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<taxonomicName id="2EF740AF4C53FFE4FCAC84C7A5C7F4B7" authority="(Rohon, 1900)" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" baseAuthorityName="Rohon" baseAuthorityYear="1900" box="[862,1508,1409,1437]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Antiarchi" pageId="4" pageNumber="101" phylum="Chordata" rank="subSpecies" species="leptocheira" status="comb. nov." subSpecies="jeremejevi">
<emphasis id="DB83E73E4C53FFE4FCAC84C7A501F4B7" box="[862,1314,1409,1437]" italics="true" pageId="4" pageNumber="101">Bothriolepis leptocheira jeremejevi</emphasis>
(
<bibRefCitation id="8D6646DD4C53FFE4FAC184C7A5FFF4B7" author="Rohon, J. V." box="[1331,1500,1409,1437]" pageId="4" pageNumber="101" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon, 1900</bibRefCitation>
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<taxonomicNameLabel id="C0B05A454C53FFE4FCAC84E3A3C8F4EB" box="[862,1003,1445,1473]" pageId="4" pageNumber="101" rank="species">comb. nov.</taxonomicNameLabel>
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<figureCitation id="71CC27A94C53FFE4FCAC848AA3E7F4CB" box="[862,964,1484,1505]" captionStart-0="Fig" captionStart-1="Fig" captionStart-10="Fig" captionStart-11="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="→" captionStart-7="Fig" captionStart-8="Fig" captionStart-9="Fig" captionStartId-0="4.[151,184,1188,1208]" captionStartId-1="5.[98,131,1308,1328]" captionStartId-10="14.[151,184,878,898]" captionStartId-11="15.[98,131,1132,1152]" captionStartId-2="6.[151,184,1321,1341]" captionStartId-3="7.[98,131,1207,1227]" captionStartId-4="8.[862,895,774,794]" captionStartId-5="8.[863,896,1656,1676]" captionStartId-6="10.[116,144,1620,1650]" captionStartId-7="10.[151,184,862,882]" captionStartId-8="11.[98,131,1825,1845]" captionStartId-9="12.[151,184,958,978]" captionTargetBox-0="[157,1530,160,1164]" captionTargetBox-1="[103,1477,164,1280]" captionTargetBox-10="[161,1533,162,851]" captionTargetBox-11="[103,764,164,1109]" captionTargetBox-2="[153,1534,165,1291]" captionTargetBox-3="[98,1478,163,1188]" captionTargetBox-4="[864,1534,161,746]" captionTargetBox-5="[891,1506,1125,1629]" captionTargetBox-6="[155,1534,159,837]" captionTargetBox-7="[155,1534,159,837]" captionTargetBox-8="[99,1479,163,1803]" captionTargetBox-9="[161,1528,166,928]" captionTargetId-0="figure-306@4.[173,1531,159,1164]" captionTargetId-1="figure-314@5.[99,1480,160,1281]" captionTargetId-10="figure-604@14.[153,1534,163,851]" captionTargetId-11="figure-870@15.[102,764,160,1109]" captionTargetId-2="figure-270@6.[153,1534,165,1294]" captionTargetId-3="figure-351@7.[98,1478,163,1189]" captionTargetId-4="figure-587@8.[863,1534,161,749]" captionTargetId-5="figure-600@8.[891,1506,1125,1630]" captionTargetId-6="figure-8@9.[99,1479,175,1936]" captionTargetId-7="figure-325@10.[152,1535,159,836]" captionTargetId-8="figure-8@11.[99,1480,163,1804]" captionTargetId-9="figure-466@12.[157,1529,161,933]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-10="14" captionTargetPageId-11="15" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="11" captionTargetPageId-9="12" captionText-0="Fig. 3. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, separate plates and parts of articulated head shields. A. Articulated Nu, left Pn and left La plates IG KSC 155/90 in dorsal view. B. Disarticulated plates of the head shield IG KSC 155/150 in dorsal view. C. Part of articulated head shield IG KSC 155/162 in dorsal view. D. Imprint of the ventral wall of trunk armour and some skeletal elements of the head armour IG KSC 155/10-3 in ventral view showing suborbital and infragnathal bones. Abbreviations: AVL, anterior ventro-lateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; ifc1, principal section of infraorbital sensory line; IG, inferognathal; La, lateral plate; Nu, nuchal plate; mp, middle pit-line groove; Pn, paranuchal plate; Pp, postpineal plate; Prm, premedian plate; SO, suborbital plate; socc, supraoccipital cross-commissural pit-line groove." captionText-1="Fig. 4. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, lateral AG), premedian (H), and left postmarginal (I) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/65 in visceral (D1) and dorsal (D2) views. E. IG KSC 155/66 in dorsal (E1), right lateral (E2), and visceral (E3) views. F. IG KSC 155/69 in dorsal view. G. IG KSC 155/64 in dorsal view. H. IG KSC 155/114 in dorsal (H1) and anterior (H2) views.. Plate disconnected from the partial head shield IG KSC 155/161, in dorsal view." captionText-10="Fig. 13. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, plates of the pectoral fin armour. A. Left Cv1 plate IG 155/6 in visceral (A1) and ventral (A2) views. B. Right Cv1 plate IG KSC 155/134 in ventral view. C. Left Ml2 plate IG KSC 155/149 in ventral view. D. Right Ml2 plate IG 155/137 in dorsal (D1) and ventral (D2) views. E. Right distal segment of the pectoral fin armour IG KSC 155/21 in dorsal (E1) and ventral (E2) views. F. Mm1 plate IG KSC 155/85 in mesial view. Abbreviations: Cd35, dorsal central plates 3 to 5; Cv4, ventral central plate 4; Ml45, lateral marginal plates 4 and 5; Mm46, mesial marginal plates 4 to 6; T, terminal plate." captionText-11="Fig. 14. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), reconstruction of the head, the trunk armour and the pectoral fins in ventral view, based on IG KSC 155/10-1, IG KSC 155/10-2, and IG KSC 155/10- 3. Abbreviations: AVL, anterior ventro-lateral plate; Cv14, ventral central plate 1 to 4; Ifg, inferognathal plate; La, lateral plate; Ml25, lateral marginal plates 2 to 5; Mm4, mesial marginal plate 4; MV, median ventral plate; Prl, prelateral plate; Prm, premedian plate; PVL, posterior ventro-lateral plate; SM, semilunar plate; SMg, submarginal plate; T, terminal plate." captionText-2="Fig. 5. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), lateral (AG, I) and premedian (H) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/74 in dorsal view. E. IG KSC 155/65 in dorsal view. F. IG KSC 155/66 in dorsal (F1) and visceral (F2) views.G. IG KSC 155/69 in dorsal view. H. IG KSC 155/114 in dorsal view. I. IG KSC 155/64 in dorsal view. Abbreviations: a1Sm and a2Sm, anterior and posterior attachment areas on lateral plate for submarginal plate; a.Prl, attachment area on lateral plate for prelateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; ifc1, principal section of infraorbital sensory line; ifc2, branch of infraorbital sensory line diverging on La; nprl, prelateral notch of head shield; p, lateral pit of head shield; prh, preorbital recess of head shield; pr.po, antero-lateral corner of otico-occipital depression; soc, anterior section of the supraorbital sensory line; spg, spiracular groove of head shield." captionText-3="Fig. 6. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, postpineal AC, E, G), nuchal (DG, I, KM), and paranuchal (H, J, L, M) plates of the head shield. A. IG KSC 155/54-2, disconnected from the Nu IG KSC 155/54-1, in dorsal (A1) and visceral (A2) views. B. IG KSC 155/146 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/155 in dorsal (C1) and visceral C2) views. D. IG KSC 155/58 in dorsal view. E. Articulated plates IG KSC 155/118 in dorsal view. F. IG KSC 155/53 in dorsal (F1), visceral (F2), and posterior (F3) views. G. Articulated plates IG KSC 155/154 in dorsal (G1) and visceral (G2) views. H. IG KSC 155/55 in dorsal (H1) and visceral (H2) views. I. IG KSC 155/54-1 from the same individual as plate in A, in dorsal view. J. IG KSC 155/119 in dorsal view. K. IG KSC 155/56 in dorsal view. L. IG KSC 155/117 in visceral (L1) and dorsal (L2) views. M. IG KSC 155/125 in dorsal view." captionText-4="Fig. 7. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), paranuchal (A) and postpineal and nuchal (B), plates of the head shield. A. IG KSC 155/117 in dorsal (A1) and visceral (A2) views. B. Articulated plates IG KSC 155/154 in dorsal (B1) and visceral (B2) views.Abbreviations: cr.o, median occipital crest of head shield; cr.pm, paramarginal crest of head shield; cr.pto, postorbital crest of head shield; cr.tv, transverse nuchal crest of head shield; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; g, paired pits on Pp; ifc1, principal section of infraorbital sensory line; mp, middle pit-line groove; nm, obtected nuchal area of head shield; Nu, nuchal plate; ood, otico-occipital depression of head shield; Pp, postpineal plate; socc, supraoccipital cross-commissural pit-line groove; sot, supraotic thickening of head shield." captionText-5="Fig. 8. Reconstruction of the head shield of antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), based on partial head shield (IG KSC 71/1-87), left (IG KSC 155/15), and right (IG KSC 155/65) lateral plates. Abbreviations: cir, semicircular pit-line groove; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; ifc1, principal section of infraorbital sensory line; ifc2, branch of infraorbital sensory line diverging on La; La, lateral plate; mp, middle pit-line groove; Nu, nuchal plate; Pmg, postmarginal plate; Pn, paranuchal plate; Pp, postpineal plate; Prm, premedian plate; prh, preorbital recess of head shield; soc, anterior section of the supraorbital sensory line; socc, supraoccipital cross-commissural pit-line groove." captionText-6="→ Fig. 9. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior median dorsal (AG) and posterior median dorsal (HM) plates of the trunk armour. A. IG KSC 155/5 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/108 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/97 in dorsal view. D. IG KSC 155/113 in dorsal (D1) and visceral (D2) views. E. IG KSC 155/140 in dorsal (E1) and visceral (E2) views. F. Impression of the dorsal surface of IG KSC 155/42.G. IG KSC 155/44 in dorsal view. H. Fragment of IG KSC 155/7 in dorsal view. I. IG KSC 155/1 in dorsal (I1) and visceral (I2) views. J. IG KSC 155/71 in dorsal view. K. Slightly deformed IG KSC 155/70 in dorsal (K1) and visceral (K2) views. L. IG KSC 155/158 in dorsal view. M. IG KSC 155/157 in dorsal (M1) and visceral (M2) views. Abbreviations: ADL, anterior dorso-lateral plate; alr, postlevator thickening; AMD, anterior median dorsal plate; cf.ADL, cf.AMD, and cf.MxL, area overlapping ADL, AMD or MxL respectively; cr.tp, posterior transversal internal crest; dlg1 and dlg2, anterior and posterior oblique dorsal sensory line groove; dma, tergal angle; dmr, dorsal median ridge; f.retr, levator fossa; grm, ventral median groove; l, lateral corner; mvr, median ventral ridge; MxL, mixilateral plate; npn, postnuchal notch; oa.ADL, oa.MxL and oa.PMD, area overlapped by ADL, MxL or PMD respectively; pa, posterior corner; pma, posterior marginal area; PMD, posterior median dorsal plate; pr.p, posterior process of AMD; pr.pl, external postlevator process; prv2, posterior ventral process of dorsal wall of trunk armour; pt1 and pt2, anterior and posterior ventral pit; pua, posterior unornamented area of PMD; rf, “round fossula”; sna, supranuchal area; tb, ventral tuberosity." captionText-7="Fig. 10. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior dorso-lateral (AD, F, left; E, right) and left mixilateral (GI, right) plates of the trunk armour. A. IG KSC 155/116 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/111. C. IG KSC 155/61. D. IG KSC 155/60. E. IG KSC 155/136. F. IG KSC 155/112. G. IG KSC 155/110. H. Incomplete IG KSC 155/144. I. IG KSC 155/115. BE, GI, in dorsal views; F, in lateral view.Abbreviations: ADL, anterior ventro-lateral plate; AMD, anterior median dorsal plate; AVL, anterior ventro-lateral plate; cf.AMD, area overlapping AMD; dlg2, posterior oblique dorsal sensory line groove; dlr, dorso-lateral ridge of trunk armour; dxp, dorso-ventral pit-line groove; lcg, main lateral line groove; oa.ADL, oa.AMD, oa.AVL, oa.PMD, area overlapped by ADL, AMD, AVL or PMD respectively; PMD, posterior median dorsal plate; pnoa, postnuchal ornamented corner of ADL; pro, processus obstans of trunk armour." captionText-8="Fig. 11. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, partial ventral wall (A, B), anterior ventro-lateral (AVL) (C, D, left; E, right), and median ventral (FJ) plates of the trunk armour, and elements of pectoral fin. A. Impression of partial ventral wall of the trunk armour, inferognathals, and suborbitals IG KSC 155/10-2 in ventral view. B. Partial ventral wall of the trunk armour with articulated pectoral fin armour IG KSC 155/10-1 in ventral view. C. IG KSC 155/151 (C1) and enlargened part of the ventral lamina with pathologies (C2, arrows). D. IG KSC 155/122 in ventral view. E. IG KSC 155/123 in ventral view. F. IG KSC 155/104 with the bite mark (arrow) → in ventral (F1) and visceral (F2) views. G. IG KSC 155/76 in ventral view. H. IG KSC 155/77 in ventral view. I. IG KSC 155/133 in ventral view. J. IG KSC 155/106 in ventral view. Abbreviations:AVL, anterior ventro-lateral plate; Cd12, dorsal central plates 1 and 2; Cv1, ventral central plate 1; cit1, crista transversalis interna anterior; cit2, transverse thickening on the visceral surface of AVL; ds, distal segment of the pectoral fin armour; Ifg, inferognathal; Ml2, lateral marginal plate 2; Mm2, mesial marginal plate 2; MV, median ventral plate; PVL, posterior ventro-lateral plate; SO, suborbital plate." captionText-9="Fig. 12. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, posterior ventro-lateral plates, left (A) and right (B, C) of the trunk armour. A. IG KSC 155/4 in ventral (A1) and lateral (A2) views. B. IG KSC 155/16 in ventral view. C. IG KSC 155/78 in ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.10980827" figureDoi-1="http://doi.org/10.5281/zenodo.10980829" figureDoi-10="http://doi.org/10.5281/zenodo.10980845" figureDoi-11="http://doi.org/10.5281/zenodo.10980847" figureDoi-2="http://doi.org/10.5281/zenodo.10980831" figureDoi-3="http://doi.org/10.5281/zenodo.10980833" figureDoi-4="http://doi.org/10.5281/zenodo.10980835" figureDoi-5="http://doi.org/10.5281/zenodo.10980837" figureDoi-6="http://doi.org/10.5281/zenodo.11061576" figureDoi-7="http://doi.org/10.5281/zenodo.10980839" figureDoi-8="http://doi.org/10.5281/zenodo.10980841" figureDoi-9="http://doi.org/10.5281/zenodo.10980843" httpUri-0="https://zenodo.org/record/10980827/files/figure.png" httpUri-1="https://zenodo.org/record/10980829/files/figure.png" httpUri-10="https://zenodo.org/record/10980845/files/figure.png" httpUri-11="https://zenodo.org/record/10980847/files/figure.png" httpUri-2="https://zenodo.org/record/10980831/files/figure.png" httpUri-3="https://zenodo.org/record/10980833/files/figure.png" httpUri-4="https://zenodo.org/record/10980835/files/figure.png" httpUri-5="https://zenodo.org/record/10980837/files/figure.png" httpUri-6="https://zenodo.org/record/11061576/files/figure.png" httpUri-7="https://zenodo.org/record/10980839/files/figure.png" httpUri-8="https://zenodo.org/record/10980841/files/figure.png" httpUri-9="https://zenodo.org/record/10980843/files/figure.png" pageId="4" pageNumber="101">Figs. 314</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1ED68A74C53FFE4FCAC84B5A582F70E" pageId="4" pageNumber="101" type="reference_group">
<paragraph id="E9483B2C4C53FFE4FCAC84B5A5C3F722" blockId="4.[862,1504,1523,1572]" box="[862,1504,1523,1544]" pageId="4" pageNumber="101">
<treatmentCitationGroup id="C9E71C024C53FFE4FCAC84B5A5C3F722" box="[862,1504,1523,1544]" pageId="4" pageNumber="101">
1900
<taxonomicName id="2EF740AF4C53FFE4FC6684B5A450F722" authority="Lukševičs &amp; Beznosov &amp; Stūris, 2017" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" box="[916,1139,1523,1544]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="101" phylum="Chordata" rank="species" species="jeremejevi" status="sp. nov.">
<emphasis id="DB83E73E4C53FFE4FC6684B5A450F722" box="[916,1139,1523,1544]" italics="true" pageId="4" pageNumber="101">Bothriolepis jeremejevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0B05A454C53FFE4FB8B84B5A4E4F722" box="[1145,1223,1523,1544]" pageId="4" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
;
<treatmentCitation id="68561D3D4C53FFE4FB2284B5A54EF722" author="Rohon, J. V." box="[1232,1389,1523,1544]" page="30" pageId="4" pageNumber="101" year="1900">
<bibRefCitation id="8D6646DD4C53FFE4FB2284B5A54EF722" author="Rohon, J. V." box="[1232,1389,1523,1544]" pageId="4" pageNumber="101" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon 1900: 30</bibRefCitation>
</treatmentCitation>
, fig. 18, 19.
</treatmentCitationGroup>
</paragraph>
<paragraph id="E9483B2C4C53FFE4FCAC8749A582F70E" blockId="4.[862,1504,1523,1572]" box="[862,1441,1551,1572]" pageId="4" pageNumber="101">
<treatmentCitationGroup id="C9E71C024C53FFE4FCAC8749A582F70E" box="[862,1441,1551,1572]" pageId="4" pageNumber="101">
1932
<taxonomicName id="2EF740AF4C53FFE4FC668749A4DDF70E" authority="Rohon, 1900" authorityName="Rohon" authorityYear="1900" box="[916,1278,1551,1572]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="101" phylum="Chordata" rank="species" species="jeremejewi">
<emphasis id="DB83E73E4C53FFE4FC668749A45BF70E" box="[916,1144,1551,1572]" italics="true" pageId="4" pageNumber="101">Bothriolepis jeremejewi</emphasis>
<bibRefCitation id="8D6646DD4C53FFE4FB8D8749A4DDF70E" author="Rohon, J. V." box="[1151,1278,1551,1572]" pageId="4" pageNumber="101" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon, 1900</bibRefCitation>
</taxonomicName>
;
<treatmentCitation id="68561D3D4C53FFE4FAF58749A5BEF70E" author="Gross, W." box="[1287,1437,1551,1572]" page="34" pageId="4" pageNumber="101" year="1932">
<bibRefCitation id="8D6646DD4C53FFE4FAF58749A5BEF70E" author="Gross, W." box="[1287,1437,1551,1572]" pageId="4" pageNumber="101" refId="ref18640" refString="Gross, W. 1932. Fossilium Catalogus. Pars 57: Antiarchi. 40 pp. W. Junk, Berlin." type="book" year="1932">Gross 1932: 34</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A1ED68A74C53FFE4FCAC8776A458F7F6" pageId="4" pageNumber="101" type="materials_examined">
<paragraph id="E9483B2C4C53FFE4FCAC8776A388F74B" blockId="4.[862,1535,1584,1633]" pageId="4" pageNumber="101">
<materialsCitation id="599F31714C53FFE4FCAC8776A388F74B" pageId="4" pageNumber="101">
<emphasis id="DB83E73E4C53FFE4FCAC8776A398F76F" box="[862,955,1584,1605]" italics="true" pageId="4" pageNumber="101">
<typeStatus id="364C858E4C53FFE4FCAC8776A398F76F" box="[862,955,1584,1605]" pageId="4" pageNumber="101" type="lectotype">Lectotype</typeStatus>
</emphasis>
: PIN 1350/12 fragmentary Cv
<subScript id="757339694C53FFE4FB13877CA4CAF763" attach="left" box="[1249,1257,1594,1609]" fontSize="6" pageId="4" pageNumber="101">1</subScript>
, illustrated by
<bibRefCitation id="8D6646DD4C53FFE4FA888776A5D8F76F" author="Rohon, J. V." box="[1402,1531,1584,1605]" pageId="4" pageNumber="101" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon (1900</bibRefCitation>
: fig. 19).
</materialsCitation>
</paragraph>
<paragraph id="E9483B2C4C53FFE4FCAC8728A506F7B5" blockId="4.[862,1535,1646,1695]" pageId="4" pageNumber="101">
<emphasis id="DB83E73E4C53FFE4FCAC8728A3F9F7A9" box="[862,986,1646,1667]" italics="true" pageId="4" pageNumber="101">Type locality</emphasis>
: Outcrop on the banks of the river Izhma, opposite Sosnogorsk, South Timan,
<collectingRegion id="2B33F5CE4C53FFE4FBCC87CCA4F7F7B5" box="[1086,1236,1674,1695]" country="Russia" name="Komi" pageId="4" pageNumber="101">Komi Republic</collectingRegion>
,
<collectingCountry id="91E07BBC4C53FFE4FB2C87CCA502F7B5" box="[1246,1313,1674,1695]" name="Russia" pageId="4" pageNumber="101">Russia</collectingCountry>
.
</paragraph>
<paragraph id="E9483B2C4C53FFE4FCAC87EDA458F7F6" blockId="4.[862,1535,1707,1756]" pageId="4" pageNumber="101">
<emphasis id="DB83E73E4C53FFE4FCAC87EDA3FBF7EA" box="[862,984,1707,1728]" italics="true" pageId="4" pageNumber="101">Type horizon</emphasis>
:
<materialsCitation id="599F31714C53FFE4FC1187EDA455F7F6" location="Upper Devonian" municipality="lower Famennian" pageId="4" pageNumber="101" specimenCount="1" stateProvince="Volgograd">
Sosnogorsk Formation,
<collectingRegion id="2B33F5CE4C53FFE4FB3787EDA50AF7EA" box="[1221,1321,1707,1728]" country="Russia" name="Volgograd" pageId="4" pageNumber="101">Volgograd</collectingRegion>
Regional Stage,
<collectingMunicipality id="092CA1564C53FFE4FA3A87EDA3E8F7F6" pageId="4" pageNumber="101">lower Famennian</collectingMunicipality>
,
<location id="EC286DF74C53FFE4FC268781A455F7F6" LSID="urn:lsid:plazi:treatment:615E8A3A4C53FFF0FCAC84C7A1C7F4F6:EC286DF74C53FFE4FC268781A455F7F6" box="[980,1142,1735,1756]" municipality="lower Famennian" name="Upper Devonian" pageId="4" pageNumber="101" stateProvince="Volgograd">Upper Devonian</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1ED68A74C53FFE5FCAC87AFA468F7F1" lastPageId="5" lastPageNumber="102" pageId="4" pageNumber="101" type="discussion">
<paragraph id="E9483B2C4C53FFE5FCAC87AFA17EF63C" blockId="4.[862,1535,1769,1958]" lastBlockId="5.[98,771,1470,1814]" lastPageId="5" lastPageNumber="102" pageId="4" pageNumber="101">
<emphasis id="DB83E73E4C53FFE4FCAC87AFA46FF7D4" box="[862,1100,1769,1790]" italics="true" pageId="4" pageNumber="101">Nomenclatorial remarks</emphasis>
:
<bibRefCitation id="8D6646DD4C53FFE4FBAF87AFA524F7D4" author="Gross, W." box="[1117,1287,1769,1790]" pageId="4" pageNumber="101" refId="ref18640" refString="Gross, W. 1932. Fossilium Catalogus. Pars 57: Antiarchi. 40 pp. W. Junk, Berlin." type="book" year="1932">Gross (1932: 26)</bibRefCitation>
selected the
<collectionCode id="8FE6A3E94C53FFE4FA7C87AFA5E5F7D4" box="[1422,1478,1769,1790]" pageId="4" pageNumber="101">AMD</collectionCode>
plate 473 from the Cilma River (North Timan) as a of
<taxonomicName id="2EF740AF4C53FFE4FACB8643A429F61C" authority="Rohon, 1900" authorityName="Rohon" authorityYear="1900" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="101" phylum="Chordata" rank="species" species="jeremejewi">
<emphasis id="DB83E73E4C53FFE4FACB8643A3A5F61C" italics="true" pageId="4" pageNumber="101">Bothriolepis jeremejewi</emphasis>
<bibRefCitation id="8D6646DD4C53FFE4FC7F8667A429F61C" author="Rohon, J. V." box="[909,1034,1825,1846]" pageId="4" pageNumber="101" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon, 1900</bibRefCitation>
</taxonomicName>
(erroneous spelling by Gross) illustrated by
<bibRefCitation id="8D6646DD4C53FFE4FA4D8667A3BAF678" author="Rohon, J. V." pageId="4" pageNumber="101" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon (1900</bibRefCitation>
: fig. 18). This specimen was most probably found in the lower Frasnian deposits and belongs to
<taxonomicName id="2EF740AF4C53FFE4FB6E861FA5C4F644" authority="(Rohon, 1900)" baseAuthorityName="Rohon" baseAuthorityYear="1900" box="[1180,1511,1881,1902]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="101" phylum="Arthropoda" rank="species" species="radiata">
<emphasis id="DB83E73E4C53FFE4FB6E861FA577F644" box="[1180,1364,1881,1902]" italics="true" pageId="4" pageNumber="101">Asterolepis radiata</emphasis>
(
<bibRefCitation id="8D6646DD4C53FFE4FA92861FA5C2F644" author="Rohon, J. V." box="[1376,1505,1881,1902]" pageId="4" pageNumber="101" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon, 1900</bibRefCitation>
)
</taxonomicName>
; it has never been found neither in CNIGR nor in PIN collections since the beginning of 1920-ies. However, the second specimen illustrated by
<bibRefCitation id="8D6646DD4C52FFE5FF7084F8A126F4F9" author="Rohon, J. V." box="[130,261,1470,1491]" pageId="5" pageNumber="102" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon (1900</bibRefCitation>
: fig. 19), 318b from the river Izhma (South Timan) has been identified by Alexander Ivanov in
<quantity id="2E0F96C94C52FFE5FDE3849AA27FF4DB" box="[529,604,1500,1521]" metricMagnitude="1" metricUnit="m" metricValue="5.06476" pageId="5" pageNumber="102" unit="in" value="1994.0">1994 in</quantity>
the collection of PIN.
<bibRefCitation id="8D6646DD4C52FFE5FF6484BFA13DF724" author="Rohon, J. V." box="[150,286,1529,1550]" pageId="5" pageNumber="102" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon (1900)</bibRefCitation>
erroneously identified it as the distal segment of the pectoral fin; in fact this is fragmentary Cv
<subScript id="757339694C52FFE5FDEC8766A205F705" attach="left" box="[542,550,1568,1583]" fontSize="6" pageId="5" pageNumber="102">1</subScript>
PIN 1350/12 from the outcrop on the banks of the river Izhma, judging from the original label. This specimen is about
<quantity id="2E0F96C94C52FFE5FEB18717A1A6F74C" box="[323,389,1617,1638]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="102" unit="mm" value="40.0">40 mm</quantity>
long light brown plate embedded in the light yellow limestone; colour and preservation of the bone, as well as character of the rock are typical for the “fish dolomite” from the locality on the right bank of the river Izhma opposite Sosnogorsk, South Timan,
<collectingCountry id="91E07BBC4C52FFE5FF598781A0CDF7F6" box="[171,238,1735,1756]" name="Russia" pageId="5" pageNumber="102">Russia</collectingCountry>
. Therefore the PIN 1350/12 illustrated by
<bibRefCitation id="8D6646DD4C52FFE5FD8C8781A2DCF7F6" author="Rohon, J. V." box="[638,767,1735,1756]" pageId="5" pageNumber="102" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon (1900</bibRefCitation>
: fig. 19) is designated here as the
<typeStatus id="364C858E4C52FFE5FE6D87A2A1D4F7D3" box="[415,503,1764,1785]" pageId="5" pageNumber="102" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="2EF740AF4C52FFE5FDEA87A2A17AF63C" authority="(Rohon, 1900)" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" baseAuthorityName="Rohon" baseAuthorityYear="1900" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="102" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="jeremejevi">
<emphasis id="DB83E73E4C52FFE5FDEA87A2A0E6F63C" italics="true" pageId="5" pageNumber="102">Bothriolepis leptocheira jeremejevi</emphasis>
(
<bibRefCitation id="8D6646DD4C52FFE5FF238647A170F63C" author="Rohon, J. V." box="[209,339,1793,1814]" pageId="5" pageNumber="102" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon, 1900</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<caption id="BD886BA44C52FFE5FF90845AA365F48A" ID-DOI="http://doi.org/10.5281/zenodo.10980829" ID-Zenodo-Dep="10980829" httpUri="https://zenodo.org/record/10980829/files/figure.png" pageId="5" pageNumber="102" startId="5.[98,131,1308,1328]" targetBox="[103,1477,164,1280]" targetPageId="5" targetType="figure">
<paragraph id="E9483B2C4C52FFE5FF90845AA365F48A" blockId="5.[98,1482,1308,1440]" pageId="5" pageNumber="102">
Fig. 4. Antiarchan fish
<taxonomicName id="2EF740AF4C52FFE5FECE845AA32CF41A" authority="(Rohon, 1900)" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" baseAuthorityName="Rohon" baseAuthorityYear="1900" box="[316,783,1308,1328]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="102" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="jeremejevi">
<emphasis id="DB83E73E4C52FFE5FECE845AA25CF41A" box="[316,639,1308,1328]" italics="true" pageId="5" pageNumber="102">Bothriolepis leptocheira jeremejevi</emphasis>
(
<bibRefCitation id="8D6646DD4C52FFE5FD7F845AA32BF41A" author="Rohon, J. V." box="[653,776,1308,1328]" pageId="5" pageNumber="102" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon, 1900</bibRefCitation>
)
</taxonomicName>
, Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, lateral AG), premedian (H), and left postmarginal (I) plates of the head shield.
<emphasis id="DB83E73E4C52FFE5FD088470A329F461" bold="true" box="[762,778,1334,1355]" pageId="5" pageNumber="102">A</emphasis>
. IG KSC 155/15 in dorsal view.
<emphasis id="DB83E73E4C52FFE5FBC38470A463F461" bold="true" box="[1073,1088,1334,1355]" pageId="5" pageNumber="102">B</emphasis>
. IG KSC 155/67 in dorsal (B
<subScript id="757339694C52FFE5FABA8407A573F465" attach="left" box="[1352,1360,1345,1359]" fontSize="6" pageId="5" pageNumber="102">1</subScript>
) and visceral B
<subScript id="757339694C52FFE5FF8A841BA0A3F441" attach="left" box="[120,128,1373,1387]" fontSize="6" pageId="5" pageNumber="102">2</subScript>
) views.
<emphasis id="DB83E73E4C52FFE5FF398414A0F8F44D" bold="true" box="[203,219,1362,1383]" pageId="5" pageNumber="102">C</emphasis>
. IG KSC 155/129 in dorsal view.
<emphasis id="DB83E73E4C52FFE5FDF58414A234F44D" bold="true" box="[519,535,1362,1383]" pageId="5" pageNumber="102">D</emphasis>
. IG KSC 155/65 in visceral (D
<subScript id="757339694C52FFE5FCD8841BA311F441" attach="left" box="[810,818,1373,1387]" fontSize="6" pageId="5" pageNumber="102">1</subScript>
) and dorsal (D
<subScript id="757339694C52FFE5FC44841BA39DF441" attach="left" box="[950,958,1373,1387]" fontSize="6" pageId="5" pageNumber="102">2</subScript>
) views.
<emphasis id="DB83E73E4C52FFE5FBFB8414A43BF44D" bold="true" box="[1033,1048,1362,1383]" pageId="5" pageNumber="102">E</emphasis>
. IG KSC 155/66 in dorsal (E
<subScript id="757339694C52FFE5FAEB841BA502F441" attach="left" box="[1305,1313,1373,1387]" fontSize="6" pageId="5" pageNumber="102">1</subScript>
), right lateral (E
<subScript id="757339694C52FFE5FA46841BA59FF441" attach="left" box="[1460,1468,1373,1387]" fontSize="6" pageId="5" pageNumber="102">2</subScript>
), and visceral (E
<subScript id="757339694C52FFE5FF1B843FA0D2F4AD" attach="left" box="[233,241,1401,1415]" fontSize="6" pageId="5" pageNumber="102">3</subScript>
) views.
<emphasis id="DB83E73E4C52FFE5FECC8428A16FF4A9" bold="true" box="[318,332,1390,1411]" pageId="5" pageNumber="102">F</emphasis>
. IG KSC 155/69 in dorsal view.
<emphasis id="DB83E73E4C52FFE5FD848428A2ABF4AE" bold="true" box="[630,648,1390,1412]" pageId="5" pageNumber="102">G</emphasis>
. IG KSC 155/64 in dorsal view.
<emphasis id="DB83E73E4C52FFE5FC408428A3E7F4A9" bold="true" box="[946,964,1390,1411]" pageId="5" pageNumber="102">H</emphasis>
. IG KSC 155/114 in dorsal (H
<subScript id="757339694C52FFE5FB28843FA4C1F4AD" attach="left" box="[1242,1250,1401,1415]" fontSize="6" pageId="5" pageNumber="102">1</subScript>
) and anterior (H
<subScript id="757339694C52FFE5FA8B843FA5A2F4AD" attach="left" box="[1401,1409,1401,1415]" fontSize="6" pageId="5" pageNumber="102">2</subScript>
) views.. Plate disconnected from the partial head shield IG KSC 155/161, in dorsal view.
</paragraph>
</caption>
<paragraph id="E9483B2C4C52FFE5FF90866DA468F7F1" blockId="5.[98,771,1835,1960]" lastBlockId="5.[808,1482,1470,1755]" pageId="5" pageNumber="102">
<emphasis id="DB83E73E4C52FFE5FF90866DA0E7F66E" box="[98,196,1835,1860]" italics="true" pageId="5" pageNumber="102">Material</emphasis>
.—
<materialsCitation id="599F31714C52FFE5FF17866DA46BF7F1" collectionCode="AMD, PVL, PMD" pageId="5" pageNumber="102" specimenCount="3">
PIN 208, part of the head shield, two fragmentary Cv
<subScript id="757339694C52FFE5FF4A8611A0E2F642" attach="left" box="[184,193,1879,1896]" fontSize="7" pageId="5" pageNumber="102">1</subScript>
, Nu,
<collectionCode id="8FE6A3E94C52FFE5FEF6860AA16AF64F" box="[260,329,1868,1893]" pageId="5" pageNumber="102">AMD</collectionCode>
, proximal segment of the pectoral appendage (on a single slab of limestone); IG KSC 155 (more than 160 disarticulated but mainly complete bones prepared out of the rock, some semiarticulated head shields, and
<specimenCount id="FFF1F0A54C52FFE5FA6284F8A3BDF4D2" count="3" pageId="5" pageNumber="102" type="generic">three specimens</specimenCount>
demonstrating almost complete ventral wall of the trunk armour on several slabs), and IG KSC 71/И (many blocks with disarticulated fish remains, usually each block containing several specimens; only small part of all specimens is prepared); CNIGR 1226/1073,
<collectionCode id="8FE6A3E94C52FFE5FAF78726A51EF753" box="[1285,1341,1632,1657]" country="Argentina" httpUri="http://grbio.org/cool/pbkd-hmeb" name="Paleontologia de Vertebrados Lillo" pageId="5" pageNumber="102">PVL</collectionCode>
; 1226/1074,
<collectionCode id="8FE6A3E94C52FFE5FCDA87C7A340F7B0" box="[808,867,1665,1690]" pageId="5" pageNumber="102">PMD</collectionCode>
; 1226/1078,
<collectionCode id="8FE6A3E94C52FFE5FBF187C7A463F7B0" box="[1027,1088,1665,1690]" pageId="5" pageNumber="102">AMD</collectionCode>
(for complete list of specimens from IG KSC see the Appendix 1). All material from the
<typeStatus id="364C858E4C52FFE5FCDA8784A37AF7F1" box="[808,857,1730,1755]" pageId="5" pageNumber="102">type</typeStatus>
locality and horizon
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1ED68A74C52FFEFFCDA87ABA594F677" lastPageId="15" lastPageNumber="112" pageId="5" pageNumber="102" type="description">
<paragraph id="E9483B2C4C52FFE6FCDA87ABA18CF789" blockId="5.[808,1481,1773,1960]" lastBlockId="6.[151,825,1543,1960]" lastPageId="6" lastPageNumber="103" pageId="5" pageNumber="102">
<emphasis id="DB83E73E4C52FFE5FCDA87ABA38DF62C" box="[808,942,1773,1798]" italics="true" pageId="5" pageNumber="102">Description</emphasis>
.—
<taxonomicName id="2EF740AF4C52FFE5FC2087ABA4CBF62C" authorityName="Traquair" authorityYear="1893" box="[978,1256,1773,1798]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="102" phylum="Chordata" rank="species" species="leptocheira">
<emphasis id="DB83E73E4C52FFE5FC2087ABA4CBF62C" box="[978,1256,1773,1798]" italics="true" pageId="5" pageNumber="102">Bothriolepis leptocheira</emphasis>
</taxonomicName>
is well-represented in limestone from the Sosnogorsk locality by some articulated head shields and ventral walls of the trunk armour, some articulated proximal segments of the pectoral fin, as well as by numerous disarticulated plates of the head and trunk shields, and pectoral appendage. Most are from individuals of moderate size; however, there are remains from a very large, as well as from very small individuals. The plates are usually slightly flattened, sometimes deformed and bear cracks, most probably due to compaction of the fossil-bearing limestone.
</paragraph>
<caption id="BD886BA44C51FFE6FF65846FA623F4CF" ID-DOI="http://doi.org/10.5281/zenodo.10980831" ID-Zenodo-Dep="10980831" httpUri="https://zenodo.org/record/10980831/files/figure.png" pageId="6" pageNumber="103" startId="6.[151,184,1321,1341]" targetBox="[153,1534,165,1291]" targetPageId="6" targetType="figure">
<paragraph id="E9483B2C4C51FFE6FF65846FA623F4CF" blockId="6.[151,1536,1319,1509]" pageId="6" pageNumber="103">
Fig. 5. Antiarchan fish
<taxonomicName id="2EF740AF4C51FFE6FE97846FA30DF417" authority="(Rohon, 1900)" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" baseAuthorityName="Rohon" baseAuthorityYear="1900" box="[357,814,1321,1341]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="103" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="jeremejevi">
<emphasis id="DB83E73E4C51FFE6FE97846FA287F417" box="[357,676,1321,1341]" italics="true" pageId="6" pageNumber="103">Bothriolepis leptocheira jeremejevi</emphasis>
(
<bibRefCitation id="8D6646DD4C51FFE6FD5D846FA304F417" author="Rohon, J. V." box="[687,807,1321,1341]" pageId="6" pageNumber="103" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon, 1900</bibRefCitation>
)
</taxonomicName>
, lateral (AG, I) and premedian (H) plates of the head shield.
<emphasis id="DB83E73E4C51FFE6FAA58461A544F416" bold="true" box="[1367,1383,1319,1340]" pageId="6" pageNumber="103">A</emphasis>
. IG KSC 155/15 in dorsal view.
<emphasis id="DB83E73E4C51FFE6FED28402A10CF473" bold="true" box="[288,303,1348,1369]" pageId="6" pageNumber="103">B</emphasis>
. IG KSC 155/67 in dorsal (B
<subScript id="757339694C51FFE6FDC78408A21EF476" attach="left" box="[565,573,1358,1372]" fontSize="6" pageId="6" pageNumber="103">1</subScript>
) and visceral (B
<subScript id="757339694C51FFE6FD238408A2FAF476" attach="left" box="[721,729,1358,1372]" fontSize="6" pageId="6" pageNumber="103">2</subScript>
) views.
<emphasis id="DB83E73E4C51FFE6FCD48405A315F473" bold="true" box="[806,822,1347,1369]" pageId="6" pageNumber="103">C</emphasis>
. IG KSC 155/129 in dorsal view.
<emphasis id="DB83E73E4C51FFE6FB948402A455F473" bold="true" box="[1126,1142,1348,1369]" pageId="6" pageNumber="103">D</emphasis>
. IG KSC 155/74 in dorsal view.
<emphasis id="DB83E73E4C51FFE6FA6E8402A588F473" bold="true" box="[1436,1451,1348,1369]" pageId="6" pageNumber="103">E</emphasis>
. IG KSC 155/65 in dorsal view.
<emphasis id="DB83E73E4C51FFE6FE908426A153F45F" bold="true" box="[354,368,1376,1397]" pageId="6" pageNumber="103">F</emphasis>
. IG KSC 155/66 in dorsal (F
<subScript id="757339694C51FFE6FD81842CA258F452" attach="left" box="[627,635,1386,1400]" fontSize="6" pageId="6" pageNumber="103">1</subScript>
) and visceral (F
<subScript id="757339694C51FFE6FCFE842CA337F452" attach="left" box="[780,788,1386,1400]" fontSize="6" pageId="6" pageNumber="103">2</subScript>
) views.
<emphasis id="DB83E73E4C51FFE6FCAD8419A352F45F" bold="true" box="[863,881,1375,1397]" pageId="6" pageNumber="103">G</emphasis>
. IG KSC 155/69 in dorsal view.
<emphasis id="DB83E73E4C51FFE6FB668426A485F45F" bold="true" box="[1172,1190,1376,1397]" pageId="6" pageNumber="103">H</emphasis>
. IG KSC 155/114 in dorsal view.
<emphasis id="DB83E73E4C51FFE6FA218426A5FFF45F" bold="true" box="[1491,1500,1376,1397]" pageId="6" pageNumber="103">I</emphasis>
. IG KSC 155/64 in dorsal view. Abbreviations: a
<subScript id="757339694C51FFE6FDD584C0A20CF4BE" attach="both" box="[551,559,1414,1428]" fontSize="6" pageId="6" pageNumber="103">1</subScript>
Sm and a
<subScript id="757339694C51FFE6FD7384C0A2AAF4BE" attach="both" box="[641,649,1414,1428]" fontSize="6" pageId="6" pageNumber="103">2</subScript>
Sm, anterior and posterior attachment areas on lateral plate for submarginal plate; a.Prl, attachment area on lateral plate for prelateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; ifc
<subScript id="757339694C51FFE6FBAF84E4A446F49A" attach="left" box="[1117,1125,1442,1456]" fontSize="6" pageId="6" pageNumber="103">1</subScript>
, principal section of infraorbital sensory line; ifc
<subScript id="757339694C51FFE6FF5D84F8A094F4E6" attach="left" box="[175,183,1470,1484]" fontSize="6" pageId="6" pageNumber="103">2</subScript>
, branch of infraorbital sensory line diverging on La; nprl, prelateral notch of head shield; p, lateral pit of head shield; prh, preorbital recess of head shield; pr.po, antero-lateral corner of otico-occipital depression; soc, anterior section of the supraorbital sensory line; spg, spiracular groove of head shield.
</paragraph>
</caption>
<paragraph id="E9483B2C4C51FFE7FF4F87ECA247F7AD" blockId="6.[151,825,1543,1960]" lastBlockId="7.[98,771,1422,1959]" lastPageId="7" lastPageNumber="104" pageId="6" pageNumber="103">
The head shield (
<figureCitation id="71CC27A94C51FFE6FE7887ECA1EEF7E9" box="[394,461,1706,1731]" captionStart="Fig" captionStartId="4.[151,184,1188,1208]" captionTargetBox="[157,1530,160,1164]" captionTargetId="figure-306@4.[173,1531,159,1164]" captionTargetPageId="4" captionText="Fig. 3. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, separate plates and parts of articulated head shields. A. Articulated Nu, left Pn and left La plates IG KSC 155/90 in dorsal view. B. Disarticulated plates of the head shield IG KSC 155/150 in dorsal view. C. Part of articulated head shield IG KSC 155/162 in dorsal view. D. Imprint of the ventral wall of trunk armour and some skeletal elements of the head armour IG KSC 155/10-3 in ventral view showing suborbital and infragnathal bones. Abbreviations: AVL, anterior ventro-lateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; ifc1, principal section of infraorbital sensory line; IG, inferognathal; La, lateral plate; Nu, nuchal plate; mp, middle pit-line groove; Pn, paranuchal plate; Pp, postpineal plate; Prm, premedian plate; SO, suborbital plate; socc, supraoccipital cross-commissural pit-line groove." figureDoi="http://doi.org/10.5281/zenodo.10980827" httpUri="https://zenodo.org/record/10980827/files/figure.png" pageId="6" pageNumber="103">Fig. 3</figureCitation>
) is moderately broad, with an average B/L index 1.24 (n = 2), slightly narrower than the head shield of
<taxonomicName id="2EF740AF4C51FFE6FEC587AAA2D8F62F" authority="Gross" authorityName="Gross" authorityYear="1942" box="[311,763,1772,1797]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="103" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="curonica">
<emphasis id="DB83E73E4C51FFE6FEC587AAA290F62F" box="[311,691,1772,1797]" italics="true" pageId="6" pageNumber="103">Bothriolepis leptocheira curonica</emphasis>
Gross
</taxonomicName>
from
<collectingCountry id="91E07BBC4C51FFE6FF65864AA0C0F60F" box="[151,227,1804,1829]" name="Latvia" pageId="6" pageNumber="103">Latvia</collectingCountry>
, but almost similar to that of
<taxonomicName id="2EF740AF4C51FFE6FDB1864AA0FAF66C" authority="(Luksevics 2001)" baseAuthorityName="Luksevics" baseAuthorityYear="2001" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="103" phylum="Chordata" rank="species" species="ornata">
<emphasis id="DB83E73E4C51FFE6FDB1864AA291F60F" box="[579,690,1804,1829]" italics="true" pageId="6" pageNumber="103">B. ornata</emphasis>
(
<bibRefCitation id="8D6646DD4C51FFE6FD31864AA0F3F66C" author="Luksevics, E." pageId="6" pageNumber="103" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
)
</taxonomicName>
. The rostral margin is moderately convex and only slightly longer than the posterior margin. The head shield is weakly vaulted both rostrocaudally and transversely, as in Scottish material (
<bibRefCitation id="8D6646DD4C51FFE6FE7C86C9A232F682" author="Miles, R. S." box="[398,529,1935,1960]" pageId="6" pageNumber="103" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles 1968</bibRefCitation>
), but the anterior part of the Prm and La plates is strongly curved. The antero-lateral corner, the prelateral notch (nprl;
<figureCitation id="71CC27A94C51FFE6FB23876EA56CF76B" box="[1233,1359,1576,1601]" captionStart="Fig" captionStartId="5.[98,131,1308,1328]" captionTargetBox="[103,1477,164,1280]" captionTargetId="figure-314@5.[99,1480,160,1281]" captionTargetPageId="5" captionText="Fig. 4. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, lateral AG), premedian (H), and left postmarginal (I) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/65 in visceral (D1) and dorsal (D2) views. E. IG KSC 155/66 in dorsal (E1), right lateral (E2), and visceral (E3) views. F. IG KSC 155/69 in dorsal view. G. IG KSC 155/64 in dorsal view. H. IG KSC 155/114 in dorsal (H1) and anterior (H2) views.. Plate disconnected from the partial head shield IG KSC 155/161, in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980829" httpUri="https://zenodo.org/record/10980829/files/figure.png" pageId="6" pageNumber="103">Figs. 4C, D</figureCitation>
<figureCitation id="71CC27A94C51FFE6FABD8775A57AF76E" box="[1359,1369,1587,1604]" captionStart="Fig" captionStartId="2.[151,184,1333,1353]" captionTargetBox="[152,823,164,1305]" captionTargetId="figure-630@2.[152,604,164,1305]" captionTargetPageId="2" captionText="Fig. 2. Lithology and stratigraphy of the Sosnogorsk locality. “Fish-dolomite”, bed No. 40 is located about 8 m above the base of the section. Abbreviations: D3f, Upper Devonian, Frasnian; D3fm, Upper Devonian, Famennian." figureDoi="http://doi.org/10.5281/zenodo.10980823" httpUri="https://zenodo.org/record/10980823/files/figure.png" pageId="6" pageNumber="103">
<subScript id="757339694C51FFE6FABD8775A57AF76E" attach="left" box="[1359,1369,1587,1604]" fontSize="7" pageId="6" pageNumber="103">2</subScript>
</figureCitation>
,
<figureCitation id="71CC27A94C51FFE6FA94876EA557F76B" box="[1382,1396,1576,1601]" captionStart="Fig" captionStartId="6.[151,184,1321,1341]" captionTargetBox="[153,1534,165,1291]" captionTargetId="figure-270@6.[153,1534,165,1294]" captionTargetPageId="6" captionText="Fig. 5. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), lateral (AG, I) and premedian (H) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/74 in dorsal view. E. IG KSC 155/65 in dorsal view. F. IG KSC 155/66 in dorsal (F1) and visceral (F2) views.G. IG KSC 155/69 in dorsal view. H. IG KSC 155/114 in dorsal view. I. IG KSC 155/64 in dorsal view. Abbreviations: a1Sm and a2Sm, anterior and posterior attachment areas on lateral plate for submarginal plate; a.Prl, attachment area on lateral plate for prelateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; ifc1, principal section of infraorbital sensory line; ifc2, branch of infraorbital sensory line diverging on La; nprl, prelateral notch of head shield; p, lateral pit of head shield; prh, preorbital recess of head shield; pr.po, antero-lateral corner of otico-occipital depression; soc, anterior section of the supraorbital sensory line; spg, spiracular groove of head shield." figureDoi="http://doi.org/10.5281/zenodo.10980831" httpUri="https://zenodo.org/record/10980831/files/figure.png" pageId="6" pageNumber="103">5</figureCitation>
) and the lateral process are usually weakly defined, as in
<taxonomicName id="2EF740AF4C51FFE6FA92870EA5DCF74B" authorityName="Traquair" authorityYear="1893" box="[1376,1535,1608,1633]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="103" phylum="Chordata" rank="species" species="leptocheira">
<emphasis id="DB83E73E4C51FFE6FA92870EA5DCF74B" box="[1376,1535,1608,1633]" italics="true" pageId="6" pageNumber="103">B. leptocheira</emphasis>
</taxonomicName>
from
<collectingCountry id="91E07BBC4C51FFE6FC6F872FA423F7A8" box="[925,1024,1641,1666]" name="United Kingdom" pageId="6" pageNumber="103">Scotland</collectingCountry>
and
<collectingCountry id="91E07BBC4C51FFE6FBCB872FA4A5F7A8" box="[1081,1158,1641,1666]" name="Latvia" pageId="6" pageNumber="103">Latvia</collectingCountry>
. The orbital fenestra is well seen in IG KSC 155/162 (
<figureCitation id="71CC27A94C51FFE6FBB687CCA4B9F789" box="[1092,1178,1674,1699]" captionStart="Fig" captionStartId="4.[151,184,1188,1208]" captionTargetBox="[157,1530,160,1164]" captionTargetId="figure-306@4.[173,1531,159,1164]" captionTargetPageId="4" captionText="Fig. 3. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, separate plates and parts of articulated head shields. A. Articulated Nu, left Pn and left La plates IG KSC 155/90 in dorsal view. B. Disarticulated plates of the head shield IG KSC 155/150 in dorsal view. C. Part of articulated head shield IG KSC 155/162 in dorsal view. D. Imprint of the ventral wall of trunk armour and some skeletal elements of the head armour IG KSC 155/10-3 in ventral view showing suborbital and infragnathal bones. Abbreviations: AVL, anterior ventro-lateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; ifc1, principal section of infraorbital sensory line; IG, inferognathal; La, lateral plate; Nu, nuchal plate; mp, middle pit-line groove; Pn, paranuchal plate; Pp, postpineal plate; Prm, premedian plate; SO, suborbital plate; socc, supraoccipital cross-commissural pit-line groove." figureDoi="http://doi.org/10.5281/zenodo.10980827" httpUri="https://zenodo.org/record/10980827/files/figure.png" pageId="6" pageNumber="103">Fig. 3C</figureCitation>
) and IG KSC 155/45 representing individual of moderate size; it is relatively small as in other subspecies of
<taxonomicName id="2EF740AF4C51FFE6FBB7878DA4CAF7CE" authorityName="Traquair" authorityYear="1893" box="[1093,1257,1739,1764]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="103" phylum="Chordata" rank="species" species="leptocheira">
<emphasis id="DB83E73E4C51FFE6FBB7878DA4CAF7CE" box="[1093,1257,1739,1764]" italics="true" pageId="6" pageNumber="103">B. leptocheira</emphasis>
</taxonomicName>
, and with L/B index of 0.53. The preorbital recess (prh) is of trifid
<typeStatus id="364C858E4C51FFE6FAAF87AAA5B1F62F" box="[1373,1426,1772,1797]" pageId="6" pageNumber="103">type</typeStatus>
, with the lateral horns less extended laterally than these in
<taxonomicName id="2EF740AF4C51FFE6FA74864AA5DCF60F" authorityName="Gross" authorityYear="1933" box="[1414,1535,1804,1829]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="103" phylum="Chordata" rank="species" species="maxima">
<emphasis id="DB83E73E4C51FFE6FA74864AA5DCF60F" box="[1414,1535,1804,1829]" italics="true" pageId="6" pageNumber="103">B. maxima</emphasis>
</taxonomicName>
or
<taxonomicName id="2EF740AF4C51FFE6FC8E866BA3DDF66C" authorityName="Traquair" authorityYear="1844" box="[892,1022,1837,1862]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="103" phylum="Chordata" rank="species" species="gigantea">
<emphasis id="DB83E73E4C51FFE6FC8E866BA3DDF66C" box="[892,1022,1837,1862]" italics="true" pageId="6" pageNumber="103">B. gigantea</emphasis>
</taxonomicName>
, but more broad at their base (
<figureCitation id="71CC27A94C51FFE6FAA6866BA581F66C" box="[1364,1442,1837,1862]" captionStart="Fig" captionStartId="5.[98,131,1308,1328]" captionTargetBox="[103,1477,164,1280]" captionTargetId="figure-314@5.[99,1480,160,1281]" captionTargetPageId="5" captionText="Fig. 4. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, lateral AG), premedian (H), and left postmarginal (I) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/65 in visceral (D1) and dorsal (D2) views. E. IG KSC 155/66 in dorsal (E1), right lateral (E2), and visceral (E3) views. F. IG KSC 155/69 in dorsal view. G. IG KSC 155/64 in dorsal view. H. IG KSC 155/114 in dorsal (H1) and anterior (H2) views.. Plate disconnected from the partial head shield IG KSC 155/161, in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980829" httpUri="https://zenodo.org/record/10980829/files/figure.png" pageId="6" pageNumber="103">Figs. 4</figureCitation>
,
<figureCitation id="71CC27A94C51FFE6FA5C866BA59DF66C" box="[1454,1470,1837,1862]" captionStart="Fig" captionStartId="6.[151,184,1321,1341]" captionTargetBox="[153,1534,165,1291]" captionTargetId="figure-270@6.[153,1534,165,1294]" captionTargetPageId="6" captionText="Fig. 5. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), lateral (AG, I) and premedian (H) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/74 in dorsal view. E. IG KSC 155/65 in dorsal view. F. IG KSC 155/66 in dorsal (F1) and visceral (F2) views.G. IG KSC 155/69 in dorsal view. H. IG KSC 155/114 in dorsal view. I. IG KSC 155/64 in dorsal view. Abbreviations: a1Sm and a2Sm, anterior and posterior attachment areas on lateral plate for submarginal plate; a.Prl, attachment area on lateral plate for prelateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; ifc1, principal section of infraorbital sensory line; ifc2, branch of infraorbital sensory line diverging on La; nprl, prelateral notch of head shield; p, lateral pit of head shield; prh, preorbital recess of head shield; pr.po, antero-lateral corner of otico-occipital depression; soc, anterior section of the supraorbital sensory line; spg, spiracular groove of head shield." figureDoi="http://doi.org/10.5281/zenodo.10980831" httpUri="https://zenodo.org/record/10980831/files/figure.png" pageId="6" pageNumber="103">5</figureCitation>
). The borders of the otico-occipital depression (ood;
<figureCitation id="71CC27A94C51FFE6FA77860BA5F4F64C" box="[1413,1495,1869,1894]" captionStart="Fig" captionStartId="7.[98,131,1207,1227]" captionTargetBox="[98,1478,163,1188]" captionTargetId="figure-351@7.[98,1478,163,1189]" captionTargetPageId="7" captionText="Fig. 6. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, postpineal AC, E, G), nuchal (DG, I, KM), and paranuchal (H, J, L, M) plates of the head shield. A. IG KSC 155/54-2, disconnected from the Nu IG KSC 155/54-1, in dorsal (A1) and visceral (A2) views. B. IG KSC 155/146 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/155 in dorsal (C1) and visceral C2) views. D. IG KSC 155/58 in dorsal view. E. Articulated plates IG KSC 155/118 in dorsal view. F. IG KSC 155/53 in dorsal (F1), visceral (F2), and posterior (F3) views. G. Articulated plates IG KSC 155/154 in dorsal (G1) and visceral (G2) views. H. IG KSC 155/55 in dorsal (H1) and visceral (H2) views. I. IG KSC 155/54-1 from the same individual as plate in A, in dorsal view. J. IG KSC 155/119 in dorsal view. K. IG KSC 155/56 in dorsal view. L. IG KSC 155/117 in visceral (L1) and dorsal (L2) views. M. IG KSC 155/125 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980833" httpUri="https://zenodo.org/record/10980833/files/figure.png" pageId="6" pageNumber="103">Figs. 6</figureCitation>
,
<figureCitation id="71CC27A94C51FFE6FA15860BA5D5F64C" box="[1511,1526,1869,1894]" captionStart="Fig" captionStartId="8.[862,895,774,794]" captionTargetBox="[864,1534,161,746]" captionTargetId="figure-587@8.[863,1534,161,749]" captionTargetPageId="8" captionText="Fig. 7. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), paranuchal (A) and postpineal and nuchal (B), plates of the head shield. A. IG KSC 155/117 in dorsal (A1) and visceral (A2) views. B. Articulated plates IG KSC 155/154 in dorsal (B1) and visceral (B2) views.Abbreviations: cr.o, median occipital crest of head shield; cr.pm, paramarginal crest of head shield; cr.pto, postorbital crest of head shield; cr.tv, transverse nuchal crest of head shield; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; g, paired pits on Pp; ifc1, principal section of infraorbital sensory line; mp, middle pit-line groove; nm, obtected nuchal area of head shield; Nu, nuchal plate; ood, otico-occipital depression of head shield; Pp, postpineal plate; socc, supraoccipital cross-commissural pit-line groove; sot, supraotic thickening of head shield." figureDoi="http://doi.org/10.5281/zenodo.10980835" httpUri="https://zenodo.org/record/10980835/files/figure.png" pageId="6" pageNumber="103">7</figureCitation>
) on the internal surface of the head shield are well defined by the paramarginal crista (cr.pm) on the paranuchal plates
<figureCitation id="71CC27A94C50FFE7FF9884C8A09AF48D" box="[106,185,1422,1447]" captionStart="Fig" captionStartId="7.[98,131,1207,1227]" captionTargetBox="[98,1478,163,1188]" captionTargetId="figure-351@7.[98,1478,163,1189]" captionTargetPageId="7" captionText="Fig. 6. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, postpineal AC, E, G), nuchal (DG, I, KM), and paranuchal (H, J, L, M) plates of the head shield. A. IG KSC 155/54-2, disconnected from the Nu IG KSC 155/54-1, in dorsal (A1) and visceral (A2) views. B. IG KSC 155/146 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/155 in dorsal (C1) and visceral C2) views. D. IG KSC 155/58 in dorsal view. E. Articulated plates IG KSC 155/118 in dorsal view. F. IG KSC 155/53 in dorsal (F1), visceral (F2), and posterior (F3) views. G. Articulated plates IG KSC 155/154 in dorsal (G1) and visceral (G2) views. H. IG KSC 155/55 in dorsal (H1) and visceral (H2) views. I. IG KSC 155/54-1 from the same individual as plate in A, in dorsal view. J. IG KSC 155/119 in dorsal view. K. IG KSC 155/56 in dorsal view. L. IG KSC 155/117 in visceral (L1) and dorsal (L2) views. M. IG KSC 155/125 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980833" httpUri="https://zenodo.org/record/10980833/files/figure.png" pageId="7" pageNumber="104">Figs. 6</figureCitation>
,
<figureCitation id="71CC27A94C50FFE7FF3484C8A0F5F48D" box="[198,214,1422,1447]" captionStart="Fig" captionStartId="8.[862,895,774,794]" captionTargetBox="[864,1534,161,746]" captionTargetId="figure-587@8.[863,1534,161,749]" captionTargetPageId="8" captionText="Fig. 7. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), paranuchal (A) and postpineal and nuchal (B), plates of the head shield. A. IG KSC 155/117 in dorsal (A1) and visceral (A2) views. B. Articulated plates IG KSC 155/154 in dorsal (B1) and visceral (B2) views.Abbreviations: cr.o, median occipital crest of head shield; cr.pm, paramarginal crest of head shield; cr.pto, postorbital crest of head shield; cr.tv, transverse nuchal crest of head shield; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; g, paired pits on Pp; ifc1, principal section of infraorbital sensory line; mp, middle pit-line groove; nm, obtected nuchal area of head shield; Nu, nuchal plate; ood, otico-occipital depression of head shield; Pp, postpineal plate; socc, supraoccipital cross-commissural pit-line groove; sot, supraotic thickening of head shield." figureDoi="http://doi.org/10.5281/zenodo.10980835" httpUri="https://zenodo.org/record/10980835/files/figure.png" pageId="7" pageNumber="104">7</figureCitation>
), which are rather low on the lateral plates (
<figureCitation id="71CC27A94C50FFE7FD3984C8A051F4ED" captionStart="Fig" captionStartId="5.[98,131,1308,1328]" captionTargetBox="[103,1477,164,1280]" captionTargetId="figure-314@5.[99,1480,160,1281]" captionTargetPageId="5" captionText="Fig. 4. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, lateral AG), premedian (H), and left postmarginal (I) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/65 in visceral (D1) and dorsal (D2) views. E. IG KSC 155/66 in dorsal (E1), right lateral (E2), and visceral (E3) views. F. IG KSC 155/69 in dorsal view. G. IG KSC 155/64 in dorsal view. H. IG KSC 155/114 in dorsal (H1) and anterior (H2) views.. Plate disconnected from the partial head shield IG KSC 155/161, in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980829" httpUri="https://zenodo.org/record/10980829/files/figure.png" pageId="7" pageNumber="104">Figs. 4</figureCitation>
,
<figureCitation id="71CC27A94C50FFE7FF7384E8A0B2F4ED" box="[129,145,1454,1479]" captionStart="Fig" captionStartId="6.[151,184,1321,1341]" captionTargetBox="[153,1534,165,1291]" captionTargetId="figure-270@6.[153,1534,165,1294]" captionTargetPageId="6" captionText="Fig. 5. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), lateral (AG, I) and premedian (H) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/74 in dorsal view. E. IG KSC 155/65 in dorsal view. F. IG KSC 155/66 in dorsal (F1) and visceral (F2) views.G. IG KSC 155/69 in dorsal view. H. IG KSC 155/114 in dorsal view. I. IG KSC 155/64 in dorsal view. Abbreviations: a1Sm and a2Sm, anterior and posterior attachment areas on lateral plate for submarginal plate; a.Prl, attachment area on lateral plate for prelateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; ifc1, principal section of infraorbital sensory line; ifc2, branch of infraorbital sensory line diverging on La; nprl, prelateral notch of head shield; p, lateral pit of head shield; prh, preorbital recess of head shield; pr.po, antero-lateral corner of otico-occipital depression; soc, anterior section of the supraorbital sensory line; spg, spiracular groove of head shield." figureDoi="http://doi.org/10.5281/zenodo.10980831" httpUri="https://zenodo.org/record/10980831/files/figure.png" pageId="7" pageNumber="104">5</figureCitation>
). The antero-lateral corners of the otico-occipital depression (the imprints of the anterior postorbital processes of the endocranium, pr.po;
<figureCitation id="71CC27A94C50FFE7FE6484A8A1C4F72D" box="[406,487,1518,1543]" captionStart="Fig" captionStartId="6.[151,184,1321,1341]" captionTargetBox="[153,1534,165,1291]" captionTargetId="figure-270@6.[153,1534,165,1294]" captionTargetPageId="6" captionText="Fig. 5. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), lateral (AG, I) and premedian (H) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/74 in dorsal view. E. IG KSC 155/65 in dorsal view. F. IG KSC 155/66 in dorsal (F1) and visceral (F2) views.G. IG KSC 155/69 in dorsal view. H. IG KSC 155/114 in dorsal view. I. IG KSC 155/64 in dorsal view. Abbreviations: a1Sm and a2Sm, anterior and posterior attachment areas on lateral plate for submarginal plate; a.Prl, attachment area on lateral plate for prelateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; ifc1, principal section of infraorbital sensory line; ifc2, branch of infraorbital sensory line diverging on La; nprl, prelateral notch of head shield; p, lateral pit of head shield; prh, preorbital recess of head shield; pr.po, antero-lateral corner of otico-occipital depression; soc, anterior section of the supraorbital sensory line; spg, spiracular groove of head shield." figureDoi="http://doi.org/10.5281/zenodo.10980831" httpUri="https://zenodo.org/record/10980831/files/figure.png" pageId="7" pageNumber="104">Fig. 5F</figureCitation>
<figureCitation id="71CC27A94C50FFE7FE1484BFA1CCF720" box="[486,495,1529,1546]" captionStart="Fig" captionStartId="2.[151,184,1333,1353]" captionTargetBox="[152,823,164,1305]" captionTargetId="figure-630@2.[152,604,164,1305]" captionTargetPageId="2" captionText="Fig. 2. Lithology and stratigraphy of the Sosnogorsk locality. “Fish-dolomite”, bed No. 40 is located about 8 m above the base of the section. Abbreviations: D3f, Upper Devonian, Frasnian; D3fm, Upper Devonian, Famennian." figureDoi="http://doi.org/10.5281/zenodo.10980823" httpUri="https://zenodo.org/record/10980823/files/figure.png" pageId="7" pageNumber="104">
<subScript id="757339694C50FFE7FE1484BFA1CCF720" attach="left" box="[486,495,1529,1546]" fontSize="7" pageId="7" pageNumber="104">2</subScript>
</figureCitation>
) are moderately narrow at the base in comparison with these in
<taxonomicName id="2EF740AF4C50FFE7FDDD8748A2B8F70D" baseAuthorityName="Luksevics" baseAuthorityYear="2001" box="[559,667,1550,1575]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="104" phylum="Chordata" rank="species" species="ornata">
<emphasis id="DB83E73E4C50FFE7FDDD8748A2B8F70D" box="[559,667,1550,1575]" italics="true" pageId="7" pageNumber="104">B. ornata</emphasis>
</taxonomicName>
. In most cases the anterior postorbital processes do not extend in front of the anterior margin of the orbital fenestra, contrary to these in Scottish material (
<bibRefCitation id="8D6646DD4C50FFE7FE5E8728A27BF7AD" author="Miles, R. S." box="[428,600,1646,1671]" pageId="7" pageNumber="104" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles 1968: 76</bibRefCitation>
).
</paragraph>
<caption id="BD886BA44C50FFE7FF9085F1A39EF459" ID-DOI="http://doi.org/10.5281/zenodo.10980833" ID-Zenodo-Dep="10980833" httpUri="https://zenodo.org/record/10980833/files/figure.png" pageId="7" pageNumber="104" startId="7.[98,131,1207,1227]" targetBox="[98,1478,163,1188]" targetPageId="7" targetType="figure">
<paragraph id="E9483B2C4C50FFE7FF9085F1A39EF459" blockId="7.[98,1482,1207,1398]" pageId="7" pageNumber="104">
Fig. 6. Antiarchan fish
<taxonomicName id="2EF740AF4C50FFE7FEC285F1A2D8F5E1" authority="(Rohon, 1900)" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" baseAuthorityName="Rohon" baseAuthorityYear="1900" box="[304,763,1207,1227]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="104" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="jeremejevi">
<emphasis id="DB83E73E4C50FFE7FEC285F1A253F5E1" box="[304,624,1207,1227]" italics="true" pageId="7" pageNumber="104">Bothriolepis leptocheira jeremejevi</emphasis>
(
<bibRefCitation id="8D6646DD4C50FFE7FD8985F1A2D0F5E1" author="Rohon, J. V." box="[635,755,1207,1227]" pageId="7" pageNumber="104" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon, 1900</bibRefCitation>
)
</taxonomicName>
, Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, postpineal AC, E, G), nuchal (DG, I, KM), and paranuchal (H, J, L, M) plates of the head shield.
<emphasis id="DB83E73E4C50FFE7FC4A8597A3EBF5CC" bold="true" box="[952,968,1233,1254]" pageId="7" pageNumber="104">A</emphasis>
. IG KSC 155/54-2, disconnected from the Nu IG KSC 155/54-1, in dorsal (A
<subScript id="757339694C50FFE7FEDA85BEA113F42C" attach="left" box="[296,304,1272,1286]" fontSize="6" pageId="7" pageNumber="104">1</subScript>
) and visceral (A
<subScript id="757339694C50FFE7FE3785BEA1EEF42C" attach="left" box="[453,461,1272,1286]" fontSize="6" pageId="7" pageNumber="104">2</subScript>
) views.
<emphasis id="DB83E73E4C50FFE7FDEB85ABA20BF428" bold="true" box="[537,552,1261,1282]" pageId="7" pageNumber="104">B</emphasis>
. IG KSC 155/146 in dorsal (B
<subScript id="757339694C50FFE7FCCA85BEA363F42C" attach="left" box="[824,832,1272,1286]" fontSize="6" pageId="7" pageNumber="104">1</subScript>
) and visceral (B
<subScript id="757339694C50FFE7FC2685BEA3FFF42C" attach="left" box="[980,988,1272,1286]" fontSize="6" pageId="7" pageNumber="104">2</subScript>
) views.
<emphasis id="DB83E73E4C50FFE7FBDA85ABA41BF429" bold="true" box="[1064,1080,1261,1283]" pageId="7" pageNumber="104">C</emphasis>
. IG KSC 155/155 in dorsal (C
<subScript id="757339694C50FFE7FABB85BEA572F42C" attach="left" box="[1353,1361,1272,1286]" fontSize="6" pageId="7" pageNumber="104">1</subScript>
) and visceral C
<subScript id="757339694C50FFE7FF8A8452A0A3F408" attach="left" box="[120,128,1300,1314]" fontSize="6" pageId="7" pageNumber="104">2</subScript>
) views.
<emphasis id="DB83E73E4C50FFE7FF3C844FA0FDF434" bold="true" box="[206,222,1289,1310]" pageId="7" pageNumber="104">D</emphasis>
. IG KSC 155/58 in dorsal view.
<emphasis id="DB83E73E4C50FFE7FDFE844FA238F434" bold="true" box="[524,539,1289,1310]" pageId="7" pageNumber="104">E</emphasis>
. Articulated plates IG KSC 155/118 in dorsal view.
<emphasis id="DB83E73E4C50FFE7FC0A844FA425F434" bold="true" box="[1016,1030,1289,1310]" pageId="7" pageNumber="104">F</emphasis>
. IG KSC 155/53 in dorsal (F
<subScript id="757339694C50FFE7FAE38452A53AF408" attach="left" box="[1297,1305,1300,1314]" fontSize="6" pageId="7" pageNumber="104">1</subScript>
), visceral (F
<subScript id="757339694C50FFE7FA7F8452A5B6F408" attach="left" box="[1421,1429,1300,1314]" fontSize="6" pageId="7" pageNumber="104">2</subScript>
), and posterior (F
<subScript id="757339694C50FFE7FF3F8476A0F6F414" attach="left" box="[205,213,1328,1342]" fontSize="6" pageId="7" pageNumber="104">3</subScript>
) views.
<emphasis id="DB83E73E4C50FFE7FED68463A115F411" bold="true" box="[292,310,1317,1339]" pageId="7" pageNumber="104">G</emphasis>
. Articulated plates IG KSC 155/154 in dorsal (G
<subScript id="757339694C50FFE7FD088476A321F414" attach="right" box="[762,770,1328,1342]" fontSize="6" pageId="7" pageNumber="104">1</subScript>
) and visceral (G
<subScript id="757339694C50FFE7FC6E8476A387F414" attach="left" box="[924,932,1328,1342]" fontSize="6" pageId="7" pageNumber="104">2</subScript>
) views.
<emphasis id="DB83E73E4C50FFE7FC068463A425F410" bold="true" box="[1012,1030,1317,1338]" pageId="7" pageNumber="104">H</emphasis>
. IG KSC 155/55 in dorsal (H
<subScript id="757339694C50FFE7FAE58476A53CF414" attach="right" box="[1303,1311,1328,1342]" fontSize="6" pageId="7" pageNumber="104">1</subScript>
) and visceral (H
<subScript id="757339694C50FFE7FA4B8476A5E2F414" attach="left" box="[1465,1473,1328,1342]" fontSize="6" pageId="7" pageNumber="104">2</subScript>
) views.
<emphasis id="DB83E73E4C50FFE7FF508407A088F47C" bold="true" box="[162,171,1345,1366]" pageId="7" pageNumber="104">I</emphasis>
. IG KSC 155/54-1 from the same individual as plate in A, in dorsal view.
<emphasis id="DB83E73E4C50FFE7FCA28407A378F47C" bold="true" box="[848,859,1345,1366]" pageId="7" pageNumber="104">J</emphasis>
. IG KSC 155/119 in dorsal view.
<emphasis id="DB83E73E4C50FFE7FB608407A487F47C" bold="true" box="[1170,1188,1345,1366]" pageId="7" pageNumber="104">K</emphasis>
. IG KSC 155/56 in dorsal view.
<emphasis id="DB83E73E4C50FFE7FF90841BA052F458" bold="true" box="[98,113,1373,1394]" pageId="7" pageNumber="104">L</emphasis>
. IG KSC 155/117 in visceral (L
<subScript id="757339694C50FFE7FE61842EA1B8F45C" attach="left" box="[403,411,1384,1398]" fontSize="6" pageId="7" pageNumber="104">1</subScript>
) and dorsal (L
<subScript id="757339694C50FFE7FDD3842EA20AF45C" attach="left" box="[545,553,1384,1398]" fontSize="6" pageId="7" pageNumber="104">2</subScript>
) views.
<emphasis id="DB83E73E4C50FFE7FD85841BA2AFF458" bold="true" box="[631,652,1373,1394]" pageId="7" pageNumber="104">M</emphasis>
. IG KSC 155/125 in dorsal view.
</paragraph>
</caption>
<paragraph id="E9483B2C4C50FFE7FF7587C8A44FF7ED" blockId="7.[98,771,1422,1959]" lastBlockId="7.[808,1481,1422,1959]" pageId="7" pageNumber="104">
The premedian plate (Prm;
<figureCitation id="71CC27A94C50FFE7FE3187C8A200F78D" box="[451,547,1678,1703]" captionStart="Fig" captionStartId="4.[151,184,1188,1208]" captionTargetBox="[157,1530,160,1164]" captionTargetId="figure-306@4.[173,1531,159,1164]" captionTargetPageId="4" captionText="Fig. 3. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, separate plates and parts of articulated head shields. A. Articulated Nu, left Pn and left La plates IG KSC 155/90 in dorsal view. B. Disarticulated plates of the head shield IG KSC 155/150 in dorsal view. C. Part of articulated head shield IG KSC 155/162 in dorsal view. D. Imprint of the ventral wall of trunk armour and some skeletal elements of the head armour IG KSC 155/10-3 in ventral view showing suborbital and infragnathal bones. Abbreviations: AVL, anterior ventro-lateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; ifc1, principal section of infraorbital sensory line; IG, inferognathal; La, lateral plate; Nu, nuchal plate; mp, middle pit-line groove; Pn, paranuchal plate; Pp, postpineal plate; Prm, premedian plate; SO, suborbital plate; socc, supraoccipital cross-commissural pit-line groove." figureDoi="http://doi.org/10.5281/zenodo.10980827" httpUri="https://zenodo.org/record/10980827/files/figure.png" pageId="7" pageNumber="104">Figs. 3C</figureCitation>
,
<figureCitation id="71CC27A94C50FFE7FDDD87C8A277F78D" box="[559,596,1678,1703]" captionStart="Fig" captionStartId="5.[98,131,1308,1328]" captionTargetBox="[103,1477,164,1280]" captionTargetId="figure-314@5.[99,1480,160,1281]" captionTargetPageId="5" captionText="Fig. 4. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, lateral AG), premedian (H), and left postmarginal (I) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/65 in visceral (D1) and dorsal (D2) views. E. IG KSC 155/66 in dorsal (E1), right lateral (E2), and visceral (E3) views. F. IG KSC 155/69 in dorsal view. G. IG KSC 155/64 in dorsal view. H. IG KSC 155/114 in dorsal (H1) and anterior (H2) views.. Plate disconnected from the partial head shield IG KSC 155/161, in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980829" httpUri="https://zenodo.org/record/10980829/files/figure.png" pageId="7" pageNumber="104">4H</figureCitation>
,
<figureCitation id="71CC27A94C50FFE7FD9287C8A2A0F78D" box="[608,643,1678,1703]" captionStart="Fig" captionStartId="6.[151,184,1321,1341]" captionTargetBox="[153,1534,165,1291]" captionTargetId="figure-270@6.[153,1534,165,1294]" captionTargetPageId="6" captionText="Fig. 5. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), lateral (AG, I) and premedian (H) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/74 in dorsal view. E. IG KSC 155/65 in dorsal view. F. IG KSC 155/66 in dorsal (F1) and visceral (F2) views.G. IG KSC 155/69 in dorsal view. H. IG KSC 155/114 in dorsal view. I. IG KSC 155/64 in dorsal view. Abbreviations: a1Sm and a2Sm, anterior and posterior attachment areas on lateral plate for submarginal plate; a.Prl, attachment area on lateral plate for prelateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; ifc1, principal section of infraorbital sensory line; ifc2, branch of infraorbital sensory line diverging on La; nprl, prelateral notch of head shield; p, lateral pit of head shield; prh, preorbital recess of head shield; pr.po, antero-lateral corner of otico-occipital depression; soc, anterior section of the supraorbital sensory line; spg, spiracular groove of head shield." figureDoi="http://doi.org/10.5281/zenodo.10980831" httpUri="https://zenodo.org/record/10980831/files/figure.png" pageId="7" pageNumber="104">5H</figureCitation>
) is slightly wider than long, with a B/L index of 1.071.37. It is slightly narrower than the Prm of
<taxonomicName id="2EF740AF4C50FFE7FE758788A204F7CD" authorityName="Traquair" authorityYear="1893" box="[391,551,1742,1767]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="104" phylum="Chordata" rank="species" species="leptocheira">
<emphasis id="DB83E73E4C50FFE7FE758788A204F7CD" box="[391,551,1742,1767]" italics="true" pageId="7" pageNumber="104">B. leptocheira</emphasis>
</taxonomicName>
from
<collectingCountry id="91E07BBC4C50FFE7FD9E8788A297F7CD" box="[620,692,1742,1767]" name="Latvia" pageId="7" pageNumber="104">Latvia</collectingCountry>
(1.23 1.48;
<bibRefCitation id="8D6646DD4C50FFE7FF5787A8A17DF62D" author="Luksevics, E." box="[165,350,1774,1799]" pageId="7" pageNumber="104" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
), but approximately coincides with proportions of Prm in Scottish material of
<taxonomicName id="2EF740AF4C50FFE7FDAC8648A101F66D" authority="Miles [1968: 76]" authorityName="Miles" authorityPageNumber="76" authorityYear="1968" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="104" phylum="Chordata" rank="species" species="leptocheira">
<emphasis id="DB83E73E4C50FFE7FDAC8648A320F60D" box="[606,771,1806,1831]" italics="true" pageId="7" pageNumber="104">B. leptocheira</emphasis>
<bibRefCitation id="8D6646DD4C50FFE7FF9B8668A101F66D" author="Miles, R. S." box="[105,290,1838,1863]" pageId="7" pageNumber="104" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles [1968: 76]</bibRefCitation>
</taxonomicName>
has mentioned B/L index of 1.14, 1.20, and 1.28). Prm is weakly arched in a lateral direction, but rather strongly vaulted in the parasagittal section with curvature in the anterior most part. The rostral margin is convex, it is 2.53.2 times longer than the slightly concave or straight orbital margin, similar to
<taxonomicName id="2EF740AF4C50FFE7FBA984E8A4DCF4ED" authorityName="Miles" authorityYear="1968" box="[1115,1279,1454,1479]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="104" phylum="Chordata" rank="species" species="leptocheira">
<emphasis id="DB83E73E4C50FFE7FBA984E8A4DCF4ED" box="[1115,1279,1454,1479]" italics="true" pageId="7" pageNumber="104">B. leptocheira</emphasis>
</taxonomicName>
from
<collectingCountry id="91E07BBC4C50FFE7FABE84E8A5B2F4ED" box="[1356,1425,1454,1479]" name="Latvia" pageId="7" pageNumber="104">Latvia</collectingCountry>
and
<collectingCountry id="91E07BBC4C50FFE7FCDA8488A3A4F4CD" box="[808,903,1486,1511]" name="United Kingdom" pageId="7" pageNumber="104">Scotland</collectingCountry>
(for comparison see
<bibRefCitation id="8D6646DD4C50FFE7FB768488A518F4CD" author="Luksevics, E." box="[1156,1339,1486,1511]" pageId="7" pageNumber="104" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
: figs. 3234 and
<bibRefCitation id="8D6646DD4C50FFE7FCAF84A8A3C7F72D" author="Miles, R. S." box="[861,996,1518,1543]" pageId="7" pageNumber="104" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles 1968</bibRefCitation>
: pl. 20: 1, 2), and bears slightly defined median rostral projection (
<collectionCode id="8FE6A3E94C50FFE7FBAB8748A45BF70D" box="[1113,1144,1550,1575]" country="China" name="Institute of Geology" pageId="7" pageNumber="104">IG</collectionCode>
KSC 155/114;
<figureCitation id="71CC27A94C50FFE7FAD78748A559F70D" box="[1317,1402,1550,1575]" captionStart="Fig" captionStartId="5.[98,131,1308,1328]" captionTargetBox="[103,1477,164,1280]" captionTargetId="figure-314@5.[99,1480,160,1281]" captionTargetPageId="5" captionText="Fig. 4. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, lateral AG), premedian (H), and left postmarginal (I) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/65 in visceral (D1) and dorsal (D2) views. E. IG KSC 155/66 in dorsal (E1), right lateral (E2), and visceral (E3) views. F. IG KSC 155/69 in dorsal view. G. IG KSC 155/64 in dorsal view. H. IG KSC 155/114 in dorsal (H1) and anterior (H2) views.. Plate disconnected from the partial head shield IG KSC 155/161, in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980829" httpUri="https://zenodo.org/record/10980829/files/figure.png" pageId="7" pageNumber="104">Fig. 4H</figureCitation>
<figureCitation id="71CC27A94C50FFE7FA89875FA5A7F700" box="[1403,1412,1561,1578]" captionStart="Fig" captionStartId="2.[151,184,1333,1353]" captionTargetBox="[152,823,164,1305]" captionTargetId="figure-630@2.[152,604,164,1305]" captionTargetPageId="2" captionText="Fig. 2. Lithology and stratigraphy of the Sosnogorsk locality. “Fish-dolomite”, bed No. 40 is located about 8 m above the base of the section. Abbreviations: D3f, Upper Devonian, Frasnian; D3fm, Upper Devonian, Famennian." figureDoi="http://doi.org/10.5281/zenodo.10980823" httpUri="https://zenodo.org/record/10980823/files/figure.png" pageId="7" pageNumber="104">
<subScript id="757339694C50FFE7FA89875FA5A7F700" attach="right" box="[1403,1412,1561,1578]" fontSize="7" pageId="7" pageNumber="104">2</subScript>
</figureCitation>
) seen only in anterior view. Also the position of the infraorbital sensory groove (ifc
<subScript id="757339694C50FFE7FC0C871FA424F740" attach="left" box="[1022,1031,1625,1642]" fontSize="7" pageId="7" pageNumber="104">1</subScript>
) that crosses the plate close to its rostral margin and the well-defined anterior section of the supraorbital sensory line (soc) are similar characteristics in all three subspecies of
<taxonomicName id="2EF740AF4C50FFE7FC3787E8A446F7ED" authorityName="Miles" authorityYear="1968" box="[965,1125,1710,1735]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="104" phylum="Chordata" rank="species" species="leptocheira">
<emphasis id="DB83E73E4C50FFE7FC3787E8A446F7ED" box="[965,1125,1710,1735]" italics="true" pageId="7" pageNumber="104">B. leptocheira</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="E9483B2C4C50FFE8FCBC8788A307F234" blockId="7.[808,1481,1422,1959]" lastBlockId="8.[151,825,160,1960]" lastPageId="8" lastPageNumber="105" pageId="7" pageNumber="104">
The lateral plate (La;
<figureCitation id="71CC27A94C50FFE7FBC98788A4A6F7CD" box="[1083,1157,1742,1767]" captionStart="Fig" captionStartId="4.[151,184,1188,1208]" captionTargetBox="[157,1530,160,1164]" captionTargetId="figure-306@4.[173,1531,159,1164]" captionTargetPageId="4" captionText="Fig. 3. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, separate plates and parts of articulated head shields. A. Articulated Nu, left Pn and left La plates IG KSC 155/90 in dorsal view. B. Disarticulated plates of the head shield IG KSC 155/150 in dorsal view. C. Part of articulated head shield IG KSC 155/162 in dorsal view. D. Imprint of the ventral wall of trunk armour and some skeletal elements of the head armour IG KSC 155/10-3 in ventral view showing suborbital and infragnathal bones. Abbreviations: AVL, anterior ventro-lateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; ifc1, principal section of infraorbital sensory line; IG, inferognathal; La, lateral plate; Nu, nuchal plate; mp, middle pit-line groove; Pn, paranuchal plate; Pp, postpineal plate; Prm, premedian plate; SO, suborbital plate; socc, supraoccipital cross-commissural pit-line groove." figureDoi="http://doi.org/10.5281/zenodo.10980827" httpUri="https://zenodo.org/record/10980827/files/figure.png" pageId="7" pageNumber="104">Figs. 3</figureCitation>
,
<figureCitation id="71CC27A94C50FFE7FB628788A4FBF7CD" box="[1168,1240,1742,1767]" captionStart="Fig" captionStartId="5.[98,131,1308,1328]" captionTargetBox="[103,1477,164,1280]" captionTargetId="figure-314@5.[99,1480,160,1281]" captionTargetPageId="5" captionText="Fig. 4. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, lateral AG), premedian (H), and left postmarginal (I) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/65 in visceral (D1) and dorsal (D2) views. E. IG KSC 155/66 in dorsal (E1), right lateral (E2), and visceral (E3) views. F. IG KSC 155/69 in dorsal view. G. IG KSC 155/64 in dorsal view. H. IG KSC 155/114 in dorsal (H1) and anterior (H2) views.. Plate disconnected from the partial head shield IG KSC 155/161, in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980829" httpUri="https://zenodo.org/record/10980829/files/figure.png" pageId="7" pageNumber="104">4AG</figureCitation>
,
<figureCitation id="71CC27A94C50FFE7FB138788A518F7CD" box="[1249,1339,1742,1767]" captionStart="Fig" captionStartId="6.[151,184,1321,1341]" captionTargetBox="[153,1534,165,1291]" captionTargetId="figure-270@6.[153,1534,165,1294]" captionTargetPageId="6" captionText="Fig. 5. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), lateral (AG, I) and premedian (H) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/74 in dorsal view. E. IG KSC 155/65 in dorsal view. F. IG KSC 155/66 in dorsal (F1) and visceral (F2) views.G. IG KSC 155/69 in dorsal view. H. IG KSC 155/114 in dorsal view. I. IG KSC 155/64 in dorsal view. Abbreviations: a1Sm and a2Sm, anterior and posterior attachment areas on lateral plate for submarginal plate; a.Prl, attachment area on lateral plate for prelateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; ifc1, principal section of infraorbital sensory line; ifc2, branch of infraorbital sensory line diverging on La; nprl, prelateral notch of head shield; p, lateral pit of head shield; prh, preorbital recess of head shield; pr.po, antero-lateral corner of otico-occipital depression; soc, anterior section of the supraorbital sensory line; spg, spiracular groove of head shield." figureDoi="http://doi.org/10.5281/zenodo.10980831" httpUri="https://zenodo.org/record/10980831/files/figure.png" pageId="7" pageNumber="104">5AG, I</figureCitation>
) is short and broad with an overall shape and proportions resembling these of
<taxonomicName id="2EF740AF4C50FFE7FCB58648A471F60D" authorityName="Gross" authorityYear="1942" box="[839,1106,1806,1831]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="104" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="curonica">
<emphasis id="DB83E73E4C50FFE7FCB58648A471F60D" box="[839,1106,1806,1831]" italics="true" pageId="7" pageNumber="104">B. leptocheira curonica</emphasis>
</taxonomicName>
. The infraorbital sensory groove crosses the plate in its anterior part, not far from the anterior margin and along its lateral margins. The path of the sensory groove significantly varies in various specimens, e.g., on La plate IG KSC 155/15 (
<figureCitation id="71CC27A94C50FFE7FBD686C8A4ABF68D" box="[1060,1160,1934,1959]" captionStart="Fig" captionStartId="5.[98,131,1308,1328]" captionTargetBox="[103,1477,164,1280]" captionTargetId="figure-314@5.[99,1480,160,1281]" captionTargetPageId="5" captionText="Fig. 4. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, lateral AG), premedian (H), and left postmarginal (I) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/65 in visceral (D1) and dorsal (D2) views. E. IG KSC 155/66 in dorsal (E1), right lateral (E2), and visceral (E3) views. F. IG KSC 155/69 in dorsal view. G. IG KSC 155/64 in dorsal view. H. IG KSC 155/114 in dorsal (H1) and anterior (H2) views.. Plate disconnected from the partial head shield IG KSC 155/161, in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980829" httpUri="https://zenodo.org/record/10980829/files/figure.png" pageId="7" pageNumber="104">Figs. 4A</figureCitation>
,
<figureCitation id="71CC27A94C50FFE7FB6786C8A495F68D" box="[1173,1206,1934,1959]" captionStart="Fig" captionStartId="6.[151,184,1321,1341]" captionTargetBox="[153,1534,165,1291]" captionTargetId="figure-270@6.[153,1534,165,1294]" captionTargetPageId="6" captionText="Fig. 5. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), lateral (AG, I) and premedian (H) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/74 in dorsal view. E. IG KSC 155/65 in dorsal view. F. IG KSC 155/66 in dorsal (F1) and visceral (F2) views.G. IG KSC 155/69 in dorsal view. H. IG KSC 155/114 in dorsal view. I. IG KSC 155/64 in dorsal view. Abbreviations: a1Sm and a2Sm, anterior and posterior attachment areas on lateral plate for submarginal plate; a.Prl, attachment area on lateral plate for prelateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; ifc1, principal section of infraorbital sensory line; ifc2, branch of infraorbital sensory line diverging on La; nprl, prelateral notch of head shield; p, lateral pit of head shield; prh, preorbital recess of head shield; pr.po, antero-lateral corner of otico-occipital depression; soc, anterior section of the supraorbital sensory line; spg, spiracular groove of head shield." figureDoi="http://doi.org/10.5281/zenodo.10980831" httpUri="https://zenodo.org/record/10980831/files/figure.png" pageId="7" pageNumber="104">5A</figureCitation>
) it displays a strong deviation from its usual path (
<figureCitation id="71CC27A94C5FFFE8FE3E81E6A267F193" box="[460,580,160,185]" captionStart="Fig" captionStartId="5.[98,131,1308,1328]" captionTargetBox="[103,1477,164,1280]" captionTargetId="figure-314@5.[99,1480,160,1281]" captionTargetPageId="5" captionText="Fig. 4. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, lateral AG), premedian (H), and left postmarginal (I) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/65 in visceral (D1) and dorsal (D2) views. E. IG KSC 155/66 in dorsal (E1), right lateral (E2), and visceral (E3) views. F. IG KSC 155/69 in dorsal view. G. IG KSC 155/64 in dorsal view. H. IG KSC 155/114 in dorsal (H1) and anterior (H2) views.. Plate disconnected from the partial head shield IG KSC 155/161, in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980829" httpUri="https://zenodo.org/record/10980829/files/figure.png" pageId="8" pageNumber="105">Fig. 4BG</figureCitation>
) because it partly follows the course of the semicircular pit-line groove, which is missing in this specimen (
<collectionCode id="8FE6A3E94C5FFFE8FE4781A6A1F7F1D3" box="[437,468,224,249]" country="China" name="Institute of Geology" pageId="8" pageNumber="105">IG</collectionCode>
KSC 155/15). The plate is rather strongly vaulted in its anterior most part in large individuals (
<collectionCode id="8FE6A3E94C5FFFE8FF6D8067A09DF010" box="[159,190,289,314]" country="China" name="Institute of Geology" pageId="8" pageNumber="105">IG</collectionCode>
KSC 155/66;
<figureCitation id="71CC27A94C5FFFE8FEAB8067A18BF010" box="[345,424,289,314]" captionStart="Fig" captionStartId="5.[98,131,1308,1328]" captionTargetBox="[103,1477,164,1280]" captionTargetId="figure-314@5.[99,1480,160,1281]" captionTargetPageId="5" captionText="Fig. 4. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, lateral AG), premedian (H), and left postmarginal (I) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/65 in visceral (D1) and dorsal (D2) views. E. IG KSC 155/66 in dorsal (E1), right lateral (E2), and visceral (E3) views. F. IG KSC 155/69 in dorsal view. G. IG KSC 155/64 in dorsal view. H. IG KSC 155/114 in dorsal (H1) and anterior (H2) views.. Plate disconnected from the partial head shield IG KSC 155/161, in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980829" httpUri="https://zenodo.org/record/10980829/files/figure.png" pageId="8" pageNumber="105">Fig. 4E</figureCitation>
<figureCitation id="71CC27A94C5FFFE8FE5A806AA192F017" box="[424,433,300,317]" captionStart="Fig" captionStartId="2.[151,184,1333,1353]" captionTargetBox="[152,823,164,1305]" captionTargetId="figure-630@2.[152,604,164,1305]" captionTargetPageId="2" captionText="Fig. 2. Lithology and stratigraphy of the Sosnogorsk locality. “Fish-dolomite”, bed No. 40 is located about 8 m above the base of the section. Abbreviations: D3f, Upper Devonian, Frasnian; D3fm, Upper Devonian, Famennian." figureDoi="http://doi.org/10.5281/zenodo.10980823" httpUri="https://zenodo.org/record/10980823/files/figure.png" pageId="8" pageNumber="105">
<subScript id="757339694C5FFFE8FE5A806AA192F017" attach="left" box="[424,433,300,317]" fontSize="7" pageId="8" pageNumber="105">2</subScript>
</figureCitation>
), but almost flat in small individuals (
<collectionCode id="8FE6A3E94C5FFFE8FF348007A0C6F070" box="[198,229,321,346]" country="China" name="Institute of Geology" pageId="8" pageNumber="105">IG</collectionCode>
KSC 155/69;
<figureCitation id="71CC27A94C5FFFE8FE718007A1F5F070" box="[387,470,321,346]" captionStart="Fig" captionStartId="5.[98,131,1308,1328]" captionTargetBox="[103,1477,164,1280]" captionTargetId="figure-314@5.[99,1480,160,1281]" captionTargetPageId="5" captionText="Fig. 4. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, lateral AG), premedian (H), and left postmarginal (I) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/65 in visceral (D1) and dorsal (D2) views. E. IG KSC 155/66 in dorsal (E1), right lateral (E2), and visceral (E3) views. F. IG KSC 155/69 in dorsal view. G. IG KSC 155/64 in dorsal view. H. IG KSC 155/114 in dorsal (H1) and anterior (H2) views.. Plate disconnected from the partial head shield IG KSC 155/161, in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980829" httpUri="https://zenodo.org/record/10980829/files/figure.png" pageId="8" pageNumber="105">Fig. 4F</figureCitation>
). A moderately small, but deep pit (p) is positioned just posterior to the spiracular groove (spg;
<figureCitation id="71CC27A94C5FFFE8FF2180C4A101F0B1" box="[211,290,386,411]" captionStart="Fig" captionStartId="6.[151,184,1321,1341]" captionTargetBox="[153,1534,165,1291]" captionTargetId="figure-270@6.[153,1534,165,1294]" captionTargetPageId="6" captionText="Fig. 5. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), lateral (AG, I) and premedian (H) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/74 in dorsal view. E. IG KSC 155/65 in dorsal view. F. IG KSC 155/66 in dorsal (F1) and visceral (F2) views.G. IG KSC 155/69 in dorsal view. H. IG KSC 155/114 in dorsal view. I. IG KSC 155/64 in dorsal view. Abbreviations: a1Sm and a2Sm, anterior and posterior attachment areas on lateral plate for submarginal plate; a.Prl, attachment area on lateral plate for prelateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; ifc1, principal section of infraorbital sensory line; ifc2, branch of infraorbital sensory line diverging on La; nprl, prelateral notch of head shield; p, lateral pit of head shield; prh, preorbital recess of head shield; pr.po, antero-lateral corner of otico-occipital depression; soc, anterior section of the supraorbital sensory line; spg, spiracular groove of head shield." figureDoi="http://doi.org/10.5281/zenodo.10980831" httpUri="https://zenodo.org/record/10980831/files/figure.png" pageId="8" pageNumber="105">Fig. 5F</figureCitation>
<figureCitation id="71CC27A94C5FFFE8FED380CBA109F0B4" box="[289,298,397,414]" captionStart="Fig" captionStartId="2.[151,184,1333,1353]" captionTargetBox="[152,823,164,1305]" captionTargetId="figure-630@2.[152,604,164,1305]" captionTargetPageId="2" captionText="Fig. 2. Lithology and stratigraphy of the Sosnogorsk locality. “Fish-dolomite”, bed No. 40 is located about 8 m above the base of the section. Abbreviations: D3f, Upper Devonian, Frasnian; D3fm, Upper Devonian, Famennian." figureDoi="http://doi.org/10.5281/zenodo.10980823" httpUri="https://zenodo.org/record/10980823/files/figure.png" pageId="8" pageNumber="105">
<subScript id="757339694C5FFFE8FED380CBA109F0B4" attach="left" box="[289,298,397,414]" fontSize="7" pageId="8" pageNumber="105">2</subScript>
</figureCitation>
). The spiracular groove is moderately long and shallow. It is widest at the lateral margin of the head shield and narrows only slightly close to the pit. The anterior and posterior crests delimiting the spiracular groove are well developed along their entire extent. The anterior attachment for the submarginal plate (a
<subScript id="757339694C5FFFE8FE568368A18EF315" attach="both" box="[420,429,558,575]" fontSize="7" pageId="8" pageNumber="105">1</subScript>
Sm;
<figureCitation id="71CC27A94C5FFFE8FE138365A213F316" box="[481,560,547,572]" captionStart="Fig" captionStartId="6.[151,184,1321,1341]" captionTargetBox="[153,1534,165,1291]" captionTargetId="figure-270@6.[153,1534,165,1294]" captionTargetPageId="6" captionText="Fig. 5. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), lateral (AG, I) and premedian (H) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/74 in dorsal view. E. IG KSC 155/65 in dorsal view. F. IG KSC 155/66 in dorsal (F1) and visceral (F2) views.G. IG KSC 155/69 in dorsal view. H. IG KSC 155/114 in dorsal view. I. IG KSC 155/64 in dorsal view. Abbreviations: a1Sm and a2Sm, anterior and posterior attachment areas on lateral plate for submarginal plate; a.Prl, attachment area on lateral plate for prelateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; ifc1, principal section of infraorbital sensory line; ifc2, branch of infraorbital sensory line diverging on La; nprl, prelateral notch of head shield; p, lateral pit of head shield; prh, preorbital recess of head shield; pr.po, antero-lateral corner of otico-occipital depression; soc, anterior section of the supraorbital sensory line; spg, spiracular groove of head shield." figureDoi="http://doi.org/10.5281/zenodo.10980831" httpUri="https://zenodo.org/record/10980831/files/figure.png" pageId="8" pageNumber="105">Fig. 5F</figureCitation>
<figureCitation id="71CC27A94C5FFFE8FDC28368A21AF315" box="[560,569,558,575]" captionStart="Fig" captionStartId="2.[151,184,1333,1353]" captionTargetBox="[152,823,164,1305]" captionTargetId="figure-630@2.[152,604,164,1305]" captionTargetPageId="2" captionText="Fig. 2. Lithology and stratigraphy of the Sosnogorsk locality. “Fish-dolomite”, bed No. 40 is located about 8 m above the base of the section. Abbreviations: D3f, Upper Devonian, Frasnian; D3fm, Upper Devonian, Famennian." figureDoi="http://doi.org/10.5281/zenodo.10980823" httpUri="https://zenodo.org/record/10980823/files/figure.png" pageId="8" pageNumber="105">
<subScript id="757339694C5FFFE8FDC28368A21AF315" attach="left" box="[560,569,558,575]" fontSize="7" pageId="8" pageNumber="105">2</subScript>
</figureCitation>
) is strongly developed transverse structure, which borders the anterior crest of the spiracular groove. This attachment is very close to the more anteriorly located triangular attachment area for the prelateral plate (a.PrL;
<figureCitation id="71CC27A94C5FFFE8FEA983E2A188F397" box="[347,427,676,701]" captionStart="Fig" captionStartId="6.[151,184,1321,1341]" captionTargetBox="[153,1534,165,1291]" captionTargetId="figure-270@6.[153,1534,165,1294]" captionTargetPageId="6" captionText="Fig. 5. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), lateral (AG, I) and premedian (H) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/74 in dorsal view. E. IG KSC 155/65 in dorsal view. F. IG KSC 155/66 in dorsal (F1) and visceral (F2) views.G. IG KSC 155/69 in dorsal view. H. IG KSC 155/114 in dorsal view. I. IG KSC 155/64 in dorsal view. Abbreviations: a1Sm and a2Sm, anterior and posterior attachment areas on lateral plate for submarginal plate; a.Prl, attachment area on lateral plate for prelateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; ifc1, principal section of infraorbital sensory line; ifc2, branch of infraorbital sensory line diverging on La; nprl, prelateral notch of head shield; p, lateral pit of head shield; prh, preorbital recess of head shield; pr.po, antero-lateral corner of otico-occipital depression; soc, anterior section of the supraorbital sensory line; spg, spiracular groove of head shield." figureDoi="http://doi.org/10.5281/zenodo.10980831" httpUri="https://zenodo.org/record/10980831/files/figure.png" pageId="8" pageNumber="105">Fig. 5F</figureCitation>
<figureCitation id="71CC27A94C5FFFE8FE5883E9A190F3EA" box="[426,435,687,704]" captionStart="Fig" captionStartId="2.[151,184,1333,1353]" captionTargetBox="[152,823,164,1305]" captionTargetId="figure-630@2.[152,604,164,1305]" captionTargetPageId="2" captionText="Fig. 2. Lithology and stratigraphy of the Sosnogorsk locality. “Fish-dolomite”, bed No. 40 is located about 8 m above the base of the section. Abbreviations: D3f, Upper Devonian, Frasnian; D3fm, Upper Devonian, Famennian." figureDoi="http://doi.org/10.5281/zenodo.10980823" httpUri="https://zenodo.org/record/10980823/files/figure.png" pageId="8" pageNumber="105">
<subScript id="757339694C5FFFE8FE5883E9A190F3EA" attach="left" box="[426,435,687,704]" fontSize="7" pageId="8" pageNumber="105">2</subScript>
</figureCitation>
). The posterior attachment for the submarginal plate (a
<subScript id="757339694C5FFFE8FE8C8389A1ABF3CA" attach="both" box="[382,392,719,736]" fontSize="7" pageId="8" pageNumber="105">2</subScript>
Sm) constitutes an elongate boss (
<figureCitation id="71CC27A94C5FFFE8FCFE8382A096F3D4" captionStart="Fig" captionStartId="6.[151,184,1321,1341]" captionTargetBox="[153,1534,165,1291]" captionTargetId="figure-270@6.[153,1534,165,1294]" captionTargetPageId="6" captionText="Fig. 5. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), lateral (AG, I) and premedian (H) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/74 in dorsal view. E. IG KSC 155/65 in dorsal view. F. IG KSC 155/66 in dorsal (F1) and visceral (F2) views.G. IG KSC 155/69 in dorsal view. H. IG KSC 155/114 in dorsal view. I. IG KSC 155/64 in dorsal view. Abbreviations: a1Sm and a2Sm, anterior and posterior attachment areas on lateral plate for submarginal plate; a.Prl, attachment area on lateral plate for prelateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; ifc1, principal section of infraorbital sensory line; ifc2, branch of infraorbital sensory line diverging on La; nprl, prelateral notch of head shield; p, lateral pit of head shield; prh, preorbital recess of head shield; pr.po, antero-lateral corner of otico-occipital depression; soc, anterior section of the supraorbital sensory line; spg, spiracular groove of head shield." figureDoi="http://doi.org/10.5281/zenodo.10980831" httpUri="https://zenodo.org/record/10980831/files/figure.png" pageId="8" pageNumber="105">Fig. 5F</figureCitation>
<figureCitation id="71CC27A94C5FFFE8FF4783B6A09DF22B" box="[181,190,752,769]" captionStart="Fig" captionStartId="2.[151,184,1333,1353]" captionTargetBox="[152,823,164,1305]" captionTargetId="figure-630@2.[152,604,164,1305]" captionTargetPageId="2" captionText="Fig. 2. Lithology and stratigraphy of the Sosnogorsk locality. “Fish-dolomite”, bed No. 40 is located about 8 m above the base of the section. Abbreviations: D3f, Upper Devonian, Frasnian; D3fm, Upper Devonian, Famennian." figureDoi="http://doi.org/10.5281/zenodo.10980823" httpUri="https://zenodo.org/record/10980823/files/figure.png" pageId="8" pageNumber="105">
<subScript id="757339694C5FFFE8FF4783B6A09DF22B" attach="left" box="[181,190,752,769]" fontSize="7" pageId="8" pageNumber="105">2</subScript>
</figureCitation>
), widening posteriorly. The attachment reaches anteriorly to the posterior crest delimiting the spiracular groove.
</paragraph>
<caption id="BD886BA44C5FFFE8FCAC8240A52EF564" ID-DOI="http://doi.org/10.5281/zenodo.10980835" ID-Zenodo-Dep="10980835" httpUri="https://zenodo.org/record/10980835/files/figure.png" pageId="8" pageNumber="105" startId="8.[862,895,774,794]" targetBox="[864,1534,161,746]" targetPageId="8" targetType="figure">
<paragraph id="E9483B2C4C5FFFE8FCAC8240A52EF564" blockId="8.[862,1536,774,1102]" pageId="8" pageNumber="105">
Fig. 7. Antiarchan fish
<taxonomicName id="2EF740AF4C5FFFE8FBC28240A5D8F230" authority="(Rohon, 1900)" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" baseAuthorityName="Rohon" baseAuthorityYear="1900" box="[1072,1531,774,794]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="105" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="jeremejevi">
<emphasis id="DB83E73E4C5FFFE8FBC28240A54CF230" box="[1072,1391,774,794]" italics="true" pageId="8" pageNumber="105">Bothriolepis leptocheira jeremejevi</emphasis>
(
<bibRefCitation id="8D6646DD4C5FFFE8FA898240A5D6F230" author="Rohon, J. V." box="[1403,1525,774,794]" pageId="8" pageNumber="105" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon, 1900</bibRefCitation>
)
</taxonomicName>
, paranuchal (A) and postpineal and nuchal (B), plates of the head shield.
<emphasis id="DB83E73E4C5FFFE8FCAC827BA34DF278" bold="true" box="[862,878,829,850]" pageId="8" pageNumber="105">A</emphasis>
. IG KSC 155/117 in dorsal (A
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) and visceral (A
<subScript id="757339694C5FFFE8FADB8201A512F27F" attach="left" box="[1321,1329,839,853]" fontSize="6" pageId="8" pageNumber="105">2</subScript>
) views.
<emphasis id="DB83E73E4C5FFFE8FA72827BA5ACF278" bold="true" box="[1408,1423,829,850]" pageId="8" pageNumber="105">B</emphasis>
. Articulated plates IG KSC 155/154 in dorsal (B
<subScript id="757339694C5FFFE8FB618225A4B8F25B" attach="left" box="[1171,1179,867,881]" fontSize="6" pageId="8" pageNumber="105">1</subScript>
) and visceral (B
<subScript id="757339694C5FFFE8FAD88225A511F25B" attach="left" box="[1322,1330,867,881]" fontSize="6" pageId="8" pageNumber="105">2</subScript>
) views.Abbreviations: cr.o, median occipital crest of head shield; cr.pm, paramarginal crest of head shield; cr.pto, postorbital crest of head shield; cr.tv, transverse nuchal crest of head shield; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; g, paired pits on Pp; ifc
<subScript id="757339694C5FFFE8FB0F8295A526F2CB" attach="left" box="[1277,1285,979,993]" fontSize="6" pageId="8" pageNumber="105">1</subScript>
, principal section of infraorbital sensory line; mp, middle pit-line groove; nm, obtected nuchal area of head shield; Nu, nuchal plate; ood, otico-occipital depression of head shield; Pp, postpineal plate; socc, supraoccipital cross-commissural pit-line groove; sot, supraotic thickening of head shield.
</paragraph>
</caption>
<paragraph id="E9483B2C4C5FFFE8FF4F8263A286F58B" blockId="8.[151,825,160,1960]" pageId="8" pageNumber="105">
The postpineal plate (Pp;
<figureCitation id="71CC27A94C5FFFE8FE208263A214F214" box="[466,567,805,830]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="7.[98,131,1207,1227]" captionStartId-1="8.[862,895,774,794]" captionStartId-2="8.[863,896,1656,1676]" captionTargetBox-0="[98,1478,163,1188]" captionTargetBox-1="[864,1534,161,746]" captionTargetBox-2="[891,1506,1125,1629]" captionTargetId-0="figure-351@7.[98,1478,163,1189]" captionTargetId-1="figure-587@8.[863,1534,161,749]" captionTargetId-2="figure-600@8.[891,1506,1125,1630]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="8" captionText-0="Fig. 6. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, postpineal AC, E, G), nuchal (DG, I, KM), and paranuchal (H, J, L, M) plates of the head shield. A. IG KSC 155/54-2, disconnected from the Nu IG KSC 155/54-1, in dorsal (A1) and visceral (A2) views. B. IG KSC 155/146 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/155 in dorsal (C1) and visceral C2) views. D. IG KSC 155/58 in dorsal view. E. Articulated plates IG KSC 155/118 in dorsal view. F. IG KSC 155/53 in dorsal (F1), visceral (F2), and posterior (F3) views. G. Articulated plates IG KSC 155/154 in dorsal (G1) and visceral (G2) views. H. IG KSC 155/55 in dorsal (H1) and visceral (H2) views. I. IG KSC 155/54-1 from the same individual as plate in A, in dorsal view. J. IG KSC 155/119 in dorsal view. K. IG KSC 155/56 in dorsal view. L. IG KSC 155/117 in visceral (L1) and dorsal (L2) views. M. IG KSC 155/125 in dorsal view." captionText-1="Fig. 7. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), paranuchal (A) and postpineal and nuchal (B), plates of the head shield. A. IG KSC 155/117 in dorsal (A1) and visceral (A2) views. B. Articulated plates IG KSC 155/154 in dorsal (B1) and visceral (B2) views.Abbreviations: cr.o, median occipital crest of head shield; cr.pm, paramarginal crest of head shield; cr.pto, postorbital crest of head shield; cr.tv, transverse nuchal crest of head shield; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; g, paired pits on Pp; ifc1, principal section of infraorbital sensory line; mp, middle pit-line groove; nm, obtected nuchal area of head shield; Nu, nuchal plate; ood, otico-occipital depression of head shield; Pp, postpineal plate; socc, supraoccipital cross-commissural pit-line groove; sot, supraotic thickening of head shield." captionText-2="Fig. 8. Reconstruction of the head shield of antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), based on partial head shield (IG KSC 71/1-87), left (IG KSC 155/15), and right (IG KSC 155/65) lateral plates. Abbreviations: cir, semicircular pit-line groove; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; ifc1, principal section of infraorbital sensory line; ifc2, branch of infraorbital sensory line diverging on La; La, lateral plate; mp, middle pit-line groove; Nu, nuchal plate; Pmg, postmarginal plate; Pn, paranuchal plate; Pp, postpineal plate; Prm, premedian plate; prh, preorbital recess of head shield; soc, anterior section of the supraorbital sensory line; socc, supraoccipital cross-commissural pit-line groove." figureDoi-0="http://doi.org/10.5281/zenodo.10980833" figureDoi-1="http://doi.org/10.5281/zenodo.10980835" figureDoi-2="http://doi.org/10.5281/zenodo.10980837" httpUri-0="https://zenodo.org/record/10980833/files/figure.png" httpUri-1="https://zenodo.org/record/10980835/files/figure.png" httpUri-2="https://zenodo.org/record/10980837/files/figure.png" pageId="8" pageNumber="105">Figs. 68</figureCitation>
) is, as usually observed in
<taxonomicName id="2EF740AF4C5FFFE8FF478203A11EF274" authorityName="Eichwald" authorityYear="1840" box="[181,317,837,862]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="105" phylum="Chordata" rank="genus">
<emphasis id="DB83E73E4C5FFFE8FF478203A11EF274" box="[181,317,837,862]" italics="true" pageId="8" pageNumber="105">Bothriolepis</emphasis>
</taxonomicName>
, slightly wider than it is long and has an L/B index varying from 0.96 to 0.97, it is slightly less elongated than in the material of
<taxonomicName id="2EF740AF4C5FFFE8FE6D82C0A21DF2B5" authorityName="Miles" authorityYear="1968" box="[415,574,902,927]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="105" phylum="Chordata" rank="species" species="leptocheira">
<emphasis id="DB83E73E4C5FFFE8FE6D82C0A21DF2B5" box="[415,574,902,927]" italics="true" pageId="8" pageNumber="105">B. leptocheira</emphasis>
</taxonomicName>
from
<collectingCountry id="91E07BBC4C5FFFE8FD7482C0A2C7F2B5" box="[646,740,902,927]" name="United Kingdom" pageId="8" pageNumber="105">Scotland</collectingCountry>
(
<bibRefCitation id="8D6646DD4C5FFFE8FD0582C0A0F3F295" author="Miles, R. S." pageId="8" pageNumber="105" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles 1968</bibRefCitation>
: pl. 20: 4) and
<collectingCountry id="91E07BBC4C5FFFE8FE7A82E0A1F3F295" box="[392,464,934,959]" name="Latvia" pageId="8" pageNumber="105">Latvia</collectingCountry>
(
<bibRefCitation id="8D6646DD4C5FFFE8FE1082E0A2BAF295" author="Luksevics, E." box="[482,665,934,959]" pageId="8" pageNumber="105" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
: fig. 34). The rounded anterior margin of Pp protrudes more into the orbital fenestra of small specimens, e.g., IG KSC 155/155 (
<figureCitation id="71CC27A94C5FFFE8FD2982A1A313F52A" box="[731,816,999,1024]" captionStart="Fig" captionStartId="7.[98,131,1207,1227]" captionTargetBox="[98,1478,163,1188]" captionTargetId="figure-351@7.[98,1478,163,1189]" captionTargetPageId="7" captionText="Fig. 6. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, postpineal AC, E, G), nuchal (DG, I, KM), and paranuchal (H, J, L, M) plates of the head shield. A. IG KSC 155/54-2, disconnected from the Nu IG KSC 155/54-1, in dorsal (A1) and visceral (A2) views. B. IG KSC 155/146 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/155 in dorsal (C1) and visceral C2) views. D. IG KSC 155/58 in dorsal view. E. Articulated plates IG KSC 155/118 in dorsal view. F. IG KSC 155/53 in dorsal (F1), visceral (F2), and posterior (F3) views. G. Articulated plates IG KSC 155/154 in dorsal (G1) and visceral (G2) views. H. IG KSC 155/55 in dorsal (H1) and visceral (H2) views. I. IG KSC 155/54-1 from the same individual as plate in A, in dorsal view. J. IG KSC 155/119 in dorsal view. K. IG KSC 155/56 in dorsal view. L. IG KSC 155/117 in visceral (L1) and dorsal (L2) views. M. IG KSC 155/125 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980833" httpUri="https://zenodo.org/record/10980833/files/figure.png" pageId="8" pageNumber="105">Fig. 6C</figureCitation>
) than in larger specimens (
<figureCitation id="71CC27A94C5FFFE8FE318541A269F50A" box="[451,586,1031,1056]" captionStart="Fig" captionStartId="7.[98,131,1207,1227]" captionTargetBox="[98,1478,163,1188]" captionTargetId="figure-351@7.[98,1478,163,1189]" captionTargetPageId="7" captionText="Fig. 6. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, postpineal AC, E, G), nuchal (DG, I, KM), and paranuchal (H, J, L, M) plates of the head shield. A. IG KSC 155/54-2, disconnected from the Nu IG KSC 155/54-1, in dorsal (A1) and visceral (A2) views. B. IG KSC 155/146 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/155 in dorsal (C1) and visceral C2) views. D. IG KSC 155/58 in dorsal view. E. Articulated plates IG KSC 155/118 in dorsal view. F. IG KSC 155/53 in dorsal (F1), visceral (F2), and posterior (F3) views. G. Articulated plates IG KSC 155/154 in dorsal (G1) and visceral (G2) views. H. IG KSC 155/55 in dorsal (H1) and visceral (H2) views. I. IG KSC 155/54-1 from the same individual as plate in A, in dorsal view. J. IG KSC 155/119 in dorsal view. K. IG KSC 155/56 in dorsal view. L. IG KSC 155/117 in visceral (L1) and dorsal (L2) views. M. IG KSC 155/125 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980833" httpUri="https://zenodo.org/record/10980833/files/figure.png" pageId="8" pageNumber="105">Figs. 6A, B</figureCitation>
). IG KSC 155/146 is an asymmetrical plate with wider right part of the bone and pointed, rather than rounded posterior angle (
<figureCitation id="71CC27A94C5FFFE8FD6E850EA2D1F54B" box="[668,754,1096,1121]" captionStart="Fig" captionStartId="7.[98,131,1207,1227]" captionTargetBox="[98,1478,163,1188]" captionTargetId="figure-351@7.[98,1478,163,1189]" captionTargetPageId="7" captionText="Fig. 6. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, postpineal AC, E, G), nuchal (DG, I, KM), and paranuchal (H, J, L, M) plates of the head shield. A. IG KSC 155/54-2, disconnected from the Nu IG KSC 155/54-1, in dorsal (A1) and visceral (A2) views. B. IG KSC 155/146 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/155 in dorsal (C1) and visceral C2) views. D. IG KSC 155/58 in dorsal view. E. Articulated plates IG KSC 155/118 in dorsal view. F. IG KSC 155/53 in dorsal (F1), visceral (F2), and posterior (F3) views. G. Articulated plates IG KSC 155/154 in dorsal (G1) and visceral (G2) views. H. IG KSC 155/55 in dorsal (H1) and visceral (H2) views. I. IG KSC 155/54-1 from the same individual as plate in A, in dorsal view. J. IG KSC 155/119 in dorsal view. K. IG KSC 155/56 in dorsal view. L. IG KSC 155/117 in visceral (L1) and dorsal (L2) views. M. IG KSC 155/125 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980833" httpUri="https://zenodo.org/record/10980833/files/figure.png" pageId="8" pageNumber="105">Fig. 6B</figureCitation>
). Two large, rounded pits (g;
<figureCitation id="71CC27A94C5FFFE8FE62852EA1C3F5AB" box="[400,480,1128,1153]" captionStart="Fig" captionStartId="8.[862,895,774,794]" captionTargetBox="[864,1534,161,746]" captionTargetId="figure-587@8.[863,1534,161,749]" captionTargetPageId="8" captionText="Fig. 7. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), paranuchal (A) and postpineal and nuchal (B), plates of the head shield. A. IG KSC 155/117 in dorsal (A1) and visceral (A2) views. B. Articulated plates IG KSC 155/154 in dorsal (B1) and visceral (B2) views.Abbreviations: cr.o, median occipital crest of head shield; cr.pm, paramarginal crest of head shield; cr.pto, postorbital crest of head shield; cr.tv, transverse nuchal crest of head shield; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; g, paired pits on Pp; ifc1, principal section of infraorbital sensory line; mp, middle pit-line groove; nm, obtected nuchal area of head shield; Nu, nuchal plate; ood, otico-occipital depression of head shield; Pp, postpineal plate; socc, supraoccipital cross-commissural pit-line groove; sot, supraotic thickening of head shield." figureDoi="http://doi.org/10.5281/zenodo.10980835" httpUri="https://zenodo.org/record/10980835/files/figure.png" pageId="8" pageNumber="105">Fig. 7B</figureCitation>
<figureCitation id="71CC27A94C5FFFE8FE138535A1C9F5AE" box="[481,490,1139,1156]" captionStart="Fig" captionStartId="2.[151,184,1333,1353]" captionTargetBox="[152,823,164,1305]" captionTargetId="figure-630@2.[152,604,164,1305]" captionTargetPageId="2" captionText="Fig. 2. Lithology and stratigraphy of the Sosnogorsk locality. “Fish-dolomite”, bed No. 40 is located about 8 m above the base of the section. Abbreviations: D3f, Upper Devonian, Frasnian; D3fm, Upper Devonian, Famennian." figureDoi="http://doi.org/10.5281/zenodo.10980823" httpUri="https://zenodo.org/record/10980823/files/figure.png" pageId="8" pageNumber="105">
<subScript id="757339694C5FFFE8FE138535A1C9F5AE" attach="right" box="[481,490,1139,1156]" fontSize="7" pageId="8" pageNumber="105">2</subScript>
</figureCitation>
) on the internal surface of the postpineal are separated by a low median ridge.
</paragraph>
<paragraph id="E9483B2C4C5FFFE8FF4F85EEA182F682" blockId="8.[151,825,160,1960]" pageId="8" pageNumber="105">
The nuchal plate (Nu;
<figureCitation id="71CC27A94C5FFFE8FE4C85EEA228F5EB" box="[446,523,1192,1217]" captionStart="Fig" captionStartId="4.[151,184,1188,1208]" captionTargetBox="[157,1530,160,1164]" captionTargetId="figure-306@4.[173,1531,159,1164]" captionTargetPageId="4" captionText="Fig. 3. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, separate plates and parts of articulated head shields. A. Articulated Nu, left Pn and left La plates IG KSC 155/90 in dorsal view. B. Disarticulated plates of the head shield IG KSC 155/150 in dorsal view. C. Part of articulated head shield IG KSC 155/162 in dorsal view. D. Imprint of the ventral wall of trunk armour and some skeletal elements of the head armour IG KSC 155/10-3 in ventral view showing suborbital and infragnathal bones. Abbreviations: AVL, anterior ventro-lateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; ifc1, principal section of infraorbital sensory line; IG, inferognathal; La, lateral plate; Nu, nuchal plate; mp, middle pit-line groove; Pn, paranuchal plate; Pp, postpineal plate; Prm, premedian plate; SO, suborbital plate; socc, supraoccipital cross-commissural pit-line groove." figureDoi="http://doi.org/10.5281/zenodo.10980827" httpUri="https://zenodo.org/record/10980827/files/figure.png" pageId="8" pageNumber="105">Figs. 3</figureCitation>
,
<figureCitation id="71CC27A94C5FFFE8FDEB85EEA266F5EB" box="[537,581,1192,1217]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="7.[98,131,1207,1227]" captionStartId-1="8.[862,895,774,794]" captionStartId-2="8.[863,896,1656,1676]" captionTargetBox-0="[98,1478,163,1188]" captionTargetBox-1="[864,1534,161,746]" captionTargetBox-2="[891,1506,1125,1629]" captionTargetId-0="figure-351@7.[98,1478,163,1189]" captionTargetId-1="figure-587@8.[863,1534,161,749]" captionTargetId-2="figure-600@8.[891,1506,1125,1630]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="8" captionText-0="Fig. 6. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, postpineal AC, E, G), nuchal (DG, I, KM), and paranuchal (H, J, L, M) plates of the head shield. A. IG KSC 155/54-2, disconnected from the Nu IG KSC 155/54-1, in dorsal (A1) and visceral (A2) views. B. IG KSC 155/146 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/155 in dorsal (C1) and visceral C2) views. D. IG KSC 155/58 in dorsal view. E. Articulated plates IG KSC 155/118 in dorsal view. F. IG KSC 155/53 in dorsal (F1), visceral (F2), and posterior (F3) views. G. Articulated plates IG KSC 155/154 in dorsal (G1) and visceral (G2) views. H. IG KSC 155/55 in dorsal (H1) and visceral (H2) views. I. IG KSC 155/54-1 from the same individual as plate in A, in dorsal view. J. IG KSC 155/119 in dorsal view. K. IG KSC 155/56 in dorsal view. L. IG KSC 155/117 in visceral (L1) and dorsal (L2) views. M. IG KSC 155/125 in dorsal view." captionText-1="Fig. 7. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), paranuchal (A) and postpineal and nuchal (B), plates of the head shield. A. IG KSC 155/117 in dorsal (A1) and visceral (A2) views. B. Articulated plates IG KSC 155/154 in dorsal (B1) and visceral (B2) views.Abbreviations: cr.o, median occipital crest of head shield; cr.pm, paramarginal crest of head shield; cr.pto, postorbital crest of head shield; cr.tv, transverse nuchal crest of head shield; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; g, paired pits on Pp; ifc1, principal section of infraorbital sensory line; mp, middle pit-line groove; nm, obtected nuchal area of head shield; Nu, nuchal plate; ood, otico-occipital depression of head shield; Pp, postpineal plate; socc, supraoccipital cross-commissural pit-line groove; sot, supraotic thickening of head shield." captionText-2="Fig. 8. Reconstruction of the head shield of antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), based on partial head shield (IG KSC 71/1-87), left (IG KSC 155/15), and right (IG KSC 155/65) lateral plates. Abbreviations: cir, semicircular pit-line groove; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; ifc1, principal section of infraorbital sensory line; ifc2, branch of infraorbital sensory line diverging on La; La, lateral plate; mp, middle pit-line groove; Nu, nuchal plate; Pmg, postmarginal plate; Pn, paranuchal plate; Pp, postpineal plate; Prm, premedian plate; prh, preorbital recess of head shield; soc, anterior section of the supraorbital sensory line; socc, supraoccipital cross-commissural pit-line groove." figureDoi-0="http://doi.org/10.5281/zenodo.10980833" figureDoi-1="http://doi.org/10.5281/zenodo.10980835" figureDoi-2="http://doi.org/10.5281/zenodo.10980837" httpUri-0="https://zenodo.org/record/10980833/files/figure.png" httpUri-1="https://zenodo.org/record/10980835/files/figure.png" httpUri-2="https://zenodo.org/record/10980837/files/figure.png" pageId="8" pageNumber="105">68</figureCitation>
) has an L/B index of about 0.83 (
<bibRefCitation id="8D6646DD4C5FFFE8FEEF858FA1BEF5C8" author="Miles, R. S." box="[285,413,1225,1250]" pageId="8" pageNumber="105" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles 1968</bibRefCitation>
mentioned an L/B index of 0.83 for Scottish material and
<bibRefCitation id="8D6646DD4C5FFFE8FE6C85AFA276F428" author="Luksevics, E." box="[414,597,1257,1282]" pageId="8" pageNumber="105" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
mentioned an L/B index of 0.84 for material from
<collectingCountry id="91E07BBC4C5FFFE8FDF2844FA26FF408" box="[512,588,1289,1314]" name="Latvia" pageId="8" pageNumber="105">Latvia</collectingCountry>
). The plate is always broadest across the lateral corners. The anterior division of the lateral margin is usually convex and a little shorter than the concave posterior division (
<figureCitation id="71CC27A94C5FFFE8FDFE842CA2E3F4A9" box="[524,704,1386,1411]" captionStart="Fig" captionStartId="7.[98,131,1207,1227]" captionTargetBox="[98,1478,163,1188]" captionTargetId="figure-351@7.[98,1478,163,1189]" captionTargetPageId="7" captionText="Fig. 6. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, postpineal AC, E, G), nuchal (DG, I, KM), and paranuchal (H, J, L, M) plates of the head shield. A. IG KSC 155/54-2, disconnected from the Nu IG KSC 155/54-1, in dorsal (A1) and visceral (A2) views. B. IG KSC 155/146 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/155 in dorsal (C1) and visceral C2) views. D. IG KSC 155/58 in dorsal view. E. Articulated plates IG KSC 155/118 in dorsal view. F. IG KSC 155/53 in dorsal (F1), visceral (F2), and posterior (F3) views. G. Articulated plates IG KSC 155/154 in dorsal (G1) and visceral (G2) views. H. IG KSC 155/55 in dorsal (H1) and visceral (H2) views. I. IG KSC 155/54-1 from the same individual as plate in A, in dorsal view. J. IG KSC 155/119 in dorsal view. K. IG KSC 155/56 in dorsal view. L. IG KSC 155/117 in visceral (L1) and dorsal (L2) views. M. IG KSC 155/125 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980833" httpUri="https://zenodo.org/record/10980833/files/figure.png" pageId="8" pageNumber="105">Figs. 6DH, K</figureCitation>
,
<figureCitation id="71CC27A94C5FFFE8FD23842CA2D0F4A9" box="[721,755,1386,1411]" captionStart="Fig" captionStartId="8.[862,895,774,794]" captionTargetBox="[864,1534,161,746]" captionTargetId="figure-587@8.[863,1534,161,749]" captionTargetPageId="8" captionText="Fig. 7. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), paranuchal (A) and postpineal and nuchal (B), plates of the head shield. A. IG KSC 155/117 in dorsal (A1) and visceral (A2) views. B. Articulated plates IG KSC 155/154 in dorsal (B1) and visceral (B2) views.Abbreviations: cr.o, median occipital crest of head shield; cr.pm, paramarginal crest of head shield; cr.pto, postorbital crest of head shield; cr.tv, transverse nuchal crest of head shield; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; g, paired pits on Pp; ifc1, principal section of infraorbital sensory line; mp, middle pit-line groove; nm, obtected nuchal area of head shield; Nu, nuchal plate; ood, otico-occipital depression of head shield; Pp, postpineal plate; socc, supraoccipital cross-commissural pit-line groove; sot, supraotic thickening of head shield." figureDoi="http://doi.org/10.5281/zenodo.10980835" httpUri="https://zenodo.org/record/10980835/files/figure.png" pageId="8" pageNumber="105">7B</figureCitation>
). The orbital facets are very short, tapering to a point in IG KSC 155/53 and IG KSC 155/154 (
<figureCitation id="71CC27A94C5FFFE8FE2F84EDA272F4EE" box="[477,593,1451,1476]" captionStart="Fig" captionStartId="7.[98,131,1207,1227]" captionTargetBox="[98,1478,163,1188]" captionTargetId="figure-351@7.[98,1478,163,1189]" captionTargetPageId="7" captionText="Fig. 6. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, postpineal AC, E, G), nuchal (DG, I, KM), and paranuchal (H, J, L, M) plates of the head shield. A. IG KSC 155/54-2, disconnected from the Nu IG KSC 155/54-1, in dorsal (A1) and visceral (A2) views. B. IG KSC 155/146 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/155 in dorsal (C1) and visceral C2) views. D. IG KSC 155/58 in dorsal view. E. Articulated plates IG KSC 155/118 in dorsal view. F. IG KSC 155/53 in dorsal (F1), visceral (F2), and posterior (F3) views. G. Articulated plates IG KSC 155/154 in dorsal (G1) and visceral (G2) views. H. IG KSC 155/55 in dorsal (H1) and visceral (H2) views. I. IG KSC 155/54-1 from the same individual as plate in A, in dorsal view. J. IG KSC 155/119 in dorsal view. K. IG KSC 155/56 in dorsal view. L. IG KSC 155/117 in visceral (L1) and dorsal (L2) views. M. IG KSC 155/125 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980833" httpUri="https://zenodo.org/record/10980833/files/figure.png" pageId="8" pageNumber="105">Fig. 6F, G</figureCitation>
). The posterior margin is usually strongly concave; however, IG KSC 155/54-1 and IG KSC 155/56 demonstrate convex posterior margin (
<figureCitation id="71CC27A94C5FFFE8FF6D874DA13AF70E" box="[159,281,1547,1572]" captionStart="Fig" captionStartId="7.[98,131,1207,1227]" captionTargetBox="[98,1478,163,1188]" captionTargetId="figure-351@7.[98,1478,163,1189]" captionTargetPageId="7" captionText="Fig. 6. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, postpineal AC, E, G), nuchal (DG, I, KM), and paranuchal (H, J, L, M) plates of the head shield. A. IG KSC 155/54-2, disconnected from the Nu IG KSC 155/54-1, in dorsal (A1) and visceral (A2) views. B. IG KSC 155/146 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/155 in dorsal (C1) and visceral C2) views. D. IG KSC 155/58 in dorsal view. E. Articulated plates IG KSC 155/118 in dorsal view. F. IG KSC 155/53 in dorsal (F1), visceral (F2), and posterior (F3) views. G. Articulated plates IG KSC 155/154 in dorsal (G1) and visceral (G2) views. H. IG KSC 155/55 in dorsal (H1) and visceral (H2) views. I. IG KSC 155/54-1 from the same individual as plate in A, in dorsal view. J. IG KSC 155/119 in dorsal view. K. IG KSC 155/56 in dorsal view. L. IG KSC 155/117 in visceral (L1) and dorsal (L2) views. M. IG KSC 155/125 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980833" httpUri="https://zenodo.org/record/10980833/files/figure.png" pageId="8" pageNumber="105">Fig. 6H, I</figureCitation>
). The median process is more pronounced in specimens with a concave posterior margin. The external openings for the endolymphatic ducts (d.end) are closely set to one another, in a distance of
<quantity id="2E0F96C94C5FFFE8FDF7872AA27FF7AF" box="[517,604,1644,1669]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="8" pageNumber="105" unit="mm" value="2.9">2.9 mm</quantity>
in IG KSC 155/53. The supraoccipital cross-commissural pit-line groove (socc) is usually well defined, however, in IG KSC 155/53 (
<figureCitation id="71CC27A94C5FFFE8FD2C87EBA313F7EC" box="[734,816,1709,1734]" captionStart="Fig" captionStartId="7.[98,131,1207,1227]" captionTargetBox="[98,1478,163,1188]" captionTargetId="figure-351@7.[98,1478,163,1189]" captionTargetPageId="7" captionText="Fig. 6. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, postpineal AC, E, G), nuchal (DG, I, KM), and paranuchal (H, J, L, M) plates of the head shield. A. IG KSC 155/54-2, disconnected from the Nu IG KSC 155/54-1, in dorsal (A1) and visceral (A2) views. B. IG KSC 155/146 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/155 in dorsal (C1) and visceral C2) views. D. IG KSC 155/58 in dorsal view. E. Articulated plates IG KSC 155/118 in dorsal view. F. IG KSC 155/53 in dorsal (F1), visceral (F2), and posterior (F3) views. G. Articulated plates IG KSC 155/154 in dorsal (G1) and visceral (G2) views. H. IG KSC 155/55 in dorsal (H1) and visceral (H2) views. I. IG KSC 155/54-1 from the same individual as plate in A, in dorsal view. J. IG KSC 155/119 in dorsal view. K. IG KSC 155/56 in dorsal view. L. IG KSC 155/117 in visceral (L1) and dorsal (L2) views. M. IG KSC 155/125 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980833" httpUri="https://zenodo.org/record/10980833/files/figure.png" pageId="8" pageNumber="105">Fig. 6F</figureCitation>
) it is seen only as a line of pores along the posterior margin. Internally, the openings for the endolymphatic ducts are larger than their external openings, divided by the large supraotic thickening (sot;
<figureCitation id="71CC27A94C5FFFE8FE318668A23BF66D" box="[451,536,1838,1863]" captionStart="Fig" captionStartId="8.[862,895,774,794]" captionTargetBox="[864,1534,161,746]" captionTargetId="figure-587@8.[863,1534,161,749]" captionTargetPageId="8" captionText="Fig. 7. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), paranuchal (A) and postpineal and nuchal (B), plates of the head shield. A. IG KSC 155/117 in dorsal (A1) and visceral (A2) views. B. Articulated plates IG KSC 155/154 in dorsal (B1) and visceral (B2) views.Abbreviations: cr.o, median occipital crest of head shield; cr.pm, paramarginal crest of head shield; cr.pto, postorbital crest of head shield; cr.tv, transverse nuchal crest of head shield; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; g, paired pits on Pp; ifc1, principal section of infraorbital sensory line; mp, middle pit-line groove; nm, obtected nuchal area of head shield; Nu, nuchal plate; ood, otico-occipital depression of head shield; Pp, postpineal plate; socc, supraoccipital cross-commissural pit-line groove; sot, supraotic thickening of head shield." figureDoi="http://doi.org/10.5281/zenodo.10980835" httpUri="https://zenodo.org/record/10980835/files/figure.png" pageId="8" pageNumber="105">Fig. 7B</figureCitation>
<figureCitation id="71CC27A94C5FFFE8FDEA867FA202F660" box="[536,545,1849,1866]" captionStart="Fig" captionStartId="2.[151,184,1333,1353]" captionTargetBox="[152,823,164,1305]" captionTargetId="figure-630@2.[152,604,164,1305]" captionTargetPageId="2" captionText="Fig. 2. Lithology and stratigraphy of the Sosnogorsk locality. “Fish-dolomite”, bed No. 40 is located about 8 m above the base of the section. Abbreviations: D3f, Upper Devonian, Frasnian; D3fm, Upper Devonian, Famennian." figureDoi="http://doi.org/10.5281/zenodo.10980823" httpUri="https://zenodo.org/record/10980823/files/figure.png" pageId="8" pageNumber="105">
<subScript id="757339694C5FFFE8FDEA867FA202F660" attach="left" box="[536,545,1849,1866]" fontSize="7" pageId="8" pageNumber="105">2</subScript>
</figureCitation>
). The transverse nuchal crista (cr.tv) is high and well developed. Posteriorly, the median occipital crest of the head shield (cro;
<figureCitation id="71CC27A94C5FFFE8FD678629A2C2F6A2" box="[661,737,1903,1928]" captionStart="Fig" captionStartId="7.[98,131,1207,1227]" captionTargetBox="[98,1478,163,1188]" captionTargetId="figure-351@7.[98,1478,163,1189]" captionTargetPageId="7" captionText="Fig. 6. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, postpineal AC, E, G), nuchal (DG, I, KM), and paranuchal (H, J, L, M) plates of the head shield. A. IG KSC 155/54-2, disconnected from the Nu IG KSC 155/54-1, in dorsal (A1) and visceral (A2) views. B. IG KSC 155/146 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/155 in dorsal (C1) and visceral C2) views. D. IG KSC 155/58 in dorsal view. E. Articulated plates IG KSC 155/118 in dorsal view. F. IG KSC 155/53 in dorsal (F1), visceral (F2), and posterior (F3) views. G. Articulated plates IG KSC 155/154 in dorsal (G1) and visceral (G2) views. H. IG KSC 155/55 in dorsal (H1) and visceral (H2) views. I. IG KSC 155/54-1 from the same individual as plate in A, in dorsal view. J. IG KSC 155/119 in dorsal view. K. IG KSC 155/56 in dorsal view. L. IG KSC 155/117 in visceral (L1) and dorsal (L2) views. M. IG KSC 155/125 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980833" httpUri="https://zenodo.org/record/10980833/files/figure.png" pageId="8" pageNumber="105">Figs. 6</figureCitation>
,
<figureCitation id="71CC27A94C5FFFE8FD1E8629A32EF6A2" box="[748,781,1903,1928]" captionStart="Fig" captionStartId="8.[862,895,774,794]" captionTargetBox="[864,1534,161,746]" captionTargetId="figure-587@8.[863,1534,161,749]" captionTargetPageId="8" captionText="Fig. 7. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), paranuchal (A) and postpineal and nuchal (B), plates of the head shield. A. IG KSC 155/117 in dorsal (A1) and visceral (A2) views. B. Articulated plates IG KSC 155/154 in dorsal (B1) and visceral (B2) views.Abbreviations: cr.o, median occipital crest of head shield; cr.pm, paramarginal crest of head shield; cr.pto, postorbital crest of head shield; cr.tv, transverse nuchal crest of head shield; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; g, paired pits on Pp; ifc1, principal section of infraorbital sensory line; mp, middle pit-line groove; nm, obtected nuchal area of head shield; Nu, nuchal plate; ood, otico-occipital depression of head shield; Pp, postpineal plate; socc, supraoccipital cross-commissural pit-line groove; sot, supraotic thickening of head shield." figureDoi="http://doi.org/10.5281/zenodo.10980835" httpUri="https://zenodo.org/record/10980835/files/figure.png" pageId="8" pageNumber="105">7B</figureCitation>
<figureCitation id="71CC27A94C5FFFE8FCFF863CA335F6A1" box="[781,790,1914,1931]" captionStart="Fig" captionStartId="2.[151,184,1333,1353]" captionTargetBox="[152,823,164,1305]" captionTargetId="figure-630@2.[152,604,164,1305]" captionTargetPageId="2" captionText="Fig. 2. Lithology and stratigraphy of the Sosnogorsk locality. “Fish-dolomite”, bed No. 40 is located about 8 m above the base of the section. Abbreviations: D3f, Upper Devonian, Frasnian; D3fm, Upper Devonian, Famennian." figureDoi="http://doi.org/10.5281/zenodo.10980823" httpUri="https://zenodo.org/record/10980823/files/figure.png" pageId="8" pageNumber="105">
<subScript id="757339694C5FFFE8FCFF863CA335F6A1" attach="left" box="[781,790,1914,1931]" fontSize="7" pageId="8" pageNumber="105">2</subScript>
</figureCitation>
) is usually well developed.
</paragraph>
<caption id="BD886BA44C5FFFE8FCAD873EA43EF68E" ID-DOI="http://doi.org/10.5281/zenodo.10980837" ID-Zenodo-Dep="10980837" httpUri="https://zenodo.org/record/10980837/files/figure.png" pageId="8" pageNumber="105" startId="8.[863,896,1656,1676]" targetBox="[891,1506,1125,1629]" targetPageId="8" targetType="figure">
<paragraph id="E9483B2C4C5FFFE8FCAD873EA43EF68E" blockId="8.[863,1535,1656,1956]" pageId="8" pageNumber="105">
Fig. 8. Reconstruction of the head shield of antiarchan fish
<taxonomicName id="2EF740AF4C5FFFE8FA7C873EA4E3F782" authority="(Rohon, 1900)" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" baseAuthorityName="Rohon" baseAuthorityYear="1900" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="105" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="jeremejevi">
<emphasis id="DB83E73E4C5FFFE8FA7C873EA40EF782" italics="true" pageId="8" pageNumber="105">Bothriolepis leptocheira jeremejevi</emphasis>
(
<bibRefCitation id="8D6646DD4C5FFFE8FBCE87D2A49AF782" author="Rohon, J. V." box="[1084,1209,1684,1704]" pageId="8" pageNumber="105" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon, 1900</bibRefCitation>
)
</taxonomicName>
, based on partial head shield (IG KSC 71/1-87), left (IG KSC 155/15), and right (IG KSC 155/65) lateral plates. Abbreviations: cir, semicircular pit-line groove; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; ifc
<subScript id="757339694C5FFFE8FA6987B7A580F7D5" attach="left" box="[1435,1443,1777,1791]" fontSize="6" pageId="8" pageNumber="105">1</subScript>
, principal section of infraorbital sensory line; ifc
<subScript id="757339694C5FFFE8FB4A864BA4E3F631" attach="left" box="[1208,1216,1805,1819]" fontSize="6" pageId="8" pageNumber="105">2</subScript>
, branch of infraorbital sensory line diverging on La; La, lateral plate; mp, middle pit-line groove; Nu, nuchal plate; Pmg, postmarginal plate; Pn, paranuchal plate; Pp, postpineal plate; Prm, premedian plate; prh, preorbital recess of head shield; soc, anterior section of the supraorbital sensory line; socc, supraoccipital cross-commissural pit-line groove.
</paragraph>
</caption>
<caption id="BD886BA44C5DFFEAFF658218A5CBF530" ID-DOI="http://doi.org/10.5281/zenodo.10980839" ID-Zenodo-Dep="10980839" httpUri="https://zenodo.org/record/10980839/files/figure.png" pageId="10" pageNumber="107" startId="10.[151,184,862,882]" targetBox="[155,1534,159,837]" targetPageId="10" targetType="figure">
<paragraph id="E9483B2C4C5DFFEAFF658218A5CBF530" blockId="10.[151,1536,862,1050]" pageId="10" pageNumber="107">
Fig. 10. Antiarchan fish
<taxonomicName id="2EF740AF4C5DFFEAFE868218A361F258" authority="(Rohon, 1900)" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" baseAuthorityName="Rohon" baseAuthorityYear="1900" box="[372,834,862,882]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="107" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="jeremejevi">
<emphasis id="DB83E73E4C5DFFEAFE868218A296F258" box="[372,693,862,882]" italics="true" pageId="10" pageNumber="107">Bothriolepis leptocheira jeremejevi</emphasis>
(
<bibRefCitation id="8D6646DD4C5DFFEAFD338218A319F258" author="Rohon, J. V." box="[705,826,862,882]" pageId="10" pageNumber="107" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon, 1900</bibRefCitation>
)
</taxonomicName>
, Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior dorso-lateral (AD, F, left; E, right) and left mixilateral (GI, right) plates of the trunk armour.
<emphasis id="DB83E73E4C5DFFEAFC1C823FA3DDF2A4" bold="true" box="[1006,1022,889,910]" pageId="10" pageNumber="107">A</emphasis>
. IG KSC 155/116 in dorsal (A
<subScript id="757339694C5DFFEAFAE382C5A53AF2BB" attach="left" box="[1297,1305,899,913]" fontSize="6" pageId="10" pageNumber="107">1</subScript>
) and visceral (A
<subScript id="757339694C5DFFEAFA5D82C5A594F2BB" attach="left" box="[1455,1463,899,913]" fontSize="6" pageId="10" pageNumber="107">2</subScript>
) views.
<emphasis id="DB83E73E4C5DFFEAFF6582D3A085F280" bold="true" box="[151,166,917,938]" pageId="10" pageNumber="107">B</emphasis>
. IG KSC 155/111.
<emphasis id="DB83E73E4C5DFFEAFEAE82D3A14FF281" bold="true" box="[348,364,917,939]" pageId="10" pageNumber="107">C</emphasis>
. IG KSC 155/61.
<emphasis id="DB83E73E4C5DFFEAFDEA82D3A20BF280" bold="true" box="[536,552,917,938]" pageId="10" pageNumber="107">D</emphasis>
. IG KSC 155/60.
<emphasis id="DB83E73E4C5DFFEAFD2682D3A2C0F280" bold="true" box="[724,739,917,938]" pageId="10" pageNumber="107">E</emphasis>
. IG KSC 155/136.
<emphasis id="DB83E73E4C5DFFEAFC6982D3A38AF280" bold="true" box="[923,937,917,938]" pageId="10" pageNumber="107">F</emphasis>
. IG KSC 155/112.
<emphasis id="DB83E73E4C5DFFEAFBAD82D3A452F281" bold="true" box="[1119,1137,917,939]" pageId="10" pageNumber="107">G</emphasis>
. IG KSC 155/110.
<emphasis id="DB83E73E4C5DFFEAFAD582D3A51AF280" bold="true" box="[1319,1337,917,938]" pageId="10" pageNumber="107">H</emphasis>
. Incomplete IG KSC 155/144.
<emphasis id="DB83E73E4C5DFFEAFF1E82F7A0D6F2EC" bold="true" box="[236,245,945,966]" pageId="10" pageNumber="107">I</emphasis>
. IG KSC 155/115. BE, GI, in dorsal views; F, in lateral view.Abbreviations: ADL, anterior ventro-lateral plate; AMD, anterior median dorsal plate; AVL, anterior ventro-lateral plate; cf.AMD, area overlapping AMD; dlg
<subScript id="757339694C5DFFEAFC968291A34FF2CF" attach="left" box="[868,876,983,997]" fontSize="6" pageId="10" pageNumber="107">2</subScript>
, posterior oblique dorsal sensory line groove; dlr, dorso-lateral ridge of trunk armour; dxp, dorso-ventral pit-line groove; lcg, main lateral line groove; oa.ADL, oa.AMD, oa.AVL, oa.PMD, area overlapped by ADL, AMD, AVL or PMD respectively; PMD, posterior median dorsal plate; pnoa, postnuchal ornamented corner of ADL; pro, processus obstans of trunk armour.
</paragraph>
</caption>
<paragraph id="E9483B2C4C5DFFEAFF4F8571A502F458" blockId="10.[151,824,1079,1588]" lastBlockId="10.[862,1535,1079,1588]" pageId="10" pageNumber="107">
The paranuchal plate (Pn;
<figureCitation id="71CC27A94C5DFFEAFE018571A263F57A" box="[499,576,1079,1104]" captionStart="Fig" captionStartId="4.[151,184,1188,1208]" captionTargetBox="[157,1530,160,1164]" captionTargetId="figure-306@4.[173,1531,159,1164]" captionTargetPageId="4" captionText="Fig. 3. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, separate plates and parts of articulated head shields. A. Articulated Nu, left Pn and left La plates IG KSC 155/90 in dorsal view. B. Disarticulated plates of the head shield IG KSC 155/150 in dorsal view. C. Part of articulated head shield IG KSC 155/162 in dorsal view. D. Imprint of the ventral wall of trunk armour and some skeletal elements of the head armour IG KSC 155/10-3 in ventral view showing suborbital and infragnathal bones. Abbreviations: AVL, anterior ventro-lateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; ifc1, principal section of infraorbital sensory line; IG, inferognathal; La, lateral plate; Nu, nuchal plate; mp, middle pit-line groove; Pn, paranuchal plate; Pp, postpineal plate; Prm, premedian plate; SO, suborbital plate; socc, supraoccipital cross-commissural pit-line groove." figureDoi="http://doi.org/10.5281/zenodo.10980827" httpUri="https://zenodo.org/record/10980827/files/figure.png" pageId="10" pageNumber="107">Figs. 3</figureCitation>
,
<figureCitation id="71CC27A94C5DFFEAFDA38571A27CF57A" box="[593,607,1079,1104]" captionStart="Fig" captionStartId="7.[98,131,1207,1227]" captionTargetBox="[98,1478,163,1188]" captionTargetId="figure-351@7.[98,1478,163,1189]" captionTargetPageId="7" captionText="Fig. 6. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, postpineal AC, E, G), nuchal (DG, I, KM), and paranuchal (H, J, L, M) plates of the head shield. A. IG KSC 155/54-2, disconnected from the Nu IG KSC 155/54-1, in dorsal (A1) and visceral (A2) views. B. IG KSC 155/146 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/155 in dorsal (C1) and visceral C2) views. D. IG KSC 155/58 in dorsal view. E. Articulated plates IG KSC 155/118 in dorsal view. F. IG KSC 155/53 in dorsal (F1), visceral (F2), and posterior (F3) views. G. Articulated plates IG KSC 155/154 in dorsal (G1) and visceral (G2) views. H. IG KSC 155/55 in dorsal (H1) and visceral (H2) views. I. IG KSC 155/54-1 from the same individual as plate in A, in dorsal view. J. IG KSC 155/119 in dorsal view. K. IG KSC 155/56 in dorsal view. L. IG KSC 155/117 in visceral (L1) and dorsal (L2) views. M. IG KSC 155/125 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980833" httpUri="https://zenodo.org/record/10980833/files/figure.png" pageId="10" pageNumber="107">6</figureCitation>
,
<figureCitation id="71CC27A94C5DFFEAFD828571A25EF57A" box="[624,637,1079,1104]" captionStart="Fig" captionStartId="8.[862,895,774,794]" captionTargetBox="[864,1534,161,746]" captionTargetId="figure-587@8.[863,1534,161,749]" captionTargetPageId="8" captionText="Fig. 7. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), paranuchal (A) and postpineal and nuchal (B), plates of the head shield. A. IG KSC 155/117 in dorsal (A1) and visceral (A2) views. B. Articulated plates IG KSC 155/154 in dorsal (B1) and visceral (B2) views.Abbreviations: cr.o, median occipital crest of head shield; cr.pm, paramarginal crest of head shield; cr.pto, postorbital crest of head shield; cr.tv, transverse nuchal crest of head shield; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; g, paired pits on Pp; ifc1, principal section of infraorbital sensory line; mp, middle pit-line groove; nm, obtected nuchal area of head shield; Nu, nuchal plate; ood, otico-occipital depression of head shield; Pp, postpineal plate; socc, supraoccipital cross-commissural pit-line groove; sot, supraotic thickening of head shield." figureDoi="http://doi.org/10.5281/zenodo.10980835" httpUri="https://zenodo.org/record/10980835/files/figure.png" pageId="10" pageNumber="107">7</figureCitation>
) is of moderate breadth, with an L/B index of about 0.92, whereas in
<taxonomicName id="2EF740AF4C5DFFEAFCD28511A1A6F5BA" authorityName="Gross" authorityYear="1942" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="107" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="curonica">
<emphasis id="DB83E73E4C5DFFEAFCD28511A1A6F5BA" italics="true" pageId="10" pageNumber="107">B. leptocheira curonica</emphasis>
</taxonomicName>
it is broader, the index reaching 0.86 (
<bibRefCitation id="8D6646DD4C5DFFEAFF6D85DEA174F59B" author="Luksevics, E." box="[159,343,1176,1201]" pageId="10" pageNumber="107" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
);
<bibRefCitation id="8D6646DD4C5DFFEAFE9F85DEA1DCF59B" author="Miles, R. S." box="[365,511,1176,1201]" pageId="10" pageNumber="107" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles (1968)</bibRefCitation>
mentioned that Pn of
<emphasis id="DB83E73E4C5DFFEAFCF585DEA13BF5FB" italics="true" pageId="10" pageNumber="107">
B. l.
<taxonomicName id="2EF740AF4C5DFFEAFF6585FEA13BF5FB" authorityName="Miles" authorityYear="1968" box="[151,280,1208,1233]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="107" phylum="Chordata" rank="species" species="leptocheira">leptocheira</taxonomicName>
</emphasis>
is poorly known and in the restoration (
<bibRefCitation id="8D6646DD4C5DFFEAFD0485FEA0F2F5DB" author="Miles, R. S." pageId="10" pageNumber="107" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles 1968</bibRefCitation>
: text-fig. 36) it has an L/B index of about 0.80. The lateral division of Pn composes 3944% (41.9 on the average) of the general breadth of the plate, thus being slightly broader than in
<taxonomicName id="2EF740AF4C5DFFEAFF02847FA223F478" authorityName="Gross" authorityYear="1942" box="[240,512,1337,1362]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="107" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="curonica">
<emphasis id="DB83E73E4C5DFFEAFF02847FA223F478" box="[240,512,1337,1362]" italics="true" pageId="10" pageNumber="107">B. leptocheira curonica</emphasis>
</taxonomicName>
with 38.7% on the average (
<bibRefCitation id="8D6646DD4C5DFFEAFF6D841FA177F458" author="Luksevics, E." box="[159,340,1369,1394]" pageId="10" pageNumber="107" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
). A long supraoccipital cross-commissural pit-line groove (socc) is present in all examined specimens; it terminates approximately at the level of the middle of the plate posterior margin.
<taxonomicName id="2EF740AF4C5DFFEAFE2884FCA31BF4F9" authority="(Agassiz, 1844)" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[474,824,1466,1491]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="107" phylum="Chordata" rank="species" species="hydrophila">
<emphasis id="DB83E73E4C5DFFEAFE2884FCA25AF4F9" box="[474,633,1466,1491]" italics="true" pageId="10" pageNumber="107">B. hydrophila</emphasis>
(Agassiz, 1844)
</taxonomicName>
from the Upper Old Red Sandstone of Dura Den,
<collectingCountry id="91E07BBC4C5DFFEAFD3C849DA317F4DE" box="[718,820,1499,1524]" name="United Kingdom" pageId="10" pageNumber="107">Scotland</collectingCountry>
, also possess a long supraoccipital groove; however, it never extends to the Pn plate. The middle pit-line groove (mp) is always present, as it is in
<taxonomicName id="2EF740AF4C5DFFEAFB6B8571A5E9F57A" box="[1177,1482,1079,1104]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="107" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="leptocheira">
<emphasis id="DB83E73E4C5DFFEAFB6B8571A5E9F57A" box="[1177,1482,1079,1104]" italics="true" pageId="10" pageNumber="107">B. leptocheira leptocheira</emphasis>
</taxonomicName>
and
<taxonomicName id="2EF740AF4C5DFFEAFCAC8511A562F55A" authority="(Luksevics 2001)" baseAuthorityName="Luksevics" baseAuthorityYear="2001" box="[862,1345,1111,1136]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="107" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="curonica">
<emphasis id="DB83E73E4C5DFFEAFCAC8511A453F55A" box="[862,1136,1111,1136]" italics="true" pageId="10" pageNumber="107">B. leptocheira curonica</emphasis>
(
<bibRefCitation id="8D6646DD4C5DFFEAFB738511A51AF55A" author="Luksevics, E." box="[1153,1337,1111,1136]" pageId="10" pageNumber="107" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
)
</taxonomicName>
; sometimes it is rather long, but it is not fused with the supraoccipital pit-line groove, contrary to
<taxonomicName id="2EF740AF4C5DFFEAFBC885DEA560F59B" baseAuthorityName="Luksevics" baseAuthorityYear="2001" box="[1082,1347,1176,1201]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="107" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="curonica">
<emphasis id="DB83E73E4C5DFFEAFBC885DEA560F59B" box="[1082,1347,1176,1201]" italics="true" pageId="10" pageNumber="107">B. leptocheira curonica</emphasis>
</taxonomicName>
where it is fused in some specimens (
<bibRefCitation id="8D6646DD4C5DFFEAFBCC85FEA4D0F5FB" author="Luksevics, E." box="[1086,1267,1208,1233]" pageId="10" pageNumber="107" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
: fig. 33A, B). Internally, the paramarginal crest (cr.pm) is very distinct; it is of semilunar shape in IG KSC 155/117 (
<figureCitation id="71CC27A94C5DFFEAFB3085BFA53BF438" box="[1218,1304,1273,1298]" captionStart="Fig" captionStartId="8.[862,895,774,794]" captionTargetBox="[864,1534,161,746]" captionTargetId="figure-587@8.[863,1534,161,749]" captionTargetPageId="8" captionText="Fig. 7. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), paranuchal (A) and postpineal and nuchal (B), plates of the head shield. A. IG KSC 155/117 in dorsal (A1) and visceral (A2) views. B. Articulated plates IG KSC 155/154 in dorsal (B1) and visceral (B2) views.Abbreviations: cr.o, median occipital crest of head shield; cr.pm, paramarginal crest of head shield; cr.pto, postorbital crest of head shield; cr.tv, transverse nuchal crest of head shield; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; g, paired pits on Pp; ifc1, principal section of infraorbital sensory line; mp, middle pit-line groove; nm, obtected nuchal area of head shield; Nu, nuchal plate; ood, otico-occipital depression of head shield; Pp, postpineal plate; socc, supraoccipital cross-commissural pit-line groove; sot, supraotic thickening of head shield." figureDoi="http://doi.org/10.5281/zenodo.10980835" httpUri="https://zenodo.org/record/10980835/files/figure.png" pageId="10" pageNumber="107">Fig. 7A</figureCitation>
). The postero-lateral angle of the otico-occipital depression is complicated by two postero-lateral projections. Paranuchal trochlea does not reach the lateral margin of the plate.
</paragraph>
<paragraph id="E9483B2C4C5DFFEAFC76843CA5D8F71E" blockId="10.[862,1535,1079,1588]" pageId="10" pageNumber="107">
The postmarginal plate IG KSC 155/161 (
<figureCitation id="71CC27A94C5DFFEAFAB0843CA5AFF4B9" box="[1346,1420,1402,1427]" captionStart="Fig" captionStartId="5.[98,131,1308,1328]" captionTargetBox="[103,1477,164,1280]" captionTargetId="figure-314@5.[99,1480,160,1281]" captionTargetPageId="5" captionText="Fig. 4. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, lateral AG), premedian (H), and left postmarginal (I) plates of the head shield. A. IG KSC 155/15 in dorsal view. B. IG KSC 155/67 in dorsal (B1) and visceral B2) views. C. IG KSC 155/129 in dorsal view. D. IG KSC 155/65 in visceral (D1) and dorsal (D2) views. E. IG KSC 155/66 in dorsal (E1), right lateral (E2), and visceral (E3) views. F. IG KSC 155/69 in dorsal view. G. IG KSC 155/64 in dorsal view. H. IG KSC 155/114 in dorsal (H1) and anterior (H2) views.. Plate disconnected from the partial head shield IG KSC 155/161, in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10980829" httpUri="https://zenodo.org/record/10980829/files/figure.png" pageId="10" pageNumber="107">Fig. 4J</figureCitation>
) is shorter and broader than it is in
<taxonomicName id="2EF740AF4C5DFFEAFB8684DCA55EF499" baseAuthorityName="Luksevics" baseAuthorityYear="2001" box="[1140,1405,1434,1459]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="107" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="curonica">
<emphasis id="DB83E73E4C5DFFEAFB8684DCA55EF499" box="[1140,1405,1434,1459]" italics="true" pageId="10" pageNumber="107">B. leptocheira curonica</emphasis>
</taxonomicName>
(
<bibRefCitation id="8D6646DD4C5DFFEAFA7E84DCA3B4F4F9" author="Luksevics, E." pageId="10" pageNumber="107" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
: figs. 32A, B, F, 33A, B, F), more closely resembling
<taxonomicName id="2EF740AF4C5DFFEAFCAC849DA4A4F4DE" box="[862,1159,1499,1524]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="107" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="leptocheira">
<emphasis id="DB83E73E4C5DFFEAFCAC849DA4A4F4DE" box="[862,1159,1499,1524]" italics="true" pageId="10" pageNumber="107">B. leptocheira leptocheira</emphasis>
</taxonomicName>
(
<bibRefCitation id="8D6646DD4C5DFFEAFB68849DA53CF4DE" author="Miles, R. S." box="[1178,1311,1499,1524]" pageId="10" pageNumber="107" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles 1968</bibRefCitation>
: text-fig. 36) by its irregular quadrangular shape with an almost straight postero-mesial margin and strongly convex antero-mesial margin.
</paragraph>
<caption id="BD886BA44C5DFFEAFF868712A5D1F68E" ID-DOI="http://doi.org/10.5281/zenodo.11061576" ID-Zenodo-Dep="11061576" httpUri="https://zenodo.org/record/11061576/files/figure.png" pageId="10" pageNumber="107" startId="10.[116,144,1620,1650]" subCaptionStartIDs="10.[151,184,1628,1648]" subCaptionStarts="Fig. 9" targetBox="[99,1479,176,1937]" targetPageId="9">
<paragraph id="E9483B2C4C5DFFEAFF868712A5D1F68E" blockId="10.[116,1535,1620,1959]" pageId="10" pageNumber="107">
<emphasis id="DB83E73E4C5DFFEAFF868712A0B3F758" bold="true" box="[116,144,1620,1650]" pageId="10" pageNumber="107"></emphasis>
Fig. 9. Antiarchan fish
<taxonomicName id="2EF740AF4C5DFFEAFE9C871AA31EF75A" authority="(Rohon, 1900)" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" baseAuthorityName="Rohon" baseAuthorityYear="1900" box="[366,829,1628,1648]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="107" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="jeremejevi">
<emphasis id="DB83E73E4C5DFFEAFE9C871AA28CF75A" box="[366,687,1628,1648]" italics="true" pageId="10" pageNumber="107">Bothriolepis leptocheira jeremejevi</emphasis>
(
<bibRefCitation id="8D6646DD4C5DFFEAFD4E871AA315F75A" author="Rohon, J. V." box="[700,822,1628,1648]" pageId="10" pageNumber="107" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon, 1900</bibRefCitation>
)
</taxonomicName>
, Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior median dorsal (AG) and posterior median dorsal (HM) plates of the trunk armour.
<emphasis id="DB83E73E4C5DFFEAFC648731A385F7A6" bold="true" box="[918,934,1655,1676]" pageId="10" pageNumber="107">A</emphasis>
. IG KSC 155/
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dorsal (A
<subScript id="757339694C5DFFEAFB5487C7A48DF7A5" attach="left" box="[1190,1198,1665,1679]" fontSize="6" pageId="10" pageNumber="107">1</subScript>
) and visceral (A
<subScript id="757339694C5DFFEAFAB787C7A56EF7A5" attach="left" box="[1349,1357,1665,1679]" fontSize="6" pageId="10" pageNumber="107">2</subScript>
) views.
<emphasis id="DB83E73E4C5DFFEAFA688731A58AF7A6" bold="true" box="[1434,1449,1655,1676]" pageId="10" pageNumber="107">B</emphasis>
. IG KSC 155/
<quantity id="2E0F96C94C5DFFEAFF3287D2A0DAF782" box="[192,249,1684,1704]" metricMagnitude="0" metricUnit="m" metricValue="2.7432" pageId="10" pageNumber="107" unit="in" value="108.0">108 in</quantity>
dorsal (B
<subScript id="757339694C5DFFEAFEA087DBA179F781" attach="left" box="[338,346,1693,1707]" fontSize="6" pageId="10" pageNumber="107">1</subScript>
) and visceral (B
<subScript id="757339694C5DFFEAFE0287DBA1DBF781" attach="left" box="[496,504,1693,1707]" fontSize="6" pageId="10" pageNumber="107">2</subScript>
) views.
<emphasis id="DB83E73E4C5DFFEAFDB787D5A276F783" bold="true" box="[581,597,1683,1705]" pageId="10" pageNumber="107">C</emphasis>
. IG KSC 155/
<quantity id="2E0F96C94C5DFFEAFD2A87D2A324F782" box="[728,775,1684,1704]" metricMagnitude="0" metricUnit="m" metricValue="2.4638" pageId="10" pageNumber="107" unit="in" value="97.0">97 in</quantity>
dorsal view.
<emphasis id="DB83E73E4C5DFFEAFC8D87D5A3ACF782" bold="true" box="[895,911,1683,1704]" pageId="10" pageNumber="107">D</emphasis>
. IG KSC 155/
<quantity id="2E0F96C94C5DFFEAFBE187D2A468F782" box="[1043,1099,1684,1704]" metricMagnitude="0" metricUnit="m" metricValue="2.8701999999999996" pageId="10" pageNumber="107" unit="in" value="113.0">113 in</quantity>
dorsal (D
<subScript id="757339694C5DFFEAFB5787DBA48EF781" attach="left" box="[1189,1197,1693,1707]" fontSize="6" pageId="10" pageNumber="107">1</subScript>
) and visceral (D
<subScript id="757339694C5DFFEAFAB787DBA56EF781" attach="left" box="[1349,1357,1693,1707]" fontSize="6" pageId="10" pageNumber="107">2</subScript>
) views.
<emphasis id="DB83E73E4C5DFFEAFA6887D5A58AF782" bold="true" box="[1434,1449,1683,1704]" pageId="10" pageNumber="107">E</emphasis>
. IG KSC 155/
<quantity id="2E0F96C94C5DFFEAFF4D87F6A0DBF7EE" box="[191,248,1712,1732]" metricMagnitude="0" metricUnit="m" metricValue="3.556" pageId="10" pageNumber="107" unit="in" value="140.0">140 in</quantity>
dorsal (E
<subScript id="757339694C5DFFEAFEBF87FFA176F7ED" attach="left" box="[333,341,1721,1735]" fontSize="6" pageId="10" pageNumber="107">1</subScript>
) and visceral (E
<subScript id="757339694C5DFFEAFE1587FFA1CCF7ED" attach="left" box="[487,495,1721,1735]" fontSize="6" pageId="10" pageNumber="107">2</subScript>
) views.
<emphasis id="DB83E73E4C5DFFEAFDC887E9A26BF7EE" bold="true" box="[570,584,1711,1732]" pageId="10" pageNumber="107">F</emphasis>
. Impression of the dorsal surface of IG KSC 155/42.
<emphasis id="DB83E73E4C5DFFEAFBD187E9A416F7EF" bold="true" box="[1059,1077,1711,1733]" pageId="10" pageNumber="107">G</emphasis>
. IG KSC 155/
<quantity id="2E0F96C94C5DFFEAFB4687F6A4C0F7EE" box="[1204,1251,1712,1732]" metricMagnitude="0" metricUnit="m" metricValue="1.1176" pageId="10" pageNumber="107" unit="in" value="44.0">44 in</quantity>
dorsal view.
<emphasis id="DB83E73E4C5DFFEAFAAA87E9A549F7EE" bold="true" box="[1368,1386,1711,1732]" pageId="10" pageNumber="107">H</emphasis>
. Fragment of IG KSC 155/
<quantity id="2E0F96C94C5DFFEAFF03878AA131F7CA" box="[241,274,1740,1760]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="10" pageNumber="107" unit="in" value="7.0">7 in</quantity>
dorsal view.
<emphasis id="DB83E73E4C5DFFEAFE7A878DA1B2F7CA" bold="true" box="[392,401,1739,1760]" pageId="10" pageNumber="107">I</emphasis>
. IG KSC 155/
<quantity id="2E0F96C94C5DFFEAFDE7878AA210F7CA" box="[533,563,1740,1760]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="10" pageNumber="107" unit="in" value="1.0">1 in</quantity>
dorsal (I
<subScript id="757339694C5DFFEAFD718793A2A8F7C9" attach="left" box="[643,651,1749,1763]" fontSize="6" pageId="10" pageNumber="107">1</subScript>
) and visceral (I
<subScript id="757339694C5DFFEAFCE48793A33DF7C9" attach="left" box="[790,798,1749,1763]" fontSize="6" pageId="10" pageNumber="107">2</subScript>
) views.
<emphasis id="DB83E73E4C5DFFEAFC98878DA356F7CA" bold="true" box="[874,885,1739,1760]" pageId="10" pageNumber="107">J</emphasis>
. IG KSC 155/
<quantity id="2E0F96C94C5DFFEAFC04878AA400F7CA" box="[1014,1059,1740,1760]" metricMagnitude="0" metricUnit="m" metricValue="1.8034" pageId="10" pageNumber="107" unit="in" value="71.0">71 in</quantity>
dorsal view.
<emphasis id="DB83E73E4C5DFFEAFB6A878DA489F7CA" bold="true" box="[1176,1194,1739,1760]" pageId="10" pageNumber="107">K</emphasis>
. Slightly deformed IG KSC 155/
<quantity id="2E0F96C94C5DFFEAFA23878AA5DCF7CA" box="[1489,1535,1740,1760]" metricMagnitude="0" metricUnit="m" metricValue="1.778" pageId="10" pageNumber="107" unit="in" value="70.0">70 in</quantity>
dorsal (K
<subScript id="757339694C5DFFEAFF1987B7A0D0F7D5" attach="left" box="[235,243,1777,1791]" fontSize="6" pageId="10" pageNumber="107">1</subScript>
) and visceral (K
<subScript id="757339694C5DFFEAFE7B87B7A1B2F7D5" attach="left" box="[393,401,1777,1791]" fontSize="6" pageId="10" pageNumber="107">2</subScript>
) views.
<emphasis id="DB83E73E4C5DFFEAFE2C87A1A1CEF7D6" bold="true" box="[478,493,1767,1788]" pageId="10" pageNumber="107">L</emphasis>
. IG KSC 155/
<quantity id="2E0F96C94C5DFFEAFD9D87AEA28BF7D6" box="[623,680,1768,1788]" metricMagnitude="0" metricUnit="m" metricValue="4.013199999999999" pageId="10" pageNumber="107" unit="in" value="158.0">158 in</quantity>
dorsal view.
<emphasis id="DB83E73E4C5DFFEAFCED87A1A317F7D6" bold="true" box="[799,820,1767,1788]" pageId="10" pageNumber="107">M</emphasis>
. IG KSC 155/
<quantity id="2E0F96C94C5DFFEAFC4487AEA3CCF7D6" box="[950,1007,1768,1788]" metricMagnitude="0" metricUnit="m" metricValue="3.9878" pageId="10" pageNumber="107" unit="in" value="157.0">157 in</quantity>
dorsal (M
<subScript id="757339694C5DFFEAFBBE87B7A477F7D5" attach="left" box="[1100,1108,1777,1791]" fontSize="6" pageId="10" pageNumber="107">1</subScript>
) and visceral (M
<subScript id="757339694C5DFFEAFB1C87B7A4D5F7D5" attach="left" box="[1262,1270,1777,1791]" fontSize="6" pageId="10" pageNumber="107">2</subScript>
) views. Abbreviations:
<collectionCode id="8FE6A3E94C5DFFEAFA3987AEA5DFF7D6" box="[1483,1532,1768,1788]" pageId="10" pageNumber="107">ADL</collectionCode>
, anterior dorso-lateral plate; alr, postlevator thickening;
<collectionCode id="8FE6A3E94C5DFFEAFD708642A29AF632" box="[642,697,1796,1816]" pageId="10" pageNumber="107">AMD</collectionCode>
, anterior median dorsal plate; cf.
<collectionCode id="8FE6A3E94C5DFFEAFC2E8642A42EF632" box="[988,1037,1796,1816]" pageId="10" pageNumber="107">ADL</collectionCode>
, cf.
<collectionCode id="8FE6A3E94C5DFFEAFBDE8642A447F632" box="[1068,1124,1796,1816]" pageId="10" pageNumber="107">AMD</collectionCode>
, and cf.
<collectionCode id="8FE6A3E94C5DFFEAFB5A8642A4FBF632" box="[1192,1240,1796,1816]" pageId="10" pageNumber="107">MxL</collectionCode>
, area overlapping
<collectionCode id="8FE6A3E94C5DFFEAFA888642A588F632" box="[1402,1451,1796,1816]" pageId="10" pageNumber="107">ADL</collectionCode>
,
<collectionCode id="8FE6A3E94C5DFFEAFA408642A5C4F632" box="[1458,1511,1796,1816]" pageId="10" pageNumber="107">AMD</collectionCode>
or
<collectionCode id="8FE6A3E94C5DFFEAFF658666A0E0F61E" box="[151,195,1824,1844]" pageId="10" pageNumber="107">MxL</collectionCode>
respectively; cr.tp, posterior transversal internal crest; dlg
<subScript id="757339694C5DFFEAFD3B866FA2F2F61D" attach="left" box="[713,721,1833,1847]" fontSize="6" pageId="10" pageNumber="107">1</subScript>
and dlg
<subScript id="757339694C5DFFEAFCEB866FA302F61D" attach="left" box="[793,801,1833,1847]" fontSize="6" pageId="10" pageNumber="107">2</subScript>
, anterior and posterior oblique dorsal sensory line groove; dma, tergal angle; dmr, dorsal median ridge; f.retr, levator fossa; grm, ventral median groove; l, lateral corner; mvr, median ventral ridge;
<collectionCode id="8FE6A3E94C5DFFEAFB68867AA4EAF67A" box="[1178,1225,1852,1872]" pageId="10" pageNumber="107">MxL</collectionCode>
, mixilateral plate; npn, postnuchal notch; oa.
<collectionCode id="8FE6A3E94C5DFFEAFF1C861EA103F646" box="[238,288,1880,1900]" pageId="10" pageNumber="107">ADL</collectionCode>
, oa.
<collectionCode id="8FE6A3E94C5DFFEAFEB3861EA153F646" box="[321,368,1880,1900]" pageId="10" pageNumber="107">MxL</collectionCode>
and oa.
<collectionCode id="8FE6A3E94C5DFFEAFE47861EA1C9F646" box="[437,490,1880,1900]" pageId="10" pageNumber="107">PMD</collectionCode>
, area overlapped by
<collectionCode id="8FE6A3E94C5DFFEAFD50861EA2F0F646" box="[674,723,1880,1900]" pageId="10" pageNumber="107">ADL</collectionCode>
,
<collectionCode id="8FE6A3E94C5DFFEAFD29861EA324F646" box="[731,775,1880,1900]" pageId="10" pageNumber="107">MxL</collectionCode>
or
<collectionCode id="8FE6A3E94C5DFFEAFCD7861EA375F646" box="[805,854,1880,1900]" pageId="10" pageNumber="107">PMD</collectionCode>
respectively; pa, posterior corner; pma, posterior marginal area;
<collectionCode id="8FE6A3E94C5DFFEAFA6E861EA5ECF646" box="[1436,1487,1880,1900]" pageId="10" pageNumber="107">PMD</collectionCode>
, posterior median dorsal plate; pr.p, posterior process of
<collectionCode id="8FE6A3E94C5DFFEAFD948632A2BEF6A2" box="[614,669,1908,1928]" pageId="10" pageNumber="107">AMD</collectionCode>
; pr.pl, external postlevator process; prv
<subScript id="757339694C5DFFEAFC08863BA421F6A1" attach="left" box="[1018,1026,1917,1931]" fontSize="6" pageId="10" pageNumber="107">2</subScript>
, posterior ventral process of dorsal wall of trunk armour; pt
<subScript id="757339694C5DFFEAFF5B86DFA092F68D" attach="left" box="[169,177,1945,1959]" fontSize="6" pageId="10" pageNumber="107">1</subScript>
and pt
<subScript id="757339694C5DFFEAFF1F86DFA0D6F68D" attach="left" box="[237,245,1945,1959]" fontSize="6" pageId="10" pageNumber="107">2</subScript>
, anterior and posterior ventral pit; pua, posterior unornamented area of
<collectionCode id="8FE6A3E94C5DFFEAFC8586D6A388F68E" box="[887,939,1936,1956]" pageId="10" pageNumber="107">PMD</collectionCode>
; rf, “round fossula”; sna, supranuchal area; tb, ventral tuberosity.
</paragraph>
</caption>
<caption id="BD886BA44C5CFFECFF908667A58EF68F" ID-DOI="http://doi.org/10.5281/zenodo.10980841" ID-Zenodo-Dep="10980841" httpUri="https://zenodo.org/record/10980841/files/figure.png" lastPageId="12" lastPageNumber="109" pageId="11" pageNumber="108" startId="11.[98,131,1825,1845]" targetBox="[99,1479,163,1803]" targetPageId="11" targetType="figure">
<paragraph id="E9483B2C4C5CFFEBFF908667A5E9F67B" blockId="11.[98,1521,1825,1960]" pageId="11" pageNumber="108">
Fig. 11. Antiarchan fish
<taxonomicName id="2EF740AF4C5CFFEBFEB08667A330F61F" authority="(Rohon, 1900)" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" baseAuthorityName="Rohon" baseAuthorityYear="1900" box="[322,787,1825,1845]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="108" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="jeremejevi">
<emphasis id="DB83E73E4C5CFFEBFEB08667A2A6F61F" box="[322,645,1825,1845]" italics="true" pageId="11" pageNumber="108">Bothriolepis leptocheira jeremejevi</emphasis>
(
<bibRefCitation id="8D6646DD4C5CFFEBFD608667A32FF61F" author="Rohon, J. V." box="[658,780,1825,1845]" pageId="11" pageNumber="108" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon, 1900</bibRefCitation>
)
</taxonomicName>
, Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, partial ventral wall (A, B), anterior ventro-lateral (AVL) (C, D, left; E, right), and median ventral (FJ) plates of the trunk armour, and elements of pectoral fin.
</paragraph>
<paragraph id="E9483B2C4C5CFFECFF908611A58EF68F" blockId="11.[98,1521,1825,1960]" lastBlockId="12.[151,1535,1852,1961]" lastPageId="12" lastPageNumber="109" pageId="11" pageNumber="108">
<emphasis id="DB83E73E4C5CFFEBFF908611A051F646" bold="true" box="[98,114,1879,1900]" pageId="11" pageNumber="108">A</emphasis>
. Impression of partial ventral wall of the trunk armour, inferognathals, and suborbitals IG KSC 155/10-2 in ventral view.
<emphasis id="DB83E73E4C5CFFEBFB358611A4F5F646" bold="true" box="[1223,1238,1879,1900]" pageId="11" pageNumber="108">B</emphasis>
. Partial ventral wall of the trunk armour with articulated pectoral fin armour IG KSC 155/10-1 in ventral view.
<emphasis id="DB83E73E4C5CFFEBFC958635A354F6A3" bold="true" box="[871,887,1907,1929]" pageId="11" pageNumber="108">C</emphasis>
. IG KSC 155/151 (C
<subScript id="757339694C5CFFEBFBC98638A460F6A6" attach="left" box="[1083,1091,1918,1932]" fontSize="6" pageId="11" pageNumber="108">1</subScript>
) and enlargened part of the ventral lamina with pathologies (C
<subScript id="757339694C5CFFEBFEE486DCA13DF682" attach="left" box="[278,286,1946,1960]" fontSize="6" pageId="11" pageNumber="108">2</subScript>
, arrows).
<emphasis id="DB83E73E4C5CFFEBFE8986C9A1A8F68E" bold="true" box="[379,395,1935,1956]" pageId="11" pageNumber="108">D</emphasis>
. IG KSC 155/122 in ventral view.
<emphasis id="DB83E73E4C5CFFEBFD3E86C9A2F8F68E" bold="true" box="[716,731,1935,1956]" pageId="11" pageNumber="108">E</emphasis>
. IG KSC 155/123 in ventral view.
<emphasis id="DB83E73E4C5CFFEBFBEE86C9A409F68E" bold="true" box="[1052,1066,1935,1956]" pageId="11" pageNumber="108">F</emphasis>
. IG KSC 155/104 with the bite mark (arrow) → in ventral (F
<subScript id="757339694C5BFFECFEFA8601A133F67F" attach="left" box="[264,272,1863,1877]" fontSize="6" pageId="12" pageNumber="109">1</subScript>
) and visceral (F
<subScript id="757339694C5BFFECFE578601A18EF67F" attach="left" box="[421,429,1863,1877]" fontSize="6" pageId="12" pageNumber="109">2</subScript>
) views.
<emphasis id="DB83E73E4C5BFFECFE09867AA22EF678" bold="true" box="[507,525,1852,1874]" pageId="12" pageNumber="109">G</emphasis>
. IG KSC 155/
<quantity id="2E0F96C94C5BFFECFD63867BA2E3F67B" box="[657,704,1853,1873]" metricMagnitude="0" metricUnit="m" metricValue="1.9304" pageId="12" pageNumber="109" unit="in" value="76.0">76 in</quantity>
ventral view.
<emphasis id="DB83E73E4C5BFFECFCB3867AA370F67B" bold="true" box="[833,851,1852,1873]" pageId="12" pageNumber="109">H</emphasis>
. IG KSC 155/
<quantity id="2E0F96C94C5BFFECFC25867BA425F67B" box="[983,1030,1853,1873]" metricMagnitude="0" metricUnit="m" metricValue="1.9558" pageId="12" pageNumber="109" unit="in" value="77.0">77 in</quantity>
ventral view.
<emphasis id="DB83E73E4C5BFFECFB75867AA4B3F67B" bold="true" box="[1159,1168,1852,1873]" pageId="12" pageNumber="109">I</emphasis>
. IG KSC 155/
<quantity id="2E0F96C94C5BFFECFAE7867BA56DF67B" box="[1301,1358,1853,1873]" metricMagnitude="0" metricUnit="m" metricValue="3.3781999999999996" pageId="12" pageNumber="109" unit="in" value="133.0">133 in</quantity>
ventral view.
<emphasis id="DB83E73E4C5BFFECFA3D867AA5F9F67B" bold="true" box="[1487,1498,1852,1873]" pageId="12" pageNumber="109">J</emphasis>
. IG KSC 155/
<quantity id="2E0F96C94C5BFFECFF03861FA10AF647" box="[241,297,1881,1901]" metricMagnitude="0" metricUnit="m" metricValue="2.6924" pageId="12" pageNumber="109" unit="in" value="106.0">106 in</quantity>
ventral view. Abbreviations:
<collectionCode id="8FE6A3E94C5BFFECFDC2861FA27DF647" box="[560,606,1881,1901]" pageId="12" pageNumber="109">AVL</collectionCode>
, anterior ventro-lateral plate; Cd
<subScript id="757339694C5BFFECFC708625A3B9F65B" attach="left" box="[898,922,1891,1905]" fontSize="6" pageId="12" pageNumber="109">12</subScript>
, dorsal central plates 1 and 2; Cv
<subScript id="757339694C5BFFECFB368625A4EFF65B" attach="left" box="[1220,1228,1891,1905]" fontSize="6" pageId="12" pageNumber="109">1</subScript>
, ventral central plate 1; cit
<subScript id="757339694C5BFFECFA498625A5E0F65B" attach="left" box="[1467,1475,1891,1905]" fontSize="6" pageId="12" pageNumber="109">1</subScript>
, crista transversalis interna anterior; cit
<subScript id="757339694C5BFFECFE4F8639A1E6F6A7" attach="left" box="[445,453,1919,1933]" fontSize="6" pageId="12" pageNumber="109">2</subScript>
, transverse thickening on the visceral surface of
<collectionCode id="8FE6A3E94C5BFFECFC708633A392F6A3" box="[898,945,1909,1929]" pageId="12" pageNumber="109">AVL</collectionCode>
; ds, distal segment of the pectoral fin armour; Ifg, inferognathal; Ml
<subScript id="757339694C5BFFECFF4086DDA099F683" attach="right" box="[178,186,1947,1961]" fontSize="6" pageId="12" pageNumber="109">2</subScript>
, lateral marginal plate 2; Mm
<subScript id="757339694C5BFFECFE3A86DDA1F3F683" attach="left" box="[456,464,1947,1961]" fontSize="6" pageId="12" pageNumber="109">2</subScript>
, mesial marginal plate 2;
<collectionCode id="8FE6A3E94C5BFFECFD4986D7A2FCF68F" box="[699,735,1937,1957]" country="0" name="University of Montana Museum" pageId="12" pageNumber="109">MV</collectionCode>
, median ventral plate;
<collectionCode id="8FE6A3E94C5BFFECFC5F86D7A3F9F68F" box="[941,986,1937,1957]" country="Argentina" httpUri="http://grbio.org/cool/pbkd-hmeb" name="Paleontologia de Vertebrados Lillo" pageId="12" pageNumber="109">PVL</collectionCode>
, posterior ventro-lateral plate;
<collectionCode id="8FE6A3E94C5BFFECFB0386D7A533F68F" box="[1265,1296,1937,1957]" pageId="12" pageNumber="109">SO</collectionCode>
, suborbital plate.
</paragraph>
</caption>
<caption id="BD886BA44C5BFFECFF6582F8A229F520" ID-DOI="http://doi.org/10.5281/zenodo.10980843" ID-Zenodo-Dep="10980843" httpUri="https://zenodo.org/record/10980843/files/figure.png" pageId="12" pageNumber="109" startId="12.[151,184,958,978]" targetBox="[161,1528,166,928]" targetPageId="12" targetType="figure">
<paragraph id="E9483B2C4C5BFFECFF6582F8A229F520" blockId="12.[151,1535,958,1035]" pageId="12" pageNumber="109">
Fig. 12. Antiarchan fish
<taxonomicName id="2EF740AF4C5BFFECFE8182F8A31EF2F8" authority="(Rohon, 1900)" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" baseAuthorityName="Rohon" baseAuthorityYear="1900" box="[371,829,958,978]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="109" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="jeremejevi">
<emphasis id="DB83E73E4C5BFFECFE8182F8A291F2F8" box="[371,690,958,978]" italics="true" pageId="12" pageNumber="109">Bothriolepis leptocheira jeremejevi</emphasis>
(
<bibRefCitation id="8D6646DD4C5BFFECFD4F82F8A315F2F8" author="Rohon, J. V." box="[701,822,958,978]" pageId="12" pageNumber="109" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon, 1900</bibRefCitation>
)
</taxonomicName>
, Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, posterior ventro-lateral plates, left (A) and right (B, C) of the trunk armour.
<emphasis id="DB83E73E4C5BFFECFD07829FA326F2C4" bold="true" box="[757,773,985,1006]" pageId="12" pageNumber="109">A</emphasis>
. IG KSC 155/4 in ventral (A
<subScript id="757339694C5BFFECFBFD82A5A434F2DB" attach="left" box="[1039,1047,995,1009]" fontSize="6" pageId="12" pageNumber="109">1</subScript>
) and lateral (A
<subScript id="757339694C5BFFECFB5082A5A489F2DB" attach="right" box="[1186,1194,995,1009]" fontSize="6" pageId="12" pageNumber="109">2</subScript>
) views.
<emphasis id="DB83E73E4C5BFFECFB0A829FA524F2C4" bold="true" box="[1272,1287,985,1006]" pageId="12" pageNumber="109">B</emphasis>
. IG KSC 155/16 in ventral view.
<emphasis id="DB83E73E4C5BFFECFF3F82B3A0FEF521" bold="true" box="[205,221,1013,1035]" pageId="12" pageNumber="109">C</emphasis>
. IG KSC 155/78 in ventral view.
</paragraph>
</caption>
<paragraph id="E9483B2C4C5BFFECFF4F8567A177F7F9" blockId="12.[151,826,1057,1813]" pageId="12" pageNumber="109">
Thesuborbitalplates(
<collectionCode id="8FE6A3E94C5BFFECFE438567A1F4F510" box="[433,471,1057,1082]" pageId="12" pageNumber="109">SO</collectionCode>
)havenotbeenpreviouslyfound in this species. The right and left
<collectionCode id="8FE6A3E94C5BFFECFDDD8504A270F571" box="[559,595,1090,1115]" pageId="12" pageNumber="109">SO</collectionCode>
are preserved only from one individual: in IG KSC 155/10-3 these are seen in visceral (internal) view (
<figureCitation id="71CC27A94C5BFFECFE5D85C3A225F5B4" box="[431,518,1157,1182]" captionStart="Fig" captionStartId="4.[151,184,1188,1208]" captionTargetBox="[157,1530,160,1164]" captionTargetId="figure-306@4.[173,1531,159,1164]" captionTargetPageId="4" captionText="Fig. 3. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, separate plates and parts of articulated head shields. A. Articulated Nu, left Pn and left La plates IG KSC 155/90 in dorsal view. B. Disarticulated plates of the head shield IG KSC 155/150 in dorsal view. C. Part of articulated head shield IG KSC 155/162 in dorsal view. D. Imprint of the ventral wall of trunk armour and some skeletal elements of the head armour IG KSC 155/10-3 in ventral view showing suborbital and infragnathal bones. Abbreviations: AVL, anterior ventro-lateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; ifc1, principal section of infraorbital sensory line; IG, inferognathal; La, lateral plate; Nu, nuchal plate; mp, middle pit-line groove; Pn, paranuchal plate; Pp, postpineal plate; Prm, premedian plate; SO, suborbital plate; socc, supraoccipital cross-commissural pit-line groove." figureDoi="http://doi.org/10.5281/zenodo.10980827" httpUri="https://zenodo.org/record/10980827/files/figure.png" pageId="12" pageNumber="109">Fig. 3D</figureCitation>
), and the IG KSC 155/10-2 (
<figureCitation id="71CC27A94C5BFFECFF6D85E0A0DCF595" box="[159,255,1190,1215]" captionStart="Fig" captionStartId="11.[98,131,1825,1845]" captionTargetBox="[99,1479,163,1803]" captionTargetId="figure-8@11.[99,1480,163,1804]" captionTargetPageId="11" captionText="Fig. 11. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, partial ventral wall (A, B), anterior ventro-lateral (AVL) (C, D, left; E, right), and median ventral (FJ) plates of the trunk armour, and elements of pectoral fin. A. Impression of partial ventral wall of the trunk armour, inferognathals, and suborbitals IG KSC 155/10-2 in ventral view. B. Partial ventral wall of the trunk armour with articulated pectoral fin armour IG KSC 155/10-1 in ventral view. C. IG KSC 155/151 (C1) and enlargened part of the ventral lamina with pathologies (C2, arrows). D. IG KSC 155/122 in ventral view. E. IG KSC 155/123 in ventral view. F. IG KSC 155/104 with the bite mark (arrow) → in ventral (F1) and visceral (F2) views. G. IG KSC 155/76 in ventral view. H. IG KSC 155/77 in ventral view. I. IG KSC 155/133 in ventral view. J. IG KSC 155/106 in ventral view. Abbreviations:AVL, anterior ventro-lateral plate; Cd12, dorsal central plates 1 and 2; Cv1, ventral central plate 1; cit1, crista transversalis interna anterior; cit2, transverse thickening on the visceral surface of AVL; ds, distal segment of the pectoral fin armour; Ifg, inferognathal; Ml2, lateral marginal plate 2; Mm2, mesial marginal plate 2; MV, median ventral plate; PVL, posterior ventro-lateral plate; SO, suborbital plate." figureDoi="http://doi.org/10.5281/zenodo.10980841" httpUri="https://zenodo.org/record/10980841/files/figure.png" pageId="12" pageNumber="109">Fig. 11A</figureCitation>
) shows their cast. The shape of the plate is rhombic with a slightly convex anterior and an almost straight posterior margin. The lateral margin is slightly concave, forming a prominent postero-lateral corner. The mesial margin is oblique therefore the split between pair of suborbital plates widens posteriorly. The ventral margin seems to bear rather small tubercles showing weakly defined denticulation. The visceral surface is dominated by a pronounced transverse ridge, which begins near the dorso-mesial corner, and extends obliquely, towards the middle of the lateral edge of the plate. It forms a posteriorly oriented convex curve. The height of this ridge varies: large close to the mesial margin and becoming lower laterally. The morphology of the visceral surface of
<collectionCode id="8FE6A3E94C5BFFECFEA28710A157F745" box="[336,372,1622,1647]" pageId="12" pageNumber="109">SO</collectionCode>
is almost similar to those described for other species of
<taxonomicName id="2EF740AF4C5BFFECFEA28731A1F8F7BA" authorityName="Eichwald" authorityYear="1840" box="[336,475,1655,1680]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="109" phylum="Chordata" rank="genus">
<emphasis id="DB83E73E4C5BFFECFEA28731A1F8F7BA" box="[336,475,1655,1680]" italics="true" pageId="12" pageNumber="109">Bothriolepis</emphasis>
</taxonomicName>
:
<taxonomicName id="2EF740AF4C5BFFECFE198731A316F7BA" authority="(Stensio 1948)" baseAuthorityName="Stensio" baseAuthorityYear="1948" box="[491,821,1655,1680]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="109" phylum="Chordata" rank="species" species="canadensis">
<emphasis id="DB83E73E4C5BFFECFE198731A2ABF7BA" box="[491,648,1655,1680]" italics="true" pageId="12" pageNumber="109">B. canadensis</emphasis>
(
<bibRefCitation id="8D6646DD4C5BFFECFD6A8731A30FF7BA" author="Stensio, E. A." box="[664,812,1655,1680]" pageId="12" pageNumber="109" pagination="1 - 622" refId="ref20500" refString="Stensio, E. A. 1948. On the Placodermi of the Upper Devonian of East Greenland. II: Antiarchi: subfamily Bothriolepinae. Palaeozoologica Groenlandica 2: 1 - 622." type="journal article" year="1948">Stensiö 1948</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="2EF740AF4C5BFFECFF6587DEA169F79B" box="[151,330,1688,1713]" family="Bothriolepididae" genus="Bothriolepis" kingdom="Animalia" pageId="12" pageNumber="109" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="DB83E73E4C5BFFECFF6587DEA100F79B" box="[151,291,1688,1713]" italics="true" pageId="12" pageNumber="109">Bothriolepis</emphasis>
sp.
</taxonomicName>
from Gogo (
<bibRefCitation id="8D6646DD4C5BFFECFE1187DEA253F79B" author="Young, G. C." box="[483,624,1688,1713]" pageId="12" pageNumber="109" pagination="635 - 661" refId="ref20753" refString="Young, G. C. 1984. Reconstruction of the jaws and braincase in the Devonian placoderm fish Bothriolepis. Palaeontology 27: 635 - 661." type="journal article" year="1984">Young 1984</bibRefCitation>
), and
<taxonomicName id="2EF740AF4C5BFFECFD4B87DEA173F7F9" authority="Johanson (1998)" authorityName="Johanson" authorityYear="1998" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="109" phylum="Chordata" rank="species" species="yeungae">
<emphasis id="DB83E73E4C5BFFECFD4B87DEA31BF79B" box="[697,824,1688,1713]" italics="true" pageId="12" pageNumber="109">B. yeungae</emphasis>
<bibRefCitation id="8D6646DD4C5BFFECFF6587FCA173F7F9" author="Johanson, Z." box="[151,336,1722,1747]" pageId="12" pageNumber="109" pagination="315 - 348" refId="ref18897" refString="Johanson, Z. 1998. The Upper Devonian fish Bothriolepis (Placodermi: Antiarchi) from near Canowindra, New South Wales, Australia. Records of the Australian Museum 50 (3): 315 - 348." type="journal article" year="1998">Johanson (1998)</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="E9483B2C4C5BFFECFF4F879DA5F8F5B4" blockId="12.[151,826,1057,1813]" lastBlockId="12.[862,1536,1057,1813]" pageId="12" pageNumber="109">
The S-shaped infragnathals of
<taxonomicName id="2EF740AF4C5BFFECFDDE879DA2F2F7DE" authorityName="Miles" authorityYear="1968" box="[556,721,1755,1780]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="109" phylum="Chordata" rank="species" species="leptocheira">
<emphasis id="DB83E73E4C5BFFECFDDE879DA2F2F7DE" box="[556,721,1755,1780]" italics="true" pageId="12" pageNumber="109">B. leptocheira</emphasis>
</taxonomicName>
are also described here for the first time; they are preserved in the IG KSC 155/10-2 and IG KSC 155/10-3 (
<figureCitation id="71CC27A94C5BFFECFB0F8567A57EF510" box="[1277,1373,1057,1082]" captionStart="Fig" captionStartId="4.[151,184,1188,1208]" captionTargetBox="[157,1530,160,1164]" captionTargetId="figure-306@4.[173,1531,159,1164]" captionTargetPageId="4" captionText="Fig. 3. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, separate plates and parts of articulated head shields. A. Articulated Nu, left Pn and left La plates IG KSC 155/90 in dorsal view. B. Disarticulated plates of the head shield IG KSC 155/150 in dorsal view. C. Part of articulated head shield IG KSC 155/162 in dorsal view. D. Imprint of the ventral wall of trunk armour and some skeletal elements of the head armour IG KSC 155/10-3 in ventral view showing suborbital and infragnathal bones. Abbreviations: AVL, anterior ventro-lateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; ifc1, principal section of infraorbital sensory line; IG, inferognathal; La, lateral plate; Nu, nuchal plate; mp, middle pit-line groove; Pn, paranuchal plate; Pp, postpineal plate; Prm, premedian plate; SO, suborbital plate; socc, supraoccipital cross-commissural pit-line groove." figureDoi="http://doi.org/10.5281/zenodo.10980827" httpUri="https://zenodo.org/record/10980827/files/figure.png" pageId="12" pageNumber="109">Figs. 3D</figureCitation>
,
<figureCitation id="71CC27A94C5BFFECFA9A8567A5BAF510" box="[1384,1433,1057,1082]" captionStart="Fig" captionStartId="11.[98,131,1825,1845]" captionTargetBox="[99,1479,163,1803]" captionTargetId="figure-8@11.[99,1480,163,1804]" captionTargetPageId="11" captionText="Fig. 11. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, partial ventral wall (A, B), anterior ventro-lateral (AVL) (C, D, left; E, right), and median ventral (FJ) plates of the trunk armour, and elements of pectoral fin. A. Impression of partial ventral wall of the trunk armour, inferognathals, and suborbitals IG KSC 155/10-2 in ventral view. B. Partial ventral wall of the trunk armour with articulated pectoral fin armour IG KSC 155/10-1 in ventral view. C. IG KSC 155/151 (C1) and enlargened part of the ventral lamina with pathologies (C2, arrows). D. IG KSC 155/122 in ventral view. E. IG KSC 155/123 in ventral view. F. IG KSC 155/104 with the bite mark (arrow) → in ventral (F1) and visceral (F2) views. G. IG KSC 155/76 in ventral view. H. IG KSC 155/77 in ventral view. I. IG KSC 155/133 in ventral view. J. IG KSC 155/106 in ventral view. Abbreviations:AVL, anterior ventro-lateral plate; Cd12, dorsal central plates 1 and 2; Cv1, ventral central plate 1; cit1, crista transversalis interna anterior; cit2, transverse thickening on the visceral surface of AVL; ds, distal segment of the pectoral fin armour; Ifg, inferognathal; Ml2, lateral marginal plate 2; Mm2, mesial marginal plate 2; MV, median ventral plate; PVL, posterior ventro-lateral plate; SO, suborbital plate." figureDoi="http://doi.org/10.5281/zenodo.10980841" httpUri="https://zenodo.org/record/10980841/files/figure.png" pageId="12" pageNumber="109">11A</figureCitation>
,
<figureCitation id="71CC27A94C5BFFECFA518567A5E3F510" box="[1443,1472,1057,1082]" captionStart="Fig" captionStartId="15.[98,131,1132,1152]" captionTargetBox="[103,764,164,1109]" captionTargetId="figure-870@15.[102,764,160,1109]" captionTargetPageId="15" captionText="Fig. 14. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), reconstruction of the head, the trunk armour and the pectoral fins in ventral view, based on IG KSC 155/10-1, IG KSC 155/10-2, and IG KSC 155/10- 3. Abbreviations: AVL, anterior ventro-lateral plate; Cv14, ventral central plate 1 to 4; Ifg, inferognathal plate; La, lateral plate; Ml25, lateral marginal plates 2 to 5; Mm4, mesial marginal plate 4; MV, median ventral plate; Prl, prelateral plate; Prm, premedian plate; PVL, posterior ventro-lateral plate; SM, semilunar plate; SMg, submarginal plate; T, terminal plate." figureDoi="http://doi.org/10.5281/zenodo.10980847" httpUri="https://zenodo.org/record/10980847/files/figure.png" pageId="12" pageNumber="109">14</figureCitation>
). The lateral projection of the right infragnathal lies very close to the tip of posterolateral corner of
<collectionCode id="8FE6A3E94C5BFFECFB248525A4D9F556" box="[1238,1274,1123,1148]" pageId="12" pageNumber="109">SO</collectionCode>
(
<figureCitation id="71CC27A94C5BFFECFAFA8525A57DF556" box="[1288,1374,1123,1148]" captionStart="Fig" captionStartId="4.[151,184,1188,1208]" captionTargetBox="[157,1530,160,1164]" captionTargetId="figure-306@4.[173,1531,159,1164]" captionTargetPageId="4" captionText="Fig. 3. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, separate plates and parts of articulated head shields. A. Articulated Nu, left Pn and left La plates IG KSC 155/90 in dorsal view. B. Disarticulated plates of the head shield IG KSC 155/150 in dorsal view. C. Part of articulated head shield IG KSC 155/162 in dorsal view. D. Imprint of the ventral wall of trunk armour and some skeletal elements of the head armour IG KSC 155/10-3 in ventral view showing suborbital and infragnathal bones. Abbreviations: AVL, anterior ventro-lateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; ifc1, principal section of infraorbital sensory line; IG, inferognathal; La, lateral plate; Nu, nuchal plate; mp, middle pit-line groove; Pn, paranuchal plate; Pp, postpineal plate; Prm, premedian plate; SO, suborbital plate; socc, supraoccipital cross-commissural pit-line groove." figureDoi="http://doi.org/10.5281/zenodo.10980827" httpUri="https://zenodo.org/record/10980827/files/figure.png" pageId="12" pageNumber="109">Fig. 3D</figureCitation>
). Infragnathal is strongly curved in the mesial part and tapers mesially.
</paragraph>
<paragraph id="E9483B2C4C5BFFEDFC7685E0A239F070" blockId="12.[862,1536,1057,1813]" lastBlockId="13.[98,771,160,1960]" lastPageId="13" lastPageNumber="110" pageId="12" pageNumber="109">
The trunk armour is narrow, relatively low, with a rather flattened dorsal wall, and the lateral wall more than three times as long as it is high. The length of the dorsal wall, estimated from the largest specimens of
<collectionCode id="8FE6A3E94C5BFFECFB09844CA51EF409" box="[1275,1341,1290,1315]" pageId="12" pageNumber="109">AMD</collectionCode>
and
<collectionCode id="8FE6A3E94C5BFFECFA86844CA593F409" box="[1396,1456,1290,1315]" pageId="12" pageNumber="109">PMD</collectionCode>
plates, probably, slightly exceeds
<quantity id="2E0F96C94C5BFFECFB75846DA4CAF46E" box="[1159,1257,1323,1348]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="109" unit="mm" value="200.0">200 mm</quantity>
. Thus it is larger than in
<taxonomicName id="2EF740AF4C5BFFECFCAC840AA422F44F" authorityName="Miles" authorityYear="1968" box="[862,1025,1356,1381]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="109" phylum="Chordata" rank="species" species="leptocheira">
<emphasis id="DB83E73E4C5BFFECFCAC840AA422F44F" box="[862,1025,1356,1381]" italics="true" pageId="12" pageNumber="109">B. leptocheira</emphasis>
</taxonomicName>
from
<collectingCountry id="91E07BBC4C5BFFECFBBD840AA488F44F" box="[1103,1195,1356,1381]" name="United Kingdom" pageId="12" pageNumber="109">Scotland</collectingCountry>
(
<bibRefCitation id="8D6646DD4C5BFFECFB36840AA570F44F" author="Miles, R. S." box="[1220,1363,1356,1381]" pageId="12" pageNumber="109" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles [1968]</bibRefCitation>
estimated the dorsal wall reached a length of some
<quantity id="2E0F96C94C5BFFECFAFE842BA54FF4AC" box="[1292,1388,1389,1414]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="109" unit="mm" value="140.0">140 mm</quantity>
), but smaller than in
<taxonomicName id="2EF740AF4C5BFFECFC4984C9A4EFF482" baseAuthorityName="Luksevics" baseAuthorityYear="2001" box="[955,1228,1423,1448]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="109" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="curonica">
<emphasis id="DB83E73E4C5BFFECFC4984C9A4EFF482" box="[955,1228,1423,1448]" italics="true" pageId="12" pageNumber="109">B. leptocheira curonica</emphasis>
</taxonomicName>
from
<collectingCountry id="91E07BBC4C5BFFECFAEB84C9A543F482" box="[1305,1376,1423,1448]" name="Latvia" pageId="12" pageNumber="109">Latvia</collectingCountry>
estimated by
<bibRefCitation id="8D6646DD4C5BFFECFCAC84F6A403F4E3" author="Luksevics, E." box="[862,1056,1456,1481]" pageId="12" pageNumber="109" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs (2001)</bibRefCitation>
as reaching
<quantity id="2E0F96C94C5BFFECFB4484F6A539F4E3" box="[1206,1306,1456,1481]" metricMagnitude="-1" metricUnit="m" metricValue="2.4" pageId="12" pageNumber="109" unit="mm" value="240.0">240 mm</quantity>
. The median dorsal ridge is weakly defined only in the posterior part of the
<collectionCode id="8FE6A3E94C5BFFECFA4E8497A5D8F4C0" box="[1468,1531,1489,1514]" pageId="12" pageNumber="109">PMD</collectionCode>
. The dorso-lateral and ventro-lateral ridges are well marked. The ventral wall of the trunk armour is not completely flat as it was demonstrated in the 3D model of
<taxonomicName id="2EF740AF4C5BFFECFAAC8773A46AF745" authority="(Bechard et al. 2014)" baseAuthorityName="Bechard" baseAuthorityYear="2014" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="109" phylum="Chordata" rank="species" species="canadensis">
<emphasis id="DB83E73E4C5BFFECFAAC8773A5DCF764" box="[1374,1535,1589,1614]" italics="true" pageId="12" pageNumber="109">B. canadensis</emphasis>
(
<bibRefCitation id="8D6646DD4C5BFFECFC978710A460F745" author="Bechard, I. &amp; Arsenault, F. &amp; Cloutier, R. &amp; Kerr, J." box="[869,1091,1622,1647]" pageId="12" pageNumber="109" pagination="1 - 19" refId="ref17888" refString="Bechard, I., Arsenault, F., Cloutier, R., and Kerr, J. 2014. The Devonian placoderm fish Bothriolepis canadensis revisited with three-dimensional digital imagery. Palaeontologia Electronica 17 (1, 2 A): 1 - 19." type="journal article" year="2014">Béchard et al. 2014</bibRefCitation>
)
</taxonomicName>
, but it is slightly convex both in transverse and longitudinal directions. The ventral wall is moderately wide with an estimated B/L index of
<quantity id="2E0F96C94C5BFFECFAA987DEA5B8F79B" box="[1371,1435,1688,1713]" metricMagnitude="-2" metricUnit="m" metricValue="1.524" pageId="12" pageNumber="109" unit="in" value="0.6">0.6 in</quantity>
IG KSC 155/10-3 (
<figureCitation id="71CC27A94C5BFFECFC3B87FCA40AF7F9" box="[969,1065,1722,1747]" captionStart="Fig" captionStartId="4.[151,184,1188,1208]" captionTargetBox="[157,1530,160,1164]" captionTargetId="figure-306@4.[173,1531,159,1164]" captionTargetPageId="4" captionText="Fig. 3. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, separate plates and parts of articulated head shields. A. Articulated Nu, left Pn and left La plates IG KSC 155/90 in dorsal view. B. Disarticulated plates of the head shield IG KSC 155/150 in dorsal view. C. Part of articulated head shield IG KSC 155/162 in dorsal view. D. Imprint of the ventral wall of trunk armour and some skeletal elements of the head armour IG KSC 155/10-3 in ventral view showing suborbital and infragnathal bones. Abbreviations: AVL, anterior ventro-lateral plate; cir, semicircular pit-line groove; csl, central sensory line groove; d.end, opening of canal for endolymphatic duct; ifc1, principal section of infraorbital sensory line; IG, inferognathal; La, lateral plate; Nu, nuchal plate; mp, middle pit-line groove; Pn, paranuchal plate; Pp, postpineal plate; Prm, premedian plate; SO, suborbital plate; socc, supraoccipital cross-commissural pit-line groove." figureDoi="http://doi.org/10.5281/zenodo.10980827" httpUri="https://zenodo.org/record/10980827/files/figure.png" pageId="12" pageNumber="109">Figs. 3D</figureCitation>
,
<figureCitation id="71CC27A94C5BFFECFBC187FCA46CF7F9" box="[1075,1103,1722,1747]" captionStart="Fig" captionStartId="15.[98,131,1132,1152]" captionTargetBox="[103,764,164,1109]" captionTargetId="figure-870@15.[102,764,160,1109]" captionTargetPageId="15" captionText="Fig. 14. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), reconstruction of the head, the trunk armour and the pectoral fins in ventral view, based on IG KSC 155/10-1, IG KSC 155/10-2, and IG KSC 155/10- 3. Abbreviations: AVL, anterior ventro-lateral plate; Cv14, ventral central plate 1 to 4; Ifg, inferognathal plate; La, lateral plate; Ml25, lateral marginal plates 2 to 5; Mm4, mesial marginal plate 4; MV, median ventral plate; Prl, prelateral plate; Prm, premedian plate; PVL, posterior ventro-lateral plate; SM, semilunar plate; SMg, submarginal plate; T, terminal plate." figureDoi="http://doi.org/10.5281/zenodo.10980847" httpUri="https://zenodo.org/record/10980847/files/figure.png" pageId="12" pageNumber="109">14</figureCitation>
), exceeding the estimated B/L index of about
<quantity id="2E0F96C94C5BFFECFC56879DA3D2F7DE" box="[932,1009,1755,1780]" metricMagnitude="-2" metricUnit="m" metricValue="1.3462" pageId="12" pageNumber="109" unit="in" value="0.53">0.53 in</quantity>
Latvian material of
<taxonomicName id="2EF740AF4C5BFFECFB29879DA383F63F" authority="(Luksevics 2001)" baseAuthorityName="Luksevics" baseAuthorityYear="2001" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="109" phylum="Chordata" rank="species" species="leptocheira">
<emphasis id="DB83E73E4C5BFFECFB29879DA558F7DE" box="[1243,1403,1755,1780]" italics="true" pageId="12" pageNumber="109">B. leptocheira</emphasis>
(
<bibRefCitation id="8D6646DD4C5BFFECFA79879DA3BBF63F" author="Luksevics, E." pageId="12" pageNumber="109" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
)
</taxonomicName>
.However, this difference could be explained by the use of separate
<collectionCode id="8FE6A3E94C5AFFEDFF1281E6A134F193" box="[224,279,160,185]" pageId="13" pageNumber="110">AVL</collectionCode>
and
<collectionCode id="8FE6A3E94C5AFFEDFEB981E6A1A2F193" box="[331,385,160,185]" country="Argentina" httpUri="http://grbio.org/cool/pbkd-hmeb" name="Paleontologia de Vertebrados Lillo" pageId="13" pageNumber="110">PVL</collectionCode>
plates of moderate size to estimate the B/L index of the ventral wall of
<taxonomicName id="2EF740AF4C5AFFEDFE018186A2D8F1F3" baseAuthorityName="Luksevics" baseAuthorityYear="2001" box="[499,763,192,217]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="110" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="curonica">
<emphasis id="DB83E73E4C5AFFEDFE018186A2D8F1F3" box="[499,763,192,217]" italics="true" pageId="13" pageNumber="110">B. leptocheira curonica</emphasis>
</taxonomicName>
. The ventral wall is slightly narrower than that restored for
<taxonomicName id="2EF740AF4C5AFFEDFD1881A6A086F030" authorityName="Gross" authorityYear="1942" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="110" phylum="Chordata" rank="species" species="prima">
<emphasis id="DB83E73E4C5AFFEDFD1881A6A086F030" italics="true" pageId="13" pageNumber="110">B. prima</emphasis>
</taxonomicName>
and
<taxonomicName id="2EF740AF4C5AFFEDFF168047A1B2F030" authorityName="Gross" authorityYear="1942" box="[228,401,257,282]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="110" phylum="Chordata" rank="species" species="obrutschewi">
<emphasis id="DB83E73E4C5AFFEDFF168047A1B2F030" box="[228,401,257,282]" italics="true" pageId="13" pageNumber="110">B. obrutschewi</emphasis>
</taxonomicName>
(
<bibRefCitation id="8D6646DD4C5AFFEDFE568047A2E1F030" author="Karatajute-Talimaa, V. N." box="[420,706,257,282]" pageId="13" pageNumber="110" pagination="191 - 279" refId="ref18937" refString="Karatajute-Talimaa, V. N. 1966. Bothriolepids of Sventoji Regional Stage of the Baltics [in Russian]. In: A. Grigelis (ed.), Paleontologia i stratigrafia Pribaltiki i Belorussji I (VI), 191 - 279. Mintis, Vilnius." type="journal article" year="1966">Karatajūte-Talimaa 1966</bibRefCitation>
: figs. 3, 10), but almost perfectly similar in the proportions to
<taxonomicName id="2EF740AF4C5AFFEDFD188067A0FCF070" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="110" phylum="Chordata" rank="species" species="canadensis">
<emphasis id="DB83E73E4C5AFFEDFD188067A0FCF070" italics="true" pageId="13" pageNumber="110">B. canadensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="8D6646DD4C5AFFEDFF1F8007A1E4F070" author="Bechard, I. &amp; Arsenault, F. &amp; Cloutier, R. &amp; Kerr, J." box="[237,455,321,346]" pageId="13" pageNumber="110" pagination="1 - 19" refId="ref17888" refString="Bechard, I., Arsenault, F., Cloutier, R., and Kerr, J. 2014. The Devonian placoderm fish Bothriolepis canadensis revisited with three-dimensional digital imagery. Palaeontologia Electronica 17 (1, 2 A): 1 - 19." type="journal article" year="2014">Béchard et al. 2014</bibRefCitation>
: fig. 3).
</paragraph>
<paragraph id="E9483B2C4C5AFFEDFF758027A2F9F4EE" blockId="13.[98,771,160,1960]" pageId="13" pageNumber="110">
The anterior median dorsal plate (
<collectionCode id="8FE6A3E94C5AFFEDFE0F8027A260F050" box="[509,579,353,378]" pageId="13" pageNumber="110">AMD</collectionCode>
;
<figureCitation id="71CC27A94C5AFFEDFDBF8027A2A8F050" box="[589,651,353,378]" captionStart="→" captionStartId="10.[116,144,1620,1650]" captionTargetId="figure-8@9.[99,1479,175,1936]" captionTargetPageId="9" captionText="→ Fig. 9. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior median dorsal (AG) and posterior median dorsal (HM) plates of the trunk armour. A. IG KSC 155/5 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/108 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/97 in dorsal view. D. IG KSC 155/113 in dorsal (D1) and visceral (D2) views. E. IG KSC 155/140 in dorsal (E1) and visceral (E2) views. F. Impression of the dorsal surface of IG KSC 155/42.G. IG KSC 155/44 in dorsal view. H. Fragment of IG KSC 155/7 in dorsal view. I. IG KSC 155/1 in dorsal (I1) and visceral (I2) views. J. IG KSC 155/71 in dorsal view. K. Slightly deformed IG KSC 155/70 in dorsal (K1) and visceral (K2) views. L. IG KSC 155/158 in dorsal view. M. IG KSC 155/157 in dorsal (M1) and visceral (M2) views. Abbreviations: ADL, anterior dorso-lateral plate; alr, postlevator thickening; AMD, anterior median dorsal plate; cf.ADL, cf.AMD, and cf.MxL, area overlapping ADL, AMD or MxL respectively; cr.tp, posterior transversal internal crest; dlg1 and dlg2, anterior and posterior oblique dorsal sensory line groove; dma, tergal angle; dmr, dorsal median ridge; f.retr, levator fossa; grm, ventral median groove; l, lateral corner; mvr, median ventral ridge; MxL, mixilateral plate; npn, postnuchal notch; oa.ADL, oa.MxL and oa.PMD, area overlapped by ADL, MxL or PMD respectively; pa, posterior corner; pma, posterior marginal area; PMD, posterior median dorsal plate; pr.p, posterior process of AMD; pr.pl, external postlevator process; prv2, posterior ventral process of dorsal wall of trunk armour; pt1 and pt2, anterior and posterior ventral pit; pua, posterior unornamented area of PMD; rf, “round fossula”; sna, supranuchal area; tb, ventral tuberosity." figureDoi="http://doi.org/10.5281/zenodo.11061576" httpUri="https://zenodo.org/record/11061576/files/figure.png" pageId="13" pageNumber="110">Fig. 9</figureCitation>
) is weakly arched, relatively narrow, with a B/L index of about
<quantity id="2E0F96C94C5AFFEDFD4480C4A321F0B1" box="[694,770,386,411]" metricMagnitude="-2" metricUnit="m" metricValue="1.9812" pageId="13" pageNumber="110" unit="in" value="0.78">0.78 in</quantity>
the Sosnogorsk material. The index is
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Latvian material (
<bibRefCitation id="8D6646DD4C5AFFEDFF658084A1A1F0F1" author="Luksevics, E." box="[151,386,450,475]" pageId="13" pageNumber="110" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001: 549</bibRefCitation>
), and about
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Scottish material (
<bibRefCitation id="8D6646DD4C5AFFEDFF6480A4A163F0D1" author="Miles, R. S." box="[150,320,482,507]" pageId="13" pageNumber="110" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles 1968: 78</bibRefCitation>
). The anterior margin is weakly convex, sometimes slightly concave in its middle (
<collectionCode id="8FE6A3E94C5AFFEDFDBB8345A24BF336" box="[585,616,515,540]" country="China" name="Institute of Geology" pageId="13" pageNumber="110">IG</collectionCode>
KSC 155/44;
<figureCitation id="71CC27A94C5AFFEDFF908365A09BF316" box="[98,184,547,572]" captionStart="→" captionStartId="10.[116,144,1620,1650]" captionTargetId="figure-8@9.[99,1479,175,1936]" captionTargetPageId="9" captionText="→ Fig. 9. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior median dorsal (AG) and posterior median dorsal (HM) plates of the trunk armour. A. IG KSC 155/5 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/108 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/97 in dorsal view. D. IG KSC 155/113 in dorsal (D1) and visceral (D2) views. E. IG KSC 155/140 in dorsal (E1) and visceral (E2) views. F. Impression of the dorsal surface of IG KSC 155/42.G. IG KSC 155/44 in dorsal view. H. Fragment of IG KSC 155/7 in dorsal view. I. IG KSC 155/1 in dorsal (I1) and visceral (I2) views. J. IG KSC 155/71 in dorsal view. K. Slightly deformed IG KSC 155/70 in dorsal (K1) and visceral (K2) views. L. IG KSC 155/158 in dorsal view. M. IG KSC 155/157 in dorsal (M1) and visceral (M2) views. Abbreviations: ADL, anterior dorso-lateral plate; alr, postlevator thickening; AMD, anterior median dorsal plate; cf.ADL, cf.AMD, and cf.MxL, area overlapping ADL, AMD or MxL respectively; cr.tp, posterior transversal internal crest; dlg1 and dlg2, anterior and posterior oblique dorsal sensory line groove; dma, tergal angle; dmr, dorsal median ridge; f.retr, levator fossa; grm, ventral median groove; l, lateral corner; mvr, median ventral ridge; MxL, mixilateral plate; npn, postnuchal notch; oa.ADL, oa.MxL and oa.PMD, area overlapped by ADL, MxL or PMD respectively; pa, posterior corner; pma, posterior marginal area; PMD, posterior median dorsal plate; pr.p, posterior process of AMD; pr.pl, external postlevator process; prv2, posterior ventral process of dorsal wall of trunk armour; pt1 and pt2, anterior and posterior ventral pit; pua, posterior unornamented area of PMD; rf, “round fossula”; sna, supranuchal area; tb, ventral tuberosity." figureDoi="http://doi.org/10.5281/zenodo.11061576" httpUri="https://zenodo.org/record/11061576/files/figure.png" pageId="13" pageNumber="110">Fig. 9G</figureCitation>
), relatively narrow, and about 1.3 times longer than the posterior margin. The antero-lateral and lateral corners, and the postlevator processes (pr.pl) are well defined, especially in larger specimens. The posterior division of the lateral margin is 1.41.6 times shorter than the anterior division. There is no median dorsal ridge. Overlap areas for
<collectionCode id="8FE6A3E94C5AFFEDFD3B8382A320F3F7" box="[713,771,708,733]" pageId="13" pageNumber="110">ADL</collectionCode>
and
<collectionCode id="8FE6A3E94C5AFFEDFF6183A3A0EFF3D4" box="[147,204,741,766]" pageId="13" pageNumber="110">MxL</collectionCode>
are sometimes normally developed as typical for
<taxonomicName id="2EF740AF4C5AFFEDFF908243A0CFF234" authorityName="Eichwald" authorityYear="1840" box="[98,236,773,798]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="110" phylum="Chordata" rank="genus">
<emphasis id="DB83E73E4C5AFFEDFF908243A0CFF234" box="[98,236,773,798]" italics="true" pageId="13" pageNumber="110">Bothriolepis</emphasis>
</taxonomicName>
(
<collectionCode id="8FE6A3E94C5AFFEDFF0E8243A138F234" box="[252,283,773,798]" country="China" name="Institute of Geology" pageId="13" pageNumber="110">IG</collectionCode>
KSC 155/44;
<figureCitation id="71CC27A94C5AFFEDFE498243A231F234" box="[443,530,773,798]" captionStart="→" captionStartId="10.[116,144,1620,1650]" captionTargetId="figure-8@9.[99,1479,175,1936]" captionTargetPageId="9" captionText="→ Fig. 9. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior median dorsal (AG) and posterior median dorsal (HM) plates of the trunk armour. A. IG KSC 155/5 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/108 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/97 in dorsal view. D. IG KSC 155/113 in dorsal (D1) and visceral (D2) views. E. IG KSC 155/140 in dorsal (E1) and visceral (E2) views. F. Impression of the dorsal surface of IG KSC 155/42.G. IG KSC 155/44 in dorsal view. H. Fragment of IG KSC 155/7 in dorsal view. I. IG KSC 155/1 in dorsal (I1) and visceral (I2) views. J. IG KSC 155/71 in dorsal view. K. Slightly deformed IG KSC 155/70 in dorsal (K1) and visceral (K2) views. L. IG KSC 155/158 in dorsal view. M. IG KSC 155/157 in dorsal (M1) and visceral (M2) views. Abbreviations: ADL, anterior dorso-lateral plate; alr, postlevator thickening; AMD, anterior median dorsal plate; cf.ADL, cf.AMD, and cf.MxL, area overlapping ADL, AMD or MxL respectively; cr.tp, posterior transversal internal crest; dlg1 and dlg2, anterior and posterior oblique dorsal sensory line groove; dma, tergal angle; dmr, dorsal median ridge; f.retr, levator fossa; grm, ventral median groove; l, lateral corner; mvr, median ventral ridge; MxL, mixilateral plate; npn, postnuchal notch; oa.ADL, oa.MxL and oa.PMD, area overlapped by ADL, MxL or PMD respectively; pa, posterior corner; pma, posterior marginal area; PMD, posterior median dorsal plate; pr.p, posterior process of AMD; pr.pl, external postlevator process; prv2, posterior ventral process of dorsal wall of trunk armour; pt1 and pt2, anterior and posterior ventral pit; pua, posterior unornamented area of PMD; rf, “round fossula”; sna, supranuchal area; tb, ventral tuberosity." figureDoi="http://doi.org/10.5281/zenodo.11061576" httpUri="https://zenodo.org/record/11061576/files/figure.png" pageId="13" pageNumber="110">Fig. 9G</figureCitation>
), but usually demonstrate the so called
<taxonomicName id="2EF740AF4C5AFFEDFEB18263A1EFF214" authorityName="Stensiö" authorityYear="1931" box="[323,460,805,830]" family="Asterolepididae" genus="Remigolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="110" phylum="Chordata" rank="genus">
<emphasis id="DB83E73E4C5AFFEDFEB18263A1EFF214" box="[323,460,805,830]" italics="true" pageId="13" pageNumber="110">Remigolepis</emphasis>
</taxonomicName>
-
<typeStatus id="364C858E4C5AFFEDFE268263A224F214" box="[468,519,805,830]" pageId="13" pageNumber="110">type</typeStatus>
overlapping: IG KSC 155/140 and IG KSC 155/42 (
<figureCitation id="71CC27A94C5AFFEDFE418203A209F274" box="[435,554,837,862]" captionStart="→" captionStartId="10.[116,144,1620,1650]" captionTargetId="figure-8@9.[99,1479,175,1936]" captionTargetPageId="9" captionText="→ Fig. 9. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior median dorsal (AG) and posterior median dorsal (HM) plates of the trunk armour. A. IG KSC 155/5 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/108 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/97 in dorsal view. D. IG KSC 155/113 in dorsal (D1) and visceral (D2) views. E. IG KSC 155/140 in dorsal (E1) and visceral (E2) views. F. Impression of the dorsal surface of IG KSC 155/42.G. IG KSC 155/44 in dorsal view. H. Fragment of IG KSC 155/7 in dorsal view. I. IG KSC 155/1 in dorsal (I1) and visceral (I2) views. J. IG KSC 155/71 in dorsal view. K. Slightly deformed IG KSC 155/70 in dorsal (K1) and visceral (K2) views. L. IG KSC 155/158 in dorsal view. M. IG KSC 155/157 in dorsal (M1) and visceral (M2) views. Abbreviations: ADL, anterior dorso-lateral plate; alr, postlevator thickening; AMD, anterior median dorsal plate; cf.ADL, cf.AMD, and cf.MxL, area overlapping ADL, AMD or MxL respectively; cr.tp, posterior transversal internal crest; dlg1 and dlg2, anterior and posterior oblique dorsal sensory line groove; dma, tergal angle; dmr, dorsal median ridge; f.retr, levator fossa; grm, ventral median groove; l, lateral corner; mvr, median ventral ridge; MxL, mixilateral plate; npn, postnuchal notch; oa.ADL, oa.MxL and oa.PMD, area overlapped by ADL, MxL or PMD respectively; pa, posterior corner; pma, posterior marginal area; PMD, posterior median dorsal plate; pr.p, posterior process of AMD; pr.pl, external postlevator process; prv2, posterior ventral process of dorsal wall of trunk armour; pt1 and pt2, anterior and posterior ventral pit; pua, posterior unornamented area of PMD; rf, “round fossula”; sna, supranuchal area; tb, ventral tuberosity." figureDoi="http://doi.org/10.5281/zenodo.11061576" httpUri="https://zenodo.org/record/11061576/files/figure.png" pageId="13" pageNumber="110">Fig. 9E, F</figureCitation>
) show the anterior ¼–⅓ of the posterior lateral margin overlapping
<collectionCode id="8FE6A3E94C5AFFEDFD698220A2F7F255" box="[667,724,870,895]" pageId="13" pageNumber="110">MxL</collectionCode>
(cf.
<collectionCode id="8FE6A3E94C5AFFEDFF9082C0A0B8F2B5" box="[98,155,902,927]" pageId="13" pageNumber="110">MxL</collectionCode>
) and remaining ¾ or ⅔ of that margin are overlapped by
<collectionCode id="8FE6A3E94C5AFFEDFF7782E0A0E3F295" box="[133,192,934,959]" pageId="13" pageNumber="110">MxL</collectionCode>
. The same feature is also seen in IG KSC 155/108, where the
<collectionCode id="8FE6A3E94C5AFFEDFF258281A13AF2CA" box="[215,281,967,992]" pageId="13" pageNumber="110">AMD</collectionCode>
overlaps
<collectionCode id="8FE6A3E94C5AFFEDFE778281A19DF2CA" box="[389,446,967,992]" pageId="13" pageNumber="110">MxL</collectionCode>
to 1/6 of the common suture
<figureCitation id="71CC27A94C5AFFEDFF9882A1A0E7F52A" box="[106,196,999,1024]" captionStart="→" captionStartId="10.[116,144,1620,1650]" captionTargetId="figure-8@9.[99,1479,175,1936]" captionTargetPageId="9" captionText="→ Fig. 9. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior median dorsal (AG) and posterior median dorsal (HM) plates of the trunk armour. A. IG KSC 155/5 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/108 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/97 in dorsal view. D. IG KSC 155/113 in dorsal (D1) and visceral (D2) views. E. IG KSC 155/140 in dorsal (E1) and visceral (E2) views. F. Impression of the dorsal surface of IG KSC 155/42.G. IG KSC 155/44 in dorsal view. H. Fragment of IG KSC 155/7 in dorsal view. I. IG KSC 155/1 in dorsal (I1) and visceral (I2) views. J. IG KSC 155/71 in dorsal view. K. Slightly deformed IG KSC 155/70 in dorsal (K1) and visceral (K2) views. L. IG KSC 155/158 in dorsal view. M. IG KSC 155/157 in dorsal (M1) and visceral (M2) views. Abbreviations: ADL, anterior dorso-lateral plate; alr, postlevator thickening; AMD, anterior median dorsal plate; cf.ADL, cf.AMD, and cf.MxL, area overlapping ADL, AMD or MxL respectively; cr.tp, posterior transversal internal crest; dlg1 and dlg2, anterior and posterior oblique dorsal sensory line groove; dma, tergal angle; dmr, dorsal median ridge; f.retr, levator fossa; grm, ventral median groove; l, lateral corner; mvr, median ventral ridge; MxL, mixilateral plate; npn, postnuchal notch; oa.ADL, oa.MxL and oa.PMD, area overlapped by ADL, MxL or PMD respectively; pa, posterior corner; pma, posterior marginal area; PMD, posterior median dorsal plate; pr.p, posterior process of AMD; pr.pl, external postlevator process; prv2, posterior ventral process of dorsal wall of trunk armour; pt1 and pt2, anterior and posterior ventral pit; pua, posterior unornamented area of PMD; rf, “round fossula”; sna, supranuchal area; tb, ventral tuberosity." figureDoi="http://doi.org/10.5281/zenodo.11061576" httpUri="https://zenodo.org/record/11061576/files/figure.png" pageId="13" pageNumber="110">Fig. 9B</figureCitation>
), and IG KSC 155/5, where the cf.
<collectionCode id="8FE6A3E94C5AFFEDFD9A82A1A281F52A" box="[616,674,999,1024]" pageId="13" pageNumber="110">MxL</collectionCode>
is even smaller (
<figureCitation id="71CC27A94C5AFFEDFF368541A138F50A" box="[196,283,1031,1056]" captionStart="→" captionStartId="10.[116,144,1620,1650]" captionTargetId="figure-8@9.[99,1479,175,1936]" captionTargetPageId="9" captionText="→ Fig. 9. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior median dorsal (AG) and posterior median dorsal (HM) plates of the trunk armour. A. IG KSC 155/5 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/108 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/97 in dorsal view. D. IG KSC 155/113 in dorsal (D1) and visceral (D2) views. E. IG KSC 155/140 in dorsal (E1) and visceral (E2) views. F. Impression of the dorsal surface of IG KSC 155/42.G. IG KSC 155/44 in dorsal view. H. Fragment of IG KSC 155/7 in dorsal view. I. IG KSC 155/1 in dorsal (I1) and visceral (I2) views. J. IG KSC 155/71 in dorsal view. K. Slightly deformed IG KSC 155/70 in dorsal (K1) and visceral (K2) views. L. IG KSC 155/158 in dorsal view. M. IG KSC 155/157 in dorsal (M1) and visceral (M2) views. Abbreviations: ADL, anterior dorso-lateral plate; alr, postlevator thickening; AMD, anterior median dorsal plate; cf.ADL, cf.AMD, and cf.MxL, area overlapping ADL, AMD or MxL respectively; cr.tp, posterior transversal internal crest; dlg1 and dlg2, anterior and posterior oblique dorsal sensory line groove; dma, tergal angle; dmr, dorsal median ridge; f.retr, levator fossa; grm, ventral median groove; l, lateral corner; mvr, median ventral ridge; MxL, mixilateral plate; npn, postnuchal notch; oa.ADL, oa.MxL and oa.PMD, area overlapped by ADL, MxL or PMD respectively; pa, posterior corner; pma, posterior marginal area; PMD, posterior median dorsal plate; pr.p, posterior process of AMD; pr.pl, external postlevator process; prv2, posterior ventral process of dorsal wall of trunk armour; pt1 and pt2, anterior and posterior ventral pit; pua, posterior unornamented area of PMD; rf, “round fossula”; sna, supranuchal area; tb, ventral tuberosity." figureDoi="http://doi.org/10.5281/zenodo.11061576" httpUri="https://zenodo.org/record/11061576/files/figure.png" pageId="13" pageNumber="110">Fig. 9A</figureCitation>
). Unusual are sutural connections of
<collectionCode id="8FE6A3E94C5AFFEDFD328541A321F50A" box="[704,770,1031,1056]" pageId="13" pageNumber="110">AMD</collectionCode>
and
<collectionCode id="8FE6A3E94C5AFFEDFF668561A0EDF56A" box="[148,206,1063,1088]" pageId="13" pageNumber="110">ADL</collectionCode>
in IG KSC 155/44, where the posterior ⅓ of the anterior lateral margin of
<collectionCode id="8FE6A3E94C5AFFEDFE7B850EA1EAF54B" box="[393,457,1096,1121]" pageId="13" pageNumber="110">AMD</collectionCode>
is overlapped by
<collectionCode id="8FE6A3E94C5AFFEDFD65850EA2ECF54B" box="[663,719,1096,1121]" pageId="13" pageNumber="110">ADL</collectionCode>
(oa.
<collectionCode id="8FE6A3E94C5AFFEDFF90852EA0BDF5AB" box="[98,158,1128,1153]" pageId="13" pageNumber="110">ADL</collectionCode>
;
<figureCitation id="71CC27A94C5AFFEDFF58852EA122F5AB" box="[170,257,1128,1153]" captionStart="→" captionStartId="10.[116,144,1620,1650]" captionTargetId="figure-8@9.[99,1479,175,1936]" captionTargetPageId="9" captionText="→ Fig. 9. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior median dorsal (AG) and posterior median dorsal (HM) plates of the trunk armour. A. IG KSC 155/5 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/108 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/97 in dorsal view. D. IG KSC 155/113 in dorsal (D1) and visceral (D2) views. E. IG KSC 155/140 in dorsal (E1) and visceral (E2) views. F. Impression of the dorsal surface of IG KSC 155/42.G. IG KSC 155/44 in dorsal view. H. Fragment of IG KSC 155/7 in dorsal view. I. IG KSC 155/1 in dorsal (I1) and visceral (I2) views. J. IG KSC 155/71 in dorsal view. K. Slightly deformed IG KSC 155/70 in dorsal (K1) and visceral (K2) views. L. IG KSC 155/158 in dorsal view. M. IG KSC 155/157 in dorsal (M1) and visceral (M2) views. Abbreviations: ADL, anterior dorso-lateral plate; alr, postlevator thickening; AMD, anterior median dorsal plate; cf.ADL, cf.AMD, and cf.MxL, area overlapping ADL, AMD or MxL respectively; cr.tp, posterior transversal internal crest; dlg1 and dlg2, anterior and posterior oblique dorsal sensory line groove; dma, tergal angle; dmr, dorsal median ridge; f.retr, levator fossa; grm, ventral median groove; l, lateral corner; mvr, median ventral ridge; MxL, mixilateral plate; npn, postnuchal notch; oa.ADL, oa.MxL and oa.PMD, area overlapped by ADL, MxL or PMD respectively; pa, posterior corner; pma, posterior marginal area; PMD, posterior median dorsal plate; pr.p, posterior process of AMD; pr.pl, external postlevator process; prv2, posterior ventral process of dorsal wall of trunk armour; pt1 and pt2, anterior and posterior ventral pit; pua, posterior unornamented area of PMD; rf, “round fossula”; sna, supranuchal area; tb, ventral tuberosity." figureDoi="http://doi.org/10.5281/zenodo.11061576" httpUri="https://zenodo.org/record/11061576/files/figure.png" pageId="13" pageNumber="110">Fig. 9G</figureCitation>
). Internally, the levator fossa (f.retr) is longer than it is wide; it is delineated by the low and narrow postlevator thickenings (alr). The low and short medial crest mc;
<figureCitation id="71CC27A94C5AFFEDFF69858FA12CF5C8" box="[155,271,1225,1250]" captionStart="→" captionStartId="10.[116,144,1620,1650]" captionTargetId="figure-8@9.[99,1479,175,1936]" captionTargetPageId="9" captionText="→ Fig. 9. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior median dorsal (AG) and posterior median dorsal (HM) plates of the trunk armour. A. IG KSC 155/5 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/108 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/97 in dorsal view. D. IG KSC 155/113 in dorsal (D1) and visceral (D2) views. E. IG KSC 155/140 in dorsal (E1) and visceral (E2) views. F. Impression of the dorsal surface of IG KSC 155/42.G. IG KSC 155/44 in dorsal view. H. Fragment of IG KSC 155/7 in dorsal view. I. IG KSC 155/1 in dorsal (I1) and visceral (I2) views. J. IG KSC 155/71 in dorsal view. K. Slightly deformed IG KSC 155/70 in dorsal (K1) and visceral (K2) views. L. IG KSC 155/158 in dorsal view. M. IG KSC 155/157 in dorsal (M1) and visceral (M2) views. Abbreviations: ADL, anterior dorso-lateral plate; alr, postlevator thickening; AMD, anterior median dorsal plate; cf.ADL, cf.AMD, and cf.MxL, area overlapping ADL, AMD or MxL respectively; cr.tp, posterior transversal internal crest; dlg1 and dlg2, anterior and posterior oblique dorsal sensory line groove; dma, tergal angle; dmr, dorsal median ridge; f.retr, levator fossa; grm, ventral median groove; l, lateral corner; mvr, median ventral ridge; MxL, mixilateral plate; npn, postnuchal notch; oa.ADL, oa.MxL and oa.PMD, area overlapped by ADL, MxL or PMD respectively; pa, posterior corner; pma, posterior marginal area; PMD, posterior median dorsal plate; pr.p, posterior process of AMD; pr.pl, external postlevator process; prv2, posterior ventral process of dorsal wall of trunk armour; pt1 and pt2, anterior and posterior ventral pit; pua, posterior unornamented area of PMD; rf, “round fossula”; sna, supranuchal area; tb, ventral tuberosity." figureDoi="http://doi.org/10.5281/zenodo.11061576" httpUri="https://zenodo.org/record/11061576/files/figure.png" pageId="13" pageNumber="110">Fig. 9A, B</figureCitation>
) is more than twice shorter than the levator fossa. The supranuchal area (sna) is well defined and slightly broader medially from the antero-lateral corner. The centre of the anterior ventral pit (pt
<subScript id="757339694C5AFFEDFE228473A1FAF46C" attach="left" box="[464,473,1333,1350]" fontSize="7" pageId="13" pageNumber="110">1</subScript>
;
<figureCitation id="71CC27A94C5AFFEDFE18846CA2A6F469" box="[490,645,1322,1347]" captionStart="→" captionStartId="10.[116,144,1620,1650]" captionTargetId="figure-8@9.[99,1479,175,1936]" captionTargetPageId="9" captionText="→ Fig. 9. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior median dorsal (AG) and posterior median dorsal (HM) plates of the trunk armour. A. IG KSC 155/5 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/108 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/97 in dorsal view. D. IG KSC 155/113 in dorsal (D1) and visceral (D2) views. E. IG KSC 155/140 in dorsal (E1) and visceral (E2) views. F. Impression of the dorsal surface of IG KSC 155/42.G. IG KSC 155/44 in dorsal view. H. Fragment of IG KSC 155/7 in dorsal view. I. IG KSC 155/1 in dorsal (I1) and visceral (I2) views. J. IG KSC 155/71 in dorsal view. K. Slightly deformed IG KSC 155/70 in dorsal (K1) and visceral (K2) views. L. IG KSC 155/158 in dorsal view. M. IG KSC 155/157 in dorsal (M1) and visceral (M2) views. Abbreviations: ADL, anterior dorso-lateral plate; alr, postlevator thickening; AMD, anterior median dorsal plate; cf.ADL, cf.AMD, and cf.MxL, area overlapping ADL, AMD or MxL respectively; cr.tp, posterior transversal internal crest; dlg1 and dlg2, anterior and posterior oblique dorsal sensory line groove; dma, tergal angle; dmr, dorsal median ridge; f.retr, levator fossa; grm, ventral median groove; l, lateral corner; mvr, median ventral ridge; MxL, mixilateral plate; npn, postnuchal notch; oa.ADL, oa.MxL and oa.PMD, area overlapped by ADL, MxL or PMD respectively; pa, posterior corner; pma, posterior marginal area; PMD, posterior median dorsal plate; pr.p, posterior process of AMD; pr.pl, external postlevator process; prv2, posterior ventral process of dorsal wall of trunk armour; pt1 and pt2, anterior and posterior ventral pit; pua, posterior unornamented area of PMD; rf, “round fossula”; sna, supranuchal area; tb, ventral tuberosity." figureDoi="http://doi.org/10.5281/zenodo.11061576" httpUri="https://zenodo.org/record/11061576/files/figure.png" pageId="13" pageNumber="110">Fig. 9A, B, D</figureCitation>
) is located slightly anteriorly to the tergal angle (dma). The median ventral ridge (mvr) is relatively low; in IG KSC 155/113 (
<figureCitation id="71CC27A94C5AFFEDFD55842CA2D9F4A9" box="[679,762,1386,1411]" captionStart="→" captionStartId="10.[116,144,1620,1650]" captionTargetId="figure-8@9.[99,1479,175,1936]" captionTargetPageId="9" captionText="→ Fig. 9. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior median dorsal (AG) and posterior median dorsal (HM) plates of the trunk armour. A. IG KSC 155/5 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/108 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/97 in dorsal view. D. IG KSC 155/113 in dorsal (D1) and visceral (D2) views. E. IG KSC 155/140 in dorsal (E1) and visceral (E2) views. F. Impression of the dorsal surface of IG KSC 155/42.G. IG KSC 155/44 in dorsal view. H. Fragment of IG KSC 155/7 in dorsal view. I. IG KSC 155/1 in dorsal (I1) and visceral (I2) views. J. IG KSC 155/71 in dorsal view. K. Slightly deformed IG KSC 155/70 in dorsal (K1) and visceral (K2) views. L. IG KSC 155/158 in dorsal view. M. IG KSC 155/157 in dorsal (M1) and visceral (M2) views. Abbreviations: ADL, anterior dorso-lateral plate; alr, postlevator thickening; AMD, anterior median dorsal plate; cf.ADL, cf.AMD, and cf.MxL, area overlapping ADL, AMD or MxL respectively; cr.tp, posterior transversal internal crest; dlg1 and dlg2, anterior and posterior oblique dorsal sensory line groove; dma, tergal angle; dmr, dorsal median ridge; f.retr, levator fossa; grm, ventral median groove; l, lateral corner; mvr, median ventral ridge; MxL, mixilateral plate; npn, postnuchal notch; oa.ADL, oa.MxL and oa.PMD, area overlapped by ADL, MxL or PMD respectively; pa, posterior corner; pma, posterior marginal area; PMD, posterior median dorsal plate; pr.p, posterior process of AMD; pr.pl, external postlevator process; prv2, posterior ventral process of dorsal wall of trunk armour; pt1 and pt2, anterior and posterior ventral pit; pua, posterior unornamented area of PMD; rf, “round fossula”; sna, supranuchal area; tb, ventral tuberosity." figureDoi="http://doi.org/10.5281/zenodo.11061576" httpUri="https://zenodo.org/record/11061576/files/figure.png" pageId="13" pageNumber="110">Fig. 9D</figureCitation>
) it is extremely short and divides into two crests which continue as a moderately deep median ventral groove (grm).
</paragraph>
<paragraph id="E9483B2C4C5AFFEDFF75848DA55EF54B" blockId="13.[98,771,160,1960]" lastBlockId="13.[808,1482,160,1960]" pageId="13" pageNumber="110">
The posterior median dorsal plate (
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;
<figureCitation id="71CC27A94C5AFFEDFD94848DA2FCF4CE" box="[614,735,1483,1508]" captionStart="→" captionStartId="10.[116,144,1620,1650]" captionTargetId="figure-8@9.[99,1479,175,1936]" captionTargetPageId="9" captionText="→ Fig. 9. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior median dorsal (AG) and posterior median dorsal (HM) plates of the trunk armour. A. IG KSC 155/5 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/108 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/97 in dorsal view. D. IG KSC 155/113 in dorsal (D1) and visceral (D2) views. E. IG KSC 155/140 in dorsal (E1) and visceral (E2) views. F. Impression of the dorsal surface of IG KSC 155/42.G. IG KSC 155/44 in dorsal view. H. Fragment of IG KSC 155/7 in dorsal view. I. IG KSC 155/1 in dorsal (I1) and visceral (I2) views. J. IG KSC 155/71 in dorsal view. K. Slightly deformed IG KSC 155/70 in dorsal (K1) and visceral (K2) views. L. IG KSC 155/158 in dorsal view. M. IG KSC 155/157 in dorsal (M1) and visceral (M2) views. Abbreviations: ADL, anterior dorso-lateral plate; alr, postlevator thickening; AMD, anterior median dorsal plate; cf.ADL, cf.AMD, and cf.MxL, area overlapping ADL, AMD or MxL respectively; cr.tp, posterior transversal internal crest; dlg1 and dlg2, anterior and posterior oblique dorsal sensory line groove; dma, tergal angle; dmr, dorsal median ridge; f.retr, levator fossa; grm, ventral median groove; l, lateral corner; mvr, median ventral ridge; MxL, mixilateral plate; npn, postnuchal notch; oa.ADL, oa.MxL and oa.PMD, area overlapped by ADL, MxL or PMD respectively; pa, posterior corner; pma, posterior marginal area; PMD, posterior median dorsal plate; pr.p, posterior process of AMD; pr.pl, external postlevator process; prv2, posterior ventral process of dorsal wall of trunk armour; pt1 and pt2, anterior and posterior ventral pit; pua, posterior unornamented area of PMD; rf, “round fossula”; sna, supranuchal area; tb, ventral tuberosity." figureDoi="http://doi.org/10.5281/zenodo.11061576" httpUri="https://zenodo.org/record/11061576/files/figure.png" pageId="13" pageNumber="110">Fig. 9HN</figureCitation>
) is broad in comparison with the
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. Its B/L index varies from 0.84 to 1.00, 0.92 on the average in the Sosnogorsk material, whereas it reaches about
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the material from
<collectingCountry id="91E07BBC4C5AFFEDFF90870AA0E7F74F" box="[98,196,1612,1637]" name="United Kingdom" pageId="13" pageNumber="110">Scotland</collectingCountry>
(
<bibRefCitation id="8D6646DD4C5AFFEDFF21870AA177F74F" author="Miles, R. S." box="[211,340,1612,1637]" pageId="13" pageNumber="110" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles 1968</bibRefCitation>
) and varies from 0.88 to 1.01, 0.95 on the average, in the material from
<collectingCountry id="91E07BBC4C5AFFEDFE14872AA20EF7AF" box="[486,557,1644,1669]" name="Latvia" pageId="13" pageNumber="110">Latvia</collectingCountry>
(
<bibRefCitation id="8D6646DD4C5AFFEDFDB2872AA2D4F7AF" author="Luksevics, E." box="[576,759,1644,1669]" pageId="13" pageNumber="110" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
). The anterior margin is rather narrow, strongly convex and pointed in small individuals, and convex but rounded in large individuals. The posterior margin is well convex, 1.72.7 times longer than the anterior margin in the material from Sosnogorsk, but only
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the Scottish and Latvian material. The posterior corner of the
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is not pronounced as in the Scottish (
<bibRefCitation id="8D6646DD4C5AFFEDFE328608A260F64D" author="Miles, R. S." box="[448,579,1870,1895]" pageId="13" pageNumber="110" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles 1968</bibRefCitation>
: text-fig. 38) and Latvian material (
<bibRefCitation id="8D6646DD4C5AFFEDFEC78629A1CCF6A2" author="Luksevics, E." box="[309,495,1903,1928]" pageId="13" pageNumber="110" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
: fig. 35C). The median dorsal ridge (dmr) is present only in the posterior third of the plate in medium-sized individuals, and only in the posterior quarter in the largest
<collectionCode id="8FE6A3E94C5AFFEDFBEE8186A47AF1F3" box="[1052,1113,192,217]" pageId="13" pageNumber="110">PMD</collectionCode>
IG KSC 155/142, or even absent in some large
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; however, in the smallest individual IG KSC 155/148 reaching only
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in length, it is seen along the whole plate. IG KSC 155/7 differs from all other specimens in possessing two distinct abnormalities which are most probably related to one another: (i) the posterior margin bears
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wide unornamented area (pua) that superficially resembles the obtected nuchal area of the head shield; (ii) a branch of the sensory line canal (dlg
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) is clearly seen along the same posterior margin of the left side of the plate (
<figureCitation id="71CC27A94C5AFFEDFC808345A3EEF336" box="[882,973,515,540]" captionStart="→" captionStartId="10.[116,144,1620,1650]" captionTargetId="figure-8@9.[99,1479,175,1936]" captionTargetPageId="9" captionText="→ Fig. 9. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior median dorsal (AG) and posterior median dorsal (HM) plates of the trunk armour. A. IG KSC 155/5 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/108 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/97 in dorsal view. D. IG KSC 155/113 in dorsal (D1) and visceral (D2) views. E. IG KSC 155/140 in dorsal (E1) and visceral (E2) views. F. Impression of the dorsal surface of IG KSC 155/42.G. IG KSC 155/44 in dorsal view. H. Fragment of IG KSC 155/7 in dorsal view. I. IG KSC 155/1 in dorsal (I1) and visceral (I2) views. J. IG KSC 155/71 in dorsal view. K. Slightly deformed IG KSC 155/70 in dorsal (K1) and visceral (K2) views. L. IG KSC 155/158 in dorsal view. M. IG KSC 155/157 in dorsal (M1) and visceral (M2) views. Abbreviations: ADL, anterior dorso-lateral plate; alr, postlevator thickening; AMD, anterior median dorsal plate; cf.ADL, cf.AMD, and cf.MxL, area overlapping ADL, AMD or MxL respectively; cr.tp, posterior transversal internal crest; dlg1 and dlg2, anterior and posterior oblique dorsal sensory line groove; dma, tergal angle; dmr, dorsal median ridge; f.retr, levator fossa; grm, ventral median groove; l, lateral corner; mvr, median ventral ridge; MxL, mixilateral plate; npn, postnuchal notch; oa.ADL, oa.MxL and oa.PMD, area overlapped by ADL, MxL or PMD respectively; pa, posterior corner; pma, posterior marginal area; PMD, posterior median dorsal plate; pr.p, posterior process of AMD; pr.pl, external postlevator process; prv2, posterior ventral process of dorsal wall of trunk armour; pt1 and pt2, anterior and posterior ventral pit; pua, posterior unornamented area of PMD; rf, “round fossula”; sna, supranuchal area; tb, ventral tuberosity." figureDoi="http://doi.org/10.5281/zenodo.11061576" httpUri="https://zenodo.org/record/11061576/files/figure.png" pageId="13" pageNumber="110">Fig. 9H</figureCitation>
) in the position where this canal has been reported in
<taxonomicName id="2EF740AF4C5AFFEDFC458365A479F316" box="[951,1114,547,572]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="110" phylum="Chordata" rank="species" species="canadensis">
<emphasis id="DB83E73E4C5AFFEDFC458365A479F316" box="[951,1114,547,572]" italics="true" pageId="13" pageNumber="110">B. canadensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="8D6646DD4C5AFFEDFB9C8365A57EF316" author="Graham-Smith, W." box="[1134,1373,547,572]" pageId="13" pageNumber="110" pagination="1 - 39" refId="ref18605" refString="Graham-Smith, W. 1978. On some variations in the latero-sensory lines of the placoderm fish Bothriolepis. Philosophical Transactions of Royal Society London B: Biological Science 282: 1 - 39." type="journal article" year="1978">Graham-Smith 1978</bibRefCitation>
: fig. 2e). The canal crosses the lateral margin of the
<collectionCode id="8FE6A3E94C5AFFEDFAFC8305A568F376" box="[1294,1355,579,604]" pageId="13" pageNumber="110">PMD</collectionCode>
close to its posterior margin, runs parallel to the posterior margin and a few milimeters from the dorsal median ridge turns posteriorly and becomes deeper than in the most part of its course, “diving” into the bone. The
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narrow belt between the canal and pua is ornamented by separate round and high tubercles, in some cases two or three of them fused into very short ridges, while the rest of the surface bears mainly vermiculating ridges and only several separate tubercles close to the canal. The ventral surface of the
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shows a relatively low posterior transverse thoracic crest (cr.tp), in some specimens (e.g., IG KSC 155/1;
<figureCitation id="71CC27A94C5AFFEDFB0B82E0A564F295" box="[1273,1351,934,959]" captionStart="→" captionStartId="10.[116,144,1620,1650]" captionTargetId="figure-8@9.[99,1479,175,1936]" captionTargetPageId="9" captionText="→ Fig. 9. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior median dorsal (AG) and posterior median dorsal (HM) plates of the trunk armour. A. IG KSC 155/5 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/108 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/97 in dorsal view. D. IG KSC 155/113 in dorsal (D1) and visceral (D2) views. E. IG KSC 155/140 in dorsal (E1) and visceral (E2) views. F. Impression of the dorsal surface of IG KSC 155/42.G. IG KSC 155/44 in dorsal view. H. Fragment of IG KSC 155/7 in dorsal view. I. IG KSC 155/1 in dorsal (I1) and visceral (I2) views. J. IG KSC 155/71 in dorsal view. K. Slightly deformed IG KSC 155/70 in dorsal (K1) and visceral (K2) views. L. IG KSC 155/158 in dorsal view. M. IG KSC 155/157 in dorsal (M1) and visceral (M2) views. Abbreviations: ADL, anterior dorso-lateral plate; alr, postlevator thickening; AMD, anterior median dorsal plate; cf.ADL, cf.AMD, and cf.MxL, area overlapping ADL, AMD or MxL respectively; cr.tp, posterior transversal internal crest; dlg1 and dlg2, anterior and posterior oblique dorsal sensory line groove; dma, tergal angle; dmr, dorsal median ridge; f.retr, levator fossa; grm, ventral median groove; l, lateral corner; mvr, median ventral ridge; MxL, mixilateral plate; npn, postnuchal notch; oa.ADL, oa.MxL and oa.PMD, area overlapped by ADL, MxL or PMD respectively; pa, posterior corner; pma, posterior marginal area; PMD, posterior median dorsal plate; pr.p, posterior process of AMD; pr.pl, external postlevator process; prv2, posterior ventral process of dorsal wall of trunk armour; pt1 and pt2, anterior and posterior ventral pit; pua, posterior unornamented area of PMD; rf, “round fossula”; sna, supranuchal area; tb, ventral tuberosity." figureDoi="http://doi.org/10.5281/zenodo.11061576" httpUri="https://zenodo.org/record/11061576/files/figure.png" pageId="13" pageNumber="110">Fig. 9J</figureCitation>
) forming a distinct shelf on the short area posterior to the crest. There is a wide median ventral groove (grm) and a peculiar posterior ventral pit (pt
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;
<figureCitation id="71CC27A94C5AFFEDFC178541A47AF50A" box="[997,1113,1031,1056]" captionStart="→" captionStartId="10.[116,144,1620,1650]" captionTargetId="figure-8@9.[99,1479,175,1936]" captionTargetPageId="9" captionText="→ Fig. 9. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior median dorsal (AG) and posterior median dorsal (HM) plates of the trunk armour. A. IG KSC 155/5 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/108 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/97 in dorsal view. D. IG KSC 155/113 in dorsal (D1) and visceral (D2) views. E. IG KSC 155/140 in dorsal (E1) and visceral (E2) views. F. Impression of the dorsal surface of IG KSC 155/42.G. IG KSC 155/44 in dorsal view. H. Fragment of IG KSC 155/7 in dorsal view. I. IG KSC 155/1 in dorsal (I1) and visceral (I2) views. J. IG KSC 155/71 in dorsal view. K. Slightly deformed IG KSC 155/70 in dorsal (K1) and visceral (K2) views. L. IG KSC 155/158 in dorsal view. M. IG KSC 155/157 in dorsal (M1) and visceral (M2) views. Abbreviations: ADL, anterior dorso-lateral plate; alr, postlevator thickening; AMD, anterior median dorsal plate; cf.ADL, cf.AMD, and cf.MxL, area overlapping ADL, AMD or MxL respectively; cr.tp, posterior transversal internal crest; dlg1 and dlg2, anterior and posterior oblique dorsal sensory line groove; dma, tergal angle; dmr, dorsal median ridge; f.retr, levator fossa; grm, ventral median groove; l, lateral corner; mvr, median ventral ridge; MxL, mixilateral plate; npn, postnuchal notch; oa.ADL, oa.MxL and oa.PMD, area overlapped by ADL, MxL or PMD respectively; pa, posterior corner; pma, posterior marginal area; PMD, posterior median dorsal plate; pr.p, posterior process of AMD; pr.pl, external postlevator process; prv2, posterior ventral process of dorsal wall of trunk armour; pt1 and pt2, anterior and posterior ventral pit; pua, posterior unornamented area of PMD; rf, “round fossula”; sna, supranuchal area; tb, ventral tuberosity." figureDoi="http://doi.org/10.5281/zenodo.11061576" httpUri="https://zenodo.org/record/11061576/files/figure.png" pageId="13" pageNumber="110">Fig. 9J, N</figureCitation>
), which is funnel-shaped, quite elongated and oblique in sagittal view so that the anterior wall is very low and the posterior wall is rather high.
</paragraph>
<paragraph id="E9483B2C4C5AFFEDFCBC852EA4B9F682" blockId="13.[808,1482,160,1960]" pageId="13" pageNumber="110">
The dorsal lamina of
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is of moderate breadth, about 2.63.3 times (2.9 on the average) as long as it is broad (
<figureCitation id="71CC27A94C5AFFEDFCC285EEA39FF5EB" box="[816,956,1192,1217]" captionStart="Fig" captionStartId="10.[151,184,862,882]" captionTargetBox="[155,1534,159,837]" captionTargetId="figure-325@10.[152,1535,159,836]" captionTargetPageId="10" captionText="Fig. 10. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior dorso-lateral (AD, F, left; E, right) and left mixilateral (GI, right) plates of the trunk armour. A. IG KSC 155/116 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/111. C. IG KSC 155/61. D. IG KSC 155/60. E. IG KSC 155/136. F. IG KSC 155/112. G. IG KSC 155/110. H. Incomplete IG KSC 155/144. I. IG KSC 155/115. BE, GI, in dorsal views; F, in lateral view.Abbreviations: ADL, anterior ventro-lateral plate; AMD, anterior median dorsal plate; AVL, anterior ventro-lateral plate; cf.AMD, area overlapping AMD; dlg2, posterior oblique dorsal sensory line groove; dlr, dorso-lateral ridge of trunk armour; dxp, dorso-ventral pit-line groove; lcg, main lateral line groove; oa.ADL, oa.AMD, oa.AVL, oa.PMD, area overlapped by ADL, AMD, AVL or PMD respectively; PMD, posterior median dorsal plate; pnoa, postnuchal ornamented corner of ADL; pro, processus obstans of trunk armour." figureDoi="http://doi.org/10.5281/zenodo.10980839" httpUri="https://zenodo.org/record/10980839/files/figure.png" pageId="13" pageNumber="110">Fig. 10AG</figureCitation>
), thus the proportions of the
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of
<taxonomicName id="2EF740AF4C5AFFEDFA8885EEA42AF5C8" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" baseAuthorityName="Rohon" baseAuthorityYear="1900" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="110" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="jeremejevi">
<emphasis id="DB83E73E4C5AFFEDFA8885EEA42AF5C8" italics="true" pageId="13" pageNumber="110">B. leptocheira jeremejevi</emphasis>
</taxonomicName>
are more similar to those of
<taxonomicName id="2EF740AF4C5AFFEDFA85858FA3D6F428" baseAuthorityName="Luksevics" baseAuthorityYear="2001" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="110" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="curonica">
<emphasis id="DB83E73E4C5AFFEDFA85858FA3D6F428" italics="true" pageId="13" pageNumber="110">B. leptocheira curonica</emphasis>
</taxonomicName>
(e.g., LDM 98/58;
<bibRefCitation id="8D6646DD4C5AFFEDFB2185AFA5A9F428" author="Luksevics, E." box="[1235,1418,1257,1282]" pageId="13" pageNumber="110" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
: figs. 36B, 37A) and
<collectionCode id="8FE6A3E94C5AFFEDFC3D844FA42AF408" box="[975,1033,1289,1314]" pageId="13" pageNumber="110">ADL</collectionCode>
is shorter than it is 3.5 times longer and broader in
<taxonomicName id="2EF740AF4C5AFFEDFC5F846CA506F469" authority="RSM" authorityName="RSM" box="[941,1317,1322,1347]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="110" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="leptocheira">
<emphasis id="DB83E73E4C5AFFEDFC5F846CA4FCF469" box="[941,1247,1322,1347]" italics="true" pageId="13" pageNumber="110">B. leptocheira leptocheira</emphasis>
RSM
</taxonomicName>
1859.33.632D (
<bibRefCitation id="8D6646DD4C5AFFEDFCC2840CA396F449" author="Miles, R. S." box="[816,949,1354,1379]" pageId="13" pageNumber="110" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles 1968</bibRefCitation>
: pl. 20: 3). The postnuchal ornamented corner (pnoa) is prominent in all studied specimens. The width of the dorsal lamina is usually slightly larger than the height of the lateral lamina. The dorso-lateral ridge (dlr) is well pronounced. Usually, the
<collectionCode id="8FE6A3E94C5AFFEDFBBE848DA4ACF4CE" box="[1100,1167,1483,1508]" pageId="13" pageNumber="110">AMD</collectionCode>
overlaps the mesial margin of the
<collectionCode id="8FE6A3E94C5AFFEDFC8484ADA390F72E" box="[886,947,1515,1540]" pageId="13" pageNumber="110">ADL</collectionCode>
, but in IG KSC 155/116 (
<figureCitation id="71CC27A94C5AFFEDFB1284ADA567F72E" box="[1248,1348,1515,1540]" captionStart="Fig" captionStartId="10.[151,184,862,882]" captionTargetBox="[155,1534,159,837]" captionTargetId="figure-325@10.[152,1535,159,836]" captionTargetPageId="10" captionText="Fig. 10. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior dorso-lateral (AD, F, left; E, right) and left mixilateral (GI, right) plates of the trunk armour. A. IG KSC 155/116 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/111. C. IG KSC 155/61. D. IG KSC 155/60. E. IG KSC 155/136. F. IG KSC 155/112. G. IG KSC 155/110. H. Incomplete IG KSC 155/144. I. IG KSC 155/115. BE, GI, in dorsal views; F, in lateral view.Abbreviations: ADL, anterior ventro-lateral plate; AMD, anterior median dorsal plate; AVL, anterior ventro-lateral plate; cf.AMD, area overlapping AMD; dlg2, posterior oblique dorsal sensory line groove; dlr, dorso-lateral ridge of trunk armour; dxp, dorso-ventral pit-line groove; lcg, main lateral line groove; oa.ADL, oa.AMD, oa.AVL, oa.PMD, area overlapped by ADL, AMD, AVL or PMD respectively; PMD, posterior median dorsal plate; pnoa, postnuchal ornamented corner of ADL; pro, processus obstans of trunk armour." figureDoi="http://doi.org/10.5281/zenodo.10980839" httpUri="https://zenodo.org/record/10980839/files/figure.png" pageId="13" pageNumber="110">Fig. 10A</figureCitation>
<figureCitation id="71CC27A94C5AFFEDFAB784B0A56DF72D" box="[1349,1358,1526,1543]" captionStart="Fig" captionStartId="2.[151,184,1333,1353]" captionTargetBox="[152,823,164,1305]" captionTargetId="figure-630@2.[152,604,164,1305]" captionTargetPageId="2" captionText="Fig. 2. Lithology and stratigraphy of the Sosnogorsk locality. “Fish-dolomite”, bed No. 40 is located about 8 m above the base of the section. Abbreviations: D3f, Upper Devonian, Frasnian; D3fm, Upper Devonian, Famennian." figureDoi="http://doi.org/10.5281/zenodo.10980823" httpUri="https://zenodo.org/record/10980823/files/figure.png" pageId="13" pageNumber="110">
<subScript id="757339694C5AFFEDFAB784B0A56DF72D" attach="right" box="[1349,1358,1526,1543]" fontSize="7" pageId="13" pageNumber="110">2</subScript>
</figureCitation>
) the
<collectionCode id="8FE6A3E94C5AFFEDFA7D84ADA5EAF72E" box="[1423,1481,1515,1540]" pageId="13" pageNumber="110">ADL</collectionCode>
overlaps the
<collectionCode id="8FE6A3E94C5AFFEDFC48874DA3DEF70E" box="[954,1021,1547,1572]" pageId="13" pageNumber="110">AMD</collectionCode>
(cf.
<collectionCode id="8FE6A3E94C5AFFEDFBDA874DA44FF70E" box="[1064,1132,1547,1572]" pageId="13" pageNumber="110">AMD</collectionCode>
) along the posterior third part of the common suture. The outline of the mesial margin of this specimen greatly complements the lateral margin of
<collectionCode id="8FE6A3E94C5AFFEDFCDA872AA348F7AF" box="[808,875,1644,1669]" pageId="13" pageNumber="110">AMD</collectionCode>
plate IG KSC 155/44 (
<figureCitation id="71CC27A94C5AFFEDFB81872AA4EDF7AF" box="[1139,1230,1644,1669]" captionStart="→" captionStartId="10.[116,144,1620,1650]" captionTargetId="figure-8@9.[99,1479,175,1936]" captionTargetPageId="9" captionText="→ Fig. 9. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior median dorsal (AG) and posterior median dorsal (HM) plates of the trunk armour. A. IG KSC 155/5 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/108 in dorsal (B1) and visceral (B2) views. C. IG KSC 155/97 in dorsal view. D. IG KSC 155/113 in dorsal (D1) and visceral (D2) views. E. IG KSC 155/140 in dorsal (E1) and visceral (E2) views. F. Impression of the dorsal surface of IG KSC 155/42.G. IG KSC 155/44 in dorsal view. H. Fragment of IG KSC 155/7 in dorsal view. I. IG KSC 155/1 in dorsal (I1) and visceral (I2) views. J. IG KSC 155/71 in dorsal view. K. Slightly deformed IG KSC 155/70 in dorsal (K1) and visceral (K2) views. L. IG KSC 155/158 in dorsal view. M. IG KSC 155/157 in dorsal (M1) and visceral (M2) views. Abbreviations: ADL, anterior dorso-lateral plate; alr, postlevator thickening; AMD, anterior median dorsal plate; cf.ADL, cf.AMD, and cf.MxL, area overlapping ADL, AMD or MxL respectively; cr.tp, posterior transversal internal crest; dlg1 and dlg2, anterior and posterior oblique dorsal sensory line groove; dma, tergal angle; dmr, dorsal median ridge; f.retr, levator fossa; grm, ventral median groove; l, lateral corner; mvr, median ventral ridge; MxL, mixilateral plate; npn, postnuchal notch; oa.ADL, oa.MxL and oa.PMD, area overlapped by ADL, MxL or PMD respectively; pa, posterior corner; pma, posterior marginal area; PMD, posterior median dorsal plate; pr.p, posterior process of AMD; pr.pl, external postlevator process; prv2, posterior ventral process of dorsal wall of trunk armour; pt1 and pt2, anterior and posterior ventral pit; pua, posterior unornamented area of PMD; rf, “round fossula”; sna, supranuchal area; tb, ventral tuberosity." figureDoi="http://doi.org/10.5281/zenodo.11061576" httpUri="https://zenodo.org/record/11061576/files/figure.png" pageId="13" pageNumber="110">Fig. 9G</figureCitation>
). Thus, this feature is not remarkably rare in
<taxonomicName id="2EF740AF4C5AFFEDFBDA87CBA561F78C" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" baseAuthorityName="Rohon" baseAuthorityYear="1900" box="[1064,1346,1677,1702]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="110" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="jeremejevi">
<emphasis id="DB83E73E4C5AFFEDFBDA87CBA561F78C" box="[1064,1346,1677,1702]" italics="true" pageId="13" pageNumber="110">B. leptocheira jeremejevi</emphasis>
</taxonomicName>
. The lateral lamina of
<collectionCode id="8FE6A3E94C5AFFEDFC6F87EBA3F5F7EC" box="[925,982,1709,1734]" pageId="13" pageNumber="110">ADL</collectionCode>
is 4.7 times as long as it is high in IG KSC IG 155/112 (
<figureCitation id="71CC27A94C5AFFEDFC47878BA438F7CC" box="[949,1051,1741,1766]" captionStart="Fig" captionStartId="10.[151,184,862,882]" captionTargetBox="[155,1534,159,837]" captionTargetId="figure-325@10.[152,1535,159,836]" captionTargetPageId="10" captionText="Fig. 10. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior dorso-lateral (AD, F, left; E, right) and left mixilateral (GI, right) plates of the trunk armour. A. IG KSC 155/116 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/111. C. IG KSC 155/61. D. IG KSC 155/60. E. IG KSC 155/136. F. IG KSC 155/112. G. IG KSC 155/110. H. Incomplete IG KSC 155/144. I. IG KSC 155/115. BE, GI, in dorsal views; F, in lateral view.Abbreviations: ADL, anterior ventro-lateral plate; AMD, anterior median dorsal plate; AVL, anterior ventro-lateral plate; cf.AMD, area overlapping AMD; dlg2, posterior oblique dorsal sensory line groove; dlr, dorso-lateral ridge of trunk armour; dxp, dorso-ventral pit-line groove; lcg, main lateral line groove; oa.ADL, oa.AMD, oa.AVL, oa.PMD, area overlapped by ADL, AMD, AVL or PMD respectively; PMD, posterior median dorsal plate; pnoa, postnuchal ornamented corner of ADL; pro, processus obstans of trunk armour." figureDoi="http://doi.org/10.5281/zenodo.10980839" httpUri="https://zenodo.org/record/10980839/files/figure.png" pageId="13" pageNumber="110">Fig. 10G</figureCitation>
) and more than 6 times as long as it is high in IG KSC IG 71/i-68; the lateral lamina of
<collectionCode id="8FE6A3E94C5AFFEDFA8687ABA58DF62C" box="[1396,1454,1773,1798]" pageId="13" pageNumber="110">ADL</collectionCode>
is slightly higher in the material from
<collectingCountry id="91E07BBC4C5AFFEDFB4B8648A521F60D" box="[1209,1282,1806,1831]" name="Latvia" pageId="13" pageNumber="110">Latvia</collectingCountry>
(
<bibRefCitation id="8D6646DD4C5AFFEDFAE28648A5E5F60D" author="Luksevics, E." box="[1296,1478,1806,1831]" pageId="13" pageNumber="110" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
: fig. 37B), and Scottish material does not show the lateral lamina well (
<bibRefCitation id="8D6646DD4C5AFFEDFC328608A452F64D" author="Miles, R. S." box="[960,1137,1870,1895]" pageId="13" pageNumber="110" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles 1968: 78</bibRefCitation>
). The processus obstans (pro) is rather deep, but the course of the main lateral line groove (lcg) conforms the usual pattern.
</paragraph>
<caption id="BD886BA44C59FFEEFF658228A506F2D8" ID-DOI="http://doi.org/10.5281/zenodo.10980845" ID-Zenodo-Dep="10980845" httpUri="https://zenodo.org/record/10980845/files/figure.png" pageId="14" pageNumber="111" startId="14.[151,184,878,898]" targetBox="[161,1533,162,851]" targetPageId="14" targetType="figure">
<paragraph id="E9483B2C4C59FFEEFF658228A506F2D8" blockId="14.[151,1535,878,1013]" pageId="14" pageNumber="111">
Fig. 13. Antiarchan fish
<taxonomicName id="2EF740AF4C59FFEEFE868228A31CF2A8" authority="(Rohon, 1900)" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" baseAuthorityName="Rohon" baseAuthorityYear="1900" box="[372,831,878,898]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="111" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="jeremejevi">
<emphasis id="DB83E73E4C59FFEEFE868228A297F2A8" box="[372,692,878,898]" italics="true" pageId="14" pageNumber="111">Bothriolepis leptocheira jeremejevi</emphasis>
(
<bibRefCitation id="8D6646DD4C59FFEEFD4D8228A314F2A8" author="Rohon, J. V." box="[703,823,878,898]" pageId="14" pageNumber="111" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon, 1900</bibRefCitation>
)
</taxonomicName>
, Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, plates of the pectoral fin armour.
<emphasis id="DB83E73E4C59FFEEFE8682CFA1A7F2B4" bold="true" box="[372,388,905,926]" pageId="14" pageNumber="111">A</emphasis>
. Left Cv
<subScript id="757339694C59FFEEFE2482D5A1FDF28B" attach="left" box="[470,478,915,929]" fontSize="6" pageId="14" pageNumber="111">1</subScript>
plate IG 155/6 in visceral (A
<subScript id="757339694C59FFEEFD1982D5A2D0F28B" attach="right" box="[747,755,915,929]" fontSize="6" pageId="14" pageNumber="111">1</subScript>
) and ventral (A
<subScript id="757339694C59FFEEFC7682D5A3AFF28B" attach="left" box="[900,908,915,929]" fontSize="6" pageId="14" pageNumber="111">2</subScript>
) views.
<emphasis id="DB83E73E4C59FFEEFC2882CFA3CAF2B4" bold="true" box="[986,1001,905,926]" pageId="14" pageNumber="111">B</emphasis>
. Right Cv
<subScript id="757339694C59FFEEFBBA82D5A473F28B" attach="left" box="[1096,1104,915,929]" fontSize="6" pageId="14" pageNumber="111">1</subScript>
plate IG KSC 155/134 in ventral view.
<emphasis id="DB83E73E4C59FFEEFA4F82CFA5EEF2B5" bold="true" box="[1469,1485,905,927]" pageId="14" pageNumber="111">C</emphasis>
. Left Ml
<subScript id="757339694C59FFEEFF4082E9A099F297" attach="left" box="[178,186,943,957]" fontSize="6" pageId="14" pageNumber="111">2</subScript>
plate IG KSC 155/149 in ventral view.
<emphasis id="DB83E73E4C59FFEEFDD782E3A216F290" bold="true" box="[549,565,933,954]" pageId="14" pageNumber="111">D</emphasis>
. Right Ml
<subScript id="757339694C59FFEEFD6682E9A2BFF297" attach="left" box="[660,668,943,957]" fontSize="6" pageId="14" pageNumber="111">2</subScript>
plate IG 155/137 in dorsal (D
<subScript id="757339694C59FFEEFC4282E9A39BF297" attach="right" box="[944,952,943,957]" fontSize="6" pageId="14" pageNumber="111">1</subScript>
) and ventral (D
<subScript id="757339694C59FFEEFBBB82E9A472F297" attach="right" box="[1097,1105,943,957]" fontSize="6" pageId="14" pageNumber="111">2</subScript>
) views.
<emphasis id="DB83E73E4C59FFEEFB6D82E3A48DF290" bold="true" box="[1183,1198,933,954]" pageId="14" pageNumber="111">E</emphasis>
. Right distal segment of the pectoral fin armour IG KSC 155/21 in dorsal (E
<subScript id="757339694C59FFEEFE0B828DA222F2F3" attach="left" box="[505,513,971,985]" fontSize="6" pageId="14" pageNumber="111">1</subScript>
) and ventral (E
<subScript id="757339694C59FFEEFD7C828DA2B5F2F3" attach="left" box="[654,662,971,985]" fontSize="6" pageId="14" pageNumber="111">2</subScript>
) views.
<emphasis id="DB83E73E4C59FFEEFD118287A2D2F2FC" bold="true" box="[739,753,961,982]" pageId="14" pageNumber="111">F</emphasis>
. Mm
<subScript id="757339694C59FFEEFCD0828DA309F2F3" attach="left" box="[802,810,971,985]" fontSize="6" pageId="14" pageNumber="111">1</subScript>
plate IG KSC 155/85 in mesial view. Abbreviations: Cd
<subScript id="757339694C59FFEEFAD5828DA51DF2F3" attach="left" box="[1319,1342,971,985]" fontSize="6" pageId="14" pageNumber="111">35</subScript>
, dorsal central plates 3 to 5; Cv
<subScript id="757339694C59FFEEFF0382A1A0DAF2DF" attach="left" box="[241,249,999,1013]" fontSize="6" pageId="14" pageNumber="111">4</subScript>
, ventral central plate 4; Ml
<subScript id="757339694C59FFEEFE0282A1A22BF2DF" attach="right" box="[496,520,999,1013]" fontSize="6" pageId="14" pageNumber="111">45</subScript>
, lateral marginal plates 4 and 5; Mm
<subScript id="757339694C59FFEEFCA482A1A34DF2DF" attach="left" box="[854,878,999,1013]" fontSize="6" pageId="14" pageNumber="111">46</subScript>
, mesial marginal plates 4 to 6; T, terminal plate.
</paragraph>
</caption>
<paragraph id="E9483B2C4C59FFEEFF4F8548A224F7ED" blockId="14.[151,825,1038,1959]" pageId="14" pageNumber="111">
The dorsal lamina of
<collectionCode id="8FE6A3E94C59FFEEFE478548A1CEF50D" box="[437,493,1038,1063]" pageId="14" pageNumber="111">MxL</collectionCode>
is slightly less than twice as long as it is broad (
<figureCitation id="71CC27A94C59FFEEFE9D8568A1CCF56D" box="[367,495,1070,1095]" captionStart="Fig" captionStartId="10.[151,184,862,882]" captionTargetBox="[155,1534,159,837]" captionTargetId="figure-325@10.[152,1535,159,836]" captionTargetPageId="10" captionText="Fig. 10. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior dorso-lateral (AD, F, left; E, right) and left mixilateral (GI, right) plates of the trunk armour. A. IG KSC 155/116 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/111. C. IG KSC 155/61. D. IG KSC 155/60. E. IG KSC 155/136. F. IG KSC 155/112. G. IG KSC 155/110. H. Incomplete IG KSC 155/144. I. IG KSC 155/115. BE, GI, in dorsal views; F, in lateral view.Abbreviations: ADL, anterior ventro-lateral plate; AMD, anterior median dorsal plate; AVL, anterior ventro-lateral plate; cf.AMD, area overlapping AMD; dlg2, posterior oblique dorsal sensory line groove; dlr, dorso-lateral ridge of trunk armour; dxp, dorso-ventral pit-line groove; lcg, main lateral line groove; oa.ADL, oa.AMD, oa.AVL, oa.PMD, area overlapped by ADL, AMD, AVL or PMD respectively; PMD, posterior median dorsal plate; pnoa, postnuchal ornamented corner of ADL; pro, processus obstans of trunk armour." figureDoi="http://doi.org/10.5281/zenodo.10980839" httpUri="https://zenodo.org/record/10980839/files/figure.png" pageId="14" pageNumber="111">Fig. 10HJ</figureCitation>
). The dorsal corner is clearly seen. The lateral lamina is moderately low, about 22.5 times as long as it is high, but relatively higher than in material from
<collectingCountry id="91E07BBC4C59FFEEFEF185C8A16FF58D" box="[259,332,1166,1191]" name="Latvia" pageId="14" pageNumber="111">Latvia</collectingCountry>
where it is about three times as long as it is high (
<bibRefCitation id="8D6646DD4C59FFEEFF2B85E8A1B3F5ED" author="Luksevics, E." box="[217,400,1198,1223]" pageId="14" pageNumber="111" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
: fig. 37E). The posterior oblique sensory line groove (dlg
<subScript id="757339694C59FFEEFE77859CA1ADF5C1" attach="left" box="[389,398,1242,1259]" fontSize="7" pageId="14" pageNumber="111">2</subScript>
) usually terminates in some distance from the lateral margin (e.g., IG KSC 155/82); as in
<taxonomicName id="2EF740AF4C59FFEEFD1985A8A188F40D" authority="LDM" authorityName="LDM" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="111" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="curonica">
<emphasis id="DB83E73E4C59FFEEFD1985A8A147F40D" italics="true" pageId="14" pageNumber="111">B. leptocheira curonica</emphasis>
LDM
</taxonomicName>
98/53 (
<bibRefCitation id="8D6646DD4C59FFEEFDF68448A29EF40D" author="Luksevics, E." box="[516,701,1294,1319]" pageId="14" pageNumber="111" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
: fig. 36C), IG KSC 155/115 shows a dlg
<subScript id="757339694C59FFEEFE3D847CA1FAF461" attach="left" box="[463,473,1338,1355]" fontSize="7" pageId="14" pageNumber="111">2</subScript>
crossing the dorso-lateral ridge (
<figureCitation id="71CC27A94C59FFEEFF6D8408A0DAF44D" box="[159,249,1358,1383]" captionStart="Fig" captionStartId="10.[151,184,862,882]" captionTargetBox="[155,1534,159,837]" captionTargetId="figure-325@10.[152,1535,159,836]" captionTargetPageId="10" captionText="Fig. 10. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior dorso-lateral (AD, F, left; E, right) and left mixilateral (GI, right) plates of the trunk armour. A. IG KSC 155/116 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/111. C. IG KSC 155/61. D. IG KSC 155/60. E. IG KSC 155/136. F. IG KSC 155/112. G. IG KSC 155/110. H. Incomplete IG KSC 155/144. I. IG KSC 155/115. BE, GI, in dorsal views; F, in lateral view.Abbreviations: ADL, anterior ventro-lateral plate; AMD, anterior median dorsal plate; AVL, anterior ventro-lateral plate; cf.AMD, area overlapping AMD; dlg2, posterior oblique dorsal sensory line groove; dlr, dorso-lateral ridge of trunk armour; dxp, dorso-ventral pit-line groove; lcg, main lateral line groove; oa.ADL, oa.AMD, oa.AVL, oa.PMD, area overlapped by ADL, AMD, AVL or PMD respectively; PMD, posterior median dorsal plate; pnoa, postnuchal ornamented corner of ADL; pro, processus obstans of trunk armour." figureDoi="http://doi.org/10.5281/zenodo.10980839" httpUri="https://zenodo.org/record/10980839/files/figure.png" pageId="14" pageNumber="111">Fig. 10I</figureCitation>
). The separate dorso-ventral pit-line groove (dxp) is present in a medium sized IG KSC 155/144 (
<figureCitation id="71CC27A94C59FFEEFD748428A2C8F4AD" box="[646,747,1390,1415]" captionStart="Fig" captionStartId="10.[151,184,862,882]" captionTargetBox="[155,1534,159,837]" captionTargetId="figure-325@10.[152,1535,159,836]" captionTargetPageId="10" captionText="Fig. 10. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, anterior dorso-lateral (AD, F, left; E, right) and left mixilateral (GI, right) plates of the trunk armour. A. IG KSC 155/116 in dorsal (A1) and visceral (A2) views. B. IG KSC 155/111. C. IG KSC 155/61. D. IG KSC 155/60. E. IG KSC 155/136. F. IG KSC 155/112. G. IG KSC 155/110. H. Incomplete IG KSC 155/144. I. IG KSC 155/115. BE, GI, in dorsal views; F, in lateral view.Abbreviations: ADL, anterior ventro-lateral plate; AMD, anterior median dorsal plate; AVL, anterior ventro-lateral plate; cf.AMD, area overlapping AMD; dlg2, posterior oblique dorsal sensory line groove; dlr, dorso-lateral ridge of trunk armour; dxp, dorso-ventral pit-line groove; lcg, main lateral line groove; oa.ADL, oa.AMD, oa.AVL, oa.PMD, area overlapped by ADL, AMD, AVL or PMD respectively; PMD, posterior median dorsal plate; pnoa, postnuchal ornamented corner of ADL; pro, processus obstans of trunk armour." figureDoi="http://doi.org/10.5281/zenodo.10980839" httpUri="https://zenodo.org/record/10980839/files/figure.png" pageId="14" pageNumber="111">Fig. 10H</figureCitation>
), similarly to
<taxonomicName id="2EF740AF4C59FFEEFF1C84C8A1FDF48D" authority="Gross, 1942" authorityName="Gross" authorityYear="1942" box="[238,478,1422,1447]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="111" phylum="Chordata" rank="species" species="prima">
<emphasis id="DB83E73E4C59FFEEFF1C84C8A173F48D" box="[238,336,1422,1447]" italics="true" pageId="14" pageNumber="111">B. prima</emphasis>
<bibRefCitation id="8D6646DD4C59FFEEFEA584C8A1FDF48D" author="Gross, W." box="[343,478,1422,1447]" pageId="14" pageNumber="111" pagination="373 - 436" refId="ref18663" refString="Gross, W. 1942. Die Fischfaunen des baltischen Devons und ihre biostratigraphische Bedeutung. Korrespondenz-blatt des Naturforscher-Vereins zu Riga 64: 373 - 436." type="journal article" year="1942">Gross, 1942</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="8D6646DD4C59FFEEFE1984C8A324F48D" author="Karatajute-Talimaa, V. N." box="[491,775,1422,1447]" pageId="14" pageNumber="111" pagination="191 - 279" refId="ref18937" refString="Karatajute-Talimaa, V. N. 1966. Bothriolepids of Sventoji Regional Stage of the Baltics [in Russian]. In: A. Grigelis (ed.), Paleontologia i stratigrafia Pribaltiki i Belorussji I (VI), 191 - 279. Mintis, Vilnius." type="journal article" year="1966">Karatajūte-Talimaa 1966</bibRefCitation>
: fig. 3),
<taxonomicName id="2EF740AF4C59FFEEFF4C84E8A1DEF4ED" authority="Gross, 1942" authorityName="Gross" authorityYear="1942" box="[190,509,1454,1479]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="111" phylum="Chordata" rank="species" species="obrutschewi">
<emphasis id="DB83E73E4C59FFEEFF4C84E8A14FF4ED" box="[190,364,1454,1479]" italics="true" pageId="14" pageNumber="111">B. obrutschewi</emphasis>
<bibRefCitation id="8D6646DD4C59FFEEFE8584E8A1DEF4ED" author="Gross, W." box="[375,509,1454,1479]" pageId="14" pageNumber="111" pagination="373 - 436" refId="ref18663" refString="Gross, W. 1942. Die Fischfaunen des baltischen Devons und ihre biostratigraphische Bedeutung. Korrespondenz-blatt des Naturforscher-Vereins zu Riga 64: 373 - 436." type="journal article" year="1942">Gross, 1942</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="8D6646DD4C59FFEEFDE684E8A317F4ED" author="Karatajute-Talimaa, V. N." box="[532,820,1454,1479]" pageId="14" pageNumber="111" pagination="191 - 279" refId="ref18937" refString="Karatajute-Talimaa, V. N. 1966. Bothriolepids of Sventoji Regional Stage of the Baltics [in Russian]. In: A. Grigelis (ed.), Paleontologia i stratigrafia Pribaltiki i Belorussji I (VI), 191 - 279. Mintis, Vilnius." type="journal article" year="1966">Karatajūte-Talimaa 1966</bibRefCitation>
: fig. 10), and
<taxonomicName id="2EF740AF4C59FFEEFEC28488A27CF4CD" authority="Lyarskaya, 1986" authorityName="Lyarskaya" authorityYear="1986" box="[304,607,1486,1511]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="111" phylum="Chordata" rank="species" species="evaldi">
<emphasis id="DB83E73E4C59FFEEFEC28488A1BBF4CD" box="[304,408,1486,1511]" italics="true" pageId="14" pageNumber="111">B. evaldi</emphasis>
<bibRefCitation id="8D6646DD4C59FFEEFE508488A27CF4CD" author="Lyarskaya, L. &amp; Larskaa, L." box="[418,607,1486,1511]" pageId="14" pageNumber="111" pagination="123 - 130" refId="ref19702" refString="Lyarskaya, L. [Larskaa, L.] 1986. A new Bothriolepis (Antiarchi) from the Upper Devonian of Baltic states [in Russian]. In: A. Brangulis (ed.), Biofacii i fauna silurijskogo i devonskih bassejnov Pribaltiki, 123 - 130. Zinatne, Riga." type="book chapter" year="1986">Lyarskaya, 1986</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="8D6646DD4C59FFEEFD838488A30FF4CD" author="Lyarskaya, L. &amp; Larskaa, L." box="[625,812,1486,1511]" pageId="14" pageNumber="111" pagination="123 - 130" refId="ref19702" refString="Lyarskaya, L. [Larskaa, L.] 1986. A new Bothriolepis (Antiarchi) from the Upper Devonian of Baltic states [in Russian]. In: A. Brangulis (ed.), Biofacii i fauna silurijskogo i devonskih bassejnov Pribaltiki, 123 - 130. Zinatne, Riga." type="book chapter" year="1986">Lyarskaya 1986</bibRefCitation>
). The dorso-lateral ridge (dlr) is well defined both in the
<collectionCode id="8FE6A3E94C59FFEEFD0C84A8A31BF72D" box="[766,824,1518,1543]" pageId="14" pageNumber="111">ADL</collectionCode>
and
<collectionCode id="8FE6A3E94C59FFEEFF3B8748A121F70D" box="[201,258,1550,1575]" pageId="14" pageNumber="111">MxL</collectionCode>
plates of small and medium sized specimens, as it is also seen in material from
<collectingCountry id="91E07BBC4C59FFEEFDE88768A241F76D" box="[538,610,1582,1607]" name="Latvia" pageId="14" pageNumber="111">Latvia</collectingCountry>
;
<bibRefCitation id="8D6646DD4C59FFEEFD858768A310F76D" author="Miles, R. S." box="[631,819,1582,1607]" pageId="14" pageNumber="111" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles (1968: 78)</bibRefCitation>
claimed that “there is no clear development of the dlr on either the
<collectionCode id="8FE6A3E94C59FFEEFEFB8728A161F7AD" box="[265,322,1646,1671]" pageId="14" pageNumber="111">MxL</collectionCode>
or
<collectionCode id="8FE6A3E94C59FFEEFE9B8728A186F7AD" box="[361,421,1646,1671]" pageId="14" pageNumber="111">ADL</collectionCode>
”; in fact dlr is not clearly seen, particularly in
<collectionCode id="8FE6A3E94C59FFEEFED087C8A17DF78D" box="[290,350,1678,1703]" pageId="14" pageNumber="111">MxL</collectionCode>
, because of preservation of the Scottish material as internal impressions.
</paragraph>
<paragraph id="E9483B2C4C59FFEEFF4F8788A33AF60D" blockId="14.[151,825,1038,1959]" pageId="14" pageNumber="111">
The semilunar plate (
<collectionCode id="8FE6A3E94C59FFEEFE488788A1C7F7CD" box="[442,484,1742,1767]" country="Malaysia" name="Sarawak Museum" pageId="14" pageNumber="111">SM</collectionCode>
) is not known, but from the configuration of the anterior margin of the
<collectionCode id="8FE6A3E94C59FFEEFD7487A8A29EF62D" box="[646,701,1774,1799]" pageId="14" pageNumber="111">AVL</collectionCode>
plates, the SM is of average proportions and normal shape (
<figureCitation id="71CC27A94C59FFEEFD4E8648A32FF60D" box="[700,780,1806,1831]" captionStart="Fig" captionStartId="15.[98,131,1132,1152]" captionTargetBox="[103,764,164,1109]" captionTargetId="figure-870@15.[102,764,160,1109]" captionTargetPageId="15" captionText="Fig. 14. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), reconstruction of the head, the trunk armour and the pectoral fins in ventral view, based on IG KSC 155/10-1, IG KSC 155/10-2, and IG KSC 155/10- 3. Abbreviations: AVL, anterior ventro-lateral plate; Cv14, ventral central plate 1 to 4; Ifg, inferognathal plate; La, lateral plate; Ml25, lateral marginal plates 2 to 5; Mm4, mesial marginal plate 4; MV, median ventral plate; Prl, prelateral plate; Prm, premedian plate; PVL, posterior ventro-lateral plate; SM, semilunar plate; SMg, submarginal plate; T, terminal plate." figureDoi="http://doi.org/10.5281/zenodo.10980847" httpUri="https://zenodo.org/record/10980847/files/figure.png" pageId="14" pageNumber="111">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="E9483B2C4C59FFEEFF4F8668A4BDF7ED" blockId="14.[151,825,1038,1959]" lastBlockId="14.[862,1535,1038,1959]" pageId="14" pageNumber="111">
The ventral lamina of the
<collectionCode id="8FE6A3E94C59FFEEFE068668A208F66D" box="[500,555,1838,1863]" pageId="14" pageNumber="111">AVL</collectionCode>
is 1.62 times as long as it is broad (
<figureCitation id="71CC27A94C59FFEEFEB38608A1B1F64D" box="[321,402,1870,1895]" captionStart="Fig" captionStartId="11.[98,131,1825,1845]" captionTargetBox="[99,1479,163,1803]" captionTargetId="figure-8@11.[99,1480,163,1804]" captionTargetPageId="11" captionText="Fig. 11. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, partial ventral wall (A, B), anterior ventro-lateral (AVL) (C, D, left; E, right), and median ventral (FJ) plates of the trunk armour, and elements of pectoral fin. A. Impression of partial ventral wall of the trunk armour, inferognathals, and suborbitals IG KSC 155/10-2 in ventral view. B. Partial ventral wall of the trunk armour with articulated pectoral fin armour IG KSC 155/10-1 in ventral view. C. IG KSC 155/151 (C1) and enlargened part of the ventral lamina with pathologies (C2, arrows). D. IG KSC 155/122 in ventral view. E. IG KSC 155/123 in ventral view. F. IG KSC 155/104 with the bite mark (arrow) → in ventral (F1) and visceral (F2) views. G. IG KSC 155/76 in ventral view. H. IG KSC 155/77 in ventral view. I. IG KSC 155/133 in ventral view. J. IG KSC 155/106 in ventral view. Abbreviations:AVL, anterior ventro-lateral plate; Cd12, dorsal central plates 1 and 2; Cv1, ventral central plate 1; cit1, crista transversalis interna anterior; cit2, transverse thickening on the visceral surface of AVL; ds, distal segment of the pectoral fin armour; Ifg, inferognathal; Ml2, lateral marginal plate 2; Mm2, mesial marginal plate 2; MV, median ventral plate; PVL, posterior ventro-lateral plate; SO, suborbital plate." figureDoi="http://doi.org/10.5281/zenodo.10980841" httpUri="https://zenodo.org/record/10980841/files/figure.png" pageId="14" pageNumber="111">Fig. 11</figureCitation>
). The anterior margin of the ventral lamina is rounded without clearly defined corners, similar to
<taxonomicName id="2EF740AF4C59FFEEFF4586C8A1C7F68D" box="[183,484,1934,1959]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="111" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="leptocheira">
<emphasis id="DB83E73E4C59FFEEFF4586C8A1C7F68D" box="[183,484,1934,1959]" italics="true" pageId="14" pageNumber="111">B. leptocheira leptocheira</emphasis>
</taxonomicName>
(
<bibRefCitation id="8D6646DD4C59FFEEFE0786C8A286F68D" author="Miles, R. S." box="[501,677,1934,1959]" pageId="14" pageNumber="111" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles 1968: 79</bibRefCitation>
, text-fig. 39) and
<taxonomicName id="2EF740AF4C59FFEEFC7D8548A4BEF50D" box="[911,1181,1038,1063]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="111" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="curonica">
<emphasis id="DB83E73E4C59FFEEFC7D8548A4BEF50D" box="[911,1181,1038,1063]" italics="true" pageId="14" pageNumber="111">B. leptocheira curonica</emphasis>
</taxonomicName>
(
<bibRefCitation id="8D6646DD4C59FFEEFB5C8548A546F50D" author="Luksevics, E." box="[1198,1381,1038,1063]" pageId="14" pageNumber="111" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
: figs. 37F, G, 38A). The subcephalic division comprises 2130% (24% on average) of the total length of the ventral lamina (about 20% in the Scottish material [
<bibRefCitation id="8D6646DD4C59FFEEFB4C8528A564F5AD" author="Miles, R. S." box="[1214,1351,1134,1159]" pageId="14" pageNumber="111" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles 1968</bibRefCitation>
]; about 2125% with 22.5% on average in the Latvian material [
<bibRefCitation id="8D6646DD4C59FFEEFA7A85C8A3ECF5ED" author="Luksevics, E." pageId="14" pageNumber="111" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001: 551</bibRefCitation>
]). The lateral lamina is not preserved within the material from
<collectingCountry id="91E07BBC4C59FFEEFBF78588A448F5CD" box="[1029,1131,1230,1255]" name="United Kingdom" pageId="14" pageNumber="111">Scotland</collectingCountry>
. The lateral lamina is low, and the ventral lamina is 3.8 times as broad as the lateral lamina is high in
<taxonomicName id="2EF740AF4C59FFEEFC278448A508F40D" authority="LDM" authorityName="LDM" box="[981,1323,1294,1319]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="111" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="curonica">
<emphasis id="DB83E73E4C59FFEEFC278448A4C7F40D" box="[981,1252,1294,1319]" italics="true" pageId="14" pageNumber="111">B. leptocheira curonica</emphasis>
LDM
</taxonomicName>
98/30. The lateral lamina is relatively higher in the Sosnogorsk material, e.g., IG KSC 155/122 (
<figureCitation id="71CC27A94C59FFEEFBDB8408A4A8F44D" box="[1065,1163,1358,1383]" captionStart="Fig" captionStartId="11.[98,131,1825,1845]" captionTargetBox="[99,1479,163,1803]" captionTargetId="figure-8@11.[99,1480,163,1804]" captionTargetPageId="11" captionText="Fig. 11. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, partial ventral wall (A, B), anterior ventro-lateral (AVL) (C, D, left; E, right), and median ventral (FJ) plates of the trunk armour, and elements of pectoral fin. A. Impression of partial ventral wall of the trunk armour, inferognathals, and suborbitals IG KSC 155/10-2 in ventral view. B. Partial ventral wall of the trunk armour with articulated pectoral fin armour IG KSC 155/10-1 in ventral view. C. IG KSC 155/151 (C1) and enlargened part of the ventral lamina with pathologies (C2, arrows). D. IG KSC 155/122 in ventral view. E. IG KSC 155/123 in ventral view. F. IG KSC 155/104 with the bite mark (arrow) → in ventral (F1) and visceral (F2) views. G. IG KSC 155/76 in ventral view. H. IG KSC 155/77 in ventral view. I. IG KSC 155/133 in ventral view. J. IG KSC 155/106 in ventral view. Abbreviations:AVL, anterior ventro-lateral plate; Cd12, dorsal central plates 1 and 2; Cv1, ventral central plate 1; cit1, crista transversalis interna anterior; cit2, transverse thickening on the visceral surface of AVL; ds, distal segment of the pectoral fin armour; Ifg, inferognathal; Ml2, lateral marginal plate 2; Mm2, mesial marginal plate 2; MV, median ventral plate; PVL, posterior ventro-lateral plate; SO, suborbital plate." figureDoi="http://doi.org/10.5281/zenodo.10980841" httpUri="https://zenodo.org/record/10980841/files/figure.png" pageId="14" pageNumber="111">Fig. 11D</figureCitation>
) shows the ventral lamina is 2.9 times as broad as the lateral lamina is high. The right
<collectionCode id="8FE6A3E94C59FFEEFA3A8428A5DCF4AD" box="[1480,1535,1390,1415]" pageId="14" pageNumber="111">AVL</collectionCode>
overlaps the left
<collectionCode id="8FE6A3E94C59FFEEFBD384C8A47BF48D" box="[1057,1112,1422,1447]" pageId="14" pageNumber="111">AVL</collectionCode>
by a very narrow overlap area. The pectoral fin attachment area is usually not well preserved in the studied Sosnogorsk material, except on
<collectionCode id="8FE6A3E94C59FFEEFA948488A5BEF4CD" box="[1382,1437,1486,1511]" pageId="14" pageNumber="111">AVL</collectionCode>
IG KSC 155/122 (
<figureCitation id="71CC27A94C59FFEEFC3B84A8A412F72D" box="[969,1073,1518,1543]" captionStart="Fig" captionStartId="11.[98,131,1825,1845]" captionTargetBox="[99,1479,163,1803]" captionTargetId="figure-8@11.[99,1480,163,1804]" captionTargetPageId="11" captionText="Fig. 11. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, partial ventral wall (A, B), anterior ventro-lateral (AVL) (C, D, left; E, right), and median ventral (FJ) plates of the trunk armour, and elements of pectoral fin. A. Impression of partial ventral wall of the trunk armour, inferognathals, and suborbitals IG KSC 155/10-2 in ventral view. B. Partial ventral wall of the trunk armour with articulated pectoral fin armour IG KSC 155/10-1 in ventral view. C. IG KSC 155/151 (C1) and enlargened part of the ventral lamina with pathologies (C2, arrows). D. IG KSC 155/122 in ventral view. E. IG KSC 155/123 in ventral view. F. IG KSC 155/104 with the bite mark (arrow) → in ventral (F1) and visceral (F2) views. G. IG KSC 155/76 in ventral view. H. IG KSC 155/77 in ventral view. I. IG KSC 155/133 in ventral view. J. IG KSC 155/106 in ventral view. Abbreviations:AVL, anterior ventro-lateral plate; Cd12, dorsal central plates 1 and 2; Cv1, ventral central plate 1; cit1, crista transversalis interna anterior; cit2, transverse thickening on the visceral surface of AVL; ds, distal segment of the pectoral fin armour; Ifg, inferognathal; Ml2, lateral marginal plate 2; Mm2, mesial marginal plate 2; MV, median ventral plate; PVL, posterior ventro-lateral plate; SO, suborbital plate." figureDoi="http://doi.org/10.5281/zenodo.10980841" httpUri="https://zenodo.org/record/10980841/files/figure.png" pageId="14" pageNumber="111">Fig. 11D</figureCitation>
), which shows the typical morphology of
<taxonomicName id="2EF740AF4C59FFEEFC728748A428F70D" authorityName="Eichwald" authorityYear="1840" box="[896,1035,1550,1575]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="111" phylum="Chordata" rank="genus">
<emphasis id="DB83E73E4C59FFEEFC728748A428F70D" box="[896,1035,1550,1575]" italics="true" pageId="14" pageNumber="111">Bothriolepis</emphasis>
</taxonomicName>
. Internally, the crista transversalis interna anterior (cit
<subScript id="757339694C59FFEEFC11877CA3CFF761" attach="left" box="[995,1004,1594,1611]" fontSize="7" pageId="14" pageNumber="111">1</subScript>
;
<figureCitation id="71CC27A94C59FFEEFC0D8768A442F76D" box="[1023,1121,1582,1607]" captionStart="Fig" captionStartId="11.[98,131,1825,1845]" captionTargetBox="[99,1479,163,1803]" captionTargetId="figure-8@11.[99,1480,163,1804]" captionTargetPageId="11" captionText="Fig. 11. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, partial ventral wall (A, B), anterior ventro-lateral (AVL) (C, D, left; E, right), and median ventral (FJ) plates of the trunk armour, and elements of pectoral fin. A. Impression of partial ventral wall of the trunk armour, inferognathals, and suborbitals IG KSC 155/10-2 in ventral view. B. Partial ventral wall of the trunk armour with articulated pectoral fin armour IG KSC 155/10-1 in ventral view. C. IG KSC 155/151 (C1) and enlargened part of the ventral lamina with pathologies (C2, arrows). D. IG KSC 155/122 in ventral view. E. IG KSC 155/123 in ventral view. F. IG KSC 155/104 with the bite mark (arrow) → in ventral (F1) and visceral (F2) views. G. IG KSC 155/76 in ventral view. H. IG KSC 155/77 in ventral view. I. IG KSC 155/133 in ventral view. J. IG KSC 155/106 in ventral view. Abbreviations:AVL, anterior ventro-lateral plate; Cd12, dorsal central plates 1 and 2; Cv1, ventral central plate 1; cit1, crista transversalis interna anterior; cit2, transverse thickening on the visceral surface of AVL; ds, distal segment of the pectoral fin armour; Ifg, inferognathal; Ml2, lateral marginal plate 2; Mm2, mesial marginal plate 2; MV, median ventral plate; PVL, posterior ventro-lateral plate; SO, suborbital plate." figureDoi="http://doi.org/10.5281/zenodo.10980841" httpUri="https://zenodo.org/record/10980841/files/figure.png" pageId="14" pageNumber="111">Fig. 11A</figureCitation>
) is developed as a high, thin ridge running at an oblique angle across half the width of the anterior portion of the plate. The transverse thickening (cit
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) is more prominent in comparison with cit
<subScript id="757339694C59FFEEFAEF87DCA504F781" attach="left" box="[1309,1319,1690,1707]" fontSize="7" pageId="14" pageNumber="111">1</subScript>
as it is seen in the IG KSC 155/10-2 (
<figureCitation id="71CC27A94C59FFEEFBDC87E8A4B2F7ED" box="[1070,1169,1710,1735]" captionStart="Fig" captionStartId="11.[98,131,1825,1845]" captionTargetBox="[99,1479,163,1803]" captionTargetId="figure-8@11.[99,1480,163,1804]" captionTargetPageId="11" captionText="Fig. 11. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, partial ventral wall (A, B), anterior ventro-lateral (AVL) (C, D, left; E, right), and median ventral (FJ) plates of the trunk armour, and elements of pectoral fin. A. Impression of partial ventral wall of the trunk armour, inferognathals, and suborbitals IG KSC 155/10-2 in ventral view. B. Partial ventral wall of the trunk armour with articulated pectoral fin armour IG KSC 155/10-1 in ventral view. C. IG KSC 155/151 (C1) and enlargened part of the ventral lamina with pathologies (C2, arrows). D. IG KSC 155/122 in ventral view. E. IG KSC 155/123 in ventral view. F. IG KSC 155/104 with the bite mark (arrow) → in ventral (F1) and visceral (F2) views. G. IG KSC 155/76 in ventral view. H. IG KSC 155/77 in ventral view. I. IG KSC 155/133 in ventral view. J. IG KSC 155/106 in ventral view. Abbreviations:AVL, anterior ventro-lateral plate; Cd12, dorsal central plates 1 and 2; Cv1, ventral central plate 1; cit1, crista transversalis interna anterior; cit2, transverse thickening on the visceral surface of AVL; ds, distal segment of the pectoral fin armour; Ifg, inferognathal; Ml2, lateral marginal plate 2; Mm2, mesial marginal plate 2; MV, median ventral plate; PVL, posterior ventro-lateral plate; SO, suborbital plate." figureDoi="http://doi.org/10.5281/zenodo.10980841" httpUri="https://zenodo.org/record/10980841/files/figure.png" pageId="14" pageNumber="111">Fig. 11A</figureCitation>
).
</paragraph>
<paragraph id="E9483B2C4C59FFEFFC768788A185F70B" blockId="14.[862,1535,1038,1959]" lastBlockId="15.[98,771,1380,1960]" lastPageId="15" lastPageNumber="112" pageId="14" pageNumber="111">
The posterior ventro-lateral plate (
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) has variable proportions (
<figureCitation id="71CC27A94C59FFEEFC0087A8A466F62D" box="[1010,1093,1774,1799]" captionStart="Fig" captionStartId="12.[151,184,958,978]" captionTargetBox="[161,1528,166,928]" captionTargetId="figure-466@12.[157,1529,161,933]" captionTargetPageId="12" captionText="Fig. 12. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, posterior ventro-lateral plates, left (A) and right (B, C) of the trunk armour. A. IG KSC 155/4 in ventral (A1) and lateral (A2) views. B. IG KSC 155/16 in ventral view. C. IG KSC 155/78 in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10980843" httpUri="https://zenodo.org/record/10980843/files/figure.png" pageId="14" pageNumber="111">Fig. 12</figureCitation>
). It is relatively narrow with a ventral lamina that is 2.12.7 times as longer than it is broad. The subanal division is rather narrow and relatively longer than in the Latvian material where it occupies about one third (2838%) of the total
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plate length. This is comparable with the 33% in
<taxonomicName id="2EF740AF4C59FFEFFB8186C8A0EEF457" authority="(Miles 1968: 80)" baseAuthorityName="Miles" baseAuthorityPageNumber="80" baseAuthorityYear="1968" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="15" lastPageNumber="112" pageId="14" pageNumber="111" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="leptocheira">
<emphasis id="DB83E73E4C59FFEEFB8186C8A584F68D" box="[1139,1447,1934,1959]" italics="true" pageId="14" pageNumber="111">B. leptocheira leptocheira</emphasis>
(
<bibRefCitation id="8D6646DD4C59FFEFFA4E86C8A0E6F457" author="Miles, R. S." lastPageId="15" lastPageNumber="112" pageId="14" pageNumber="111" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles 1968: 80</bibRefCitation>
)
</taxonomicName>
, but slightly different from
<taxonomicName id="2EF740AF4C58FFEFFDFF8422A0A7F4B4" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="112" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="curonica">
<emphasis id="DB83E73E4C58FFEFFDFF8422A0A7F4B4" italics="true" pageId="15" pageNumber="112">B. leptocheira curonica</emphasis>
</taxonomicName>
(
<bibRefCitation id="8D6646DD4C58FFEFFF6684C3A16AF4B4" author="Luksevics, E." box="[148,329,1413,1438]" pageId="15" pageNumber="112" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Luksevics 2001</bibRefCitation>
: figs. 38B, C, 39C) where it varies between 22% and 32%. The lateral lamina is moderately high in IG KSC 155/4 (
<figureCitation id="71CC27A94C58FFEFFECA8480A183F4F5" box="[312,416,1478,1503]" captionStart="Fig" captionStartId="12.[151,184,958,978]" captionTargetBox="[161,1528,166,928]" captionTargetId="figure-466@12.[157,1529,161,933]" captionTargetPageId="12" captionText="Fig. 12. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, posterior ventro-lateral plates, left (A) and right (B, C) of the trunk armour. A. IG KSC 155/4 in ventral (A1) and lateral (A2) views. B. IG KSC 155/16 in ventral view. C. IG KSC 155/78 in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10980843" httpUri="https://zenodo.org/record/10980843/files/figure.png" pageId="15" pageNumber="112">Fig. 12A</figureCitation>
). The ventro-lateral ridge (vlr) is clearly defined both in the
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and
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. The left
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overlaps the right
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plate.
</paragraph>
<caption id="BD886BA44C58FFEFFF90852AA2BDF46E" ID-DOI="http://doi.org/10.5281/zenodo.10980847" ID-Zenodo-Dep="10980847" httpUri="https://zenodo.org/record/10980847/files/figure.png" pageId="15" pageNumber="112" startId="15.[98,131,1132,1152]" targetBox="[103,764,164,1109]" targetPageId="15" targetType="figure">
<paragraph id="E9483B2C4C58FFEFFF90852AA2BDF46E" blockId="15.[98,771,1132,1348]" pageId="15" pageNumber="112">
Fig. 14. Antiarchan fish
<taxonomicName id="2EF740AF4C58FFEFFEC8852AA2DCF5AA" authority="(Rohon, 1900)" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" baseAuthorityName="Rohon" baseAuthorityYear="1900" box="[314,767,1132,1152]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="112" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="jeremejevi">
<emphasis id="DB83E73E4C58FFEFFEC8852AA255F5AA" box="[314,630,1132,1152]" italics="true" pageId="15" pageNumber="112">Bothriolepis leptocheira jeremejevi</emphasis>
(
<bibRefCitation id="8D6646DD4C58FFEFFD72852AA2DBF5AA" author="Rohon, J. V." box="[640,760,1132,1152]" pageId="15" pageNumber="112" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon, 1900</bibRefCitation>
)
</taxonomicName>
, reconstruction of the head, the trunk armour and the pectoral fins in ventral view, based on IG KSC 155/10-1, IG KSC 155/10-2, and IG KSC 155/10- 3. Abbreviations: AVL, anterior ventro-lateral plate; Cv
<subScript id="757339694C58FFEFFDA9858FA250F5FD" attach="left" box="[603,627,1225,1239]" fontSize="6" pageId="15" pageNumber="112">14</subScript>
, ventral central plate 1 to 4; Ifg, inferognathal plate; La, lateral plate; Ml
<subScript id="757339694C58FFEFFDA685A3A24FF5D9" attach="left" box="[596,620,1253,1267]" fontSize="6" pageId="15" pageNumber="112">25</subScript>
, lateral marginal plates 2 to 5; Mm
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, mesial marginal plate 4; MV, median ventral plate; Prl, prelateral plate; Prm, premedian plate; PVL, posterior ventro-lateral plate; SM, semilunar plate; SMg, submarginal plate; T, terminal plate.
</paragraph>
</caption>
<paragraph id="E9483B2C4C58FFEFFF75876EA4A6F013" blockId="15.[98,771,1380,1960]" lastBlockId="15.[808,1483,160,1888]" pageId="15" pageNumber="112">
The median ventral plate (
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;
<figureCitation id="71CC27A94C58FFEFFE15876EA246F76B" box="[487,613,1576,1601]" captionStart="Fig" captionStartId="11.[98,131,1825,1845]" captionTargetBox="[99,1479,163,1803]" captionTargetId="figure-8@11.[99,1480,163,1804]" captionTargetPageId="11" captionText="Fig. 11. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, partial ventral wall (A, B), anterior ventro-lateral (AVL) (C, D, left; E, right), and median ventral (FJ) plates of the trunk armour, and elements of pectoral fin. A. Impression of partial ventral wall of the trunk armour, inferognathals, and suborbitals IG KSC 155/10-2 in ventral view. B. Partial ventral wall of the trunk armour with articulated pectoral fin armour IG KSC 155/10-1 in ventral view. C. IG KSC 155/151 (C1) and enlargened part of the ventral lamina with pathologies (C2, arrows). D. IG KSC 155/122 in ventral view. E. IG KSC 155/123 in ventral view. F. IG KSC 155/104 with the bite mark (arrow) → in ventral (F1) and visceral (F2) views. G. IG KSC 155/76 in ventral view. H. IG KSC 155/77 in ventral view. I. IG KSC 155/133 in ventral view. J. IG KSC 155/106 in ventral view. Abbreviations:AVL, anterior ventro-lateral plate; Cd12, dorsal central plates 1 and 2; Cv1, ventral central plate 1; cit1, crista transversalis interna anterior; cit2, transverse thickening on the visceral surface of AVL; ds, distal segment of the pectoral fin armour; Ifg, inferognathal; Ml2, lateral marginal plate 2; Mm2, mesial marginal plate 2; MV, median ventral plate; PVL, posterior ventro-lateral plate; SO, suborbital plate." figureDoi="http://doi.org/10.5281/zenodo.10980841" httpUri="https://zenodo.org/record/10980841/files/figure.png" pageId="15" pageNumber="112">Fig. 11FK</figureCitation>
) is not known in the material from
<collectingCountry id="91E07BBC4C58FFEFFEA6870FA194F748" box="[340,439,1609,1634]" name="United Kingdom" pageId="15" pageNumber="112">Scotland</collectingCountry>
(
<bibRefCitation id="8D6646DD4C58FFEFFE3A870FA255F748" author="Miles, R. S." box="[456,630,1609,1634]" pageId="15" pageNumber="112" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles 1968: 80</bibRefCitation>
) and
<collectingCountry id="91E07BBC4C58FFEFFD4B870FA321F748" box="[697,770,1609,1634]" name="Latvia" pageId="15" pageNumber="112">Latvia</collectingCountry>
<bibRefCitation id="8D6646DD4C58FFEFFF9B872FA176F7A8" author="Luksevics, E." box="[105,341,1641,1666]" pageId="15" pageNumber="112" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001: 552</bibRefCitation>
), whereas it is well represented in the Sosnogorsk material by several disarticulated specimens and also found in partially articulated ventral walls of the trunk armour. This plate is slightly elongated with an L/W index of about 1.2 on average. The shape of
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and
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plates in
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<emphasis id="DB83E73E4C58FFEFFF8D864AA1A9F60F" box="[127,394,1804,1829]" italics="true" pageId="15" pageNumber="112">B. leptocheira curonica</emphasis>
(
<bibRefCitation id="8D6646DD4C58FFEFFE6B864AA2A2F60F" author="Luksevics, E." box="[409,641,1804,1829]" pageId="15" pageNumber="112" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001: 552</bibRefCitation>
)
</taxonomicName>
suggests a moderately small size of the
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; whereas in
<taxonomicName id="2EF740AF4C58FFEFFD90866BA0F4F64D" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" baseAuthorityName="Rohon" baseAuthorityYear="1900" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="112" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="jeremejevi">
<emphasis id="DB83E73E4C58FFEFFD90866BA0F4F64D" italics="true" pageId="15" pageNumber="112">B. leptocheira jeremejevi</emphasis>
</taxonomicName>
, it is rather large (
<figureCitation id="71CC27A94C58FFEFFE4D8608A24CF64D" box="[447,623,1870,1895]" captionStart="Fig" captionStartId="11.[98,131,1825,1845]" captionTargetBox="[99,1479,163,1803]" captionTargetId="figure-8@11.[99,1480,163,1804]" captionTargetPageId="11" captionText="Fig. 11. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, partial ventral wall (A, B), anterior ventro-lateral (AVL) (C, D, left; E, right), and median ventral (FJ) plates of the trunk armour, and elements of pectoral fin. A. Impression of partial ventral wall of the trunk armour, inferognathals, and suborbitals IG KSC 155/10-2 in ventral view. B. Partial ventral wall of the trunk armour with articulated pectoral fin armour IG KSC 155/10-1 in ventral view. C. IG KSC 155/151 (C1) and enlargened part of the ventral lamina with pathologies (C2, arrows). D. IG KSC 155/122 in ventral view. E. IG KSC 155/123 in ventral view. F. IG KSC 155/104 with the bite mark (arrow) → in ventral (F1) and visceral (F2) views. G. IG KSC 155/76 in ventral view. H. IG KSC 155/77 in ventral view. I. IG KSC 155/133 in ventral view. J. IG KSC 155/106 in ventral view. Abbreviations:AVL, anterior ventro-lateral plate; Cd12, dorsal central plates 1 and 2; Cv1, ventral central plate 1; cit1, crista transversalis interna anterior; cit2, transverse thickening on the visceral surface of AVL; ds, distal segment of the pectoral fin armour; Ifg, inferognathal; Ml2, lateral marginal plate 2; Mm2, mesial marginal plate 2; MV, median ventral plate; PVL, posterior ventro-lateral plate; SO, suborbital plate." figureDoi="http://doi.org/10.5281/zenodo.10980841" httpUri="https://zenodo.org/record/10980841/files/figure.png" pageId="15" pageNumber="112">Fig. 11A, FK</figureCitation>
). Almost all
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plates are from relatively large individuals, the plate reaching
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in length. Some plates are asymmetrical in shape, e.g., IG KSC 155/77 (
<figureCitation id="71CC27A94C58FFEFFB3981E6A50CF193" box="[1227,1327,160,185]" captionStart="Fig" captionStartId="11.[98,131,1825,1845]" captionTargetBox="[99,1479,163,1803]" captionTargetId="figure-8@11.[99,1480,163,1804]" captionTargetPageId="11" captionText="Fig. 11. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, partial ventral wall (A, B), anterior ventro-lateral (AVL) (C, D, left; E, right), and median ventral (FJ) plates of the trunk armour, and elements of pectoral fin. A. Impression of partial ventral wall of the trunk armour, inferognathals, and suborbitals IG KSC 155/10-2 in ventral view. B. Partial ventral wall of the trunk armour with articulated pectoral fin armour IG KSC 155/10-1 in ventral view. C. IG KSC 155/151 (C1) and enlargened part of the ventral lamina with pathologies (C2, arrows). D. IG KSC 155/122 in ventral view. E. IG KSC 155/123 in ventral view. F. IG KSC 155/104 with the bite mark (arrow) → in ventral (F1) and visceral (F2) views. G. IG KSC 155/76 in ventral view. H. IG KSC 155/77 in ventral view. I. IG KSC 155/133 in ventral view. J. IG KSC 155/106 in ventral view. Abbreviations:AVL, anterior ventro-lateral plate; Cd12, dorsal central plates 1 and 2; Cv1, ventral central plate 1; cit1, crista transversalis interna anterior; cit2, transverse thickening on the visceral surface of AVL; ds, distal segment of the pectoral fin armour; Ifg, inferognathal; Ml2, lateral marginal plate 2; Mm2, mesial marginal plate 2; MV, median ventral plate; PVL, posterior ventro-lateral plate; SO, suborbital plate." figureDoi="http://doi.org/10.5281/zenodo.10980841" httpUri="https://zenodo.org/record/10980841/files/figure.png" pageId="15" pageNumber="112">Fig. 11H</figureCitation>
) and 155/106 (
<figureCitation id="71CC27A94C58FFEFFCC28186A3B6F1F3" box="[816,917,192,217]" captionStart="Fig" captionStartId="11.[98,131,1825,1845]" captionTargetBox="[99,1479,163,1803]" captionTargetId="figure-8@11.[99,1480,163,1804]" captionTargetPageId="11" captionText="Fig. 11. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, partial ventral wall (A, B), anterior ventro-lateral (AVL) (C, D, left; E, right), and median ventral (FJ) plates of the trunk armour, and elements of pectoral fin. A. Impression of partial ventral wall of the trunk armour, inferognathals, and suborbitals IG KSC 155/10-2 in ventral view. B. Partial ventral wall of the trunk armour with articulated pectoral fin armour IG KSC 155/10-1 in ventral view. C. IG KSC 155/151 (C1) and enlargened part of the ventral lamina with pathologies (C2, arrows). D. IG KSC 155/122 in ventral view. E. IG KSC 155/123 in ventral view. F. IG KSC 155/104 with the bite mark (arrow) → in ventral (F1) and visceral (F2) views. G. IG KSC 155/76 in ventral view. H. IG KSC 155/77 in ventral view. I. IG KSC 155/133 in ventral view. J. IG KSC 155/106 in ventral view. Abbreviations:AVL, anterior ventro-lateral plate; Cd12, dorsal central plates 1 and 2; Cv1, ventral central plate 1; cit1, crista transversalis interna anterior; cit2, transverse thickening on the visceral surface of AVL; ds, distal segment of the pectoral fin armour; Ifg, inferognathal; Ml2, lateral marginal plate 2; Mm2, mesial marginal plate 2; MV, median ventral plate; PVL, posterior ventro-lateral plate; SO, suborbital plate." figureDoi="http://doi.org/10.5281/zenodo.10980841" httpUri="https://zenodo.org/record/10980841/files/figure.png" pageId="15" pageNumber="112">Fig. 11K</figureCitation>
). Usually all margins of the plate are slightly convex or almost straight, but in IG KSC 155/133 both posterior lateral margins are concave (
<figureCitation id="71CC27A94C58FFEFFB9B8046A4E0F033" box="[1129,1219,256,281]" captionStart="Fig" captionStartId="11.[98,131,1825,1845]" captionTargetBox="[99,1479,163,1803]" captionTargetId="figure-8@11.[99,1480,163,1804]" captionTargetPageId="11" captionText="Fig. 11. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, partial ventral wall (A, B), anterior ventro-lateral (AVL) (C, D, left; E, right), and median ventral (FJ) plates of the trunk armour, and elements of pectoral fin. A. Impression of partial ventral wall of the trunk armour, inferognathals, and suborbitals IG KSC 155/10-2 in ventral view. B. Partial ventral wall of the trunk armour with articulated pectoral fin armour IG KSC 155/10-1 in ventral view. C. IG KSC 155/151 (C1) and enlargened part of the ventral lamina with pathologies (C2, arrows). D. IG KSC 155/122 in ventral view. E. IG KSC 155/123 in ventral view. F. IG KSC 155/104 with the bite mark (arrow) → in ventral (F1) and visceral (F2) views. G. IG KSC 155/76 in ventral view. H. IG KSC 155/77 in ventral view. I. IG KSC 155/133 in ventral view. J. IG KSC 155/106 in ventral view. Abbreviations:AVL, anterior ventro-lateral plate; Cd12, dorsal central plates 1 and 2; Cv1, ventral central plate 1; cit1, crista transversalis interna anterior; cit2, transverse thickening on the visceral surface of AVL; ds, distal segment of the pectoral fin armour; Ifg, inferognathal; Ml2, lateral marginal plate 2; Mm2, mesial marginal plate 2; MV, median ventral plate; PVL, posterior ventro-lateral plate; SO, suborbital plate." figureDoi="http://doi.org/10.5281/zenodo.10980841" httpUri="https://zenodo.org/record/10980841/files/figure.png" pageId="15" pageNumber="112">Fig. 11J</figureCitation>
). Overlap areas usually are rather narrow (
<figureCitation id="71CC27A94C58FFEFFC098066A45BF013" box="[1019,1144,288,313]" captionStart="Fig" captionStartId="11.[98,131,1825,1845]" captionTargetBox="[99,1479,163,1803]" captionTargetId="figure-8@11.[99,1480,163,1804]" captionTargetPageId="11" captionText="Fig. 11. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, partial ventral wall (A, B), anterior ventro-lateral (AVL) (C, D, left; E, right), and median ventral (FJ) plates of the trunk armour, and elements of pectoral fin. A. Impression of partial ventral wall of the trunk armour, inferognathals, and suborbitals IG KSC 155/10-2 in ventral view. B. Partial ventral wall of the trunk armour with articulated pectoral fin armour IG KSC 155/10-1 in ventral view. C. IG KSC 155/151 (C1) and enlargened part of the ventral lamina with pathologies (C2, arrows). D. IG KSC 155/122 in ventral view. E. IG KSC 155/123 in ventral view. F. IG KSC 155/104 with the bite mark (arrow) → in ventral (F1) and visceral (F2) views. G. IG KSC 155/76 in ventral view. H. IG KSC 155/77 in ventral view. I. IG KSC 155/133 in ventral view. J. IG KSC 155/106 in ventral view. Abbreviations:AVL, anterior ventro-lateral plate; Cd12, dorsal central plates 1 and 2; Cv1, ventral central plate 1; cit1, crista transversalis interna anterior; cit2, transverse thickening on the visceral surface of AVL; ds, distal segment of the pectoral fin armour; Ifg, inferognathal; Ml2, lateral marginal plate 2; Mm2, mesial marginal plate 2; MV, median ventral plate; PVL, posterior ventro-lateral plate; SO, suborbital plate." figureDoi="http://doi.org/10.5281/zenodo.10980841" httpUri="https://zenodo.org/record/10980841/files/figure.png" pageId="15" pageNumber="112">Fig. 11H, J</figureCitation>
).
</paragraph>
<paragraph id="E9483B2C4C58FFEFFCBC8006A49DF591" blockId="15.[808,1483,160,1888]" pageId="15" pageNumber="112">
The pectoral fin is represented in the Sosnogorsk material by several disarticulated bones, at least
<specimenCount id="FFF1F0A54C58FFEFFAE68026A5EAF053" box="[1300,1481,352,377]" count="3" pageId="15" pageNumber="112" type="generic">three specimens</specimenCount>
showing articulated plates of the proximal segment associated with the
<collectionCode id="8FE6A3E94C58FFEFFC7E80E6A3E1F093" box="[908,962,416,441]" pageId="15" pageNumber="112">AVL</collectionCode>
(e.g.,
<figureCitation id="71CC27A94C58FFEFFBF480E6A447F093" box="[1030,1124,416,441]" captionStart="Fig" captionStartId="11.[98,131,1825,1845]" captionTargetBox="[99,1479,163,1803]" captionTargetId="figure-8@11.[99,1480,163,1804]" captionTargetPageId="11" captionText="Fig. 11. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, partial ventral wall (A, B), anterior ventro-lateral (AVL) (C, D, left; E, right), and median ventral (FJ) plates of the trunk armour, and elements of pectoral fin. A. Impression of partial ventral wall of the trunk armour, inferognathals, and suborbitals IG KSC 155/10-2 in ventral view. B. Partial ventral wall of the trunk armour with articulated pectoral fin armour IG KSC 155/10-1 in ventral view. C. IG KSC 155/151 (C1) and enlargened part of the ventral lamina with pathologies (C2, arrows). D. IG KSC 155/122 in ventral view. E. IG KSC 155/123 in ventral view. F. IG KSC 155/104 with the bite mark (arrow) → in ventral (F1) and visceral (F2) views. G. IG KSC 155/76 in ventral view. H. IG KSC 155/77 in ventral view. I. IG KSC 155/133 in ventral view. J. IG KSC 155/106 in ventral view. Abbreviations:AVL, anterior ventro-lateral plate; Cd12, dorsal central plates 1 and 2; Cv1, ventral central plate 1; cit1, crista transversalis interna anterior; cit2, transverse thickening on the visceral surface of AVL; ds, distal segment of the pectoral fin armour; Ifg, inferognathal; Ml2, lateral marginal plate 2; Mm2, mesial marginal plate 2; MV, median ventral plate; PVL, posterior ventro-lateral plate; SO, suborbital plate." figureDoi="http://doi.org/10.5281/zenodo.10980841" httpUri="https://zenodo.org/record/10980841/files/figure.png" pageId="15" pageNumber="112">Fig. 11B</figureCitation>
), and one articulated distal segment (
<figureCitation id="71CC27A94C58FFEFFC9C8086A3EDF0F3" box="[878,974,448,473]" captionStart="Fig" captionStartId="14.[151,184,878,898]" captionTargetBox="[161,1533,162,851]" captionTargetId="figure-604@14.[153,1534,163,851]" captionTargetPageId="14" captionText="Fig. 13. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, plates of the pectoral fin armour. A. Left Cv1 plate IG 155/6 in visceral (A1) and ventral (A2) views. B. Right Cv1 plate IG KSC 155/134 in ventral view. C. Left Ml2 plate IG KSC 155/149 in ventral view. D. Right Ml2 plate IG 155/137 in dorsal (D1) and ventral (D2) views. E. Right distal segment of the pectoral fin armour IG KSC 155/21 in dorsal (E1) and ventral (E2) views. F. Mm1 plate IG KSC 155/85 in mesial view. Abbreviations: Cd35, dorsal central plates 3 to 5; Cv4, ventral central plate 4; Ml45, lateral marginal plates 4 and 5; Mm46, mesial marginal plates 4 to 6; T, terminal plate." figureDoi="http://doi.org/10.5281/zenodo.10980845" httpUri="https://zenodo.org/record/10980845/files/figure.png" pageId="15" pageNumber="112">Fig. 13E</figureCitation>
). The proximal segment bears small rarely set lateral and mesial spines. The lateral spines are larger than the mesial ones (which are even not seen in figured specimens) as in
<taxonomicName id="2EF740AF4C58FFEFFCB48367A46EF310" baseAuthorityName="Luksevics" baseAuthorityYear="2001" box="[838,1101,545,570]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="112" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="curonica">
<emphasis id="DB83E73E4C58FFEFFCB48367A46EF310" box="[838,1101,545,570]" italics="true" pageId="15" pageNumber="112">B. leptocheira curonica</emphasis>
</taxonomicName>
(
<bibRefCitation id="8D6646DD4C58FFEFFBA98367A52CF310" author="Luksevics, E." box="[1115,1295,545,570]" pageId="15" pageNumber="112" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
: figs. 38A, 39B);
<bibRefCitation id="8D6646DD4C58FFEFFCDA8307A390F370" author="Miles, R. S." box="[808,947,577,602]" pageId="15" pageNumber="112" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles (1968)</bibRefCitation>
suggested the mesial spines were perhaps absent in
<taxonomicName id="2EF740AF4C58FFEFFCB48327A44AF350" baseAuthorityName="Miles" baseAuthorityYear="1968" box="[838,1129,609,634]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="112" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="leptocheira">
<emphasis id="DB83E73E4C58FFEFFCB48327A44AF350" box="[838,1129,609,634]" italics="true" pageId="15" pageNumber="112">B. leptocheira leptocheira</emphasis>
</taxonomicName>
. The proximal segment is very long and narrow, it is 4.6 times longer than it is broad in IG KSC 155/10. It is slightly less elongated than in the Scottish material (5.5 to 6 times as long as it is broad;
<bibRefCitation id="8D6646DD4C58FFEFFAEF8387A5E2F3F0" author="Miles, R. S." box="[1309,1473,705,730]" pageId="15" pageNumber="112" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles 1968: 80</bibRefCitation>
) and in the Latvian material (4.86 [5.5 on the average] times as long as it is broad;
<bibRefCitation id="8D6646DD4C58FFEFFBD38247A52BF230" author="Luksevics, E." box="[1057,1288,769,794]" pageId="15" pageNumber="112" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001: 553</bibRefCitation>
). The Cd
<subScript id="757339694C58FFEFFA9C824AA55BF237" attach="left" box="[1390,1400,780,797]" fontSize="7" pageId="15" pageNumber="112">1</subScript>
has an L/B index varying from 2.6 to 2.9 (2.73.7, 3.3 on average in the Latvian material;
<bibRefCitation id="8D6646DD4C58FFEFFBD78207A530F270" author="Luksevics, E." box="[1061,1299,833,858]" pageId="15" pageNumber="112" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001: 553</bibRefCitation>
); measurements were made during the field work in 2009. The Cv
<subScript id="757339694C58FFEFFAAA822BA541F254" attach="left" box="[1368,1378,877,894]" fontSize="7" pageId="15" pageNumber="112">1</subScript>
is gently longer than the Cd
<subScript id="757339694C58FFEFFC0482CBA423F2B4" attach="left" box="[1014,1024,909,926]" fontSize="7" pageId="15" pageNumber="112">1</subScript>
and slightly more elongated with an L/B index of 3.54.1 (3.9 on the average) (
<figureCitation id="71CC27A94C58FFEFFB2082E4A575F291" box="[1234,1366,930,955]" captionStart="Fig" captionStartId="14.[151,184,878,898]" captionTargetBox="[161,1533,162,851]" captionTargetId="figure-604@14.[153,1534,163,851]" captionTargetPageId="14" captionText="Fig. 13. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, plates of the pectoral fin armour. A. Left Cv1 plate IG 155/6 in visceral (A1) and ventral (A2) views. B. Right Cv1 plate IG KSC 155/134 in ventral view. C. Left Ml2 plate IG KSC 155/149 in ventral view. D. Right Ml2 plate IG 155/137 in dorsal (D1) and ventral (D2) views. E. Right distal segment of the pectoral fin armour IG KSC 155/21 in dorsal (E1) and ventral (E2) views. F. Mm1 plate IG KSC 155/85 in mesial view. Abbreviations: Cd35, dorsal central plates 3 to 5; Cv4, ventral central plate 4; Ml45, lateral marginal plates 4 and 5; Mm46, mesial marginal plates 4 to 6; T, terminal plate." figureDoi="http://doi.org/10.5281/zenodo.10980845" httpUri="https://zenodo.org/record/10980845/files/figure.png" pageId="15" pageNumber="112">Fig. 13A, B</figureCitation>
). The Cd
<subScript id="757339694C58FFEFFA4D82EBA5EAF294" attach="left" box="[1471,1481,941,958]" fontSize="7" pageId="15" pageNumber="112">2</subScript>
is slightly, about 1.2 times, longer than it is broad in IG KSC 155/10. Similarly to that of
<taxonomicName id="2EF740AF4C58FFEFFBA082A4A575F2D1" baseAuthorityName="Luksevics" baseAuthorityYear="2001" box="[1106,1366,994,1019]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="112" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="curonica">
<emphasis id="DB83E73E4C58FFEFFBA082A4A575F2D1" box="[1106,1366,994,1019]" italics="true" pageId="15" pageNumber="112">B. leptocheira curonica</emphasis>
</taxonomicName>
, the distal segment (
<figureCitation id="71CC27A94C58FFEFFC678544A3D7F531" box="[917,1012,1026,1051]" captionStart="Fig" captionStartId="14.[151,184,878,898]" captionTargetBox="[161,1533,162,851]" captionTargetId="figure-604@14.[153,1534,163,851]" captionTargetPageId="14" captionText="Fig. 13. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, plates of the pectoral fin armour. A. Left Cv1 plate IG 155/6 in visceral (A1) and ventral (A2) views. B. Right Cv1 plate IG KSC 155/134 in ventral view. C. Left Ml2 plate IG KSC 155/149 in ventral view. D. Right Ml2 plate IG 155/137 in dorsal (D1) and ventral (D2) views. E. Right distal segment of the pectoral fin armour IG KSC 155/21 in dorsal (E1) and ventral (E2) views. F. Mm1 plate IG KSC 155/85 in mesial view. Abbreviations: Cd35, dorsal central plates 3 to 5; Cv4, ventral central plate 4; Ml45, lateral marginal plates 4 and 5; Mm46, mesial marginal plates 4 to 6; T, terminal plate." figureDoi="http://doi.org/10.5281/zenodo.10980845" httpUri="https://zenodo.org/record/10980845/files/figure.png" pageId="15" pageNumber="112">Fig. 13E</figureCitation>
) is adorned with small and rounded marginal spines; the lateral spines are large, sharp and proximally directed. The distal segment is very long and narrow, with an L/B index of
<quantity id="2E0F96C94C58FFEFFC4D8524A3DEF551" box="[959,1021,1122,1147]" metricMagnitude="-1" metricUnit="m" metricValue="1.3716" pageId="15" pageNumber="112" unit="in" value="5.4">5.4 in</quantity>
both IG KSC 155/21 and IG KSC 155/22 (L/B index of
<quantity id="2E0F96C94C58FFEFFC3B85C4A424F5B1" box="[969,1031,1154,1179]" metricMagnitude="-1" metricUnit="m" metricValue="1.4478" pageId="15" pageNumber="112" unit="in" value="5.7">5.7 in</quantity>
LDM 89/4). The Cd5 plate is present in the material from all three countries.
</paragraph>
<paragraph id="E9483B2C4C58FFEFFCBC8584A594F677" blockId="15.[808,1483,160,1888]" pageId="15" pageNumber="112">
<location id="EC286DF74C58FFEFFCBC8584A359F5F1" LSID="urn:lsid:plazi:treatment:615E8A3A4C53FFF0FCAC84C7A1C7F4F6:EC286DF74C58FFEFFCBC8584A359F5F1" box="[846,890,1218,1243]" country="Latvia" county="On" municipality="The" name="The" pageId="15" pageNumber="112">The</location>
ornamentation is fine and basically of the reticular
<typeStatus id="364C858E4C58FFEFFCDA85A4A379F5D1" box="[808,858,1250,1275]" pageId="15" pageNumber="112">type</typeStatus>
(
<bibRefCitation id="8D6646DD4C58FFEFFC9F85A4A43DF5D1" author="Miles, R. S." box="[877,1054,1250,1275]" pageId="15" pageNumber="112" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles 1968: 80</bibRefCitation>
) in small individuals, changing into vermiculating
<typeStatus id="364C858E4C58FFEFFC3D8444A422F431" box="[975,1025,1282,1307]" pageId="15" pageNumber="112">type</typeStatus>
in the medium-sized fish, and even in the tubercular
<typeStatus id="364C858E4C58FFEFFC3D8465A422F416" box="[975,1025,1315,1340]" pageId="15" pageNumber="112">type</typeStatus>
on the largest specimens (e.g., Prm; IG 155/114), instead of becoming smooth in quite large individuals from
<collectingCountry id="91E07BBC4C58FFEFFC4D8425A428F456" box="[959,1035,1379,1404]" name="Latvia" pageId="15" pageNumber="112">Latvia</collectingCountry>
.
<collectingMunicipality id="092CA1564C58FFEFFBE78425A462F456" box="[1045,1089,1379,1404]" pageId="15" pageNumber="112">The</collectingMunicipality>
network of anastomosing ridges is usually broken into radially arranged shorter ridges on the head shield plates.
<collectingCounty id="002943A04C58FFEFFC0D84E5A401F496" box="[1023,1058,1443,1468]" pageId="15" pageNumber="112">On</collectingCounty>
the posterior margin of the
<collectionCode id="8FE6A3E94C58FFEFFA9084E5A583F496" box="[1378,1440,1443,1468]" pageId="15" pageNumber="112">PMD</collectionCode>
, on the subcephalic and subanal divisions of the
<collectionCode id="8FE6A3E94C58FFEFFAEC8485A576F4F6" box="[1310,1365,1475,1500]" pageId="15" pageNumber="112">AVL</collectionCode>
and
<collectionCode id="8FE6A3E94C58FFEFFA7E8485A5E7F4F6" box="[1420,1476,1475,1500]" country="Argentina" httpUri="http://grbio.org/cool/pbkd-hmeb" name="Paleontologia de Vertebrados Lillo" pageId="15" pageNumber="112">PVL</collectionCode>
, the anastomoses between the ridges reduce, and nodose short ridges are present. The ornamentation on
<collectionCode id="8FE6A3E94C58FFEFFB038745A53CF736" box="[1265,1311,1539,1564]" country="0" name="University of Montana Museum" pageId="15" pageNumber="112">MV</collectionCode>
depends on the size of the plate: it is of reticular
<typeStatus id="364C858E4C58FFEFFB538765A4F2F716" box="[1185,1233,1571,1596]" pageId="15" pageNumber="112">type</typeStatus>
in the smallest plates and becomes of the vermiculating
<typeStatus id="364C858E4C58FFEFFB368705A4DBF776" box="[1220,1272,1603,1628]" pageId="15" pageNumber="112">type</typeStatus>
, i.e., short radiating nodose ridges, along the margins of the largest plates, whereas it remains of the reticular
<typeStatus id="364C858E4C58FFEFFB4887C5A4CFF7B6" box="[1210,1260,1667,1692]" pageId="15" pageNumber="112">type</typeStatus>
in the central part. The ornamentation of the pectoral appendage is reticulate in general. On the anterior part of the Cd
<subScript id="757339694C58FFEFFB018789A4DEF7CA" attach="left" box="[1267,1277,1743,1760]" fontSize="7" pageId="15" pageNumber="112">1</subScript>
and Cv
<subScript id="757339694C58FFEFFAA08789A57FF7CA" attach="left" box="[1362,1372,1743,1760]" fontSize="7" pageId="15" pageNumber="112">1</subScript>
the ornament is radially arranged, whereas on the distal part of the proximal segment the ornamentation becomes smooth. The longitudinal striation in the ornament of the dorsal side of the distal segment is well shown in IG KSC 155/21 (
<figureCitation id="71CC27A94C58FFEFFAB18602A5BCF677" box="[1347,1439,1860,1885]" captionStart="Fig" captionStartId="14.[151,184,878,898]" captionTargetBox="[161,1533,162,851]" captionTargetId="figure-604@14.[153,1534,163,851]" captionTargetPageId="14" captionText="Fig. 13. Antiarchan fish Bothriolepis leptocheira jeremejevi (Rohon, 1900), Sosnogorsk locality, Sosnogorsk Formation, lowermost Famennian, plates of the pectoral fin armour. A. Left Cv1 plate IG 155/6 in visceral (A1) and ventral (A2) views. B. Right Cv1 plate IG KSC 155/134 in ventral view. C. Left Ml2 plate IG KSC 155/149 in ventral view. D. Right Ml2 plate IG 155/137 in dorsal (D1) and ventral (D2) views. E. Right distal segment of the pectoral fin armour IG KSC 155/21 in dorsal (E1) and ventral (E2) views. F. Mm1 plate IG KSC 155/85 in mesial view. Abbreviations: Cd35, dorsal central plates 3 to 5; Cv4, ventral central plate 4; Ml45, lateral marginal plates 4 and 5; Mm46, mesial marginal plates 4 to 6; T, terminal plate." figureDoi="http://doi.org/10.5281/zenodo.10980845" httpUri="https://zenodo.org/record/10980845/files/figure.png" pageId="15" pageNumber="112">Fig. 13E</figureCitation>
<figureCitation id="71CC27A94C58FFEFFA6D8609A58BF64A" box="[1439,1448,1871,1888]" captionStart="Fig" captionStartId="1.[98,131,1054,1074]" captionTargetBox="[98,1481,161,1028]" captionTargetId="figure-390@1.[614,1481,162,1027]" captionTargetPageId="1" captionText="Fig. 1. Geographical and geological setting of the type locality of Bothriolepis leptocheira jeremejevi (Rohon, 1900): sketch-maps of Russia (A) and Komi Republic (B) and geological map of Ukhta-Sosnogorsk area, from Yukhtanov et al. (2008), amended (C). The Natural Geological Monument Sosnovskiy” (Sosnogorsk fossil site) is marked by the black square. Abbreviations: D3f1, Upper Devonian, lower Frasnian; D3f2, Upper Devonian, middle Frasnian; D3f3, Upper Devonian, upper Frasnian; D3fm1, Upper Devonian, lower Famennian; C1, Carboniferous, Mississippian; J2, Middle Jurassic." figureDoi="http://doi.org/10.5281/zenodo.10980821" httpUri="https://zenodo.org/record/10980821/files/figure.png" pageId="15" pageNumber="112">
<subScript id="757339694C58FFEFFA6D8609A58BF64A" attach="left" box="[1439,1448,1871,1888]" fontSize="7" pageId="15" pageNumber="112">1</subScript>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A1ED68A74C58FFF0FCDA8629A1C7F4F6" lastPageId="16" lastPageNumber="113" pageId="15" pageNumber="112" type="discussion">
<paragraph id="E9483B2C4C58FFF0FCDA8629A2C6F20E" blockId="15.[808,1481,1903,1960]" lastBlockId="16.[151,825,160,1424]" lastPageId="16" lastPageNumber="113" pageId="15" pageNumber="112">
<emphasis id="DB83E73E4C58FFEFFCDA8629A3A9F6A2" box="[808,906,1903,1928]" italics="true" pageId="15" pageNumber="112">Remarks</emphasis>
.—This species was established by
<bibRefCitation id="8D6646DD4C58FFEFFAE98629A5EAF6A2" author="Traquair, R. H." box="[1307,1481,1903,1928]" pageId="15" pageNumber="112" pagination="283 - 286" refId="ref20607" refString="Traquair, R. H. 1893. Achanarras revisited. Proceedings of the Royal Phylosophical Society of Edinburgh 11: 283 - 286." type="journal article" year="1893">Traquair (1893)</bibRefCitation>
based on the material from
<collectingCountry id="91E07BBC4C58FFEFFB8E86C9A4FCF682" box="[1148,1247,1935,1960]" name="United Kingdom" pageId="15" pageNumber="112">Scotland</collectingCountry>
and well-described by
<bibRefCitation id="8D6646DD4C47FFF0FF4C81E6A140F193" author="Stensio, E. A." box="[190,355,160,185]" pageId="16" pageNumber="113" pagination="1 - 622" refId="ref20500" refString="Stensio, E. A. 1948. On the Placodermi of the Upper Devonian of East Greenland. II: Antiarchi: subfamily Bothriolepinae. Palaeozoologica Groenlandica 2: 1 - 622." type="journal article" year="1948">Stensiö (1948)</bibRefCitation>
and
<bibRefCitation id="8D6646DD4C47FFF0FE5181E6A21BF193" author="Miles, R. S." box="[419,568,160,185]" pageId="16" pageNumber="113" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles (1968)</bibRefCitation>
.
<bibRefCitation id="8D6646DD4C47FFF0FDB581E6A2F8F193" author="Gross, W." box="[583,731,160,185]" pageId="16" pageNumber="113" pagination="373 - 436" refId="ref18663" refString="Gross, W. 1942. Die Fischfaunen des baltischen Devons und ihre biostratigraphische Bedeutung. Korrespondenz-blatt des Naturforscher-Vereins zu Riga 64: 373 - 436." type="journal article" year="1942">Gross (1942)</bibRefCitation>
erected
<taxonomicName id="2EF740AF4C47FFF0FF658186A1B1F1F3" authorityName="Gross" authorityYear="1942" box="[151,402,192,217]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="113" phylum="Chordata" rank="species" species="curonica">
<emphasis id="DB83E73E4C47FFF0FF658186A1B1F1F3" box="[151,402,192,217]" italics="true" pageId="16" pageNumber="113">Bothriolepis curonica</emphasis>
</taxonomicName>
as a separate species from the lowermost Famennian of
<collectingCountry id="91E07BBC4C47FFF0FE6681A7A1FEF1D0" box="[404,477,225,250]" name="Latvia" pageId="16" pageNumber="113">Latvia</collectingCountry>
based on a small set of bones collected in
<quantity id="2E0F96C94C47FFF0FED98044A1A0F031" box="[299,387,258,283]" metricMagnitude="1" metricUnit="m" metricValue="4.91236" pageId="16" pageNumber="113" unit="in" value="1934.0">1934 in</quantity>
the Bienes locality. This description was repeated by
<bibRefCitation id="8D6646DD4C47FFF0FE928064A227F011" author="Stensio, E. A." box="[352,516,290,315]" pageId="16" pageNumber="113" pagination="1 - 622" refId="ref20500" refString="Stensio, E. A. 1948. On the Placodermi of the Upper Devonian of East Greenland. II: Antiarchi: subfamily Bothriolepinae. Palaeozoologica Groenlandica 2: 1 - 622." type="journal article" year="1948">Stensiö (1948)</bibRefCitation>
in Addenda to his monograph without further comments. Since then, the collection of the remains of this fish was significantly added by Vitaliy Sorokin and Lyubov Lyarskaya in 19751982 and by EL in 2013, who collected specimens from the Kalnamuiža locality in western
<collectingCountry id="91E07BBC4C47FFF0FE8B8083A1E5F0F4" box="[377,454,453,478]" name="Latvia" pageId="16" pageNumber="113">Latvia</collectingCountry>
. Specimens described by
<bibRefCitation id="8D6646DD4C47FFF0FD048083A0FEF0D5" author="Gross, W." pageId="16" pageNumber="113" pagination="373 - 436" refId="ref18663" refString="Gross, W. 1942. Die Fischfaunen des baltischen Devons und ihre biostratigraphische Bedeutung. Korrespondenz-blatt des Naturforscher-Vereins zu Riga 64: 373 - 436." type="journal article" year="1942">Gross (1942)</bibRefCitation>
and material collected from the Kalnamuiža site in 197080ies were briefly described and figured (
<bibRefCitation id="8D6646DD4C47FFF0FD318340A0F2F36A" author="Luksevics, E." pageId="16" pageNumber="113" pagination="90 - 97" refId="ref19208" refString="Luksevics, E. 1987. New information on Bothriolepis curonica Gross from the Upper Devonian of Latvia [in Russian with summaries in Latvian and English]. Priroda i muzej - muzeu prirody Latvijskoj SSR- 140, 90 - 97. Zinatne, Riga." type="journal article" year="1987">Lukševičs 1987</bibRefCitation>
), accenting the close morphological resemblance of
<taxonomicName id="2EF740AF4C47FFF0FF65830EA102F34B" authorityName="Gross" authorityYear="1942" box="[151,289,584,609]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="113" phylum="Chordata" rank="species" species="curonica">
<emphasis id="DB83E73E4C47FFF0FF65830EA102F34B" box="[151,289,584,609]" italics="true" pageId="16" pageNumber="113">B. curonica</emphasis>
</taxonomicName>
with
<taxonomicName id="2EF740AF4C47FFF0FE99830EA22CF34B" baseAuthorityName="Luksevics" baseAuthorityYear="2001" box="[363,527,584,609]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="113" phylum="Chordata" rank="species" species="leptocheira">
<emphasis id="DB83E73E4C47FFF0FE99830EA22CF34B" box="[363,527,584,609]" italics="true" pageId="16" pageNumber="113">B. leptocheira</emphasis>
</taxonomicName>
. A direct comparison of the specimens belonging to these species has indicated that they are conspecific (
<bibRefCitation id="8D6646DD4C47FFF0FE6483CFA26CF388" author="Luksevics, E." box="[406,591,649,674]" pageId="16" pageNumber="113" pagination="489 - 609" refId="ref19408" refString="Luksevics, E. 2001. Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. Geodiversitas 23: 489 - 609." type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
). Nevertheless, several features supported designation of the Latvian material to the subspecies
<taxonomicName id="2EF740AF4C47FFF0FE91838CA0F3F22E" authority="Gross, 1942" authorityName="Gross" authorityYear="1942" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="113" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="curonica">
<emphasis id="DB83E73E4C47FFF0FE91838CA2C5F3C9" box="[355,742,714,739]" italics="true" pageId="16" pageNumber="113">Bothriolepis leptocheira curonica</emphasis>
<bibRefCitation id="8D6646DD4C47FFF0FD1D838CA0F3F22E" author="Gross, W." pageId="16" pageNumber="113" pagination="373 - 436" refId="ref18663" refString="Gross, W. 1942. Die Fischfaunen des baltischen Devons und ihre biostratigraphische Bedeutung. Korrespondenz-blatt des Naturforscher-Vereins zu Riga 64: 373 - 436." type="journal article" year="1942">Gross, 1942</bibRefCitation>
</taxonomicName>
, which is morphologically very close to
<taxonomicName id="2EF740AF4C47FFF0FD5E83ADA183F20E" baseAuthorityName="Miles" baseAuthorityYear="1968" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="113" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="leptocheira">
<emphasis id="DB83E73E4C47FFF0FD5E83ADA183F20E" italics="true" pageId="16" pageNumber="113">Bothriolepis leptocheira leptocheira</emphasis>
</taxonomicName>
from
<collectingCountry id="91E07BBC4C47FFF0FE17824DA26BF20E" box="[485,584,779,804]" name="United Kingdom" pageId="16" pageNumber="113">Scotland</collectingCountry>
(
<bibRefCitation id="8D6646DD4C47FFF0FDA4824DA2FAF20E" author="Miles, R. S." box="[598,729,779,804]" pageId="16" pageNumber="113" pagination="1 - 130" refId="ref19937" refString="Miles, R. S. 1968. The Old Red Sandstone antiarchs of Scotland: family Bothriolepididae. Palaeontographical Society Monograph 122: 1 - 130." type="journal article" year="1968">Miles 1968</bibRefCitation>
).
</paragraph>
<paragraph id="E9483B2C4C47FFF0FF4F826AA2A8F4BA" blockId="16.[151,825,160,1424]" pageId="16" pageNumber="113">
<taxonomicName id="2EF740AF4C47FFF0FF4F826AA2BFF26F" authority="(Rohon, 1900)" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" baseAuthorityName="Rohon" baseAuthorityYear="1900" box="[189,668,812,837]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="113" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="jeremejevi">
<emphasis id="DB83E73E4C47FFF0FF4F826AA1C5F26F" box="[189,486,812,837]" italics="true" pageId="16" pageNumber="113">B. leptocheira jeremejevi</emphasis>
(
<bibRefCitation id="8D6646DD4C47FFF0FE09826AA2B7F26F" author="Rohon, J. V." box="[507,660,812,837]" pageId="16" pageNumber="113" pagination="3 - 77" refId="ref20379" refString="Rohon, J. V. 1900. Die Devonischen Fische von Timan in Russland. Sitzungsberichte der Konigliche Bohmischen Gesellschaft der Wissenschaften. Mathematisch-naturwissenschaltliche Classe (1899) 8: 3 - 77." type="journal article" year="1900">Rohon, 1900</bibRefCitation>
)
</taxonomicName>
differs from the nominal subspecies
<taxonomicName id="2EF740AF4C47FFF0FE49820AA2D1F24F" baseAuthorityName="Miles" baseAuthorityYear="1968" box="[443,754,844,869]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="113" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="leptocheira">
<emphasis id="DB83E73E4C47FFF0FE49820AA2D1F24F" box="[443,754,844,869]" italics="true" pageId="16" pageNumber="113">B. leptocheira leptocheira</emphasis>
</taxonomicName>
from
<collectingCountry id="91E07BBC4C47FFF0FF65822BA0D9F2AC" box="[151,250,877,902]" name="United Kingdom" pageId="16" pageNumber="113">Scotland</collectingCountry>
in its: (i) larger size; (ii) slightly narrower Pn plate; (iii) more acute posterior angle of
<collectionCode id="8FE6A3E94C47FFF0FDD582C8A247F28D" box="[551,612,910,935]" pageId="16" pageNumber="113">PMD</collectionCode>
plate; (iv) slightly shorter
<collectionCode id="8FE6A3E94C47FFF0FF1F82E8A108F2ED" box="[237,299,942,967]" pageId="16" pageNumber="113">ADL</collectionCode>
. It differs from
<emphasis id="DB83E73E4C47FFF0FE2F82E8A254F2ED" box="[477,631,942,967]" italics="true" pageId="16" pageNumber="113">
B. l.
<taxonomicName id="2EF740AF4C47FFF0FDE082E8A254F2ED" authorityName="Gross" authorityYear="1942" box="[530,631,942,967]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="113" phylum="Chordata" rank="species" species="curonica">curonica</taxonomicName>
</emphasis>
in its: (i) slightly narrower Prm and Pn plates, shorter and broader Pmg plate; (ii) slightly narrower
<collectionCode id="8FE6A3E94C47FFF0FE7F82B6A1F3F523" box="[397,464,1008,1033]" pageId="16" pageNumber="113">AMD</collectionCode>
plate; (iii) relatively higher lateral wall of
<collectionCode id="8FE6A3E94C47FFF0FEEE8556A176F503" box="[284,341,1040,1065]" pageId="16" pageNumber="113">MxL</collectionCode>
and
<collectionCode id="8FE6A3E94C47FFF0FE788556A1E2F503" box="[394,449,1040,1065]" pageId="16" pageNumber="113">AVL</collectionCode>
plates; (iv) relatively shorter pectoral fin armour and separate plates of the fin; (v) coarser ornamentation. Overall proportions of the plates of the trunk armour and of the bones of the pectoral fin, distribution and path of sensory line canals as well as ornamentation of
<taxonomicName id="2EF740AF4C47FFF0FF6585F5A198F5E6" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" baseAuthorityName="Rohon" baseAuthorityYear="1900" box="[151,443,1203,1228]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="113" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="jeremejevi">
<emphasis id="DB83E73E4C47FFF0FF6585F5A198F5E6" box="[151,443,1203,1228]" italics="true" pageId="16" pageNumber="113">B. leptocheira jeremejevi</emphasis>
</taxonomicName>
are rather similar to those of
<taxonomicName id="2EF740AF4C47FFF0FCD285F5A13BF5C7" baseAuthorityName="Luksevics" baseAuthorityYear="2001" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="113" phylum="Chordata" rank="species" species="leptocheira">
<emphasis id="DB83E73E4C47FFF0FCD285F5A13BF5C7" italics="true" pageId="16" pageNumber="113">B. leptocheira</emphasis>
</taxonomicName>
from
<collectingCountry id="91E07BBC4C47FFF0FEAC8592A1E3F5C7" box="[350,448,1236,1261]" name="United Kingdom" pageId="16" pageNumber="113">Scotland</collectingCountry>
and from the earliest Famennian of
<collectingCountry id="91E07BBC4C47FFF0FF4485B2A120F427" box="[182,259,1268,1293]" name="Latvia" pageId="16" pageNumber="113">Latvia</collectingCountry>
. The dissimilarities between the three subspecies mainly concern ornamentation, slightly differing proportions of some plates, relatively shorter but higher trunk armour, and shorter pectoral fins of
<taxonomicName id="2EF740AF4C47FFF0FDE98410A31BF445" authorityName="Lukševičs &amp; Beznosov &amp; Stūris" authorityYear="2017" baseAuthorityName="Rohon" baseAuthorityYear="1900" box="[539,824,1366,1391]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="113" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="jeremejevi">
<emphasis id="DB83E73E4C47FFF0FDE98410A31BF445" box="[539,824,1366,1391]" italics="true" pageId="16" pageNumber="113">B. leptocheira jeremejevi</emphasis>
</taxonomicName>
in comparison with
<taxonomicName id="2EF740AF4C47FFF0FE8B8431A2A7F4BA" authorityName="Gross" authorityYear="1942" box="[377,644,1399,1424]" family="Bothriolepididae" genus="Bothriolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="113" phylum="Chordata" rank="subSpecies" species="leptocheira" subSpecies="curonica">
<emphasis id="DB83E73E4C47FFF0FE8B8431A2A7F4BA" box="[377,644,1399,1424]" italics="true" pageId="16" pageNumber="113">B. leptocheira curonica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="E9483B2C4C47FFF0FF6584E4A1C7F4F6" blockId="16.[151,824,1442,1500]" pageId="16" pageNumber="113">
<emphasis id="DB83E73E4C47FFF0FF6584E4A21CF491" box="[151,575,1442,1467]" italics="true" pageId="16" pageNumber="113">Stratigraphic and geographic range</emphasis>
.—Known only from the
<typeStatus id="364C858E4C47FFF0FF338485A0D0F4F6" box="[193,243,1475,1500]" pageId="16" pageNumber="113">type</typeStatus>
locality and horizon.
</paragraph>
</subSubSection>
</treatment>
</document>