.6391690" httpUri="https://zenodo.org/record/6391690/files/figure.png" pageId="25" pageNumber="24">
Fig.
<specimenCount id="76FE64CCFFC5EF656E52FB27FB85FB1A" box="[1048,1097,1252,1278]" pageId="25" pageNumber="24" type="generic">
10
<taxonomicName id="A7F8D4C6FFC5EF656E70FB26FB85FB1A" box="[1082,1097,1253,1278]" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="24" phylum="Arthropoda" rank="species" species="knowltoni">
<emphasis id="528C7357FFC5EF656E70FB26FB85FB1A" box="[1082,1097,1253,1278]" italics="true" pageId="25" pageNumber="24">L</emphasis>
</taxonomicName>
</specimenCount>
</figureCitation>
.
<emphasis id="528C7357FFC5EF656E1CFB27FB0BFB1A" box="[1110,1223,1252,1278]" italics="true" pageId="25" pageNumber="24">knowltoni</emphasis>
); palpal segment 3 sensory pit as deep as wide, broadening internally on female (as in
<figureCitation id="F8C3B3C0FFC5EF6569F7FAC5FBD2FAC4" box="[957,1054,1286,1312]" captionStart="Figures 2732" captionStartId="167.[192,279,1833,1859]" captionTargetBox="[266,1363,216,1768]" captionTargetId="figure-71@167.[192,1440,216,1770]" captionTargetPageId="167" captionText="Figures 2732. Female Leptoconops. 27) L. knowltoni flagellomeres 911, median black seta (s). Spermathecae (sp), diverticulum (d), neck (spn). 28) L. foulki. 29) L. americanus. 30) L. sublettei flagellomeres 36, flagellomere 4 seta bases (sb), hyaline sensory seta (hss). Palpal segment 3. 31) L. sublettei. 32) L. foulki." figureDoi="http://doi.org/10.5281/zenodo.6391698" httpUri="https://zenodo.org/record/6391698/files/figure.png" pageId="25" pageNumber="24">
Fig.
<specimenCount id="76FE64CCFFC5EF6569A4FAC5FBD2FAC4" box="[1006,1054,1286,1312]" pageId="25" pageNumber="24" type="generic">
32
<taxonomicName id="A7F8D4C6FFC5EF656E45FAC4FBD2FAC4" box="[1039,1054,1287,1312]" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="24" phylum="Arthropoda" rank="species" species="foulki">
<emphasis id="528C7357FFC5EF656E45FAC4FBD2FAC4" box="[1039,1054,1287,1312]" italics="true" pageId="25" pageNumber="24">L</emphasis>
</taxonomicName>
</specimenCount>
</figureCitation>
.
<emphasis id="528C7357FFC5EF656E66FAC5FBA1FAC4" box="[1068,1133,1286,1312]" italics="true" pageId="25" pageNumber="24">foulki</emphasis>
); mid tarsomere 1 without submedian spine. Female: stigma triangular, pointed; clypeus with one to three smaller proximal setae in addition to the four distal setae, distomedian pair out-of-line distad of lateral setae by ~0.7 their distance apart; antenna with 11 flagellomeres; flagellomere 11 without submedian black seta; flagellomere 4 dorsal hyaline sensory seta distal and medial to long black seta, out of axial alignment with corresponding sensory setae on flagellomeres 510 (as in
<figureCitation id="F8C3B3C0FFC5EF656805FA6DFD79FA2C" box="[591,693,1454,1480]" captionStart="Figures 2732" captionStartId="167.[192,279,1833,1859]" captionTargetBox="[266,1363,216,1768]" captionTargetId="figure-71@167.[192,1440,216,1770]" captionTargetPageId="167" captionText="Figures 2732. Female Leptoconops. 27) L. knowltoni flagellomeres 911, median black seta (s). Spermathecae (sp), diverticulum (d), neck (spn). 28) L. foulki. 29) L. americanus. 30) L. sublettei flagellomeres 36, flagellomere 4 seta bases (sb), hyaline sensory seta (hss). Palpal segment 3. 31) L. sublettei. 32) L. foulki." figureDoi="http://doi.org/10.5281/zenodo.6391698" httpUri="https://zenodo.org/record/6391698/files/figure.png" pageId="25" pageNumber="24">
Fig.
<specimenCount id="76FE64CCFFC5EF6568C8FA6DFD79FA2C" box="[642,693,1454,1480]" pageId="25" pageNumber="24" type="generic">
30
<taxonomicName id="A7F8D4C6FFC5EF6568EDFA6CFD79FA2C" box="[679,693,1455,1480]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="24" phylum="Arthropoda" rank="species" species="sublettei">
<emphasis id="528C7357FFC5EF6568EDFA6CFD79FA2C" box="[679,693,1455,1480]" italics="true" pageId="25" pageNumber="24">L</emphasis>
</taxonomicName>
</specimenCount>
</figureCitation>
.
<emphasis id="528C7357FFC5EF65688EFA6DFCE8FA2C" box="[708,804,1454,1480]" italics="true" pageId="25" pageNumber="24">sublettei</emphasis>
); hind tarsomere 3 ~1.7× longer than 5; spermathecae somewhat pyriform, with caplike diverticulum (as in
<figureCitation id="F8C3B3C0FFC5EF656972FA0CFC51FA0D" box="[824,925,1487,1513]" captionStart="Figures 2732" captionStartId="167.[192,279,1833,1859]" captionTargetBox="[266,1363,216,1768]" captionTargetId="figure-71@167.[192,1440,216,1770]" captionTargetPageId="167" captionText="Figures 2732. Female Leptoconops. 27) L. knowltoni flagellomeres 911, median black seta (s). Spermathecae (sp), diverticulum (d), neck (spn). 28) L. foulki. 29) L. americanus. 30) L. sublettei flagellomeres 36, flagellomere 4 seta bases (sb), hyaline sensory seta (hss). Palpal segment 3. 31) L. sublettei. 32) L. foulki." figureDoi="http://doi.org/10.5281/zenodo.6391698" httpUri="https://zenodo.org/record/6391698/files/figure.png" pageId="25" pageNumber="24">
Fig.
<specimenCount id="76FE64CCFFC5EF656921FA0CFC51FA0D" box="[875,925,1487,1513]" pageId="25" pageNumber="24" type="generic">
28
<taxonomicName id="A7F8D4C6FFC5EF6569C5FA13FC51FA0D" box="[911,925,1488,1513]" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="24" phylum="Arthropoda" rank="species" species="foulki">
<emphasis id="528C7357FFC5EF6569C5FA13FC51FA0D" box="[911,925,1488,1513]" italics="true" pageId="25" pageNumber="24">L</emphasis>
</taxonomicName>
</specimenCount>
</figureCitation>
.
<emphasis id="528C7357FFC5EF6569E6FA0CFC21FA0D" box="[940,1005,1487,1513]" italics="true" pageId="25" pageNumber="24">foulki</emphasis>
); cerci&gt;3× longer than wide (
<figureCitation id="F8C3B3C0FFC5EF656F1AFA0CFA5DFA0D" box="[1360,1425,1487,1513]" captionStart="Figures 12" captionStartId="162.[192,279,1866,1892]" captionTargetBox="[491,1142,213,1804]" captionTargetId="figure-38@162.[194,1442,213,1806]" captionTargetPageId="162" captionText="Figures 12. Female lateral habitus (in alcohol). 1) Leptoconops (Holoconops) Kerteszi group. 2) Culicoides (Monoculicoides) sonorensis." figureDoi="http://doi.org/10.5281/zenodo.6391686" httpUri="https://zenodo.org/record/6391686/files/figure.png" pageId="25" pageNumber="24">Fig. 1</figureCitation>
). Male: tergite 9 with distal shoulders abruptly narrowed to base of adjacent apicolateral processes, without dorsal process, ventro-posterior setae separated by ~2× as much as separation of apicolateral processes (as in
<figureCitation id="F8C3B3C0FFC5EF656A8AF9F7FEE8F9AA" box="[192,292,1588,1614]" captionStart="Figures 2026. 20" captionStartId="166.[192,279,1799,1825]" captionTargetBox="[266,1365,214,1767]" captionTargetId="figure-87@166.[194,1442,214,1769]" captionTargetPageId="166" captionText="Figures 2026. 20) Leptoconops knowltoni male genitalia, ventral view, aedeagus (A), parameres (P), tergite 9 ventro-posterior setae (s), apicolateral processes (T9alp), gonostylus apical lamelliform expansion (Gsle). 21) L. foulki male genitalia, ventral view. 22) L. knowltoni gonostylus, ventral setae (vs). 23 L. foulki gonostylus. 24) L. foulki parameres (P), proximal tooth (t). Male tarsomere 5, basal seta (bS). 25) L. knowltoni. 26) L. foulki." figureDoi="http://doi.org/10.5281/zenodo.6391696" httpUri="https://zenodo.org/record/6391696/files/figure.png" pageId="25" pageNumber="24">
Fig.
<specimenCount id="76FE64CCFFC5EF656AB8F9F7FEE8F9AA" box="[242,292,1588,1614]" pageId="25" pageNumber="24" type="generic">
20
<taxonomicName id="A7F8D4C6FFC5EF656B5FF9F6FEE8F9AA" box="[277,292,1589,1614]" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="24" phylum="Arthropoda" rank="species" species="knowltoni">
<emphasis id="528C7357FFC5EF656B5FF9F6FEE8F9AA" box="[277,292,1589,1614]" italics="true" pageId="25" pageNumber="24">L</emphasis>
</taxonomicName>
</specimenCount>
</figureCitation>
.
<emphasis id="528C7357FFC5EF656B7BF9F7FE6EF9AA" box="[305,418,1588,1614]" italics="true" pageId="25" pageNumber="24">knowltoni</emphasis>
); gonostylus with three ventral setae within 0.40.6 of gonostylus length, apical lamelliform expansion barely covering apical tooth (as in
<figureCitation id="F8C3B3C0FFC5EF65694AF995FC93F994" box="[768,863,1622,1648]" captionStart="Figures 2026. 20" captionStartId="166.[192,279,1799,1825]" captionTargetBox="[266,1365,214,1767]" captionTargetId="figure-87@166.[194,1442,214,1769]" captionTargetPageId="166" captionText="Figures 2026. 20) Leptoconops knowltoni male genitalia, ventral view, aedeagus (A), parameres (P), tergite 9 ventro-posterior setae (s), apicolateral processes (T9alp), gonostylus apical lamelliform expansion (Gsle). 21) L. foulki male genitalia, ventral view. 22) L. knowltoni gonostylus, ventral setae (vs). 23 L. foulki gonostylus. 24) L. foulki parameres (P), proximal tooth (t). Male tarsomere 5, basal seta (bS). 25) L. knowltoni. 26) L. foulki." figureDoi="http://doi.org/10.5281/zenodo.6391696" httpUri="https://zenodo.org/record/6391696/files/figure.png" pageId="25" pageNumber="24">
Fig.
<specimenCount id="76FE64CCFFC5EF65697AF995FC93F994" box="[816,863,1622,1648]" pageId="25" pageNumber="24" type="generic">
23
<taxonomicName id="A7F8D4C6FFC5EF65691AF994FC93F994" box="[848,863,1623,1648]" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="24" phylum="Arthropoda" rank="species" species="foulki">
<emphasis id="528C7357FFC5EF65691AF994FC93F994" box="[848,863,1623,1648]" italics="true" pageId="25" pageNumber="24">L</emphasis>
</taxonomicName>
</specimenCount>
</figureCitation>
.
<emphasis id="528C7357FFC5EF656926F995FC61F994" box="[876,941,1622,1648]" italics="true" pageId="25" pageNumber="24">foulki</emphasis>
); tarsomere 5 basal seta short, erect, curved (as in
<figureCitation id="F8C3B3C0FFC5EF656B4EF9B4FEA4F975" box="[260,360,1655,1681]" captionStart="Figures 2026. 20" captionStartId="166.[192,279,1799,1825]" captionTargetBox="[266,1365,214,1767]" captionTargetId="figure-87@166.[194,1442,214,1769]" captionTargetPageId="166" captionText="Figures 2026. 20) Leptoconops knowltoni male genitalia, ventral view, aedeagus (A), parameres (P), tergite 9 ventro-posterior setae (s), apicolateral processes (T9alp), gonostylus apical lamelliform expansion (Gsle). 21) L. foulki male genitalia, ventral view. 22) L. knowltoni gonostylus, ventral setae (vs). 23 L. foulki gonostylus. 24) L. foulki parameres (P), proximal tooth (t). Male tarsomere 5, basal seta (bS). 25) L. knowltoni. 26) L. foulki." figureDoi="http://doi.org/10.5281/zenodo.6391696" httpUri="https://zenodo.org/record/6391696/files/figure.png" pageId="25" pageNumber="24">
Fig.
<specimenCount id="76FE64CCFFC5EF656B7FF9B4FEA4F975" box="[309,360,1655,1681]" pageId="25" pageNumber="24" type="generic">
26
<taxonomicName id="A7F8D4C6FFC5EF656B13F9BBFEA4F975" box="[345,360,1656,1681]" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="24" phylum="Arthropoda" rank="species" species="foulki">
<emphasis id="528C7357FFC5EF656B13F9BBFEA4F975" box="[345,360,1656,1681]" italics="true" pageId="25" pageNumber="24">L</emphasis>
</taxonomicName>
</specimenCount>
</figureCitation>
.
<emphasis id="528C7357FFC5EF656B3CF9B4FE7BF975" box="[374,439,1655,1681]" italics="true" pageId="25" pageNumber="24">foulki</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="28E2FCCEFFC5EF656A8AF962FB88F8E2" pageId="25" pageNumber="24" type="distribution">
<paragraph id="6047AF45FFC5EF656A8AF962FC83F938" blockId="25.[192,1440,1697,1757]" pageId="25" pageNumber="24">
<emphasis id="528C7357FFC5EF656A8AF962FE90F95F" bold="true" box="[192,348,1697,1723]" pageId="25" pageNumber="24">Distribution.</emphasis>
<collectingRegion id="A23C61A7FFC5EF656B2EF962FDE0F95F" box="[356,556,1697,1723]" country="Canada" name="British Columbia" pageId="25" pageNumber="24">British Columbia</collectingRegion>
,
<collectingRegion id="A23C61A7FFC5EF656872F962FD14F95F" box="[568,728,1697,1723]" country="Canada" name="Saskatchewan" pageId="25" pageNumber="24">Saskatchewan</collectingRegion>
, south through
<collectingRegion id="A23C61A7FFC5EF6569D8F962FC1AF95F" box="[914,982,1697,1723]" country="United States of America" name="Idaho" pageId="25" pageNumber="24">Idaho</collectingRegion>
,
<collectingRegion id="A23C61A7FFC5EF6569A9F962FB9BF95F" box="[995,1111,1697,1723]" country="United States of America" name="Wyoming" pageId="25" pageNumber="24">Wyoming</collectingRegion>
,
<collectingRegion id="A23C61A7FFC5EF656E2EF962FB51F95F" box="[1124,1181,1697,1723]" country="United States of America" name="Utah" pageId="25" pageNumber="24">Utah</collectingRegion>
(Box Elder, Juab, Millard, Salt Lake, Toole counties),
<collectingRegion id="A23C61A7FFC5EF65686CF901FD5EF938" box="[550,658,1730,1756]" country="United States of America" name="Colorado" pageId="25" pageNumber="24">Colorado</collectingRegion>
, to
<collectingRegion id="A23C61A7FFC5EF6568F0F900FC87F938" box="[698,843,1730,1757]" country="United States of America" name="New Mexico" pageId="25" pageNumber="24">New Mexico</collectingRegion>
.
</paragraph>
<paragraph id="6047AF45FFC5EF656A8AF92FFB88F8E2" blockId="25.[192,1092,1772,1799]" box="[192,1092,1772,1799]" pageId="25" pageNumber="24">
<emphasis id="528C7357FFC5EF656A8AF92FFEBBF8E2" bold="true" box="[192,375,1772,1798]" pageId="25" pageNumber="24">Adult behavior.</emphasis>
Known hosts are human and sheep (
<bibRefCitation id="0469D2B4FFC5EF656952F92FFBF4F8E2" author="Clastrier J &amp; Wirth WW" box="[792,1080,1772,1798]" pageId="25" pageNumber="24" pagination="1 - 58" refId="ref128879" refString="Clastrier J, Wirth WW. 1978. The Leptoconops kerteszi complex in North America (Diptera: Ceratopogonidae). United States Department of Agriculture Technical Bulletin 1573: 1 - 58." type="journal article" year="1978">Clastrier and Wirth 1978</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="28E2FCCEFFC5EF656A8AF8D5FDABF8D4" box="[192,615,1814,1840]" pageId="25" pageNumber="24" type="discussion">
<paragraph id="6047AF45FFC5EF656A8AF8D5FDABF8D4" blockId="25.[192,615,1814,1840]" box="[192,615,1814,1840]" pageId="25" pageNumber="24">
<emphasis id="528C7357FFC5EF656A8AF8D5FEFCF8D4" bold="true" box="[192,304,1814,1840]" pageId="25" pageNumber="24">Remarks.</emphasis>
No
<taxonomicName id="A7F8D4C6FFC5EF656B15F8D4FE62F8D4" box="[351,430,1814,1840]" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="24" phylum="Arthropoda" rank="species" species="reesi">
<emphasis id="528C7357FFC5EF656B15F8D4FEA2F8D4" box="[351,366,1815,1840]" italics="true" pageId="25" pageNumber="24">L</emphasis>
.
<emphasis id="528C7357FFC5EF656B36F8D4FE62F8D4" box="[380,430,1815,1840]" italics="true" pageId="25" pageNumber="24">reesi</emphasis>
</taxonomicName>
were examined.
</paragraph>
</subSubSection>
</treatment>
</document>