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<document id="7EE276AB9A20B108F7473C4E21A70E83" ID-DOI="10.5281/zenodo.201850" ID-GBIF-Dataset="5f7c25af-fcfd-4abd-a391-2139fba0c57d" ID-ISSN="1175-5326" ID-Zenodo-Dep="201850" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460226495050" checkinUser="plazi" docAuthor="Lonsdale, Owen &amp; Marshall, Stephen A." docDate="2011" docId="AF6987A7452490171FF8FCF2FA32392A" docLanguage="en" docName="zt02748p017.pdf" docOrigin="Zootaxa 2748" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hendelia saopaulo Caloren &amp; Marshall 1998" docType="treatment" docVersion="9" lastPageNumber="16" masterDocId="5350FFDF452B90181F6FFFF6FFB03F22" masterDocTitle="Revision of the New World Hendelia (Diptera: Clusiidae: Clusiodinae)" masterLastPageNumber="17" masterPageNumber="1" pageNumber="16" updateTime="1698238470997" updateUser="plazi">
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<mods:title id="BDF90EB32EBD0F61A1C534D61063C87F">Revision of the New World Hendelia (Diptera: Clusiidae: Clusiodinae)</mods:title>
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<mods:namePart id="E8362D5A22530C0D3FB443B8AB112574">Lonsdale, Owen</mods:namePart>
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<mods:namePart id="57254E793C9FC7E79D2F587AB499A560">Marshall, Stephen A.</mods:namePart>
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<mods:date id="F038C7DD915FBC6F01368AF22F0819E8">2011</mods:date>
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<mods:number id="E7110C3FE4E30677E94EF0C2A436086A">2748</mods:number>
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<mods:identifier id="EDFBA00F6C62E504054ACB5EFE6D637F" type="DOI">10.5281/zenodo.201850</mods:identifier>
<mods:identifier id="B5E53DBB5EEA781D397B6EB25673D879" type="GBIF-Dataset">5f7c25af-fcfd-4abd-a391-2139fba0c57d</mods:identifier>
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<treatment id="AF6987A7452490171FF8FCF2FA32392A" ID-DOI="http://doi.org/10.5281/zenodo.6193494" ID-GBIF-Taxon="119386494" ID-Zenodo-Dep="6193494" LSID="urn:lsid:plazi:treatment:AF6987A7452490171FF8FCF2FA32392A" httpUri="http://treatment.plazi.org/id/AF6987A7452490171FF8FCF2FA32392A" lastPageNumber="16" pageId="15" pageNumber="16">
<subSubSection id="6FDA653A452490171FF8FCF2FE973C1D" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="277F36B1452490171FF8FCF2FD6B3C3F" blockId="15.[151,731,771,831]" box="[151,731,771,798]" pageId="15" pageNumber="16">
<heading id="7C3781DD452490171FF8FCF2FD6B3C3F" bold="true" box="[151,731,771,798]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<taxonomicName id="E0C04D32452490171FF8FCF2FD6B3C3F" authority="Caloren &amp; Marshall, 1998" authorityName="Caloren &amp; Marshall" authorityYear="1998" box="[151,731,771,798]" class="Insecta" family="Clusiidae" genus="Hendelia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="saopaulo">
<emphasis id="15B4EAA3452490171FF8FCF2FD6B3C3F" bold="true" box="[151,731,771,798]" pageId="15" pageNumber="16">
<emphasis id="15B4EAA3452490171FF8FCF2FECA3C3C" bold="true" box="[151,378,772,798]" italics="true" pageId="15" pageNumber="16">Hendelia saopaulo</emphasis>
(
<bibRefCitation id="43514B40452490171EE6FCF5FD633C3F" author="Caloren" box="[393,723,771,797]" pageId="15" pageNumber="17" refString="Caloren, D. C. &amp; Marshall, S. A. (1998) A revision of the New World species of Clusiodes Coquillett (Diptera: Clusiidae). Studia Dipterologica, 5, 261 - 321." type="journal article" year="1998">
Caloren &amp;
<collectingCountry id="5FD77621452490171D79FCF5FD3A3C3F" box="[534,650,771,797]" name="Marshall Islands" pageId="15" pageNumber="16">Marshall</collectingCountry>
, 1998
</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="277F36B1452490171FF8FCD1FE973C1D" blockId="15.[151,731,771,831]" box="[151,295,807,831]" pageId="15" pageNumber="16">
(
<figureCitation id="BFFB2A34452490171FF0FCD1FEAF3C1D" box="[159,287,807,831]" captionStart="FIGURES 33 – 58" captionStartId="13.[151,264,1701,1724]" captionTargetBox="[205,1368,199,1646]" captionTargetId="figure@13.[197,1373,193,1657]" captionTargetPageId="13" captionText="FIGURES 33 – 58: Genitalic figures of Hendelia mirabilis (Frey) and those species described by Caloren &amp; Marshall (1998). Figs 33 – 36: H. kinetrolicros (Caloren &amp; Marshall); 33: male external components, left lateral; 34: surstylus, posterior surface; 35: hypandrial complex, left lateral; 36: female internal genitalia. Figs 37 – 40: H. mirabilis; 37: male external components, left lateral; 38: surstylus, posterior surface; 39: hypandrial complex, left lateral; 40: female internal genitalia. Figs 41 – 44: H. napo (Caloren &amp; Marshall); 41: male external components, left lateral; 42: surstylus, posterior surface; 43: hypandrial complex, left lateral; 44: female internal genitalia. Figs 45 – 48: H. phrenzinus (Caloren &amp; Marshall); 45: male external components, left lateral; 46: surstylus, posterior surface; 47: hypandrial complex, left lateral; 48: female internal genitalia. Figs 49 – 51: H. pterygion (Caloren &amp; Marshall); 49: male external components, left lateral; 50: surstylus, posterior surface; 51: hypandrial complex, left lateral. Figs 52 – 54: H. stimulator (Caloren &amp; Marshall); 52: male external components, left lateral; 53: surstylus, posterior surface; 54: hypandrial complex, left lateral. Figs 55 – 58: H. saopaulo (Caloren &amp; Marshall); 55: male external components, left lateral; 56: surstylus, posterior surface; 57: hypandrial complex, left lateral; 58: female internal genitalia." httpUri="https://zenodo.org/record/201856/files/figure.png" pageId="15" pageNumber="16">Figs 55–58</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="6FDA653A452490171FF8FC86FD1E3C86" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="277F36B1452490171FF8FC86FD1E3C86" blockId="15.[151,686,880,933]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="07D0119F452490171FF8FC86FD1E3C86" pageId="15" pageNumber="16">
<taxonomicName id="E0C04D32452490171FF8FC86FD2E3CA7" ID-CoL="WDFT" authority="Caloren &amp; Marshall 1998: 303" authorityName="Caloren &amp; Marshall" authorityPageNumber="303" authorityYear="1998" box="[151,670,880,902]" class="Insecta" family="Clusiidae" genus="Clusiodes" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="saopaulo">
<emphasis id="15B4EAA3452490171FF8FC86FEEE3CA4" box="[151,350,880,902]" italics="true" pageId="15" pageNumber="16">Clusiodes saopaulo</emphasis>
<treatmentCitation id="A66110A0452490171E0BFC86FD2E3CA7" author="Caloren" box="[356,670,880,902]" page="303" pageId="15" pageNumber="16" year="1998">
<bibRefCitation id="43514B40452490171E0BFC86FDDB3CA7" author="Caloren" box="[356,619,880,902]" pageId="15" pageNumber="17" refString="Caloren, D. C. &amp; Marshall, S. A. (1998) A revision of the New World species of Clusiodes Coquillett (Diptera: Clusiidae). Studia Dipterologica, 5, 261 - 321." type="journal article" year="1998">
Caloren &amp;
<collectingCountry id="5FD77621452490171EBBFC86FD9E3CA4" box="[468,558,880,902]" name="Marshall Islands" pageId="15" pageNumber="16">Marshall</collectingCountry>
1998
</bibRefCitation>
: 303
</treatmentCitation>
</taxonomicName>
.
<taxonomicName id="E0C04D32452490171FF8FC79FEE73C87" box="[151,343,911,933]" class="Insecta" family="Clusiidae" genus="Hendelia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="saopaulo">
<emphasis id="15B4EAA3452490171FF8FC79FEE73C87" box="[151,343,911,933]" italics="true" pageId="15" pageNumber="16">Hendelia saopaulo</emphasis>
</taxonomicName>
.
<treatmentCitation id="A66110A0452490171E0CFC78FD1A3C86" author="Lonsdale" box="[355,682,910,932]" page="147" pageId="15" pageNumber="16" year="2007">
<bibRefCitation id="43514B40452490171E0CFC78FDC73C86" author="Lonsdale" box="[355,631,910,932]" pageId="15" pageNumber="17" refString="Lonsdale, O. &amp; Marshall, S. A. (2007) Redefinition of the genera Clusiodes and Hendelia (Diptera: Clusiidae, Clusiodinae), with a review of Clusiodes. Studia Dipterologica, 14, 117 - 159." type="journal article" year="2007">
Lonsdale &amp;
<collectingCountry id="5FD77621452490171E8EFC78FD8B3C86" box="[481,571,910,932]" name="Marshall Islands" pageId="15" pageNumber="16">Marshall</collectingCountry>
2007
</bibRefCitation>
: 147
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="6FDA653A452490171FF8FC24FA2D3BE0" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="277F36B1452490171FF8FC24FA2D3BE0" blockId="15.[151,1437,978,1545]" pageId="15" pageNumber="16">
<emphasis id="15B4EAA3452490171FF8FC24FEA13CC9" bold="true" box="[151,273,978,1003]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
As described for
<taxonomicName id="E0C04D32452490171EB6FC22FDC13CC9" box="[473,625,979,1003]" class="Insecta" family="Clusiidae" genus="Hendelia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="heliconiae">
<emphasis id="15B4EAA3452490171EB6FC22FDC13CC9" box="[473,625,979,1003]" italics="true" pageId="15" pageNumber="16">H. heliconiae</emphasis>
</taxonomicName>
except as follows: body length
<quantity id="E0389B54452490171CB9FC24FBE93CC8" box="[982,1113,978,1003]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="3.8" metricValueMin="3.6" pageId="15" pageNumber="16" unit="mm" value="3.7" valueMax="3.8" valueMin="3.6">
3.6
<emphasis id="15B4EAA3452490171C96FC25FBB73CC9" bold="true" box="[1017,1031,979,1003]" pageId="15" pageNumber="16">–</emphasis>
3.8mm
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; bristles brown with vibrissa light brown to yellow; posterior fronto-orbital smaller than anterior bristle or absent; frons always entirely yellow with ocellar tubercle brown; base colour of thorax light yellow with venter of pleuron often white and subnotal stripe thinner, and postpronotum light yellow to white, often strongly contrasting with scutum; female darker with base colour of scutum orange, anterior margin of first flagellomere lightly infuscated, and pigment on fore leg more strongly contrasting; anterodistal margin of epandrium broadly rounded, not strongly produced; surstylus broader and rounder; base of pregonite smooth; spermatheca globose and tuberculate. Non-type male paler (see comments).
</paragraph>
</subSubSection>
<subSubSection id="6FDA653A452490171FA8FB38FD523BC5" box="[199,738,1230,1255]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="277F36B1452490171FA8FB38FD523BC5" blockId="15.[151,1437,978,1545]" box="[199,738,1230,1255]" pageId="15" pageNumber="16">
<emphasis id="15B4EAA3452490171FA8FB38FED03BC5" bold="true" box="[199,352,1230,1255]" pageId="15" pageNumber="16">Distribution.</emphasis>
Southeast
<collectingCountry id="5FD77621452490171EB4FB39FDAF3BC5" box="[475,543,1231,1255]" name="Brazil" pageId="15" pageNumber="16">Brazil</collectingCountry>
(Rio de Janiero).
</paragraph>
</subSubSection>
<subSubSection id="6FDA653A452490171FA8FB05FD483A0D" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="277F36B1452490171FA8FB05FD483A0D" blockId="15.[151,1437,978,1545]" pageId="15" pageNumber="16">
<emphasis id="15B4EAA3452490171FA8FB05FD923A29" bold="true" box="[199,546,1266,1291]" pageId="15" pageNumber="16">Non-type material examined.</emphasis>
<collectingCountry id="5FD77621452490171D47FB05FD213A29" box="[552,657,1267,1291]" name="Brazil" pageId="15" pageNumber="16">BRAZIL</collectingCountry>
. Rio de Janiero: Teresópolis, Serra dos Orgáos Natl. Pk., trail to Pedra do Sino,
<quantity id="E0389B54452490171F94FAE1FE213A0D" box="[251,401,1303,1328]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="15" pageNumber="16" unit="m" value="1500.0" valueMax="1700.0" valueMin="1300.0">
1300
<emphasis id="15B4EAA3452490171E5CFAEEFEF13A12" bold="true" box="[307,321,1304,1328]" pageId="15" pageNumber="16">–</emphasis>
1700m
</quantity>
, sweep,
<date id="537E1071452490171E9EFAE1FDEB3A0D" box="[497,603,1303,1327]" pageId="15" pageNumber="16" value="1990-01-25">25.i.1990</date>
(1Ψ,
<collectionCode id="41D1AE74452490171DF4FAE1FD5C3A0D" box="[667,748,1303,1327]" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="15" pageNumber="16">DEBU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="6FDA653A452490171FA8FACDFA32392A" pageId="15" pageNumber="16" type="discussion">
<paragraph id="277F36B1452490171FA8FACDFA32392A" blockId="15.[151,1437,978,1545]" pageId="15" pageNumber="16">
<emphasis id="15B4EAA3452490171FA8FACDFEFF3A76" bold="true" box="[199,335,1339,1364]" pageId="15" pageNumber="16">Comments.</emphasis>
<bibRefCitation id="43514B40452490171E38FACAFD3B3A76" author="Caloren" box="[343,651,1340,1364]" pageId="15" pageNumber="17" refString="Caloren, D. C. &amp; Marshall, S. A. (1998) A revision of the New World species of Clusiodes Coquillett (Diptera: Clusiidae). Studia Dipterologica, 5, 261 - 321." type="journal article" year="1998">
Caloren &amp;
<collectingCountry id="5FD77621452490171EBAFACAFD883A76" box="[469,568,1340,1364]" name="Marshall Islands" pageId="15" pageNumber="16">Marshall</collectingCountry>
(1998)
</bibRefCitation>
state that
<taxonomicName id="E0C04D32452490171D90FACAFC653A76" box="[767,981,1340,1364]" class="Insecta" family="Clusiidae" genus="Hendelia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="saopaulo">
<emphasis id="15B4EAA3452490171D90FACAFC653A76" box="[767,981,1340,1364]" italics="true" pageId="15" pageNumber="16">Hendelia saopaulo</emphasis>
</taxonomicName>
has three fronto-orbital bristles, but this character appears to be variable as the posterior bristles are reduced in the
<typeStatus id="F87B8813452490171CBFFA96FB853A5A" box="[976,1077,1376,1400]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
, two
<typeStatus id="F87B8813452490171B1FFA97FB6C3A5B" box="[1136,1244,1377,1401]" pageId="15" pageNumber="16" type="paratype">paratypes</typeStatus>
and the non-type male, and absent in three
<typeStatus id="F87B8813452490171ED2FA73FD9C3ABF" box="[445,556,1413,1437]" pageId="15" pageNumber="16" type="paratype">paratypes</typeStatus>
. The non-type male differs from the
<typeStatus id="F87B8813452490171CBBFA73FBB43ABF" box="[980,1028,1413,1437]" pageId="15" pageNumber="16">type</typeStatus>
specimens in that it is entirely pale yellow except for the wing spots, and has a faint notal stripe, a dark ocellar tubercle, and an orange tint on the frons and anepimeron. The male genitalia (which are useful in separating this species from congeners), however, are concordant with those of the dissected
<typeStatus id="F87B8813452490171D4DFA07FDD0392B" box="[546,608,1521,1545]" pageId="15" pageNumber="16">types</typeStatus>
, and the collection locality falls within the known range of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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