219 lines
29 KiB
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219 lines
29 KiB
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<document id="4717F64BD9B9239A26C2C715BEDF5C09" ID-DOI="10.11646/zootaxa.5575.1.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="14746615" ID-ZooBank="A3C53F51-EDF9-4E25-9D29-9D1579E60EDF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737970758015" checkinUser="plazi" docAuthor="Maruyama, Munetoshi, Liang, Wei-Ren, Sokh, Heng & Ando, Kiyoshi" docDate="2025" docId="7D391F31FFA9FFCB1C9E78ACFD55FD32" docLanguage="en" docName="zootaxa.5575.1.8.pdf" docOrigin="Zootaxa 5575 (1)" docSource="https://doi.org/10.11646/zootaxa.5575.1.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microblattellus kakizoei Maruyama, Liang, Sokh & Ando, 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="171" masterDocId="81006749FFA8FFCF1C097D3AFFD0FFFF" masterDocTitle="The second species of the enigmatic genus Microblattellus Ferrer, 2006 (Coleoptera, Tenebrionidae) from Cambodia: A possibility of termitophily in the genus" masterLastPageNumber="172" masterPageNumber="167" pageNumber="168" updateTime="1737971943641" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A1C7C2D040B6989C80400A9065A362B6">The second species of the enigmatic genus Microblattellus Ferrer, 2006 (Coleoptera, Tenebrionidae) from Cambodia: A possibility of termitophily in the genus</mods:title>
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<mods:namePart id="D59361576441F5A55E081674DA5B4C8B">Maruyama, Munetoshi</mods:namePart>
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<mods:affiliation id="3F859926029F2BC23D833428F7E785DA">The Kyushu University Museum, Hakozaki 6 - 10 - 1, Higashi-ku, Fukuoka, 812 - 8581 Japan.</mods:affiliation>
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<mods:roleTerm id="D629B089CDF2A2FE792DFF1445192554">Author</mods:roleTerm>
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<mods:namePart id="974F1611752D8E99917A860A2E715A4D">Liang, Wei-Ren</mods:namePart>
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<mods:nameIdentifier id="AAA66FD46F6ACD9B74A98FB65E75E4F7" type="ORCID">0000-0003-0529-6907</mods:nameIdentifier>
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<mods:affiliation id="5146F9DE8755EA56EC1D5ACDCA039CD3">The Kyushu University Museum, Hakozaki 6 - 10 - 1, Higashi-ku, Fukuoka, 812 - 8581 Japan.</mods:affiliation>
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<mods:nameIdentifier id="30D51227BA912B104145BA8C4536C960" type="email">leif2051@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="B54BB196F69EB99BC54E7A6163F914BB">Author</mods:roleTerm>
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<mods:namePart id="7216D63276F2D2A3C91DD487FA5D875D">Sokh, Heng</mods:namePart>
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<mods:affiliation id="F4460DF300111AB9B1E6990C1574BC32">Forest-Wildlife Research and Development Institute, Forestry Administration, 40 Preah Norodom Boulevard, Phnom Penh, Cambodia.</mods:affiliation>
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<mods:namePart id="26AF1F30C9AACB8D999AA778A465ED47">Ando, Kiyoshi</mods:namePart>
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<mods:affiliation id="7E1529660B32E7B053D2702137504C82">Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi 3 - 5 - 7, Matsuyama, 790 - 8566 Japan.</mods:affiliation>
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<mods:typeOfResource id="FDDA3FFE2BD76FE3F458DCF3CF47B0EC">text</mods:typeOfResource>
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<mods:title id="6EC65554053710F8FEF2CE77D89AC170">Zootaxa</mods:title>
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<mods:part id="51A7D5196EC8438BE6C47D8F40D1E238">
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<mods:date id="BF007D19D648AA6EB1BB3D265234363D">2025</mods:date>
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<mods:number id="0CE0C377CB5A866FBC9791E0E085FE83">2025-01-22</mods:number>
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<mods:url id="00CDEAE8FE6F2623B529BDF3DA173334">https://doi.org/10.11646/zootaxa.5575.1.8</mods:url>
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<paragraph id="F52FAE27FFA9FFCE1C9E78ACFDCFFA4E" blockId="1.[151,543,1430,1493]" box="[151,543,1430,1457]" pageId="1" pageNumber="168">
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<heading id="AE67194BFFA9FFCE1C9E78ACFDCFFA4E" bold="true" box="[151,543,1430,1457]" fontSize="11" level="1" pageId="1" pageNumber="168" reason="1">
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<emphasis id="C7E47235FFA9FFCE1C9E78ACFDCFFA4E" bold="true" box="[151,543,1430,1457]" pageId="1" pageNumber="168">
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<taxonomicName id="3290D5A4FFA9FFCE1C9E78ACFE6AFA4E" authorityName="Maruyama & Liang & Sokh & Ando" authorityYear="2025" box="[151,442,1430,1457]" class="Insecta" family="Tenebrionidae" genus="Microblattellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="species" species="kakizoei" status="sp. nov.">
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<emphasis id="C7E47235FFA9FFCE1C9E78ACFE6AFA4E" bold="true" box="[151,442,1430,1457]" italics="true" pageId="1" pageNumber="168">Microblattellus kakizoei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="DCD7CF4EFFA9FFCE1DC878ADFDCFFA4E" box="[449,543,1431,1457]" pageId="1" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="BD8AFDACFFA9FFCB1C9E7881FD55FD32" lastPageId="4" lastPageNumber="171" pageId="1" pageNumber="168" type="description">
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<paragraph id="F52FAE27FFA9FFCE1C9E7881FECFFA2A" blockId="1.[151,543,1430,1493]" box="[151,287,1467,1493]" pageId="1" pageNumber="168">
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(
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<figureCitation id="6DABB2A2FFA9FFCE1C967881FEC6FA2A" box="[159,278,1467,1493]" captionStart-0="FIGURES 1–9" captionStart-1="FIGURES 10–16" captionStartId-0="2.[151,264,1919,1943]" captionStartId-1="3.[151,264,1700,1724]" captionTargetBox-0="[151,1435,270,1893]" captionTargetBox-1="[151,1435,181,1675]" captionTargetId-0="figure-48@2.[151,1435,270,1893]" captionTargetId-1="figure-20@3.[151,1435,181,1677]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–9. Microblattellus kakizoei sp. nov., holotype.1, Habitus, dorsal view; 2, ditto, ventral view; 3, head and pronotum, ventral view; 4, habitus, micro-CT scan, dorsal view; 5, ditto, lateral view; 6, ditto, ventral view; 7, head, micro-CT scan, dorsal view; 8, ditto, lateral view; 9, ditto, ventral view." captionText-1="FIGURES 10–16. 10, Aedeagus of Microblattellus kakizoei sp. nov., holotype, dorsal view; 11, ditto, lateral view; 12. Microblattellus lecongmani Ferrer, 2006, holotype male; 13, ditto, paratype female; 14, Blatticephalus adelotopus Heller, 1917, habitus, dorsal view; 15, ditto, lateral view; 16, ditto, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.14746617" figureDoi-1="http://doi.org/10.5281/zenodo.14746619" httpUri-0="https://zenodo.org/record/14746617/files/figure.png" httpUri-1="https://zenodo.org/record/14746619/files/figure.png" pageId="1" pageNumber="168">Figs. 1–11</figureCitation>
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)
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</paragraph>
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<paragraph id="F52FAE27FFA9FFCE1C9E7B39FB85F9BE" blockId="1.[151,1437,1538,2033]" pageId="1" pageNumber="168">
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<emphasis id="C7E47235FFA9FFCE1C9E7B39FEC6F9E3" box="[151,278,1539,1564]" italics="true" pageId="1" pageNumber="168">Type series</emphasis>
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.
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<materialsCitation id="45F8A47AFFA9FFCE1D2E7B39FB85F9BE" collectingDate="2015-10-15" collectorName="S. Kakizoe. Deposited & M. Maruyama's & Kyushu University Museum & Fukuoka." country="Cambodia" location="Angkor Watt" pageId="1" pageNumber="168" specimenCount="1" stateProvince="Siem Reap" typeStatus="holotype">
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<typeStatus id="2A2B1085FFA9FFCE1D2E7B39FE42F9E2" box="[295,402,1539,1565]" pageId="1" pageNumber="168" type="holotype">Holotype</typeStatus>
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, male, near the exit gate of
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<location id="F04FF8FCFFA9FFCE1ED17B39FCBDF9E2" LSID="urn:lsid:plazi:treatment:7D391F31FFA9FFCB1C9E78ACFD55FD32:F04FF8FCFFA9FFCE1ED17B39FCBDF9E2" box="[728,877,1539,1565]" country="Cambodia" name="Angkor Watt" pageId="1" pageNumber="168" stateProvince="Siem Reap">Angkor Watt</location>
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,
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<collectingRegion id="375460C5FFA9FFCE1F727B39FC2AF9E2" box="[891,1018,1539,1565]" country="Cambodia" name="Siem Reap" pageId="1" pageNumber="168">Siem Reap</collectingRegion>
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,
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<collectingCountry id="8D87EEB7FFA9FFCE18017B39FBAEF9E2" box="[1032,1150,1539,1565]" name="Cambodia" pageId="1" pageNumber="168">Cambodia</collectingCountry>
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,
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<date id="812E88E7FFA9FFCE18857B38FADAF9E2" box="[1164,1290,1538,1565]" pageId="1" pageNumber="168" value="2015-10-15">
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<collectingDate id="916A710FFFA9FFCE18857B38FADAF9E2" box="[1164,1290,1538,1565]" pageId="1" pageNumber="168" value="2015-10-15">15 X 2015</collectingDate>
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</date>
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,
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<collectorName id="5865CBF1FFA9FFCE19117B39FED9F9BE" pageId="1" pageNumber="168">S. Kakizoe. Deposited</collectorName>
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in
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<collectorName id="5865CBF1FFA9FFCE1D247B1DFE0FF9BE" box="[301,479,1575,1601]" pageId="1" pageNumber="168">M. Maruyama’s</collectorName>
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collection at the
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<collectorName id="5865CBF1FFA9FFCE1EA97B1DFC31F9BE" box="[672,993,1575,1601]" pageId="1" pageNumber="168">Kyushu University Museum</collectorName>
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,
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<collectorName id="5865CBF1FFA9FFCE1FE57B1DFB85F9BE" box="[1004,1109,1575,1601]" pageId="1" pageNumber="168">Fukuoka.</collectorName>
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</materialsCitation>
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</paragraph>
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<paragraph id="F52FAE27FFA9FFCE1CCE7B71FB79F92E" blockId="1.[151,1437,1538,2033]" pageId="1" pageNumber="168">
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<emphasis id="C7E47235FFA9FFCE1CCE7B71FEE9F99B" box="[199,313,1611,1636]" italics="true" pageId="1" pageNumber="168">Diagnosis</emphasis>
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.
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<taxonomicName id="3290D5A4FFA9FFCE1D417B71FD88F99A" authorityName="Maruyama & Liang & Sokh & Ando" authorityYear="2025" box="[328,600,1611,1637]" class="Insecta" family="Tenebrionidae" genus="Microblattellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="species" species="kakizoei" status="sp. nov.">
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<emphasis id="C7E47235FFA9FFCE1D417B71FD88F99A" box="[328,600,1611,1637]" italics="true" pageId="1" pageNumber="168">Microblattellus kakizoei</emphasis>
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</taxonomicName>
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<emphasis id="C7E47235FFA9FFCE1E697B71FD6AF99A" bold="true" box="[608,698,1611,1637]" pageId="1" pageNumber="168">
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<taxonomicNameLabel id="DCD7CF4EFFA9FFCE1E697B71FD6AF99A" box="[608,698,1611,1637]" pageId="1" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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is easily distinguished from
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<taxonomicName id="3290D5A4FFA9FFCE180A7B71FB7DF99A" authorityName="Ferrer" authorityYear="2006" box="[1027,1197,1611,1637]" class="Insecta" family="Tenebrionidae" genus="Microblattellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="species" species="lecongmani">
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<emphasis id="C7E47235FFA9FFCE180A7B71FBCAF99B" box="[1027,1050,1611,1636]" italics="true" pageId="1" pageNumber="168">M</emphasis>
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.
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<emphasis id="C7E47235FFA9FFCE18237B71FB7DF99A" box="[1066,1197,1611,1637]" italics="true" pageId="1" pageNumber="168">lecongmani</emphasis>
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</taxonomicName>
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(
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<figureCitation id="6DABB2A2FFA9FFCE18B47B71FA94F99A" box="[1213,1348,1611,1637]" captionStart="FIGURES 10–16" captionStartId="3.[151,264,1700,1724]" captionTargetBox="[151,1435,181,1675]" captionTargetId="figure-20@3.[151,1435,181,1677]" captionTargetPageId="3" captionText="FIGURES 10–16. 10, Aedeagus of Microblattellus kakizoei sp. nov., holotype, dorsal view; 11, ditto, lateral view; 12. Microblattellus lecongmani Ferrer, 2006, holotype male; 13, ditto, paratype female; 14, Blatticephalus adelotopus Heller, 1917, habitus, dorsal view; 15, ditto, lateral view; 16, ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14746619" httpUri="https://zenodo.org/record/14746619/files/figure.png" pageId="1" pageNumber="168">Figs. 12, 13</figureCitation>
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) by the more elongate and less robust body, more loosely articulate antennal segments, clear and transverse scutellum instead of reduced and invisible one in the latter, obviously produced humeral corners of elytra, finely and irregularly arranged strial punctures on elytra, and less punctate body surface. (See, also key to species.)
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</paragraph>
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<paragraph id="F52FAE27FFA9FFCD1CCE7BE1FC80FF2A" blockId="1.[151,1437,1538,2033]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="169" pageId="1" pageNumber="168">
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<emphasis id="C7E47235FFA9FFCE1CCE7BE1FE9BF90B" box="[199,331,1755,1780]" italics="true" pageId="1" pageNumber="168">Description</emphasis>
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. Male. Body length: ca.
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<quantity id="326803C2FFA9FFCE1E6F7BE1FD10F90A" box="[614,704,1755,1781]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="168" unit="mm" value="2.1">2.1 mm</quantity>
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. Body (
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<figureCitation id="6DABB2A2FFA9FFCE1F107BE1FC6EF90A" box="[793,958,1755,1781]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1919,1943]" captionTargetBox="[151,1435,270,1893]" captionTargetId="figure-48@2.[151,1435,270,1893]" captionTargetPageId="2" captionText="FIGURES 1–9. Microblattellus kakizoei sp. nov., holotype.1, Habitus, dorsal view; 2, ditto, ventral view; 3, head and pronotum, ventral view; 4, habitus, micro-CT scan, dorsal view; 5, ditto, lateral view; 6, ditto, ventral view; 7, head, micro-CT scan, dorsal view; 8, ditto, lateral view; 9, ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14746617" httpUri="https://zenodo.org/record/14746617/files/figure.png" pageId="1" pageNumber="168">Figs. 1, 2, 4–6</figureCitation>
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) subparallel-sided, reddish brown overall. Head (
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<figureCitation id="6DABB2A2FFA9FFCE1CED7BC5FEA1F8E6" box="[228,369,1791,1817]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1919,1943]" captionTargetBox="[151,1435,270,1893]" captionTargetId="figure-48@2.[151,1435,270,1893]" captionTargetPageId="2" captionText="FIGURES 1–9. Microblattellus kakizoei sp. nov., holotype.1, Habitus, dorsal view; 2, ditto, ventral view; 3, head and pronotum, ventral view; 4, habitus, micro-CT scan, dorsal view; 5, ditto, lateral view; 6, ditto, ventral view; 7, head, micro-CT scan, dorsal view; 8, ditto, lateral view; 9, ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14746617" httpUri="https://zenodo.org/record/14746617/files/figure.png" pageId="1" pageNumber="168">Figs. 3, 7–9</figureCitation>
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) slightly longer than wide; traces of eyes slightly convex laterally; dorsum almost glabrous; venter with coarse and piligerous punctures; antennae long, somewhat loosely articulate and all antennal segments wider than long except for scape; distal three antennal segments distinctly dilated and forming a club; antennal segments III to XI visible in dorsal view; segment I long, large; segment II almost as long as III, slightly wider than long; segments IV to VIII short and transverse; segment IX widest; segment X slightly shorter and narrower than IX; segment XI 2/5 as wide as X; labrum semicircular; mandibles thick, slightly asymmetric, apically pointed; mentum obtrapezoidal, deeply emarginate on anterior margin; labial palpus with segment I minute, as long as wide; segment II widened apically; III large, oval; maxillary palpus with segment I large, spherical; segment II short, triangular in ventral view; segment III large, oval, slightly truncate at apex.
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</paragraph>
|
||
<caption id="A1EFFEAFFFAAFFCD1C9E7A45FD57F820" ID-DOI="http://doi.org/10.5281/zenodo.14746617" ID-Zenodo-Dep="14746617" httpUri="https://zenodo.org/record/14746617/files/figure.png" pageId="2" pageNumber="169" startId="2.[151,264,1919,1943]" targetBox="[151,1435,270,1893]" targetPageId="2" targetType="figure">
|
||
<paragraph id="F52FAE27FFAAFFCD1C9E7A45FD57F820" blockId="2.[151,1437,1919,2016]" pageId="2" pageNumber="169">
|
||
<emphasis id="C7E47235FFAAFFCD1C9E7A45FEE9F868" bold="true" box="[151,313,1919,1943]" pageId="2" pageNumber="169">FIGURES 1–9.</emphasis>
|
||
<taxonomicName id="3290D5A4FFAAFFCD1D377A45FDE1F868" authorityName="Maruyama & Liang & Sokh & Ando" authorityYear="2025" box="[318,561,1919,1943]" class="Insecta" family="Tenebrionidae" genus="Microblattellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="169" phylum="Arthropoda" rank="species" species="kakizoei" status="sp. nov.">
|
||
<emphasis id="C7E47235FFAAFFCD1D377A45FDE1F868" box="[318,561,1919,1943]" italics="true" pageId="2" pageNumber="169">Microblattellus kakizoei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C7E47235FFAAFFCD1E3C7ABAFD54F867" bold="true" box="[565,644,1920,1944]" pageId="2" pageNumber="169">
|
||
<taxonomicNameLabel id="DCD7CF4EFFAAFFCD1E3C7ABAFD54F867" box="[565,644,1920,1944]" pageId="2" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype. 1, Habitus, dorsal view; 2, ditto, ventral view; 3, head and pronotum, ventral view; 4, habitus, micro-CT scan, dorsal view; 5, ditto, lateral view; 6, ditto, ventral view; 7, head, micro-CT scan, dorsal view; 8, ditto, lateral view; 9, ditto, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="A1EFFEAFFFABFFCC1C9E7B9EFCEAF8FA" ID-DOI="http://doi.org/10.5281/zenodo.14746619" ID-Zenodo-Dep="14746619" httpUri="https://zenodo.org/record/14746619/files/figure.png" pageId="3" pageNumber="170" startId="3.[151,264,1700,1724]" targetBox="[151,1435,181,1675]" targetPageId="3" targetType="figure">
|
||
<paragraph id="F52FAE27FFABFFCC1C9E7B9EFCEAF8FA" blockId="3.[151,1437,1700,1797]" pageId="3" pageNumber="170">
|
||
<emphasis id="C7E47235FFABFFCC1C9E7B9EFE89F942" bold="true" box="[151,345,1700,1725]" pageId="3" pageNumber="170">FIGURES 10–16.</emphasis>
|
||
10, Aedeagus of
|
||
<taxonomicName id="3290D5A4FFABFFCC1E177B9FFCC8F942" authorityName="Maruyama & Liang & Sokh & Ando" authorityYear="2025" box="[542,792,1701,1725]" class="Insecta" family="Tenebrionidae" genus="Microblattellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="170" phylum="Arthropoda" rank="species" species="kakizoei" status="sp. nov.">
|
||
<emphasis id="C7E47235FFABFFCC1E177B9FFCC8F942" box="[542,792,1701,1725]" italics="true" pageId="3" pageNumber="170">Microblattellus kakizoei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C7E47235FFABFFCC1F2D7B9FFCA9F942" bold="true" box="[804,889,1701,1725]" pageId="3" pageNumber="170">
|
||
<taxonomicNameLabel id="DCD7CF4EFFABFFCC1F2D7B9FFCA9F942" box="[804,889,1701,1725]" pageId="3" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, dorsal view; 11, ditto, lateral view; 12.
|
||
<taxonomicName id="3290D5A4FFABFFCC1C9E7BF3FDE2F91E" authority="Ferrer, 2006" authorityName="Ferrer" authorityYear="2006" box="[151,562,1737,1761]" class="Insecta" family="Tenebrionidae" genus="Microblattellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="170" phylum="Arthropoda" rank="species" species="lecongmani">
|
||
<emphasis id="C7E47235FFABFFCC1C9E7BF3FE7EF91E" box="[151,430,1737,1761]" italics="true" pageId="3" pageNumber="170">Microblattellus lecongmani</emphasis>
|
||
<bibRefCitation id="9101D3D6FFABFFCC1DBD7BF3FDE2F91E" author="Ferrer, J." box="[436,562,1737,1761]" pageId="3" pageNumber="170" pagination="75 - 83" refId="ref3077" refString="Ferrer, J. (2006) Constitution du groupe indo-african des Falsocossyphini, tribu nova, et description d'un nouveau genre hypogee du Vietnam. Coleoptera, Tenebrionidae. Cahiers Scientifiques. Centre de Conservation et d'Etude des collections. Museum d'Histoire Naturelle de Lyon, 10, 75 - 83. https: // doi. org / 10.3406 / mhnly. 2006.1349" type="journal article" year="2006">Ferrer, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype male; 13, ditto, paratype female; 14,
|
||
<taxonomicName id="3290D5A4FFABFFCC18057BF3FA48F91E" authority="Heller, 1917" authorityName="Heller" authorityYear="1917" box="[1036,1432,1737,1761]" class="Insecta" family="Tenebrionidae" genus="Blatticephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="170" phylum="Arthropoda" rank="species" species="adelotopus">
|
||
<emphasis id="C7E47235FFABFFCC18057BF3FAC3F91E" box="[1036,1299,1737,1761]" italics="true" pageId="3" pageNumber="170">Blatticephalus adelotopus</emphasis>
|
||
Heller, 1917
|
||
</taxonomicName>
|
||
, habitus, dorsal view; 15, ditto, lateral view; 16, ditto, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F52FAE27FFABFFCC1CCE7A6DFBB5F822" blockId="3.[151,1436,1879,2013]" pageId="3" pageNumber="170">
|
||
Pronotum (
|
||
<figureCitation id="6DABB2A2FFABFFCC1D4E7A6DFE04F88E" box="[327,468,1879,1905]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1919,1943]" captionTargetBox="[151,1435,270,1893]" captionTargetId="figure-48@2.[151,1435,270,1893]" captionTargetPageId="2" captionText="FIGURES 1–9. Microblattellus kakizoei sp. nov., holotype.1, Habitus, dorsal view; 2, ditto, ventral view; 3, head and pronotum, ventral view; 4, habitus, micro-CT scan, dorsal view; 5, ditto, lateral view; 6, ditto, ventral view; 7, head, micro-CT scan, dorsal view; 8, ditto, lateral view; 9, ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14746617" httpUri="https://zenodo.org/record/14746617/files/figure.png" pageId="3" pageNumber="170">Figs. 1, 3, 4</figureCitation>
|
||
) elongate, very feebly bisinuate at sides in basal two-thirds, almost twice as long as wide, widest around anterior 2/5; surface moderately, uniformly punctate, the punctures shallower on apical 1/5, each bearing a seta; basal margin almost straight though faintly produced posteriad; scutellum short and transverse, impunctate, with fine microsculpture; prosternal process oblong-oval, rounded at apex.
|
||
</paragraph>
|
||
<paragraph id="F52FAE27FFACFFCB1CCE7DADFEEEFEE2" blockId="4.[151,1437,151,717]" pageId="4" pageNumber="171">
|
||
Elytra (
|
||
<figureCitation id="6DABB2A2FFACFFCB1D127DADFE55FF4E" box="[283,389,151,177]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1919,1943]" captionTargetBox="[151,1435,270,1893]" captionTargetId="figure-48@2.[151,1435,270,1893]" captionTargetPageId="2" captionText="FIGURES 1–9. Microblattellus kakizoei sp. nov., holotype.1, Habitus, dorsal view; 2, ditto, ventral view; 3, head and pronotum, ventral view; 4, habitus, micro-CT scan, dorsal view; 5, ditto, lateral view; 6, ditto, ventral view; 7, head, micro-CT scan, dorsal view; 8, ditto, lateral view; 9, ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14746617" httpUri="https://zenodo.org/record/14746617/files/figure.png" pageId="4" pageNumber="171">Figs. 1, 4</figureCitation>
|
||
) elongate, more than twice as long as wide, widest around middle; humeral corners distinctly projected antero-laterally, and their apices rounded; surface moderately convex, finely, irregularly punctate; each puncture bearing a short seta; lateral margins feebly serrated, and each serration with a short seta; hindwing reduced, brachypterous.
|
||
</paragraph>
|
||
<paragraph id="F52FAE27FFACFFCB1CCE7C1DFA91FE9A" blockId="4.[151,1437,151,717]" pageId="4" pageNumber="171">
|
||
Legs (
|
||
<figureCitation id="6DABB2A2FFACFFCB1D077C1DFEAAFEBE" box="[270,378,295,321]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1919,1943]" captionTargetBox="[151,1435,270,1893]" captionTargetId="figure-48@2.[151,1435,270,1893]" captionTargetPageId="2" captionText="FIGURES 1–9. Microblattellus kakizoei sp. nov., holotype.1, Habitus, dorsal view; 2, ditto, ventral view; 3, head and pronotum, ventral view; 4, habitus, micro-CT scan, dorsal view; 5, ditto, lateral view; 6, ditto, ventral view; 7, head, micro-CT scan, dorsal view; 8, ditto, lateral view; 9, ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14746617" httpUri="https://zenodo.org/record/14746617/files/figure.png" pageId="4" pageNumber="171">Figs. 2, 6</figureCitation>
|
||
) short and robust; femora and tibiae sparsely covered with minute setae, posterior margins of meso- and metafemora ancipital for receiving corresponding tibia; tibiae scarcely incurved, dilated apicad.
|
||
</paragraph>
|
||
<paragraph id="F52FAE27FFACFFCB1CCE7C55FE7AFE52" blockId="4.[151,1437,151,717]" pageId="4" pageNumber="171">
|
||
Male genitalia as in
|
||
<figureCitation id="6DABB2A2FFACFFCB1DA67C55FDB1FE76" box="[431,609,367,393]" captionStart="FIGURES 10–16" captionStartId="3.[151,264,1700,1724]" captionTargetBox="[151,1435,181,1675]" captionTargetId="figure-20@3.[151,1435,181,1677]" captionTargetPageId="3" captionText="FIGURES 10–16. 10, Aedeagus of Microblattellus kakizoei sp. nov., holotype, dorsal view; 11, ditto, lateral view; 12. Microblattellus lecongmani Ferrer, 2006, holotype male; 13, ditto, paratype female; 14, Blatticephalus adelotopus Heller, 1917, habitus, dorsal view; 15, ditto, lateral view; 16, ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14746619" httpUri="https://zenodo.org/record/14746619/files/figure.png" pageId="4" pageNumber="171">Figs. 10 and 11</figureCitation>
|
||
; basale more than four times as long as apicale, strongly curved ventrad; apicale short and robust.
|
||
</paragraph>
|
||
<paragraph id="F52FAE27FFACFFCB1CCE7C8DFE4EFE2E" blockId="4.[151,1437,151,717]" box="[199,414,439,465]" pageId="4" pageNumber="171">Female. Unknown.</paragraph>
|
||
<paragraph id="F52FAE27FFACFFCB1CCE7CE1FD69FDC2" blockId="4.[151,1437,151,717]" pageId="4" pageNumber="171">Measurements. Head length (from base to labral apex), 486.80; head width, 417.05; antennal length, 582.31; pronotal length, 987.59; pronotal width, 570.43; elytral length, 1242.71; elytral width, 573.08; hind tibial length, 316.02; hind tibial width, 81.03. In micrometers.</paragraph>
|
||
<paragraph id="F52FAE27FFACFFCB1CCE7F7DFD55FD32" blockId="4.[151,1437,151,717]" pageId="4" pageNumber="171">
|
||
Remarks.
|
||
<typeStatus id="2A2B1085FFACFFCB1D317F7DFE70FD9E" box="[312,416,583,609]" pageId="4" pageNumber="171" type="holotype">Holotype</typeStatus>
|
||
was collected from a fungus garden of
|
||
<taxonomicName id="3290D5A4FFACFFCB1F5B7F7DFB09FD9E" authority="(Hagen, 1858)" baseAuthorityName="Hagen" baseAuthorityYear="1858" box="[850,1241,582,609]" class="Insecta" family="Termitidae" genus="Macrotermes" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="171" phylum="Arthropoda" rank="species" species="gilvus">
|
||
<emphasis id="C7E47235FFACFFCB1F5B7F7DFBFFFD9E" box="[850,1071,583,609]" italics="true" pageId="4" pageNumber="171">Macrotermes gilvus</emphasis>
|
||
(Hagen, 1858)
|
||
</taxonomicName>
|
||
. The morphology of the mouthparts is probably mostly common to that of
|
||
<taxonomicName id="3290D5A4FFACFFCB1F137F51FC14FD7A" authorityName="Ferrer" authorityYear="2006" box="[794,964,619,645]" class="Insecta" family="Tenebrionidae" genus="Microblattellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="species" species="lecongmani">
|
||
<emphasis id="C7E47235FFACFFCB1F137F51FCE1FD7B" box="[794,817,619,644]" italics="true" pageId="4" pageNumber="171">M</emphasis>
|
||
.
|
||
<emphasis id="C7E47235FFACFFCB1F487F51FC14FD7A" box="[833,964,619,645]" italics="true" pageId="4" pageNumber="171">lecongmani</emphasis>
|
||
</taxonomicName>
|
||
due to the close resemblance of the other character states; thus, it could be the character states delimiting the genus. The absence of the hindwings was confirmed by micro-CT data of
|
||
<taxonomicName id="3290D5A4FFACFFCB1DF47F89FDAEFD32" authorityName="Maruyama & Liang & Sokh & Ando" authorityYear="2025" box="[509,638,691,717]" class="Insecta" family="Tenebrionidae" genus="Microblattellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="species" species="kakizoei">
|
||
<emphasis id="C7E47235FFACFFCB1DF47F89FDAEFD32" box="[509,638,691,717]" italics="true" pageId="4" pageNumber="171">M. kakizoei</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |